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<mods:title id="9B551E6B2E250D1366DC62EFF374A123">The genus Cephaloleia Chevrolat, 1836 (Coleoptera, Chrysomelidae, Cassidinae)</mods:title>
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<mods:namePart id="63843AA583CDC24C8B6A18CBD7B69175">Staines, Charles L.</mods:namePart>
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<mods:namePart id="0CF4B0296CD83DD2ADB6C303DE8B49A8">Garcia-Robledo, Carlos</mods:namePart>
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<mods:identifier id="416E14B9941D36C0A8E00D4536ED7D7C" type="DOI">http://dx.doi.org/10.3897/zookeys.436.5766</mods:identifier>
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<treatment id="29CE1C25FF83B8EF15C88B173322A004" ID-GBIF-Taxon="152055348" LSID="urn:lsid:plazi:treatment:29CE1C25FF83B8EF15C88B173322A004" httpUri="http://treatment.plazi.org/id/29CE1C25FF83B8EF15C88B173322A004" lastPageNumber="60" pageId="59" pageNumber="60">
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<paragraph id="11CF719BE7BEE7EFCC15FB8A9FBCFA6E" pageId="59" pageNumber="60">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
</subSubSection>
<subSubSection id="6E3D782B6B95F201C5B5E98DFB70CE1E" pageId="59" pageNumber="60" type="nomenclature">
<paragraph id="96DB9EF2658B3B9C5BED14C23668EBDB" pageId="59" pageNumber="60">
<taxonomicName id="031072B85E4705118708391A90929E93" ID-CoL="SDBL" LSID="http://species-id.net/wiki/Cephaloleia_antennalis" authority="Donckier, 1899" authorityName="Donckier" authorityYear="1899" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia antennalis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="antennalis">Cephaloleia antennalis Donckier, 1899</taxonomicName>
Fig. 72
</paragraph>
</subSubSection>
<subSubSection id="0B3846FDAD830BA8D1E5C92F4806B44E" pageId="59" pageNumber="60" type="reference_group">
<paragraph id="870DF7836714C3FAE93E7C468F1BB09B" pageId="59" pageNumber="60">
<taxonomicName id="0A92A634FAF3E91DF110DA633EA989AC" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia antennata" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="antennata">Cephaloleia antennata</taxonomicName>
<bibRefCitation id="C48F41D9594CA9E2C7ECEE81462A243D" author="Baly, JS" editor="Godman, FD" journalOrPublisher="London" pageId="284" pageNumber="285" title="Biologia CentraliAmericana, Zoology, Insecta, Coleoptera, Phytophaga" year="1885">Baly 1885</bibRefCitation>
: 11 (homonym of
<taxonomicName id="8A9294BE011E6BB596ADBFDBFCE32156" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia antennata" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="antennata">Cephaloleia antennata</taxonomicName>
Waterhouse, 1881).
<bibRefCitation id="5F8019003D88A1255ABEF184A483AAC3" author="Staines, CL" journalOrPublisher="Beitraege zur Entomologie" pageId="293" pageNumber="294" pagination="489 - 530" title="Joseph Sugar Baly: The man and his entomological works." volume="49" year="1999">Staines and Staines 1999</bibRefCitation>
: 523 (Baly species list).
</paragraph>
<paragraph id="FED8C2B27EFC9F8AD79CBE5B123A8D20" pageId="59" pageNumber="60">
<taxonomicName id="01AADCD9B57060B824850E4A03392C7A" class="Insecta" family="Chrysomelidae" genus="Cephalolia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cephalolia antennalis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="antennalis">Cephalolia antennalis</taxonomicName>
<bibRefCitation id="72495635D0805DCC4B60A1DCE1B5207E" author="Donckier, H" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="286" pageNumber="287" pagination="540 - 615" title="Catalogue systematique des Hispides." volume="68" year="1899">Donckier 1899</bibRefCitation>
: 547 (replacement name for
<taxonomicName id="E3A61AE5CA80F005ADDA912CFF528548" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia antennata" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="antennata">Cephaloleia antennata</taxonomicName>
Baly).
<bibRefCitation id="616457D9E388E4BA25AD9B1E8A064DB8" author="Weise, J" journalOrPublisher="W. Junk" pageId="297" pageNumber="298" pagination="1 - 94" title="Coleopterorum Catalogus, Chrysomelidae: Hispinae" year="1911 a">Weise 1911a</bibRefCitation>
: 7 (catalog),
<bibRefCitation id="CDA5C51EF1713161152C01F8C0742468" author="Weise, J" editor="Wytsman, P" journalOrPublisher="Genera Insectorum, Brussels" pageId="297" pageNumber="298" pagination="1 - 123" title="Coleoptera Phytophaga fam. Chrysomelidae, subfam. Hispinae." volume="fasc. 125" year="1911 b">1911b</bibRefCitation>
: 10 (catalog).
</paragraph>
<paragraph id="818253F7E9C3F4A1DCEFF2462AB3A82B" pageId="59" pageNumber="60">
<taxonomicName id="453DDDD00AA1318A6BC6B111DC8F1125" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia antennalis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="antennalis">Cephaloleia antennalis</taxonomicName>
Donckier.
