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<document ID-DOI="http://dx.doi.org/10.3897/zse.97.60254" ID-GBIF-Dataset="3a0c3603-6f6e-4e87-bc22-a6dddb9686d9" ID-GBIF-Taxon="182236426" ID-Pensoft-Pub="1860-0743-1-111" ID-Pensoft-UUID="3C29E619B8CF57C6B036DBC8DA393395" ID-ZooBank="FE7CB4589459409CB254DA0A1BA65B86" LSID="urn:lsid:zoobank.org:act:4F1D8C21-1EEF-4248-8299-012692C20D79" ModsDocID="1860-0743-97-1-111" checkinTime="1621423269654" checkinUser="pensoft" docAuthor="Radea, Canella, Lampri, Paraskevi Niki, Bakolitsas, Konstantinos &amp; Parmakelis, Aristeidis" docDate="2021" docId="FC52C90716CC53C38428EC62536F8885" docLanguage="en" docName="ZoosystEvol 97(1): 111-119" docOrigin="Zoosystematics and Evolution 97 (1)" docPubDate="2021-02-05" docSource="http://dx.doi.org/10.3897/zse.97.60254" docTitle="Daphniola dione Radea &amp; Lampri &amp; Bakolitsas &amp; Parmakelis 2021, sp. nov." docType="treatment" docUuid="4F1D8C21-1EEF-4248-8299-012692C20D79" docUuidSource="ZooBank" docVersion="3" id="3C29E619B8CF57C6B036DBC8DA393395" lastPageNumber="111" masterDocId="3C29E619B8CF57C6B036DBC8DA393395" masterDocTitle="A new hydrobiid species (Caenogastropoda, Truncatelloidea) from insular Greece" masterLastPageNumber="119" masterPageNumber="111" pageId="0" pageNumber="111" updateTime="1643427679679" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new hydrobiid species (Caenogastropoda, Truncatelloidea) from insular Greece</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Radea, Canella</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1592-7306</mods:nameIdentifier>
<mods:affiliation>Section of Ecology and Systematics, Department of Biology, National and Kapodistrian University of Athens, 15784 Panepistimiopolis, Greece</mods:affiliation>
<mods:nameIdentifier type="email">kradea@biol.uoa.gr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lampri, Paraskevi Niki</mods:namePart>
<mods:affiliation>Section of Ecology and Systematics, Department of Biology, National and Kapodistrian University of Athens, 15784 Panepistimiopolis, Greece &amp; Institute of Marine Biological Resources and Inland Waters, Hellenic Centre for Marine Research, 46.7 km of Athens - Sounio ave., 19013 Anavissos Attica, Greece</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bakolitsas, Konstantinos</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3137-4024</mods:nameIdentifier>
<mods:affiliation>High School, Agrinion, 3 rd Parodos Kolokotroni 11, 30133 Agrinion, Greece</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Parmakelis, Aristeidis</mods:namePart>
<mods:affiliation>Section of Ecology and Systematics, Department of Biology, National and Kapodistrian University of Athens, 15784 Panepistimiopolis, Greece</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoosystematics and Evolution</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>97</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>111</mods:start>
<mods:end>119</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zse.97.60254</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.60254</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-1-111</mods:identifier>
<mods:identifier type="ZooBank">FE7CB4589459409CB254DA0A1BA65B86</mods:identifier>
<mods:identifier type="Pensoft-UUID">3C29E619B8CF57C6B036DBC8DA393395</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182236426" LSID="urn:lsid:zoobank.org:act:4F1D8C21-1EEF-4248-8299-012692C20D79" httpUri="http://treatment.plazi.org/id/FC52C90716CC53C38428EC62536F8885" lastPageNumber="111" pageId="0" pageNumber="111">
<subSubSection pageId="0" pageNumber="111" type="nomenclature">
<paragraph pageId="0" pageNumber="111">
<taxonomicName LSID="http://zoobank.org/4F1D8C21-1EEF-4248-8299-012692C20D79" authority="Radea &amp; Lampri &amp; Bakolitsas &amp; Parmakelis, 2021" authorityName="Radea &amp; Lampri &amp; Bakolitsas &amp; Parmakelis" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Daphniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Daphniola dione" order="Littorinimorpha" pageId="0" pageNumber="111" phylum="Mollusca" rank="species" species="dione" status="sp. nov.">Daphniola dione</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="111">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Daphniola dione sp. nov. A., E. Dorsal view; B., F., K. Ventral view; C., D., G., H., L. Lateral view; I., J. Shape of rectum in females and males respectively. Scale bar 1 mm." figureDoi="10.3897/zse.97.60254.figure3" httpUri="https://binary.pensoft.net/fig/505899" pageId="0" pageNumber="111">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A., B. Penis (dorsal aspect); C. Female genitalia. Abbreviations are given in the section of Materials and Methods. Scale bar 0.5 mm." figureDoi="10.3897/zse.97.60254.figure4" httpUri="https://binary.pensoft.net/fig/505900" pageId="0" pageNumber="111">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="111" type="etymology">
