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<document id="EBEBD56C4E80BC53AEFC2712544ACC00" ID-CLB-Dataset="4641" ID-DOI="10.5281/zenodo.5321656" ID-GBIF-Dataset="eeb8f590-30c6-4f6a-bca2-bf8a06ac6fa0" ID-ISSN="0374-1036" ID-Zenodo-Dep="5321656" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630137224295" checkinUser="felipe" docAuthor="Short, Andrew Edward Z." docDate="2009" docId="E12C8212FFC8FF9976D5FA92FC08FD36" docLanguage="en" docName="ActaEntMusNatPra.2009.49.2.625-630.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Oocyclus madidus Short 2009, sp. nov." docType="treatment" docVersion="7" lastPageNumber="630" masterDocId="1D15FA6AFFCBFF9C7712FFCAFFA5FFE5" masterDocTitle="Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae)" masterLastPageNumber="630" masterPageNumber="625" pageNumber="628" updateTime="1699168334751" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D789EAFFC336C7AE9AB6F6C1ED360006">Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae)</mods:title>
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<mods:namePart id="D26D72D9B61C281E1AAFBDA6AB4412C6">Short, Andrew Edward Z.</mods:namePart>
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<mods:title id="830F7A4231F2DED8C29B6BA58B1E5B3F">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="2061117A2C611AEF4FC0883D5963BDF5">2009</mods:date>
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<mods:number id="B999F4E7706CDF8C5BF501CBAE57A48B">2009-12-15</mods:number>
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<treatment id="E12C8212FFC8FF9976D5FA92FC08FD36" ID-DOI="http://doi.org/10.5281/zenodo.5342792" ID-GBIF-Taxon="185202430" ID-Zenodo-Dep="5342792" LSID="urn:lsid:plazi:treatment:E12C8212FFC8FF9976D5FA92FC08FD36" httpUri="http://treatment.plazi.org/id/E12C8212FFC8FF9976D5FA92FC08FD36" lastPageId="5" lastPageNumber="630" pageId="3" pageNumber="628">
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<paragraph id="693A3304FFC8FF9F76D5FA92FD49FA94" blockId="3.[455,748,1368,1393]" box="[455,748,1368,1393]" pageId="3" pageNumber="628">
<heading id="32728468FFC8FF9F76D5FA92FD49FA94" bold="true" box="[455,748,1368,1393]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="628" reason="3">
<emphasis id="5BF1EF16FFC8FF9F76D5FA92FD49FA94" bold="true" box="[455,748,1368,1393]" pageId="3" pageNumber="628">
<taxonomicName id="AE854887FFC8FF9F76D5FA92FD2AFA94" ID-CoL="49RW5" authority="Short, 2009" authorityName="Short" authorityYear="2009" box="[455,655,1368,1393]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="628" phylum="Arthropoda" rank="species" species="madidus" status="sp. nov.">
<emphasis id="5BF1EF16FFC8FF9F76D5FA92FD2AFA94" bold="true" box="[455,655,1368,1393]" italics="true" pageId="3" pageNumber="628">Oocyclus madidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40C2526DFFC8FF9F7584FA93FD49FA94" box="[662,748,1369,1393]" pageId="3" pageNumber="628" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="219F608FFFC8FF9F7538FABDFD22FA6F" box="[554,647,1399,1418]" pageId="3" pageNumber="628" type="description">
<paragraph id="693A3304FFC8FF9F7538FABDFD22FA6F" blockId="3.[554,647,1399,1418]" box="[554,647,1399,1418]" pageId="3" pageNumber="628">
(
