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<document id="B92CF203CFD6723EFE0988A1DF3CDB7F" ID-DOI="10.5852/ejt.2020.669" ID-GBIF-Dataset="47b43e12-37bc-4b2c-b654-0e668351cc97" ID-ISSN="2118-9773" ID-Zenodo-Dep="3900298" ID-ZooBank="06FEC792-67DE-46F1-A5D3-4B0F17AF3B73" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1592505667257" checkinUser="valdenar" docAuthor="Haran, Julien M., Hansen, Steffan, Benoit, Laure &amp; Addison, Pia" docDate="2020" docId="BD7C0F46FFCB5305FDD2FB10FAC209D7" docLanguage="en" docName="ejt-669_haran_hansen_addison.pdf.imf" docOrigin="European Journal of Taxonomy 669 (669)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phlyctinus littoralis Haran &amp; Hansen &amp; Benoit &amp; Cbgp 2020, sp. nov." docType="treatment" docUuid="49467405-4829-4E45-A2A5-70130EDA95E8" docUuidSource="ZooBank" docVersion="11" lastPageNumber="21" masterDocId="4145773EFFD95311FF99FF97FFB50A6A" masterDocTitle="Description of five new species in the genus Phlyctinus Schoenherr (Coleoptera, Curculionidae): a first step in deciphering the P. callosus complex" masterLastPageNumber="29" masterPageNumber="1" pageNumber="19" updateTime="1733426279942" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="015995CB8BAF22F3F584E0A1FB29F70C">Description of five new species in the genus Phlyctinus Schoenherr (Coleoptera, Curculionidae): a first step in deciphering the P. callosus complex</mods:title>
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<mods:name id="B93A8C5BB1F940C69EAD53E365B00DF4" type="personal">
<mods:role id="74438CEA6F666142680D9EF0A13197EA">
<mods:roleTerm id="C9E9F659181A083388C9B48246FA7CDF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1340BE1E2D520729CB6104029D42574A">Haran, Julien M.</mods:namePart>
<mods:nameIdentifier id="916EF55A5E3C2AE1CAC8982E7483751C" type="LSID">A04E1722-994A-44AD-8FD2-28DC0F220805</mods:nameIdentifier>
<mods:affiliation id="72C1538988FF3EBF1C01B06556B42FF8">CBGP, CIRAD, Montpellier SupAgro, INRA, IRD, Montpellier University, Montpellier, France. &amp; Department of Conservation Ecology and Entomology, Stellenbosch University, Stellenbosch, South Africa.</mods:affiliation>
<mods:nameIdentifier id="B3004993B4909BD35F6F819AB0875963" type="email">julien.haran@cirad.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="1AB4F638B51DD94CB042C5EAAA3B9BE0" type="personal">
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</mods:role>
<mods:namePart id="8F38BFAA97A275AE8CFFF2D6131E93BC">Hansen, Steffan</mods:namePart>
<mods:nameIdentifier id="04EB113EC82494DFF5D5D8D69938E698" type="LSID">52F0FC34-03F6-4F04-A1AA-FE7607B0EFBC</mods:nameIdentifier>
<mods:affiliation id="4D4FCD01ADA6F0AE0FFAA6C36929E022">Department of Conservation Ecology and Entomology, Stellenbosch University, Stellenbosch, South Africa.</mods:affiliation>
<mods:nameIdentifier id="66F4A4565163C9DC5C803FEE4C002145" type="email">shansen.rsa@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="5CF1B0F4B4E72759538925FA01561126" type="personal">
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<mods:roleTerm id="5A2EB817EBF206791E4DA3D9E109F030">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8A30D083683877C8BDC0E648309AC396">Benoit, Laure</mods:namePart>
<mods:nameIdentifier id="7086EC1196A119FEEADF20AFCC06807F" type="LSID">61963F74-724B-4174-9E9A-8817A3516B0E</mods:nameIdentifier>
<mods:affiliation id="32A236DE895C8F125C01C7B3C16C7E87">CBGP, CIRAD, Montpellier SupAgro, INRA, IRD, Montpellier University, Montpellier, France.</mods:affiliation>
<mods:nameIdentifier id="A533D2911F457C87F00A2800E117D0EF" type="email">laure.benoit@cirad.fr</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="B4D02ABAC72832AD9E3363C587B16AD2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C57CC84F1B883EC3C954921358FFE2E9">Addison, Pia</mods:namePart>
<mods:nameIdentifier id="01556F955A6A33DF6769884D98731F09" type="LSID">A84CE8F2-E423-4ABF-815C-35169E6E42E5</mods:nameIdentifier>
<mods:affiliation id="BE5932B643E92AD4AD6078E3F942E167">Department of Conservation Ecology and Entomology, Stellenbosch University, Stellenbosch, South Africa.</mods:affiliation>
<mods:nameIdentifier id="1035598B209974CDFA97AAE11A8F94CE" type="email">pia@sun.ac.za</mods:nameIdentifier>
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<mods:title id="018669F7D3D48058A41537F8014B65F1">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="7A4B3378F2BF605F93561EB058193C26">
<mods:date id="9D64996D5DB3638B91DEE70E0CC9A180">2020</mods:date>
<mods:detail id="3A2C8785CC3289AF20A0D1328D9F0544" type="pubDate">
<mods:number id="7217DB1E174B82B2519B129CC6FAB603">2020-06-18</mods:number>
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<mods:identifier id="7F4926DE3A178FCD669D2DF224039592" type="GBIF-Dataset">47b43e12-37bc-4b2c-b654-0e668351cc97</mods:identifier>
<mods:identifier id="4A83B21D4C3736EC9E4F29F3ACDD61D8" type="ISSN">2118-9773</mods:identifier>
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<treatment id="BD7C0F46FFCB5305FDD2FB10FAC209D7" ID-GBIF-Taxon="164615713" LSID="urn:lsid:zoobank.org:act:49467405-4829-4E45-A2A5-70130EDA95E8" httpUri="http://treatment.plazi.org/id/BD7C0F46FFCB5305FDD2FB10FAC209D7" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" scope_family="Curculionidae" scope_order="Coleoptera" scope_subFamily="Entiminae" scope_superFamily="Curculionoidea" scope_tribe="Oosomini">