<paragraph pageId="0" pageNumber="111">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="111">The specific name (in apposition) derives from Greek mythology: Dione, (Διώνη in Greek), was the mother of the goddess Aphrodite according to the Greek poet Homer, author of Iliad and Odyssey.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="111" type="diagnosis">
<paragraph pageId="0" pageNumber="111">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="111">
Shell minute (maximum height 0.98 mm, maximum width 1.31 mm), valvatiform; soft body without any pigmentation; eyes small; penis long, narrow, tapered, with wider wrinkled proximal portion, filamentous distal portion and an obtuse outgrowth on the left side forming an acute angle with the penis distal portion; female genitalia with large pyriform bursa copulatrix, renal oviduct coiled in a equilateral triangle. Differentiated from
<taxonomicName genus="D." lsidName="D. exigua" pageId="0" pageNumber="111" rank="species" species="exigua">
<emphasis italics="true" pageId="0" pageNumber="111">D. exigua</emphasis>
</taxonomicName>
by its smaller size, wider and more open umbilicus, paler operculum, lack of pigmentation in soft body, pyriform-shaped bursa copulatrix, wider bursal duct, triangle-shaped oviduct and obtuse rather than pointed outgrowth at the distal penis portion. Differentiated from
<taxonomicName genus="D." lsidName="D. louisi" pageId="0" pageNumber="111" rank="species" species="louisi">
<emphasis italics="true" pageId="0" pageNumber="111">D. louisi</emphasis>
</taxonomicName>
by its smaller size, lack of pigmentation in soft body, large, pyriform and protruding bursa copulatrix, triangle-shaped oviduct, nearly centered penial duct, and more prominent penial outgrowth. Differentiated from
<taxonomicName genus="D." lsidName="D. eptalophos" pageId="0" pageNumber="111" rank="species" species="eptalophos">
<emphasis italics="true" pageId="0" pageNumber="111">D. eptalophos</emphasis>
</taxonomicName>
by its smaller size, paler operculum, lack of pigmentation in soft body, protruding bursa copulatrix, wider bursal duct, triangle-shaped oviduct, more pointed penial apex, outgrowth on distal rather than proximal penial portion and nearly centered penial duct. Differentiated from
<taxonomicName genus="D." lsidName="D. hadei" pageId="0" pageNumber="111" rank="species" species="hadei">
<emphasis italics="true" pageId="0" pageNumber="111">D. hadei</emphasis>
</taxonomicName>
by its larger size, wider and more open umbilicus, pyriform and protruding bursa copulatrix, more pointed penial apex and more prominent penial outgrowth. Differentiated from
<taxonomicName genus="D." lsidName="D. magdalenae" pageId="0" pageNumber="111" rank="species" species="magdalenae">
<emphasis italics="true" pageId="0" pageNumber="111">D. magdalenae</emphasis>
</taxonomicName>
by its smaller size, wider and more open umbilicus, paler operculum, protruding bursa copulatrix, more pointed penial apex and presence of eyes.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="111" type="type-locality">
<paragraph pageId="0" pageNumber="111">Type-locality.</paragraph>
<paragraph pageId="0" pageNumber="111">
Cave spring on Lefkada Island, Ionian Sea, Greece,
<geoCoordinate degrees="38.722532" direction="north" orientation="latitude" precision="1" value="38.72253">38.722532°N</geoCoordinate>
,
<geoCoordinate degrees="20.682713" direction="east" orientation="longitude" precision="1" value="20.682713">20.682713°E</geoCoordinate>
, 240 m a.s.l., 15.V.2017, K. Bakolitsas leg.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="111" type="type material">
<paragraph pageId="0" pageNumber="111">Type material.</paragraph>
<paragraph pageId="0" pageNumber="111">
<emphasis bold="true" italics="true" pageId="0" pageNumber="111">Holotype</emphasis>
. Ethanol-fixed specimen, GNHM 39587.