<figureCitation id="F1BE2F81FFC8FF9F7522FABDFDC8FA6F" box="[560,621,1399,1418]" captionStart="Figs" captionStartId="2.[152,191,985,1004]" captionTargetBox="[152,1127,217,955]" captionTargetId="figure-232@2.[152,1129,216,955]" captionTargetPageId="2" captionText="Figs. 12. Dorsal habitus. 1 O. armstrongorum sp. nov., 2 O. madidus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5321660" httpUri="https://zenodo.org/record/5321660/files/figure.png" pageId="3" pageNumber="628">Figs. 2</figureCitation>
,
<figureCitation id="F1BE2F81FFC8FF9F7564FABDFD24FA6F" box="[630,641,1399,1418]" captionStart="Figs" captionStartId="3.[651,692,858,877]" captionTargetBox="[114,605,223,928]" captionTargetId="figure-241@3.[114,605,219,930]" captionTargetPageId="3" captionText="Figs. 34. Aedeagus, dorsal view. 3 Oocyclus armstrongorum sp. nov., 4 O. madidus sp. nov. Scale bar = 0.33 mm." figureDoi="http://doi.org/10.5281/zenodo.5321662" httpUri="https://zenodo.org/record/5321662/files/figure.png" pageId="3" pageNumber="628">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="219F608FFFC8FF9F7763FA6BFCB1F9E1" pageId="3" pageNumber="628" type="materials_examined">
<paragraph id="693A3304FFC8FF9F7763FA6BFCB1F9E1" blockId="3.[113,1090,1440,1541]" pageId="3" pageNumber="628">
<emphasis id="5BF1EF16FFC8FF9F7763FA6BFF51FA51" bold="true" box="[113,244,1440,1460]" pageId="3" pageNumber="628">Type material.</emphasis>
<materialsCitation id="D9ED3959FFC8FF9F77EAFA68FB9BFA0F" ID-GBIF-Occurrence="3339965302" collectionCode="NMPC" collectorName="E Khasi Hills" country="India" elevation="810" latitude="25.216667" location="Meghalaya State" longLatPrecision="1247" longitude="91.65" pageId="3" pageNumber="628" specimenCount="1" stateProvince="Meghalaya" typeStatus="holotype">
<typeStatus id="B63E8DA6FFC8FF9F77EAFA68FEEEFA56" box="[248,331,1442,1461]" pageId="3" pageNumber="628" type="holotype">HOLOTYPE</typeStatus>
: male,
<collectingCountry id="11927394FFC8FF9F7699FA6BFE68FA51" box="[395,461,1441,1460]" name="India" pageId="3" pageNumber="628">INDIA</collectingCountry>
,
<collectingRegion id="AB41FDE6FFC8FF9F76C6FA6BFDC7FA51" box="[468,610,1441,1460]" country="India" name="Meghalaya" pageId="3" pageNumber="628">Meghalaya State</collectingRegion>
(10) /
<collectorName id="C47056D2FFC8FF9F7584FA68FCADFA51" box="[662,776,1441,1461]" pageId="3" pageNumber="628">E Khasi Hills</collectorName>
,
<quantity id="AE7D9EE1FFC8FF9F741DFA6BFCE5FA51" box="[783,832,1441,1460]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="628" unit="km" value="11.0">11km</quantity>
SW Cherra- / punjee, Laitkynsew,
<date id="1D3B15C4FFC8FF9F7789FA76FE81FA2A" box="[155,292,1468,1487]" pageId="3" pageNumber="628" value="2008-04-21" valueMax="2008-04-24" valueMin="2008-04-21">21-24.iv. / 2008</date>
,
<geoCoordinate id="0CB155C3FFC8FF9F763EFA76FED3FA2A" box="[300,374,1468,1487]" degrees="25" direction="north" minutes="13" orientation="latitude" pageId="3" pageNumber="628" precision="925" value="25.216667">25°13N</geoCoordinate>
<geoCoordinate id="0CB155C3FFC8FF9F7669FA76FE61FA2A" box="[379,452,1468,1487]" degrees="91" direction="east" minutes="39" orientation="longitude" pageId="3" pageNumber="628" precision="925" value="91.65">91°39E</geoCoordinate>
,
<quantity id="AE7D9EE1FFC8FF9F76DFFA76FE58FA2A" box="[461,509,1468,1487]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="3" pageNumber="628" unit="m" value="810.0">
<elevation id="E2A8D437FFC8FF9F76DFFA76FE58FA2A" box="[461,509,1468,1487]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="3" pageNumber="628" unit="m" value="810.0">810m</elevation>
</quantity>
/ Fikáček, Podskalská, Šípek lgt. // seepage: wet rocks algae / blue algae/moss ca.