<subSubSection id="7DCFEDDBFFCB5303FDD2FB10FC5C0EC8" box="[587,1001,1159,1186]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="356ABE50FFCB5303FDD2FB10FC5C0EC8" blockId="18.[375,1212,1159,1256]" box="[587,1001,1159,1186]" pageId="18" pageNumber="19">
<heading id="6E22093CFFCB5303FDD2FB10FC5C0EC8" box="[587,1001,1159,1186]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<taxonomicName id="F2D5C5D3FFCB5303FDD2FB10FC320EC8" ID-CoL="8KBN4" authority="Haran" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[587,903,1159,1186]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="littoralis" status="sp. nov.">
<emphasis id="07A16242FFCB5303FDD2FB10FC8D0EC8" bold="true" box="[587,824,1159,1186]" italics="true" pageId="18" pageNumber="19">Phlyctinus littoralis</emphasis>
Haran
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCB5303FC17FB1FFC5C0EC8" box="[910,1001,1160,1186]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7DCFEDDBFFCB5303FEEEFB3CFCC30E82" pageId="18" pageNumber="19" type="description">
<paragraph id="356ABE50FFCB5303FEEEFB3CFB090EAC" blockId="18.[375,1212,1159,1256]" box="[375,1212,1195,1222]" pageId="18" pageNumber="19">
<uri id="4144B252FFCB5303FEEEFB3CFB090EAC" box="[375,1212,1195,1222]" pageId="18" pageNumber="19">
urn:lsid:zoobank.org:act:
<uuid id="41738485FFCB5303FD0AFB3CFB090EAC" box="[659,1212,1195,1222]" pageId="18" pageNumber="19">49467405-4829-4E45-A2A5-70130EDA95E8</uuid>
</uri>
</paragraph>
<paragraph id="356ABE50FFCB5303FD27FB59FCC30E82" blockId="18.[375,1212,1159,1256]" box="[702,886,1230,1256]" pageId="18" pageNumber="19">
<figureCitation id="ADEEA2D5FFCB5303FD27FB59FCAF0E82" box="[702,794,1230,1256]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[192,1395,350,1949]" captionTargetId="figure@7.[189,1398,319,1964]" captionTargetPageId="7" captionText="Fig. 1 (opposite page). Habitus of species of Phlyctinus Schoenherr, 1826 males (left) and females (right). A. P. callosus (Schoenherr, 1826). B. P. grootbosensis Haran sp. nov. C. P. xerophilus Haran sp. nov., holotype ♂. D. P. planithorax Haran sp. nov., holotype ♂. E. P. littoralis Haran sp. nov., holotype ♂. F. P. aloevorus Haran sp. nov., holotype ♂. Scale bars = 3 mm ." figureDoi="http://doi.org/10.5281/zenodo.14285225" httpUri="https://zenodo.org/record/14285225/files/figure.png" pageId="18" pageNumber="19">Figs 1E</figureCitation>
,
<figureCitation id="ADEEA2D5FFCB5303FCBEFB59FCFF0E82" box="[807,842,1230,1256]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[252,1379,330,1800]" captionTargetId="figure@10.[189,1398,308,1806]" captionTargetPageId="10" captionText="Fig. 2. Penis of Phlyctinus Schoenherr, 1826, in dorsal (left) and lateral (right) view, and copulatory sclerite (top right). A. P. callosus (Schoenherr, 1826). B. P. grootbosensis Haran sp. nov. C. P. xerophilus Haran sp. nov., holotype ♂. D. P. planithorax Haran sp. nov., holotype ♂. E. P. littoralis Haran sp. nov., holotype ♂. F. P. aloevorus Haran sp. nov., holotype ♂. Scale bars = 500 µm. Copulatory sclerite not to scale." figureDoi="http://doi.org/10.5281/zenodo.3900302" httpUri="https://zenodo.org/record/3900302/files/figure.png" pageId="18" pageNumber="19">2E</figureCitation>
,
<figureCitation id="ADEEA2D5FFCB5303FCCFFB59FCC30E82" box="[854,886,1230,1256]" captionStart="Fig" captionStartId="14.[189,232,1855,1881]" captionTargetBox="[249,1342,301,1975]" captionTargetId="figure@13.[226,1361,265,2019]" captionTargetPageId="13" captionText="Fig. 3 (opposite page). AB. Ventrites of males of Phlyctinus Schoenherr, 1826 (left: ventrites 15, right: ventrite 5). A. P. callosus (Schoenherr, 1826). B. P. grootbosensis Haran sp. nov. CD. Ventrite 5. C. P. xerophilus Haran sp. nov., holotype ♂. D. P. planithorax Haran sp. nov. EF. Vestiture of basal right side of elytra. E. P. callosus (Schoenherr, 1826) (left). F. P. littoralis Haran sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3900304" httpUri="https://zenodo.org/record/3900304/files/figure.png" pageId="18" pageNumber="19">3F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7DCFEDDBFFCB5303FF24FA87FD9C0F11" pageId="18" pageNumber="19" type="etymology">
<paragraph id="356ABE50FFCB5303FF24FA87FEF10F40" blockId="18.[189,324,1296,1322]" box="[189,324,1296,1322]" pageId="18" pageNumber="19">
<heading id="6E22093CFFCB5303FF24FA87FEF10F40" bold="true" box="[189,324,1296,1322]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="07A16242FFCB5303FF24FA87FEF10F40" bold="true" box="[189,324,1296,1322]" pageId="18" pageNumber="19">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCB5303FF24FAA9FD9C0F11" blockId="18.[189,1399,1342,1404]" pageId="18" pageNumber="19">
<taxonomicName id="F2D5C5D3FFCB5303FF24FAA9FE120F32" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[189,423,1342,1368]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="littoralis" status="sp. nov.">
<emphasis id="07A16242FFCB5303FF24FAA9FE120F32" box="[189,423,1342,1368]" italics="true" pageId="18" pageNumber="19">Phlyctinus littoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCB5303FE29FAA8FDB90F33" box="[432,524,1343,1369]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
is named in reference to the coastal habitat in which it was found and to which it seems to be restricted.