<emphasis bold="true" italics="true" pageId="0" pageNumber="111">Paratypes</emphasis>
(from the same lot). Two ethanol-fixed specimens, GNHM 39588. Seven ethanol-fixed mature specimens dissected for anatomical study. Three mature and seven immature specimens are in the personal collection of C. Radea deposited in the Section of Ecology and Systematics, Department of Biology, NKUA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="111" type="description">
<paragraph pageId="0" pageNumber="111">Description.</paragraph>
<paragraph pageId="0" pageNumber="111">
<emphasis italics="true" pageId="0" pageNumber="111">Shell</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Daphniola dione sp. nov. A., E. Dorsal view; B., F., K. Ventral view; C., D., G., H., L. Lateral view; I., J. Shape of rectum in females and males respectively. Scale bar 1 mm." figureDoi="10.3897/zse.97.60254.figure3" httpUri="https://binary.pensoft.net/fig/505899" pageId="0" pageNumber="111">3A-L</figureCitation>
). Valvatiform with up to 3.25 whorls, thin, colorless, transparent and crystalline when fresh, finely striated; spire more or less depressed; whorls rounded, regularly growing with shallow sutures. Measurements are given in Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Shell morphometry of Daphniola dione sp. nov. Measurements are in mm. For abbreviations see the section of Materials and methods." httpUri="http://table.plazi.org/id/06C5F923CF6516B19D2DDCFA1AFD9299" pageId="0" pageNumber="111" tableUuid="06C5F923CF6516B19D2DDCFA1AFD9299">3</tableCitation>
. Periostracum light cream-colored; aperture adhering to the last whorl, prosocline, roundish to ovate; peristome continuous, slightly thickened at columellar margin, the outer margin simple; umbilicus open, deep, wide so that the first whorls can be seen through it (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Daphniola dione sp. nov. A., E. Dorsal view; B., F., K. Ventral view; C., D., G., H., L. Lateral view; I., J. Shape of rectum in females and males respectively. Scale bar 1 mm." figureDoi="10.3897/zse.97.60254.figure3" httpUri="https://binary.pensoft.net/fig/505899" pageId="0" pageNumber="111">3B, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.97.60254.figure3" httpUri="https://binary.pensoft.net/fig/505899" pageId="0" pageNumber="111" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="111">
<emphasis bold="true" pageId="0" pageNumber="111">Figure 3.</emphasis>
Shells of
<taxonomicName authorityName="Radea &amp; Lampri &amp; Bakolitsas &amp; Parmakelis" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Daphniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Daphniola dione" order="Littorinimorpha" pageId="0" pageNumber="111" phylum="Mollusca" rank="species" species="dione">
<emphasis italics="true" pageId="0" pageNumber="111">Daphniola dione</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="111">A.</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="111">E.</emphasis>
Dorsal view;
<emphasis bold="true" pageId="0" pageNumber="111">B.</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="111">F.</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="111">K.</emphasis>
Ventral view;
<emphasis bold="true" pageId="0" pageNumber="111">C.</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="111">D.</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="111">G.</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="111">H.</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="111">L.</emphasis>
Lateral view;
<emphasis bold="true" pageId="0" pageNumber="111">I.</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="111">J.</emphasis>
Shape of rectum in females and males respectively. Scale bar 1 mm.