<quantity id="AE7D9EE1FFC8FF9F77E1FA1CFEE5FA0C" box="[243,320,1494,1513]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="3" pageNumber="628" unit="km" value="1.75" valueMax="2.0" valueMin="1.5">1.5-2 km</quantity>
/ via rd. from Cherrapunjee Holid. / Resort in direct. Cherrapunjee, / exposed (
<collectionCode id="0F94ABC1FFC8FF9F74E5FA1DFB9DFA0F" box="[1015,1080,1495,1514]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="628">NMPC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="B63E8DA6FFC8FF9F7763FA38FF6CF9E6" box="[113,201,1522,1541]" pageId="3" pageNumber="628" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="7F83F88DFFC8FF9F77C6FA3BFEB8F9E0" box="[212,285,1521,1541]" pageId="3" pageNumber="628" type="generic">24 spec.</specimenCount>
, same data as
<typeStatus id="B63E8DA6FFC8FF9F7684FA3BFE44F9E1" box="[406,481,1521,1540]" pageId="3" pageNumber="628" type="holotype">holotype</typeStatus>
(
<collectionCode id="0F94ABC1FFC8FF9F76FEFA3BFD8EF9E1" box="[492,555,1521,1540]" pageId="3" pageNumber="628">KSEM</collectionCode>
,
<collectionCode id="0F94ABC1FFC8FF9F7521FA38FDDCF9E0" box="[563,633,1522,1541]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="628" type="Museum">NHMW</collectionCode>
,
<collectionCode id="0F94ABC1FFC8FF9F7593FA3BFD65F9E1" box="[641,704,1521,1540]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="628">NMPC</collectionCode>
,
<collectionCode id="0F94ABC1FFC8FF9F75DAFA3BFCAFF9E1" box="[712,778,1521,1540]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="628" type="Museum">USNM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="219F608FFFC8FF987763F9D6FCB1FD31" lastPageId="4" lastPageNumber="629" pageId="3" pageNumber="628" type="diagnosis">
<paragraph id="693A3304FFC8FF987763F9D6FCB1FD31" blockId="3.[113,1089,1564,1621]" lastBlockId="4.[151,1127,220,1620]" lastPageId="4" lastPageNumber="629" pageId="3" pageNumber="628">
<emphasis id="5BF1EF16FFC8FF9F7763F9D6FF43F9D1" bold="true" box="[113,230,1564,1588]" pageId="3" pageNumber="628">Diagnosis.</emphasis>
Posterolateral corners of pronotum evenly rounded. Procoxae without distinct spines. Abdominal ventrites finely and evenly pubescent. Elytra without detectable rows of systematic punctures. This species is externally very similar to
<taxonomicName id="AE854887FFCFFF987459FF17FBC3FF11" authority="SATO, 1979" authorityName="SATO" authorityYear="1979" box="[843,1126,220,244]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="bhutanicus">
<emphasis id="5BF1EF16FFCFFF987459FF17FC40FF11" box="[843,997,221,244]" italics="true" pageId="4" pageNumber="629">O. bhutanicus</emphasis>
<bibRefCitation id="0D144EF5FFCFFF9874FFFF16FBC3FF11" author="SATO M." box="[1005,1126,220,244]" pageId="4" pageNumber="629" pagination="43 - 67" refId="ref2890" refString="SATO M. 1979: Ergebnisse der Bhutan-Expedition 1972 und Indien-Nepal-Expeditionen 1975 - 1977 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Georissidae, Hydraenidae, Hydrophilidae und Ptilodactylidae. Entomologica Basiliensia 4: 43 - 67." type="journal article" year="1979">SATÔ, 1979</bibRefCitation>
</taxonomicName>
but differs in several subtle characters: although the dorsal punctation on the elytra of both species consists of punctures of mixed sizes, in
<taxonomicName id="AE854887FFCFFF98758BFED7FCB2FED1" authority="Short, 2009" authorityName="Short" authorityYear="2009" box="[665,791,285,308]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="madidus" status="sp. nov.">