</paragraph>
</subSubSection>
<subSubSection id="7DCFEDDBFFCB5303FF24FA34FC040CD0" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="356ABE50FFCB5303FF24FA34FE1F0FD7" blockId="18.[189,426,1443,1469]" box="[189,426,1443,1469]" pageId="18" pageNumber="19">
<heading id="6E22093CFFCB5303FF24FA34FE1F0FD7" bold="true" box="[189,426,1443,1469]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="07A16242FFCB5303FF24FA34FE1F0FD7" bold="true" box="[189,426,1443,1469]" pageId="18" pageNumber="19">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCB5303FF73FA45FEE90F86" blockId="18.[189,1398,1490,1586]" box="[234,348,1490,1516]" pageId="18" pageNumber="19">
<emphasis id="07A16242FFCB5303FF73FA45FEE90F86" bold="true" box="[234,348,1490,1516]" pageId="18" pageNumber="19">
<typeStatus id="EA6E00F2FFCB5303FF73FA45FEE90F86" box="[234,348,1490,1516]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="356ABE50FFCB5303FF24FA62FA9F0C58" blockId="18.[189,1398,1490,1586]" pageId="18" pageNumber="19">
<materialsCitation id="85BDB40DFFCB5303FF24FA62FA930C58" collectingDate="2018-05-12" collectionCode="CBGP" country="South Africa" latitude="-34.243618" location="Kogel Bay" longLatPrecision="1" longitude="18.8529" municipality="Kogel Bay" pageId="18" pageNumber="19" specimenCode="JHAR00973" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape province">
<collectingCountry id="4DC2FEC0FFCB5303FF24FA62FDFA0C65" box="[189,591,1525,1551]" name="South Africa" pageId="18" pageNumber="19">REPUBLIC OF SOUTH AFRICA</collectingCountry>
<specimenCount id="23D375D9FFCB5303FDFEFA62FDCA0C65" box="[615,639,1525,1551]" pageId="18" pageNumber="19" type="male"></specimenCount>
;
<collectingRegion id="F71170B2FFCB5303FD12FA62FC2B0C65" box="[651,926,1525,1551]" country="South Africa" name="Western Cape" pageId="18" pageNumber="19">Western Cape province</collectingRegion>
,
<collectingMunicipality id="D50E242AFFCB5303FC30FA62FB9F0C65" box="[937,1066,1525,1551]" pageId="18" pageNumber="19">Kogel Bay</collectingMunicipality>
;
<geoCoordinate id="50E1D897FFCB5303FBACFA64FB790C65" box="[1077,1228,1523,1551]" degrees="34" direction="south" minutes="14.617" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="-34.243618">34º14.617ʹ S</geoCoordinate>
,
<geoCoordinate id="50E1D897FFCB5303FB41FA64FAC40C65" box="[1240,1393,1523,1551]" degrees="18" direction="east" minutes="51.174" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="18.8529">18º51.174ʹ E</geoCoordinate>
;
<date id="416B9890FFCB5303FF24F98FFEE80C58" box="[189,349,1560,1586]" pageId="18" pageNumber="19" value="2018-05-12">
<collectingDate id="512F6178FFCB5303FF24F98FFEE80C58" box="[189,349,1560,1586]" pageId="18" pageNumber="19" value="2018-05-12">12 May 2018</collectingDate>
</date>
; at base of
<taxonomicName id="F2D5C5D3FFCB5303FE78F98FFD650C58" box="[481,720,1560,1586]" class="Magnoliopsida" family="Asteraceae" genus="Arctotis" kingdom="Plantae" order="Asterales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="angustifolia">
<emphasis id="07A16242FFCB5303FE78F98FFD650C58" box="[481,720,1560,1586]" italics="true" pageId="18" pageNumber="19">Arctotis angustifolia</emphasis>
</taxonomicName>
L. (
<taxonomicName id="F2D5C5D3FFCB5303FC99F98FFC3D0C58" box="[768,904,1560,1586]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
);
<collectionCode id="53C42695FFCB5303FC05F98FFC5D0C58" box="[924,1000,1560,1586]" pageId="18" pageNumber="19">CBGP</collectionCode>
code
<specimenCode id="6573162BFFCB5303FBB4F98FFB760C58" box="[1069,1219,1560,1586]" pageId="18" pageNumber="19">JHAR00973</specimenCode>
; SAMC
</materialsCitation>
.