</paragraph>
</caption>
<caption ID-Table-UUID="06C5F923CF6516B19D2DDCFA1AFD9299" httpUri="http://table.plazi.org/id/06C5F923CF6516B19D2DDCFA1AFD9299" pageId="0" pageNumber="111" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="111">
<emphasis bold="true" pageId="0" pageNumber="111">Table 3.</emphasis>
Shell morphometry of
<taxonomicName authorityName="Radea &amp; Lampri &amp; Bakolitsas &amp; Parmakelis" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Daphniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Daphniola dione" order="Littorinimorpha" pageId="0" pageNumber="111" phylum="Mollusca" rank="species" species="dione">
<emphasis italics="true" pageId="0" pageNumber="111">Daphniola dione</emphasis>
</taxonomicName>
sp. nov. Measurements are in mm. For abbreviations see the section of Materials and methods.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="111">
<table pageId="0" pageNumber="111">
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">n=10</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">SH</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">SW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">AH</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">AW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">SH/SW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">AH/AW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">SH/AH</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">SW/AW</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">Min</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.83</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.14</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.52</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.68</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.79</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.62</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2.05</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">Max</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.98</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.31</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.53</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.61</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.80</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.02</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2.08</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2.21</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="111">X̄</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.91</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.21</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.51</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.57</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.75</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.90</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.78</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2.13</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">SD</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.04</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.05</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.03</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.03</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.03</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.07</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.12</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.06</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">CV</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.05</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.05</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.06</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.06</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.05</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.08</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.07</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.03</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="111">
<emphasis italics="true" pageId="0" pageNumber="111">Operculum</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Daphniola dione sp. nov. A., E. Dorsal view; B., F., K. Ventral view; C., D., G., H., L. Lateral view; I., J. Shape of rectum in females and males respectively. Scale bar 1 mm." figureDoi="10.3897/zse.97.60254.figure3" httpUri="https://binary.pensoft.net/fig/505899" pageId="0" pageNumber="111">3F</figureCitation>
). Ovate, thin, pliable, corneous, paucispiral, yellowish darker at the nucleus, with weakly convex inner face without any peg, nucleus sub-central.
</paragraph>
<paragraph pageId="0" pageNumber="111">
<emphasis italics="true" pageId="0" pageNumber="111">Soft body pigmentation</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Daphniola dione sp. nov. A., E. Dorsal view; B., F., K. Ventral view; C., D., G., H., L. Lateral view; I., J. Shape of rectum in females and males respectively. Scale bar 1 mm." figureDoi="10.3897/zse.97.60254.figure3" httpUri="https://binary.pensoft.net/fig/505899" pageId="0" pageNumber="111">3A-L</figureCitation>
). Soft body totally unpigmented and visible under the transparent shell; snout longer than wide, parallel-sided with medium distal lobation; eye spots very small.
</paragraph>
<paragraph pageId="0" pageNumber="111">
<emphasis italics="true" pageId="0" pageNumber="111">Nervous system</emphasis>
. Cerebral ganglia of the same size, white-colored; supraoesophageal and suboesophageal ganglia of the same size, smaller than cerebral ganglia, white-colored; supraoesophageal connective longer than suboesophageal connective.
</paragraph>
<paragraph pageId="0" pageNumber="111">
<emphasis italics="true" pageId="0" pageNumber="111">Ctenidium-Osphradium</emphasis>
. Ctenidium with ca 8-12 lamellae. Osphradium of intermediate width, opposite approximate middle of ctenidium.
