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<mods:title id="EC194A2A6B1D9C067C73F5914A06B5C1">Clarification of the identity of deep-sea crab Tymolus uncifer (Ortmann, 1892) and validation of Tymolus melanommus (Doflein, 1904) (Decapoda: Brachyura: Cyclodorippidae)</mods:title>
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<treatment id="E40287F3397CFFFBFF749F8907D9FCD6" ID-DOI="http://doi.org/10.5281/zenodo.8412113" ID-GBIF-Taxon="213730616" ID-Zenodo-Dep="8412113" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6" httpUri="http://treatment.plazi.org/id/E40287F3397CFFFBFF749F8907D9FCD6" lastPageId="11" lastPageNumber="57" pageId="3" pageNumber="49">
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<heading id="375C8189397CFFF3FF749F890619F9E1" bold="true" box="[151,567,1534,1561]" fontSize="11" level="1" pageId="3" pageNumber="49" reason="1">
<taxonomicName id="ABAB4D66397CFFF3FF749F890619F9E1" ID-CoL="7DBPK" ID-ENA="2545257" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,567,1534,1561]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="uncifer">
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<emphasis id="5EDFEAF7397CFFF3FF749F890570F9E1" bold="true" box="[151,350,1534,1561]" italics="true" pageId="3" pageNumber="49">Tymolus uncifer</emphasis>
(
<bibRefCitation id="083A4B14397CFFF3FE8E9F890601F9E1" author="Ortmann, A. E." box="[365,559,1534,1560]" pageId="3" pageNumber="49" pagination="532 - 588" refId="ref17259" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit bedonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit Strassburger Museum aufwbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
)
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</paragraph>
<paragraph id="6C1436E5397CFFF3FF749C5406BBF9C4" blockId="3.[151,661,1534,1633]" box="[151,661,1571,1597]" pageId="3" pageNumber="49">[Japanese name: Ashinaga-mame-heike-gani]</paragraph>
</subSubSection>
<subSubSection id="24B1656E397CFFF3FF749C300542F998" box="[151,364,1607,1633]" pageId="3" pageNumber="49" type="description">
<paragraph id="6C1436E5397CFFF3FF749C300542F998" blockId="3.[151,661,1534,1633]" box="[151,364,1607,1633]" pageId="3" pageNumber="49">
(
<figureCitation id="F4902A60397CFFF3FF7C9C3004C1F998" box="[159,239,1607,1633]" captionStart="FIGURE 2" captionStartId="4.[151,250,1052,1076]" captionTargetBox="[151,1436,322,1029]" captionTargetId="figure-71@4.[151,1436,322,1029]" captionTargetPageId="4" captionText="FIGURE 2. Tymolus uncifer (Ortmann, 1892), lectotype, male (5.0×5.0 mm), MZS Cru 3187. A, cephalothorax and pleon, dorsal view; B, same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8406962" httpUri="https://zenodo.org/record/8406962/files/figure.png" pageId="3" pageNumber="49">Figs. 2</figureCitation>
,
<figureCitation id="F4902A60397CFFF3FF189C300509F998" box="[251,295,1607,1633]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStartId-0="5.[151,250,1696,1720]" captionStartId-1="6.[151,250,1290,1314]" captionStartId-2="7.[151,250,1439,1463]" captionStartId-3="9.[151,250,1909,1933]" captionStartId-4="10.[151,250,1808,1832]" captionTargetBox-0="[156,1431,181,1672]" captionTargetBox-1="[151,1436,417,1267]" captionTargetBox-2="[159,1428,189,1407]" captionTargetBox-3="[206,1380,189,1880]" captionTargetBox-4="[159,1428,857,1777]" captionTargetId-0="figure-21@5.[156,1431,181,1672]" captionTargetId-1="figure-127@6.[151,1436,417,1267]" captionTargetId-2="figure-21@7.[151,1436,181,1415]" captionTargetId-3="figure-21@9.[198,1388,181,1885]" captionTargetId-4="figure-298@10.[151,1436,849,1785]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="FIGURE 4. Tymolus uncifer (Ortmann, 1892), habitus in dorsal view. A, male (9.8×9.8 mm), CBM-ZC 16951; B, ovigerous female (7.6×7.6 mm), CBM-ZC 16943. In preservative." captionText-1="FIGURE 5. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, carapace and cephalic appendages, dorsal view; B, thoracic sternum, ventral view. In preservative." captionText-2="FIGURE 6. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted)." captionText-3="FIGURE 7. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951.A, right pereopod 2, dorsal view; B, same, distal part of merus, dorsal view; C, right pereopod 3, dorsal view (setae omitted); D, same, distal part of merus, dorsal view; E, right pereopod 4, dorsal view; F, right pereopod 5, dorsal view; G, left gonopod 1, mesial view; H, same, ventral view; I, same, lateral view; J, same, dorsal view; K, left gonopod 2, lateral view." captionText-4="FIGURE 8. Tymolus uncifer (Ortmann, 1892), ovigerous female 7.6×7.6 mm), CBM-ZC 16943. A, thoracic sternum, ventral view; B, right chela, outer view; C, pleon, external view. Setae omitted." figureDoi-0="http://doi.org/10.5281/zenodo.8406966" figureDoi-1="http://doi.org/10.5281/zenodo.8406969" figureDoi-2="http://doi.org/10.5281/zenodo.8406971" figureDoi-3="http://doi.org/10.5281/zenodo.8406973" figureDoi-4="http://doi.org/10.5281/zenodo.8406975" httpUri-0="https://zenodo.org/record/8406966/files/figure.png" httpUri-1="https://zenodo.org/record/8406969/files/figure.png" httpUri-2="https://zenodo.org/record/8406971/files/figure.png" httpUri-3="https://zenodo.org/record/8406973/files/figure.png" httpUri-4="https://zenodo.org/record/8406975/files/figure.png" pageId="3" pageNumber="49">48</figureCitation>