</paragraph>
<paragraph id="356ABE50FFCB5303FF73F9CDFED20C1E" blockId="18.[189,1399,1626,1722]" box="[234,359,1626,1652]" pageId="18" pageNumber="19">
<emphasis id="07A16242FFCB5303FF73F9CDFED20C1E" bold="true" box="[234,359,1626,1652]" pageId="18" pageNumber="19">
<typeStatus id="EA6E00F2FFCB5303FF73F9CDFED20C1E" box="[234,359,1626,1652]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="356ABE50FFCB5303FF24F9EAFC040CD0" blockId="18.[189,1399,1626,1722]" pageId="18" pageNumber="19">
<materialsCitation id="85BDB40DFFCB5303FF24F9EAFEBC0CD0" ID-GBIF-Occurrence="2640284367" collectingDate="2018-09-16" collectionCode="CBGP" country="South Africa" location="REPUBLIC OF SOUTH AFRICA" pageId="18" pageNumber="19" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="holotype">
<collectingCountry id="4DC2FEC0FFCB5303FF24F9EAFDE10CFD" box="[189,596,1661,1687]" name="South Africa" pageId="18" pageNumber="19">REPUBLIC OF SOUTH AFRICA</collectingCountry>
<specimenCount id="23D375D9FFCB5303FDF6F9EAFD010CFD" box="[623,692,1661,1687]" count="2" pageId="18" pageNumber="19" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="23D375D9FFCB5303FD5BF9EAFCB30CFD" box="[706,774,1661,1687]" count="2" pageId="18" pageNumber="19" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<date id="416B9890FFCB5303FB4BF9EBFAC70CFD" box="[1234,1394,1660,1687]" pageId="18" pageNumber="19" value="2018-09-16">
<collectingDate id="512F6178FFCB5303FB4BF9EBFAC70CFD" box="[1234,1394,1660,1687]" pageId="18" pageNumber="19" value="2018-09-16">16 Sep. 2018</collectingDate>
</date>
;
<collectionCode id="53C42695FFCB5303FF24F937FEBC0CD0" box="[189,265,1696,1722]" pageId="18" pageNumber="19">CBGP</collectionCode>
</materialsCitation>
<materialsCitation id="85BDB40DFFCB5303FEB8F937FC180CD0" ID-GBIF-Occurrence="2640284340" box="[289,941,1696,1722]" collectingDate="2018-09-16" collectionCode="SANC" country="South Africa" location="REPUBLIC OF SOUTH AFRICA" pageId="18" pageNumber="19" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="23D375D9FFCB5303FEB8F937FEFB0CD0" box="[289,334,1696,1722]" count="1" pageId="18" pageNumber="19" type="male">1 ♂</specimenCount>
,
<specimenCount id="23D375D9FFCB5303FEC5F937FE3C0CD0" box="[348,393,1696,1722]" count="1" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="53C42695FFCB5303FCC2F937FC180CD0" box="[859,941,1696,1722]" collectionName="South Africa, Pretoria, South African National Collection of Insects" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="18" pageNumber="19" type="Museum">SANC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="61AAEED8FFCB5303FF24F88EFD7B0D89" ID-DOI="http://doi.org/10.5281/zenodo.14285227" ID-Zenodo-Dep="14285227" httpUri="https://zenodo.org/record/14285227/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1817,1843]" targetBox="[264,1322,286,2013]" targetPageId="17">
<paragraph id="356ABE50FFCB5303FF24F88EFD7B0D89" blockId="18.[189,1399,1817,2019]" pageId="18" pageNumber="19">
<emphasis id="07A16242FFCB5303FF24F88EFEB20D59" bold="true" box="[189,263,1817,1843]" pageId="18" pageNumber="19">Fig. 4</emphasis>
(opposite page). Habitus
<emphasis id="07A16242FFCB5303FDDCF88DFD000D59" box="[581,693,1818,1843]" italics="true" pageId="18" pageNumber="19">in natura</emphasis>
, host plants and biotopes of
<taxonomicName id="F2D5C5D3FFCB5303FB87F88EFAC20D59" authority="Schoenherr, 1826" authorityName="Schoenherr" authorityYear="1826" box="[1054,1399,1817,1843]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="07A16242FFCB5303FB87F88EFB2F0D59" box="[1054,1178,1817,1843]" italics="true" pageId="18" pageNumber="19">Phlyctinus</emphasis>
<bibRefCitation id="5144C3A1FFCB5303FB3CF88EFAC20D59" author="Schoenherr C. J." box="[1189,1399,1817,1843]" pageId="18" pageNumber="19" refId="ref16598" refString="Schoenherr C. J. 1826. Curculionidum Dispositio Methodica, cum Generum Characteribus, Descriptionibus atque Observationibus variis, seu Prodromus ad Synonymiae Insectorum, Partem IV. Fleischer, Leipzig [Lipsiae]. https: // doi. org / 10.5962 / bhl. title. 9327" type="book" year="1826">Schoenherr, 1826</bibRefCitation>
</taxonomicName>
species.
<emphasis id="07A16242FFCB5303FEB9F8ABFE800D3C" bold="true" box="[288,309,1852,1878]" pageId="18" pageNumber="19">A</emphasis>
.
<taxonomicName id="F2D5C5D3FFCB5303FEDAF8AAFD1E0D3D" authority="(Schoenherr, 1826)" baseAuthorityName="Schoenherr" baseAuthorityYear="1826" box="[323,683,1852,1879]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="callosus">
<emphasis id="07A16242FFCB5303FEDAF8AAFE0A0D3C" box="[323,447,1852,1878]" italics="true" pageId="18" pageNumber="19">P. callosus</emphasis>
(
<bibRefCitation id="5144C3A1FFCB5303FE57F8ABFD140D3D" author="Schoenherr C. J." box="[462,673,1852,1879]" pageId="18" pageNumber="19" refId="ref16598" refString="Schoenherr C. J. 1826. Curculionidum Dispositio Methodica, cum Generum Characteribus, Descriptionibus atque Observationibus variis, seu Prodromus ad Synonymiae Insectorum, Partem IV. Fleischer, Leipzig [Lipsiae]. https: // doi. org / 10.5962 / bhl. title. 9327" type="book" year="1826">Schoenherr, 1826</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="07A16242FFCB5303FD2CF8ABFD7C0D3C" bold="true" box="[693,713,1852,1878]" pageId="18" pageNumber="19">B</emphasis>
.