</paragraph>
<paragraph pageId="0" pageNumber="111">
<emphasis italics="true" pageId="0" pageNumber="111">Digestive system</emphasis>
. Radula very fragile, not investigated; stomach unpigmented. Style sac smaller than stomach, not protruding to the intestinal loop; rectum (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Daphniola dione sp. nov. A., E. Dorsal view; B., F., K. Ventral view; C., D., G., H., L. Lateral view; I., J. Shape of rectum in females and males respectively. Scale bar 1 mm." figureDoi="10.3897/zse.97.60254.figure3" httpUri="https://binary.pensoft.net/fig/505899" pageId="0" pageNumber="111">3I, J</figureCitation>
) with V-shaped bend, in some specimens with irregular V-shaped bend. The V-shaped bend is narrower in males (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Daphniola dione sp. nov. A., E. Dorsal view; B., F., K. Ventral view; C., D., G., H., L. Lateral view; I., J. Shape of rectum in females and males respectively. Scale bar 1 mm." figureDoi="10.3897/zse.97.60254.figure3" httpUri="https://binary.pensoft.net/fig/505899" pageId="0" pageNumber="111">3J</figureCitation>
) than in females (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Daphniola dione sp. nov. A., E. Dorsal view; B., F., K. Ventral view; C., D., G., H., L. Lateral view; I., J. Shape of rectum in females and males respectively. Scale bar 1 mm." figureDoi="10.3897/zse.97.60254.figure3" httpUri="https://binary.pensoft.net/fig/505899" pageId="0" pageNumber="111">3I</figureCitation>
). Faecal pellets are longitudinally packed.
</paragraph>
<paragraph pageId="0" pageNumber="111">
<emphasis italics="true" pageId="0" pageNumber="111">Male reproductive system</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A., B. Penis (dorsal aspect); C. Female genitalia. Abbreviations are given in the section of Materials and Methods. Scale bar 0.5 mm." figureDoi="10.3897/zse.97.60254.figure4" httpUri="https://binary.pensoft.net/fig/505900" pageId="0" pageNumber="111">4A, B</figureCitation>
) Penis whitish, medium-sized, narrow, long, gradually tapering bearing an obtuse outgrowth on the left side forming an acute angle with the penis distal portion; distal portion filamentous and usually bent; proximal portion bent upon itself and wrinkled; base of penis of intermediate width, its attachment area behind the right eye; penial duct strongly undulating especially in the proximal penis portion, near outer edge positioned, penial opening terminal; prostate like an elongate bean.
</paragraph>
<caption doi="10.3897/zse.97.60254.figure4" httpUri="https://binary.pensoft.net/fig/505900" pageId="0" pageNumber="111" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="111">
<emphasis bold="true" pageId="0" pageNumber="111">Figure 4.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="111">A</emphasis>
.,
<emphasis bold="true" pageId="0" pageNumber="111">B</emphasis>
. Penis (dorsal aspect);
<emphasis bold="true" pageId="0" pageNumber="111">C</emphasis>
. Female genitalia. Abbreviations are given in the section of Materials and Methods. Scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="111">
<emphasis italics="true" pageId="0" pageNumber="111">Female reproductive system</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A., B. Penis (dorsal aspect); C. Female genitalia. Abbreviations are given in the section of Materials and Methods. Scale bar 0.5 mm." figureDoi="10.3897/zse.97.60254.figure4" httpUri="https://binary.pensoft.net/fig/505900" pageId="0" pageNumber="111">4C</figureCitation>
). Albumen and capsule glands very small, total mean height 0.38 mm (three specimens); bursa copulatrix large-sized, pyriform, posteriorly positioned and fully protruding from the posterior end of the albumen gland with longitudinal orientation relative to albumen gland; bursal duct anterior, wide and longer than bursa length; coiled oviduct unpigmented, well-developed forming an equilateral triangle with a vertical, V-shaped primary loop; two seminal receptacles; distal seminal receptacle (rs1) very small, globular without duct; proximal seminal receptacle (rs2) larger than distal one and smaller than bursa copulatrix, pyriform, with very short duct and a pink pearly shine, lying over the renal oviduct and against bursa copulatrix; an egg capsule with a single egg was found inside the umbilicus of one specimen (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells of Daphniola dione sp. nov. A., E. Dorsal view; B., F., K. Ventral view; C., D., G., H., L. Lateral view; I., J. Shape of rectum in females and males respectively. Scale bar 1 mm." figureDoi="10.3897/zse.97.60254.figure3" httpUri="https://binary.pensoft.net/fig/505899" pageId="0" pageNumber="111">3K, L</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="111">
<normalizedToken originalValue="Μorphometric">Μorphometric</normalizedToken>
data along with conchological and anatomical characters of the nominal
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Daphniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Daphniola" order="Littorinimorpha" pageId="0" pageNumber="111" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="111">Daphniola</emphasis>