<emphasis id="5BF1EF16FFCFFF98758BFED7FCB2FED1" box="[665,791,285,308]" italics="true" pageId="4" pageNumber="629">O. madidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40C2526DFFCFFF98740EFED6FCD5FED6" box="[796,880,284,307]" pageId="4" pageNumber="629" rank="species">sp. nov.</taxonomicNameLabel>
the smallest punctures are much finer and smaller than the smallest punctures seen on
<taxonomicName id="AE854887FFCFFF987452FEF7FC45FEB1" box="[832,992,317,340]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="bhutanticus">
<emphasis id="5BF1EF16FFCFFF987452FEF7FC45FEB1" box="[832,992,317,340]" italics="true" pageId="4" pageNumber="629">O. bhutanticus</emphasis>
</taxonomicName>
. The largest punctures on both species are relatively similar in size. Consequently, the size range of the punctures observed in
<taxonomicName id="AE854887FFCFFF9876AFFEB7FD98FE71" authority="Short, 2009" authorityName="Short" authorityYear="2009" box="[445,573,381,404]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="madidus" status="sp. nov.">
<emphasis id="5BF1EF16FFCFFF9876AFFEB7FD98FE71" box="[445,573,381,404]" italics="true" pageId="4" pageNumber="629">O. madidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40C2526DFFCFFF987557FEB6FD39FE76" box="[581,668,380,403]" pageId="4" pageNumber="629" rank="species">sp. nov.</taxonomicNameLabel>
is more extensive. In addition, the metafemora in
<taxonomicName id="AE854887FFCFFF987615FE57FE20FE51" authority="Short, 2009" authorityName="Short" authorityYear="2009" box="[263,389,413,436]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="madidus" status="sp. nov.">
<emphasis id="5BF1EF16FFCFFF987615FE57FE20FE51" box="[263,389,413,436]" italics="true" pageId="4" pageNumber="629">O. madidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40C2526DFFCFFF98769EFE56FE45FE56" box="[396,480,412,435]" pageId="4" pageNumber="629" rank="species">sp. nov.</taxonomicNameLabel>
are slightly broader (ca. 2.1× as long as wide, compared to ca. 2.3× for
<taxonomicName id="AE854887FFCFFF98760EFE77FE13FE31" authorityName="SATO" authorityYear="1979" box="[284,438,445,468]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="bhutanicus">
<emphasis id="5BF1EF16FFCFFF98760EFE77FE13FE31" box="[284,438,445,468]" italics="true" pageId="4" pageNumber="629">O. bhutanicus</emphasis>
</taxonomicName>
). Additionally, the anteromedial tooth of the prosternal carina, while still small in
<taxonomicName id="AE854887FFCFFF98767DFE17FE4AFE11" authority="Short, 2009" authorityName="Short" authorityYear="2009" box="[367,495,477,500]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="madidus" status="sp. nov.">
<emphasis id="5BF1EF16FFCFFF98767DFE17FE4AFE11" box="[367,495,477,500]" italics="true" pageId="4" pageNumber="629">O. madidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40C2526DFFCFFF9876E5FE16FDF5FE16" box="[503,592,476,499]" pageId="4" pageNumber="629" rank="species">sp. nov.</taxonomicNameLabel>
, is larger than in the
<typeStatus id="B63E8DA6FFCFFF98742FFE16FC38FE11" box="[829,925,476,500]" pageId="4" pageNumber="629" type="holotype">holotype</typeStatus>
of
<taxonomicName id="AE854887FFCFFF9874D6FE17FBFAFE11" authorityName="SATO" authorityYear="1979" box="[964,1119,477,500]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="bhutanicus">
<emphasis id="5BF1EF16FFCFFF9874D6FE17FBFAFE11" box="[964,1119,477,500]" italics="true" pageId="4" pageNumber="629">O. bhutanicus</emphasis>