<taxonomicName id="F2D5C5D3FFCB5303FD4FF8AAFC090D3D" authority="Haran" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[726,956,1852,1879]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="xerophilus" status="sp. nov.">
<emphasis id="07A16242FFCB5303FD4FF8AAFCDB0D3C" box="[726,878,1852,1878]" italics="true" pageId="18" pageNumber="19">P. xerophilus</emphasis>
Haran
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCB5303FC5AF8AAFBA80D3D" box="[963,1053,1853,1879]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="07A16242FFCB5303FBBAF8ABFBE90D3C" bold="true" box="[1059,1116,1852,1878]" pageId="18" pageNumber="19">CD</emphasis>
.
<taxonomicName id="F2D5C5D3FFCB5303FBF3F8AAFAC20D3D" authority="Haran" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[1130,1399,1852,1879]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="grootbosensis" status="sp. nov.">
<emphasis id="07A16242FFCB5303FBF3F8AAFA9D0D3C" box="[1130,1320,1852,1878]" italics="true" pageId="18" pageNumber="19">P. grootbosensis</emphasis>
Haran
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCB5303FF24F8F7FEA30D10" box="[189,278,1888,1914]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
and its host plant,
<taxonomicName id="F2D5C5D3FFCB5303FE77F8F7FC860D13" box="[494,819,1887,1914]" class="Magnoliopsida" family="Asteraceae" genus="Osteospermum" kingdom="Plantae" order="Asterales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="moniliferum">
<emphasis id="07A16242FFCB5303FE77F8F7FC860D13" box="[494,819,1887,1914]" italics="true" pageId="18" pageNumber="19">Osteospermum moniliferum</emphasis>
</taxonomicName>
L.
<emphasis id="07A16242FFCB5303FCC0F8F7FC380D10" bold="true" box="[857,909,1888,1914]" pageId="18" pageNumber="19">EF</emphasis>
.
<taxonomicName id="F2D5C5D3FFCB5303FC02F8F7FB3B0D10" authority="Haran" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[923,1166,1887,1914]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="planithorax" status="sp. nov.">
<emphasis id="07A16242FFCB5303FC02F8F7FBF50D13" box="[923,1088,1887,1913]" italics="true" pageId="18" pageNumber="19">P. planithorax</emphasis>
Haran
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCB5303FB0DF8F7FB580D10" box="[1172,1261,1888,1914]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
and its host plant,
<taxonomicName id="F2D5C5D3FFCB5303FE9EF814FD870DF7" box="[263,562,1923,1949]" class="Cestoda" family="Hymenolepididae" genus="Hymenolepis" kingdom="Animalia" order="Cyclophyllidea" pageId="18" pageNumber="19" phylum="Platyhelminthes" rank="species" species="crithmifolia">
<emphasis id="07A16242FFCB5303FE9EF814FD870DF7" box="[263,562,1923,1949]" italics="true" pageId="18" pageNumber="19">Hymenolepis crithmifolia</emphasis>
</taxonomicName>
(L.) Greuter.
<emphasis id="07A16242FFCB5303FD4FF815FCA70DF6" bold="true" box="[726,786,1922,1948]" pageId="18" pageNumber="19">GH</emphasis>
. Biotope and host plant (
<taxonomicName id="F2D5C5D3FFCB5303FBA5F814FA990DF7" box="[1084,1324,1923,1949]" class="Magnoliopsida" family="Asteraceae" genus="Arctotis" kingdom="Plantae" order="Asterales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="angustifolia">
<emphasis id="07A16242FFCB5303FBA5F814FA990DF7" box="[1084,1324,1923,1949]" italics="true" pageId="18" pageNumber="19">Arctotis angustifolia</emphasis>
</taxonomicName>
L.) of
<taxonomicName id="F2D5C5D3FFCB5303FF24F830FE3E0DAA" authority="Haran" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[189,395,1958,1984]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="littoralis" status="sp. nov.">
<emphasis id="07A16242FFCB5303FF24F830FE880DAA" box="[189,317,1958,1984]" italics="true" pageId="18" pageNumber="19">P. littoralis</emphasis>
Haran
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCB5303FE08F831FE5F0DAA" box="[401,490,1958,1984]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="07A16242FFCB5303FE69F831FDAC0DAA" bold="true" box="[496,537,1958,1984]" pageId="18" pageNumber="19">IJ</emphasis>
.
<taxonomicName id="F2D5C5D3FFCB5303FDBFF830FCB60DAA" authority="Haran" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[550,771,1958,1984]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aloevorus" status="sp. nov.">
<emphasis id="07A16242FFCB5303FDBFF830FD000DAA" box="[550,693,1958,1984]" italics="true" pageId="18" pageNumber="19">P. aloevorus</emphasis>
Haran
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCB5303FC90F831FCD70DAA" box="[777,866,1958,1984]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
on its host plant,
<taxonomicName id="F2D5C5D3FFCB5303FBB2F831FA8D0DAA" authority="Mill." authorityName="Mill." box="[1067,1336,1958,1984]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Aloe" kingdom="Plantae" order="Asparagales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="07A16242FFCB5303FBB2F831FB400DAA" box="[1067,1269,1958,1984]" italics="true" pageId="18" pageNumber="19">Aloe arborescens</emphasis>
Mill.