</taxonomicName>
species are provided in Tables
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphometry of Daphniola species. Measurements are in mm. For abbreviations see Materials and methods." httpUri="http://table.plazi.org/id/DC57F8627B675683FC1D1803FFDC7787" pageId="0" pageNumber="111" tableUuid="DC57F8627B675683FC1D1803FFDC7787">4</tableCitation>
and
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Morphological and anatomical diagnostic characters of D. dione sp. nov. and its congeneric species." httpUri="http://table.plazi.org/id/E65389ACFD951F5945A70C034DCB0D66" pageId="0" pageNumber="111" tableUuid="E65389ACFD951F5945A70C034DCB0D66">5</tableCitation>
, respectively.
</paragraph>
<caption ID-Table-UUID="DC57F8627B675683FC1D1803FFDC7787" httpUri="http://table.plazi.org/id/DC57F8627B675683FC1D1803FFDC7787" pageId="0" pageNumber="111" start="Table 4" startId="T4">
<paragraph pageId="0" pageNumber="111">
<emphasis bold="true" pageId="0" pageNumber="111">Table 4.</emphasis>
Morphometry of
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Daphniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Daphniola" order="Littorinimorpha" pageId="0" pageNumber="111" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="111">Daphniola</emphasis>
</taxonomicName>
species. Measurements are in mm. For abbreviations see Materials and methods.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="111">
<table pageId="0" pageNumber="111">
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">Species</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">NW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">SH</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">SW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">AH</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="111">AW</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. dione" pageId="0" pageNumber="111" rank="species" species="dione">
<emphasis italics="true" pageId="0" pageNumber="111">D. dione</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2.75-3.25</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.83-0.98</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.14-1.31</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.47-0.55</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.52-0.61</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. eptalophos" pageId="0" pageNumber="111" rank="species" species="eptalophos">
<emphasis italics="true" pageId="0" pageNumber="111">D. eptalophos</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">3.25-3.50</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.90-1.25</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.10-1.90</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.50-0.80</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.50-0.75</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. exigua" pageId="0" pageNumber="111" rank="species" species="exigua">
<emphasis italics="true" pageId="0" pageNumber="111">D. exigua</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">3.00-3.50</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.10-1.52</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.10-1.40</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.60-0.80</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.60-0.76</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. hadei" pageId="0" pageNumber="111" rank="species" species="hadei">
<emphasis italics="true" pageId="0" pageNumber="111">D. hadei</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2.25-2.50</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.84-0.85</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.14-1.15</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.55-0.57</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.52-0.54</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. louisi" pageId="0" pageNumber="111" rank="species" species="louisi">
<emphasis italics="true" pageId="0" pageNumber="111">D. louisi</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">3.50</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.09-1.45</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.17-1.69</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.59-0.98</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0.59-0.79</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. magdalenae" pageId="0" pageNumber="111" rank="species" species="magdalenae">
<emphasis italics="true" pageId="0" pageNumber="111">D. magdalenae</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">3.50-3.75</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2.34</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.35</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1.28</td>
</tr>
</table>
</paragraph>
<caption ID-Table-UUID="E65389ACFD951F5945A70C034DCB0D66" httpUri="http://table.plazi.org/id/E65389ACFD951F5945A70C034DCB0D66" pageId="0" pageNumber="111" start="Table 5" startId="T5">
<paragraph pageId="0" pageNumber="111">
<emphasis bold="true" pageId="0" pageNumber="111">Table 5.</emphasis>
Morphological and anatomical diagnostic characters of