<bibRefCitation id="F526B104479AF1D0DDD40E76063E8B7B" author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="284" pageNumber="285" pagination="551 - 763" title="Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1946">Blackwelder 1946</bibRefCitation>
: 718 (catalog);
<bibRefCitation id="68321873DF80BC225FDED9825884D48F" author="Papp, CS" journalOrPublisher="Portugaliae Acta Biologica (B)" pageId="291" pageNumber="292" pagination="1 - 147" title="The Hispinae of America. 3 rd Contribution for promoting the scientifical results of the International Hylean Amazon Institute in Manaos, Brazil." volume="4" year="1953">Papp 1953</bibRefCitation>
: 13 (catalog);
<bibRefCitation id="6D473AF1401E8B96CBD7526B0153E7D5" author="Uhmann, E" journalOrPublisher="W. Junk, Gravenhage" pageId="296" pageNumber="297" pagination="1 - 153" title="Coleopterorum Catalogus. Supplementa. Chrysomelidae: Hispinae, Hispinae Americanae" year="1957 a">Uhmann 1957a</bibRefCitation>
: 15 (catalog);
<bibRefCitation id="F43DEF879CF9B525B398D81C45D5C83B" author="Wilcox, JA" journalOrPublisher="Biological Research Institute of America, Latham, NY" pageId="297" pageNumber="298" title="Checklist of the beetles of Canada, United States, Mexico, Central America and the West Indies" year="1983">Wilcox 1983</bibRefCitation>
: 136 (catalog);
<bibRefCitation id="B7956384DDEE683BCECD4E11FEF2B324" pageId="59" pageNumber="60">Staines 1996</bibRefCitation>
: 16 (Central America species);
<bibRefCitation id="FBDCAF6ACADAB7630117F78E7D4749FA" author="McKenna, DD" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="290" pageNumber="291" pagination="117 - 131" title="Molecular phylogenetics and evolution of host plant use in the Neotropical rolled leaf ' hispine' beetle genus Cephaloleia (Chevrolat) (Coleoptera: Cassidinae)." url="10.1016/j.ympev.2005.06.011" volume="37" year="2005">McKenna and Farrell 2005</bibRefCitation>
: 119 (phylogeny),
<bibRefCitation id="39CBB7AC992455FA6D68C52299937F4D" author="McKenna, DD" journalOrPublisher="Proceedings of the National Academy of Sciences" pageId="290" pageNumber="291" pagination="10947 - 10951" title="Tropical forests are both evolutionary cradles and museums of leaf beetle diversity." url="10.1073/pnas.0602712103" volume="103" year="2006">2006</bibRefCitation>
: 10949 (phylogeny).
</paragraph>
</subSubSection>
<subSubSection id="C816F639404378004B835182F6935491" pageId="59" pageNumber="60" type="description">
<paragraph id="D9F009E53A3B80F974C70ABCB958B566" pageId="59" pageNumber="60">Description.</paragraph>
<paragraph id="674728A195096EE5DD1444B43D0CFD78" pageId="59" pageNumber="60">
Elongate; subdepressed; shining; reddish-brown; antennomeres 1-4 yellow, 5-11 darker; eyes darker. Head: vertex sparsely punctate, medial sulcus present; frons not projecting; not depressed between eyes. Antenna: reaches to humerus; slender; antennomeres 1-4 laterally compressed, with inner anterior angle produced into acute tooth in male, elongate in female; 1 large, clavate; 2-4 subequal in length, each shorter than 1; 5-10 transverse, decreasing in length, each shorter than 4; 11 pointed at apex, subequal to 1; 1-2 punctate with scattered setae; 3-11 setose. Pronotum: transverse; lateral margin straight for basal
<normalizedToken id="D84C0212CAC446142F8F9B2601D0801D" originalValue="¾">3/4</normalizedToken>
then convergent, margined; anterior angle produced, obtuse; posterior angle angulate; anterior margin emarginate behind head; surface finely, sparsely punctate; basal impression absent; pronotal length 1.3-1.8 mm; pronotal width 2.0-2.6 mm. Scutellum: triangular; impunctate. Elytron: lateral margin straight, smooth, margined; apex rounded; sutural angle without tooth; humerus rounded, not produced; slightly constricted behind humerus; weakly punctate-striate; elytral length 5.1-6.4 mm; elytral width 2.7-3.3 mm. Venter: pro-, meso- and metasterna punctate; abdominal sterna punctate, each puncture with pale seta; suture between sterna 1 and 2 complete; last sternite with apical margin truncate in female, emarginate medially in male. Leg: femur and tibia punctate, each puncture with pale seta; femur robust; tibia expanded to apex, with fringe of setae on inner margin of apex. Total length: 6.9-8.4 mm.