</taxonomicName>
, with details of feeding damage on floral peduncle.
</paragraph>
</caption>
<subSubSection id="7DCFEDDBFFCA5305FF24FE9CFD5E0844" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
<paragraph id="356ABE50FFCA5302FF24FE9CFEFB0B4F" blockId="19.[189,334,267,293]" box="[189,334,267,293]" pageId="19" pageNumber="20">
<heading id="6E22093CFFCA5302FF24FE9CFEFB0B4F" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="07A16242FFCA5302FF24FE9CFEFB0B4F" bold="true" box="[189,334,267,293]" pageId="19" pageNumber="20">Description</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCA5302FF24FEADFE4F0B3E" blockId="19.[189,506,313,340]" box="[189,506,313,340]" pageId="19" pageNumber="20">
BODY LENGTH.
<quantity id="F22D13B5FFCA5302FEFCFEAEFE400B3E" box="[357,501,313,340]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.4" metricValueMin="5.2" pageId="19" pageNumber="20" unit="mm" value="5.8" valueMax="6.4" valueMin="5.2">5.26.4 mm</quantity>
.
</paragraph>
<paragraph id="356ABE50FFCA5302FF24FEECFAE40BB1" blockId="19.[189,1399,379,476]" pageId="19" pageNumber="20">COLOUR. Body integument dark red, head and prothorax darker, usually black; vestiture of elytra with elliptic elongate scales, twice as long as wide, not concealing integument, greyish, forming a homogeneous grey appearance, usually with an ill-defined paler band on apical ⅔ of interstriae 13.</paragraph>
<paragraph id="356ABE50FFCA5302FF24FD94FC7F098C" blockId="19.[189,1399,515,999]" pageId="19" pageNumber="20">HEAD. Rostrum longer than wide (w/l ratio: 0.63) in both sexes, at base as wide as at apex, sides slightly concave near middle of length; epifrons slightly narrower in middle of length than width of eye in dorsal view, at base narrower than distance between eyes, sides subparallel, slightly widening apicad, upper face with a longitudinal carina extending from basal transverse groove to nasal plate, visible through scales; nasal plate V-shaped, reaching level of antennal insertion basally, bare of scales, with only minute setae inserted in scattered punctures and 3 pairs of long setae apically near insertion of mandibles (2 short and 1 very long); antennal scrobes slightly curved in lateral view, directed to middle of eye, separated from it by a strip of scales half as narrow as diameter of eyes; mandibles trisetose, bare of scales. Forehead with a short longitudinal furrow between eyes, scales and setae arranged centripetally towards a point below base of eyes; eyes protruding, margin rounded, surface slightly conical in dorsal view. Antennae slender, scape 0.8× as long as funicle, slightly bisinuate, subclavate at apex, exceeding anterior margin of prothorax in repose; funicle with segments 12 elongate, 1 slightly shorter (0.85 ×) than 2, segments 36 longer than wide, conical, shortening apicad, segment 7 longer and slightly wider than 6; club spindle-shaped, segment 1 longer than 2, margins slightly sinuous.</paragraph>
<paragraph id="356ABE50FFCA5302FF24FB99FAD00EFB" blockId="19.[189,1399,1038,1170]" pageId="19" pageNumber="20">PROTHORAX. Slightly wider than long (w/l ratio: 1.20), widest near middle of length, apical margin 0.6× as wide as basal margin, sides rounded, apical half slightly concave; integument densely punctate, spaces between punctures forming ill-defined, concentric smooth ridges and a small, smooth and shiny median carina visible in approximately 50% of individuals; upper face regularly convex, lacking depressions.</paragraph>
<paragraph id="356ABE50FFCA5302FF24FB2EFBC00F56" blockId="19.[189,1398,1209,1341]" pageId="19" pageNumber="20">ELYTRA. Sides convex, widest near middle of length (w/l ratio: 0.760.83), integument finely punctate and reticulate, shiny, interstria raised, 34 × as wide as width of striae in basal half; large declivital callosities absent or very reduced in apical ¼ of interstriae 3, 5 and 7; each interstria with a series of semi-erect elongate whitish scales, slightly more condensed in spots on declivity.</paragraph>
<paragraph id="356ABE50FFCA5302FF24FAF3FC560C44" blockId="19.[189,1398,1380,1582]" pageId="19" pageNumber="20">
ABDOMEN. Underside with scattered scales similar to those on elytra, not concealing integument; ventrite 1 as long as 2+ 3+4, slightly convex in middle (
<specimenCount id="23D375D9FFCA5302FCEEFA10FC3F0FCB" box="[887,906,1415,1441]" pageId="19" pageNumber="20" type="female"></specimenCount>
), or concave with erect setae as long as claws (♂), ventrite 5 longer than 3 +4, with recumbent elongate setae in middle and near apex and a weak longitudinal carina near apex (
<specimenCount id="23D375D9FFCA5302FDF7FA5AFD340F8D" box="[622,641,1485,1511]" pageId="19" pageNumber="20" type="female"></specimenCount>
) or with a large cavity extending over apical ⅔, with long erect setae mainly condensed on middle of basal margin of cavity and a brush on either side of apical margin; bottom of cavity with a few short recumbent or semi-erect setae (♂).