<taxonomicName genus="D." lsidName="D. dione" pageId="0" pageNumber="111" rank="species" species="dione">
<emphasis italics="true" pageId="0" pageNumber="111">D. dione</emphasis>
</taxonomicName>
sp. nov. and its congeneric species.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="111">
<table pageId="0" pageNumber="111">
<tr pageId="0" pageNumber="111">
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">Species</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">1</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">2</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">3</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">4</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">5</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">6</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">7</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">8</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">9</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">10</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">11</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">12</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">13</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">14</th>
<th colspan="1" pageId="0" pageNumber="111" rowspan="1">15</th>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. dione" pageId="0" pageNumber="111" rank="species" species="dione">
<emphasis italics="true" pageId="0" pageNumber="111">D. dione</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. eptalophos" pageId="0" pageNumber="111" rank="species" species="eptalophos">
<emphasis italics="true" pageId="0" pageNumber="111">D. eptalophos</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1,2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. exigua" pageId="0" pageNumber="111" rank="species" species="exigua">
<emphasis italics="true" pageId="0" pageNumber="111">D. exigua</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0,1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. hadei" pageId="0" pageNumber="111" rank="species" species="hadei">
<emphasis italics="true" pageId="0" pageNumber="111">D. hadei</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0,1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. louisi" pageId="0" pageNumber="111" rank="species" species="louisi">
<emphasis italics="true" pageId="0" pageNumber="111">D. louisi</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
</tr>
<tr pageId="0" pageNumber="111">
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">
<taxonomicName genus="D." lsidName="D. magdalenae" pageId="0" pageNumber="111" rank="species" species="magdalenae">
<emphasis italics="true" pageId="0" pageNumber="111">D. magdalenae</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">--</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="111" rowspan="1">1</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="111">
<tableNote pageId="0" pageNumber="111">Umbilicus: wide, open (0); semi-open (1) Operculum: light yellowish darker on the nucleus (0); orange; darker on the nucleus (1); yellow; orange on the nucleus (2); pink reddish (3) Soft body pigmentation: no pigmentation (0); light pigmentation (1); pigmentation (2) Bursa copulatrix shape: globular (0); ovoid (1); pyriform (2) Bursa copulatrix size: medium (0); large (1) Bursa copulatrix position relative to posterior end of albumen gland: non-protruding (0); protruding (1) Bursal duct width: narrow (0); medium wide (1); wide (2) Coiled oviduct shape: circular-ovoid (0); Ovoid (1); ellipsoid, transverse relative to albumen gland (2); triangular (3) Primary oviduct loop shape: V (0); U (1); circular (2) Penis pigmentation and shape: unpigmented with pointed apex (0); unpigmented with long, narrow filament (1); pigmented with pointed apex or short filament (2) Penial outgrowth shape: prominent; forming a right angle with the distal part of penis (0); prominent; forming an acute angle with the distal part of penis (1); slightly prominent (2) Penial outgrowth position: distal penis portion (0); medial (1); basal penis portion (2) Penial duct: near center (0); near outer edge (1) Eyes: absent (0); present (1) Habitat: spring (0); cave spring (1) -: No data</tableNote>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="111" type="distribution">
<paragraph pageId="0" pageNumber="111">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="111">
So far, the distribution of
<taxonomicName authorityName="Radea &amp; Lampri &amp; Bakolitsas &amp; Parmakelis" authorityYear="2021" class="Gastropoda" family="Hydrobiidae" genus="Daphniola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Daphniola dione" order="Littorinimorpha" pageId="0" pageNumber="111" phylum="Mollusca" rank="species" species="dione">
<emphasis italics="true" pageId="0" pageNumber="111">Daphniola dione</emphasis>
</taxonomicName>
sp. nov., is restricted to the type locality. The live specimens of the new species were found crawling on the roots of woody plants reaching the bottom of the spring.
</paragraph>
</subSubSection>
</treatment>
</document>
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