</paragraph>
</subSubSection>
<subSubSection id="D3A9D1887A002E486CFD32008731F5F5" pageId="59" pageNumber="60" type="diagnosis">
<paragraph id="68EC5BC090254444F21B000D814B5791" pageId="59" pageNumber="60">Diagnosis.</paragraph>
<paragraph id="FA7D3F2DDE747BD41A9E93A63D091E26" pageId="59" pageNumber="60">This is one of the reddish-brown immaculate species. It can be distinguished from the other reddish-brown species by the suture between abdominal sterna 1 and 2 being complete and by antennomere 1 being as long as 2 to 4 combined.</paragraph>
</subSubSection>
<subSubSection id="466A33FC131E8309B10FA2BEC7D73AE2" pageId="59" pageNumber="60" type="distribution">
<paragraph id="E92285FF54BCA5698D45289C52EF7208" pageId="59" pageNumber="60">Distribution.</paragraph>
<paragraph id="F8DC38DE02711059EF937157B4D2714D" pageId="59" pageNumber="60">Costa Rica, Guatemala, Panama.</paragraph>
</subSubSection>
<subSubSection id="9DC7DA1B08074EA43B20399F89416BC2" pageId="59" pageNumber="60" type="type material examined">
<paragraph id="3BD81DEBAAFAE80B44829FE5749FC850" pageId="59" pageNumber="60">Type material examined.</paragraph>
<paragraph id="035EA3E4F57B51F4A97BE3D14FD3AFD3" pageId="59" pageNumber="60">
Holotype male: Type H. T. [white disk with red border]/ Panama, Volcan de
<normalizedToken id="B38F4F550B68447D6267D38CBFFA78B1" originalValue="Chiriquí">Chiriqui</normalizedToken>
, Champion/ B. C. A., Col. VI, 2. Cephaloleia antennata, Baly/ Cephaloleia/ Cephalolia antennata Baly, Nicaragua [blue handwritten label] (BMNH).
</paragraph>
</subSubSection>
<subSubSection id="FD3C807E24629E22E6FB310A419087BA" pageId="59" pageNumber="60" type="specimens examined">
<paragraph id="E7E46865BD1D948E2248F2580A1EA649" pageId="59" pageNumber="60">Specimens examined.</paragraph>
<paragraph id="79F89AA530862546DA676A16068ECD65" pageId="59" pageNumber="60">
COSTA RICA: Alajuela- Res. For. San
<normalizedToken id="B969A73AC59421118890E536588799C3" originalValue="Ramón">Ramon</normalizedToken>
, 900 m, 12 March 1990 (INBIO). Cartago- Quebrada Segunda, Ref. Nac. Fauna Silv.
<normalizedToken id="317296D9C2CDCA310F4158C446D349E1" originalValue="Tapantí">Tapanti</normalizedToken>
, 1250 m, April 1992 (INBIO); Turrialba (USNM). Guanacaste-
<normalizedToken id="96CE567BD340B5EA434607ADD7173577" originalValue="Río">Rio</normalizedToken>
San Lorenzo, 1050 m, Tierras Morenas, Z.P. Tenorio, 28
<normalizedToken id="73D0DB4150F5C10F1538A57F1F38BDA8" originalValue="March">March-</normalizedToken>
21 April 1992, July 1991, 10-20 February 1992, October 1992 (INBIO). Heredia- La Selva, 18 May 1993 (SEMC).
<normalizedToken id="8E6301A4C681C6D73F7B7B4B3C86C7FE" originalValue="Limón-">Limon-</normalizedToken>
Sector Cerro
<normalizedToken id="218E5508A8E7D1F68610C91D9EB635AB" originalValue="Cocorí">Cocori</normalizedToken>
, Fca. de E. Rojas, 150 m, 28
<normalizedToken id="B9AA59D7F86D17EA5B20BF52DABBD3C4" originalValue="May">May-</normalizedToken>
17 June 1992 (INBIO); Esta. Cuatro Esquinas, 0 m, P. N. Tortuguero, 27
<normalizedToken id="B23C60D74FD455AE226FA392498A3CFC" originalValue="March">March-</normalizedToken>
29 April 1992 (INBIO); Hamburg Farm,
<normalizedToken id="C437F9CB7BF8064CC7A726F64D9C5F01" originalValue="Reventazón">Reventazon</normalizedToken>
, Ebene
<normalizedToken id="81789E320D0306DDEB923A17B7815265" originalValue="Limón">Limon</normalizedToken>
, 27 April 1934, 23 February 1934 (USNM). Puntarenas- Est. Biol. Las Alturas, 1500 m, Coto Brus, 23 March- 2 May 1992 (INBIO); Golfito, 10-200 m, 28 May 1993 (SEMC); Golfito, 3 July 1985 (EMEC); Rancho Quemado, 200 m, Peninsula de Osa, April 1992, 21
<normalizedToken id="E0C9BA6982E5B8F917122EF596EEC583" originalValue="March">March-</normalizedToken>
7 April 1992 (INBIO); Est. Sirena, P.N. Corcovado, September 1991, November 1989, February 1990, 21 March- 21 April 1992, 28
<normalizedToken id="C28CDE4FA0CEDBFB2BBD2652DD5BF811" originalValue="May">May-</normalizedToken>
17 June 1992 (INBIO). GUATEMALA: Verapaz- Chacoj (USNM). Total: 46.
</paragraph>
</subSubSection>
</treatment>
</document>