</paragraph>
<paragraph id="356ABE50FFCA5302FF24F9C1FABD0CF8" blockId="19.[189,1399,1621,1683]" pageId="19" pageNumber="20">LEGS. Femora clavate, unarmed, with a double vestiture of recumbent and semi-erect greyish scales; tibiae straight, unarmed, slightly bisinuate on ventral side; claws simple, free, equal in length.</paragraph>
<paragraph id="356ABE50FFCA5302FF24F92DFE910D0B" blockId="19.[189,1399,1722,1889]" pageId="19" pageNumber="20">
GENITALIA. Body of penis elongate, 0.75 × as long as apodemes, sides almost straight, converging regularly towards apex, more abruptly near apex, widest at base (w/l ratio: 0.40), in lateral view curvature weak, slightly stronger near middle of length (
<figureCitation id="ADEEA2D5FFCA5302FD16F897FD5F0D70" box="[655,746,1792,1818]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[252,1379,330,1800]" captionTargetId="figure@10.[189,1398,308,1806]" captionTargetPageId="10" captionText="Fig. 2. Penis of Phlyctinus Schoenherr, 1826, in dorsal (left) and lateral (right) view, and copulatory sclerite (top right). A. P. callosus (Schoenherr, 1826). B. P. grootbosensis Haran sp. nov. C. P. xerophilus Haran sp. nov., holotype ♂. D. P. planithorax Haran sp. nov., holotype ♂. E. P. littoralis Haran sp. nov., holotype ♂. F. P. aloevorus Haran sp. nov., holotype ♂. Scale bars = 500 µm. Copulatory sclerite not to scale." figureDoi="http://doi.org/10.5281/zenodo.3900302" httpUri="https://zenodo.org/record/3900302/files/figure.png" pageId="19" pageNumber="20">Fig. 2E</figureCitation>
); copulatory sclerite with left arm longer than body, not bearing setae at base; comb of right arm with setae converging; right area of body slightly wider than left area.
</paragraph>
<paragraph id="356ABE50FFCD5305FF24FE9CFE040B4F" blockId="20.[189,433,267,293]" box="[189,433,267,293]" pageId="20" pageNumber="21">
<heading id="6E22093CFFCD5305FF24FE9CFE040B4F" bold="true" box="[189,433,267,293]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="07A16242FFCD5305FF24FE9CFE040B4F" bold="true" box="[189,433,267,293]" pageId="20" pageNumber="21">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCD5305FF24FEAEFD4C0B1D" blockId="20.[189,1398,313,375]" pageId="20" pageNumber="21">
Males may be distinguished externally from females by the slightly more slender elytra (w/l ratio: 0.76,
<quantity id="F22D13B5FFCD5305FF24FECAFEBA0B1D" box="[189,271,349,375]" metricMagnitude="-2" metricUnit="m" metricValue="2.1082" pageId="20" pageNumber="21" unit="in" value="0.83">0.83 in</quantity>
<specimenCount id="23D375D9FFCD5305FE8FFECAFE9F0B1D" box="[278,298,349,375]" pageId="20" pageNumber="21" type="female"></specimenCount>
) and the cavity and setae on ventrite 5.
</paragraph>
<paragraph id="356ABE50FFCD5305FF24FE09FEFA0BD2" blockId="20.[189,335,414,440]" box="[189,335,414,440]" pageId="20" pageNumber="21">
<heading id="6E22093CFFCD5305FF24FE09FEFA0BD2" bold="true" box="[189,335,414,440]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="07A16242FFCD5305FF24FE09FEFA0BD2" bold="true" box="[189,335,414,440]" pageId="20" pageNumber="21">Life history</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCD5305FF24FE5AFD5E0844" blockId="20.[189,1399,461,558]" pageId="20" pageNumber="21">
The type series of
<taxonomicName id="F2D5C5D3FFCD5305FE0CFE5AFDA20B8D" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[405,535,461,487]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="littoralis" status="sp. nov.">
<emphasis id="07A16242FFCD5305FE0CFE5AFDA20B8D" box="[405,535,461,487]" italics="true" pageId="20" pageNumber="21">P. littoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCD5305FD87FE5AFDCC0B8D" box="[542,633,461,487]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
was collected from the base of
<taxonomicName id="F2D5C5D3FFCD5305FC75FE5AFAC20B8D" box="[1004,1399,461,487]" class="Magnoliopsida" family="Asteraceae" genus="Arctotis" kingdom="Plantae" order="Asterales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="angustifolia">
<emphasis id="07A16242FFCD5305FC75FE5AFB6E0B8D" box="[1004,1243,461,487]" italics="true" pageId="20" pageNumber="21">Arctotis angustifolia</emphasis>
(Asteraceae)
</taxonomicName>
plants growing on the gravel parts of Kogel Bay beach (
<figureCitation id="ADEEA2D5FFCD5305FCDBFE67FC320860" box="[834,903,496,522]" captionStart="Fig" captionStartId="18.[189,232,1817,1843]" captionTargetBox="[264,1322,286,2013]" captionTargetId="figure@17.[244,1344,265,2019]" captionTargetPageId="17" captionText="Fig. 4 (opposite page). Habitus in natura, host plants and biotopes of Phlyctinus Schoenherr, 1826 species. A. P. callosus (Schoenherr, 1826). B. P. xerophilus Haran sp. nov. CD. P. grootbosensis Haran sp. nov. and its host plant, Osteospermum moniliferum L. EF. P. planithorax Haran sp. nov. and its host plant, Hymenolepis crithmifolia (L.) Greuter. GH. Biotope and host plant (Arctotis angustifolia L.) of P. littoralis Haran sp. nov. IJ. P. aloevorus Haran sp. nov. on its host plant, Aloe arborescens Mill., with details of feeding damage on floral peduncle ." figureDoi="http://doi.org/10.5281/zenodo.14285227" httpUri="https://zenodo.org/record/14285227/files/figure.png" pageId="20" pageNumber="21">Fig. 4</figureCitation>
GH). Adults were collected between May and September but gravid females only in May.