</taxonomicName>
. Unfortunately, the aedeagus of the
<typeStatus id="B63E8DA6FFCFFF987509FE36FDDEFDF1" box="[539,635,508,532]" pageId="4" pageNumber="629" type="holotype">holotype</typeStatus>
of
<taxonomicName id="AE854887FFCFFF9875B3FE37FC99FDF1" authorityName="SATO" authorityYear="1979" box="[673,828,509,532]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="bhutanicus">
<emphasis id="5BF1EF16FFCFFF9875B3FE37FC99FDF1" box="[673,828,509,532]" italics="true" pageId="4" pageNumber="629">O. bhutanicus</emphasis>
</taxonomicName>
has been missing for more than 25 years (see
<bibRefCitation id="0D144EF5FFCFFF98764CFDD6FDF9FDD6" author="SHORT A. E. Z. &amp; SWANSON A. P." box="[350,604,539,564]" pageId="4" pageNumber="629" pagination="1 - 24" refId="ref2999" refString="SHORT A. E. Z. &amp; SWANSON A. P. 2005. A revision of the Oocyclus Sharp of Thailand with description of six new species (Coleoptera: Hydrophilidae). Zootaxa 1078: 1 - 24." type="journal article" year="2005">SHORT &amp; SWANSON 2005</bibRefCitation>
for discussion) and so I was not able to compare these structures directly; based on the figure by
<bibRefCitation id="0D144EF5FFCFFF9875B7FDF6FC80FDB1" author="SATO M." box="[677,805,572,596]" pageId="4" pageNumber="629" pagination="43 - 67" refId="ref2890" refString="SATO M. 1979: Ergebnisse der Bhutan-Expedition 1972 und Indien-Nepal-Expeditionen 1975 - 1977 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Georissidae, Hydraenidae, Hydrophilidae und Ptilodactylidae. Entomologica Basiliensia 4: 43 - 67." type="journal article" year="1979">SATÔ (1979</bibRefCitation>
: Fig. 16), the inner margin of the parameres are sinuate in that species (straight in
<taxonomicName id="AE854887FFCFFF9875CAFD97FCF2FD91" authorityName="Short" authorityYear="2009" box="[728,855,605,628]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="madidus">
<emphasis id="5BF1EF16FFCFFF9875CAFD97FCF2FD91" box="[728,855,605,628]" italics="true" pageId="4" pageNumber="629">O. madidus</emphasis>
</taxonomicName>
). SATÔ s (1979) drawing is not symmetrical, with the left paramere indicating a small apical constriction and the right evenly rounded; thus, I cannot determine which is correct for comparison with
<taxonomicName id="AE854887FFCFFF98735EFD57FF51FD31" authority="Short, 2009" authorityName="Short" authorityYear="2009" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="madidus" status="sp. nov.">
<emphasis id="5BF1EF16FFCFFF98735EFD57FF51FD31" italics="true" pageId="4" pageNumber="629">O. madidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40C2526DFFCFFF9877EEFD76FEF1FD36" box="[252,340,700,723]" pageId="4" pageNumber="629" rank="species">sp. nov.</taxonomicNameLabel>
, which is closer to the former condition.
</paragraph>
</subSubSection>
<subSubSection id="219F608FFFCFFF997785FD11FEC1FDB6" lastPageId="5" lastPageNumber="630" pageId="4" pageNumber="629" type="description">
<paragraph id="693A3304FFCFFF987785FD11FE3FFCF1" blockId="4.[151,1127,220,1620]" pageId="4" pageNumber="629">
<emphasis id="5BF1EF16FFCFFF987785FD11FE87FD16" bold="true" box="[151,290,731,755]" pageId="4" pageNumber="629">Description.</emphasis>
Size and form. Total body length =
<quantity id="AE7D9EE1FFCFFF9875A8FD16FCE4FD16" box="[698,833,732,756]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="4" pageNumber="629" unit="mm" value="3.95" valueMax="4.2" valueMin="3.7">3.74.2 mm</quantity>
. Oval, moderately convex. Elytra longer than wide.
</paragraph>
<paragraph id="693A3304FFCFFF9877A7FCD6FD18FC51" blockId="4.[151,1127,220,1620]" pageId="4" pageNumber="629">Color. Dorsum black; head, pronotum, and elytra with distinct greenish sheen. Maxillary and labial palps yellow; apex of maxillary palps dark brown. Mentum and stipes light to reddish brown, distinctly paler than ventral face of head. Legs, epipleura, lateral margins of prosternum, and posterior margin of each ventrite yellow to light brown, with remainder of venter slightly to moderately darker reddish brown.</paragraph>
<paragraph id="693A3304FFCFFF9877A7FC76FE6DFAB6" blockId="4.[151,1127,220,1620]" pageId="4" pageNumber="629">Head. General punctation on labrum, clypeus and frons slightly variable in size from very fine to moderately fine; densely distributed, distance between punctures 1.01.5× the width of one puncture. Systematic row of punctures on labrum moderately dense, forming a nearly continuous lateral row set with moderately long setae, but each puncture still fairly distinguishable, not forming a groove. Frons with an irregular row of systematic punctures mesad of each eye, bearing fine erect setae. Clypeus with a few very indistinct systematic punctures along anterolateral margins, slightly larger than surrounding punctation and bearing short setae. Antennae with scape subequal in length to segments 25; first two segments of club subequal in length and apical segment subequal in length to the two preceding segments combined. Maxillary palps short, about as long as width of labrum; segment 2 slightly bulbous, apical segment slightly longer than penultimate. Labial palps one-half to three-fourths as long as width of mentum. Mentum quadrate, anterior margin slightly convex, bearing very fine and scattered punctures.</paragraph>
<paragraph id="693A3304FFCFFF9977A7FA96FCA3FE36" blockId="4.[151,1127,220,1620]" lastBlockId="5.[113,1089,219,724]" lastPageId="5" lastPageNumber="630" pageId="4" pageNumber="629">Thorax. General punctation on pronotum and elytra composed of extremely fine and coarse punctures, evenly mixed and distributed. Pronotal systematic punctures present, ca. 1.5× the size of general punctation and set with a fine seta, sometimes partially blending with larger general punctures; anterior and posterior series each forming an irregular row. Lateral margins of pronotum set with a few sparsely distributed setiferous punctures; with punctures very small as to almost appear absent. Posterolateral corners of pronotum evenly rounded. Sutural punctation on elytra absent or unmodified from general punctation. Elytra without distinct rows of larger punctures, as they blend uniformly with general punctation; some of these rows (of systematic punctures) can be detected by the presence of fine, short setae. Prosternum with median carina along entire length, with small anteromedial tooth; without long spines or hairs anteriorly. Process of the mesoventrite with lateral extensions sloping evenly downward; apex set with a few long fine setae. Metaventrite with small oval glabrous area posteromedially, slightly longer than wide, length of glabrous area about half the total length of metaventrite. Pro- and mesocoxae densely pubescent; without distinct spines. Protibiae with 10 spines on dorsal face. Protarsal segments 14 small, subequal in length; apical segment ca. as long as segments 14 combined.</paragraph>
<paragraph id="693A3304FFCEFF997782FE11FEC1FDB6" blockId="5.[113,1089,219,724]" pageId="5" pageNumber="630">
Abdomen. Ventrites 15 with uniform pubescence, longest setae about as long as the setae around the metasternal glabrous area. Fifth ventrite entire. Aedeagus with parameres slightly attenuated apically; inner margins straight. Median lobe (
<figureCitation id="F1BE2F81FFCEFF99742FFDD1FCDAFDD1" box="[829,895,539,564]" captionStart="Figs" captionStartId="3.[651,692,858,877]" captionTargetBox="[114,605,223,928]" captionTargetId="figure-241@3.[114,605,219,930]" captionTargetPageId="3" captionText="Figs. 34. Aedeagus, dorsal view. 3 Oocyclus armstrongorum sp. nov., 4 O. madidus sp. nov. Scale bar = 0.33 mm." figureDoi="http://doi.org/10.5281/zenodo.5321662" httpUri="https://zenodo.org/record/5321662/files/figure.png" pageId="5" pageNumber="630">Fig. 4</figureCitation>
) not reaching the apex of the parameres.