</paragraph>
</subSubSection>
<subSubSection id="7DCFEDDBFFCD5305FF24FDC3FD4808F7" box="[189,765,596,670]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="356ABE50FFCD5305FF24FDC3FEE30804" blockId="20.[189,342,596,622]" box="[189,342,596,622]" pageId="20" pageNumber="21">
<heading id="6E22093CFFCD5305FF24FDC3FEE30804" bold="true" box="[189,342,596,622]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="07A16242FFCD5305FF24FDC3FEE30804" bold="true" box="[189,342,596,622]" pageId="20" pageNumber="21">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCD5305FF24FD14FD4808F7" blockId="20.[189,765,643,670]" box="[189,765,643,670]" pageId="20" pageNumber="21">
This species is only known from its
<typeStatus id="EA6E00F2FFCD5305FDFFFD13FD2C08F4" box="[614,665,644,670]" pageId="20" pageNumber="21">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="7DCFEDDBFFCD5305FF24FD52FAC209D7" pageId="20" pageNumber="21" type="discussion">
<paragraph id="356ABE50FFCD5305FF24FD52FE9A08B5" blockId="20.[189,303,709,735]" box="[189,303,709,735]" pageId="20" pageNumber="21">
<heading id="6E22093CFFCD5305FF24FD52FE9A08B5" bold="true" box="[189,303,709,735]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="07A16242FFCD5305FF24FD52FE9A08B5" bold="true" box="[189,303,709,735]" pageId="20" pageNumber="21">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="356ABE50FFCD5305FF24FD64FAC209D7" blockId="20.[189,1399,755,958]" pageId="20" pageNumber="21">
<taxonomicName id="F2D5C5D3FFCD5305FF24FD64FE1D0967" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[189,424,755,781]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="littoralis" status="sp. nov.">
<emphasis id="07A16242FFCD5305FF24FD64FE1D0967" box="[189,424,755,781]" italics="true" pageId="20" pageNumber="21">Phlyctinus littoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCD5305FE2AFD63FDA60964" box="[435,531,756,782]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="F2D5C5D3FFCD5305FDBBFD63FD7E0967" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[546,715,755,781]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="planithorax" status="sp. nov.">
<emphasis id="07A16242FFCD5305FDBBFD63FD7E0967" box="[546,715,755,781]" italics="true" pageId="20" pageNumber="21">P. planithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCD5305FD4CFD63FC860964" box="[725,819,756,782]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="F2D5C5D3FFCD5305FCEBFD63FBB30967" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[882,1030,755,781]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aloevorus" status="sp. nov.">
<emphasis id="07A16242FFCD5305FCEBFD63FBB30967" box="[882,1030,755,781]" italics="true" pageId="20" pageNumber="21">P. aloevorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCD5305FB89FD63FBDB0964" box="[1040,1134,756,782]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
form a species group characterized by the presence of only very small or no tubercles on the elytral declivity and short setae on ventrite 1 of the males. Preliminary phylogenetic analysis of
<emphasis id="07A16242FFCD5305FC05FCADFC7A093E" box="[924,975,826,852]" italics="true" pageId="20" pageNumber="21">COI</emphasis>
sequences indicated that they form a distinct genetic cluster (
<figureCitation id="ADEEA2D5FFCD5305FE48FCCBFDA2091D" box="[465,535,860,887]" captionStart="Fig" captionStartId="22.[189,232,1923,1949]" captionTargetBox="[249,1378,546,1865]" captionTargetId="figure@22.[189,1398,514,1882]" captionTargetPageId="22" captionText="Fig. 5. PhyML tree of Phlyctinus Schoenherr, 1826 species inferred from mitochondrial Cytochrome oxidase I (COI, 658 base pairs). Bootstrap support values as obtained from 1000 replicates (*above 80, ** above 95). Specimens codes were shortened for representation: JHAR0 = J and _0101 = -1." figureDoi="http://doi.org/10.5281/zenodo.3900308" httpUri="https://zenodo.org/record/3900308/files/figure.png" pageId="20" pageNumber="21">Fig. 5</figureCitation>
). They have so far only been found in coastal habitats.
<taxonomicName id="F2D5C5D3FFCD5305FB09FCCAFAC2091D" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[1168,1399,861,887]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="littoralis" status="sp. nov.">
<emphasis id="07A16242FFCD5305FB09FCCAFAC2091D" box="[1168,1399,861,887]" italics="true" pageId="20" pageNumber="21">Phlyctinus littoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCD5305FF24FC17FEA209F0" box="[189,279,896,922]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
is most closely related to
<taxonomicName id="F2D5C5D3FFCD5305FDDAFC16FD5209F0" authority="Haran &amp; Hansen &amp; Benoit &amp; Cbgp, 2020" authorityName="Haran &amp; Hansen &amp; Benoit &amp; Cbgp" authorityYear="2020" box="[579,743,896,922]" class="Insecta" family="Curculionidae" genus="Phlyctinus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="planithorax" status="sp. nov.">
<emphasis id="07A16242FFCD5305FDDAFC16FD5209F0" box="[579,743,896,922]" italics="true" pageId="20" pageNumber="21">P. planithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C92DF39FFCD5305FD77FC17FCFF09F0" box="[750,842,896,922]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
, from which it can easily be distinguished by its homogeneous grey vestiture. The two species show an interspecific genetic p-distance on
<emphasis id="07A16242FFCD5305FB40FC34FAB909D7" box="[1241,1292,931,957]" italics="true" pageId="20" pageNumber="21">COI</emphasis>
of 4.6%.
</paragraph>
</subSubSection>
</treatment>
</document>