</paragraph>
</subSubSection>
<subSubSection id="219F608FFFCEFF997763FD96FF48FD76" pageId="5" pageNumber="630" type="etymology">
<paragraph id="693A3304FFCEFF997763FD96FF48FD76" blockId="5.[113,1089,219,724]" pageId="5" pageNumber="630">
<emphasis id="5BF1EF16FFCEFF997763FD96FF57FD91" bold="true" box="[113,242,604,628]" pageId="5" pageNumber="630">Etymology.</emphasis>
<taxonomicName id="AE854887FFCEFF9977E8FD97FE04FD96" authority=", Latin" authorityName="Latin" box="[250,417,603,628]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="630" phylum="Arthropoda" rank="species" species="madidus">
<emphasis id="5BF1EF16FFCEFF9977E8FD97FEF2FD91" box="[250,343,605,628]" italics="true" pageId="5" pageNumber="630">Madidus</emphasis>
, Latin
</taxonomicName>
, meaning soaked or drenched, referring to the wet habitats of the species.
</paragraph>
</subSubSection>
<subSubSection id="219F608FFFCEFF997763FD51FB92FD56" box="[113,1079,667,692]" pageId="5" pageNumber="630" type="biology_ecology">
<paragraph id="693A3304FFCEFF997763FD51FB92FD56" blockId="5.[113,1089,219,724]" box="[113,1079,667,692]" pageId="5" pageNumber="630">
<emphasis id="5BF1EF16FFCEFF997763FD51FF68FD56" bold="true" box="[113,205,667,691]" pageId="5" pageNumber="630">Biology.</emphasis>
Collected with and presumed identical to
<taxonomicName id="AE854887FFCEFF99759DFD57FCF3FD51" authority="Short, 2009" authorityName="Short" authorityYear="2009" box="[655,854,669,692]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="630" phylum="Arthropoda" rank="species" species="armstrongorum" status="sp. nov.">
<emphasis id="5BF1EF16FFCEFF99759DFD57FCF3FD51" box="[655,854,669,692]" italics="true" pageId="5" pageNumber="630">O. armstrongorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40C2526DFFCEFF99744FFD56FC15FD56" box="[861,944,668,691]" pageId="5" pageNumber="630" rank="species">sp. nov.</taxonomicNameLabel>
(see above).
</paragraph>
</subSubSection>
<subSubSection id="219F608FFFCEFF997763FD71FC08FD36" box="[113,941,699,724]" pageId="5" pageNumber="630" type="distribution">
<paragraph id="693A3304FFCEFF997763FD71FC08FD36" blockId="5.[113,1089,219,724]" box="[113,941,699,724]" pageId="5" pageNumber="630">
<emphasis id="5BF1EF16FFCEFF997763FD71FEA1FD36" bold="true" box="[113,260,699,723]" pageId="5" pageNumber="630">Distribution.</emphasis>
Known only from the
<typeStatus id="B63E8DA6FFCEFF9976E5FD76FD80FD31" box="[503,549,700,724]" pageId="5" pageNumber="630">type</typeStatus>
locality in
<collectingRegion id="AB41FDE6FFCEFF99758CFD71FCE9FD36" box="[670,844,699,723]" country="India" name="Meghalaya" pageId="5" pageNumber="630">Meghalaya state</collectingRegion>
of
<collectingCountry id="11927394FFCEFF99747DFD71FC0CFD36" box="[879,937,699,723]" name="India" pageId="5" pageNumber="630">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>