,
<figureCitation id="F4902A60397CFFF3FED09C30054AF998" box="[307,356,1607,1633]" captionStart="FIGURE 14" captionStartId="18.[151,250,1691,1715]" captionTargetBox="[151,1436,181,1668]" captionTargetId="figure-15@18.[151,1436,181,1668]" captionTargetPageId="18" captionText="FIGURE 14. Habitus showing living colouration.A, Tymolus uncifer (Ortmann, 1892), ovigerous female (7.8×8.1 mm), CBM- ZC 16841; B, Tymolus melanommus (Doflein, 1904), male (13.0×13.0 mm), CBM-ZC 17323." figureDoi="http://doi.org/10.5281/zenodo.8406990" httpUri="https://zenodo.org/record/8406990/files/figure.png" pageId="3" pageNumber="49">14A</figureCitation>
)
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<paragraph id="6C1436E5397CFFF3FF749CFC017CF95A" blockId="3.[151,1437,1675,2036]" box="[151,1362,1675,1699]" pageId="3" pageNumber="49">
<treatmentCitationGroup id="4CBB11CB397CFFF3FF749CFC017CF95A" box="[151,1362,1675,1699]" pageId="3" pageNumber="49">
<treatmentCitation id="ED0A10F4397CFFF3FF749CFC067FF95A" author="Ortmann, A. E." box="[151,593,1675,1699]" page="560" pageId="3" pageNumber="49" year="1892">
<taxonomicName id="ABAB4D66397CFFF3FF749CFC067FF95A" ID-CoL="6BS8X" authority="Ortmann, 1892: 560" authorityName="Ortmann" authorityPageNumber="560" authorityYear="1892" box="[151,593,1675,1699]" class="Malacostraca" family="Cyclodorippidae" genus="Cyclodorippe" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="uncifera">
<emphasis id="5EDFEAF7397CFFF3FF749CFC0555F95A" box="[151,379,1675,1699]" italics="true" pageId="3" pageNumber="49">Cyclodorippe uncifera</emphasis>
<bibRefCitation id="083A4B14397CFFF3FE619CFC067FF95A" author="Ortmann, A. E." box="[386,593,1675,1699]" pageId="3" pageNumber="49" pagination="532 - 588" refId="ref17259" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit bedonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit Strassburger Museum aufwbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26." type="journal article" year="1892">Ortmann, 1892: 560</bibRefCitation>
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, pl. 26, fig. 6.—
<treatmentCitation id="ED0A10F4397CFFF3FD179CFC078AF95A" author="Doflein, F." box="[756,932,1675,1699]" page="34" pageId="3" pageNumber="49" year="1904">
<bibRefCitation id="083A4B14397CFFF3FD179CFC078AF95A" author="Doflein, F." box="[756,932,1675,1699]" pageId="3" pageNumber="49" pagination="1 - 6" refId="ref16210" refString="Doflein, F. (1904) Brachyura. Wissenschaftliche Ergebnisse der Deutchen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899, 6 (1 - 6), i - xiv + 1 - 314, atlas (pls. 311 - 358)." type="journal article" year="1904">Doflein 1904: 34</bibRefCitation>
</treatmentCitation>
(in part).—
<treatmentCitation id="ED0A10F4397CFFF3FBF89CFC00EBF95A" author="Parisi, B." box="[1051,1221,1675,1699]" page="197" pageId="3" pageNumber="49" year="1914">
<bibRefCitation id="083A4B14397CFFF3FBF89CFC00EBF95A" author="Parisi, B." box="[1051,1221,1675,1699]" pageId="3" pageNumber="49" pagination="282 - 311" refId="ref17332" refString="Parisi, B. (1914) I Decapodi giappnesi del Museo di Milano, I. Oxystomata. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 53, 282 - 311, pls. 11 - 13." type="journal article" year="1914">Parisi 1914: 197</bibRefCitation>
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, pl. 13, fig. 2.
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</paragraph>
<paragraph id="6C1436E5397CFFF3FF749CDD04D0F918" blockId="3.[151,1437,1675,2036]" pageId="3" pageNumber="49">
<treatmentCitationGroup id="4CBB11CB397CFFF3FF749CDD04D0F918" pageId="3" pageNumber="49">
<taxonomicName id="ABAB4D66397CFFF3FF749CDD0517F93B" ID-CoL="7DBPK" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,313,1706,1730]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF7397CFFF3FF749CDD0517F93B" box="[151,313,1706,1730]" italics="true" pageId="3" pageNumber="49">Tymolus uncifer</emphasis>
</taxonomicName>
.—
<treatmentCitation id="ED0A10F4397CFFF3FEB49CDD05C1F93B" author="Sakai, T." box="[343,495,1706,1730]" page="10" pageId="3" pageNumber="49" year="1965">
<bibRefCitation id="083A4B14397CFFF3FEB49CDD05C1F93B" author="Sakai, T." box="[343,495,1706,1730]" pageId="3" pageNumber="49" refId="ref17473" refString="Sakai, T. (1965) The Crabs of Sagami Bay, Collected by His Majesty the Emperor of Japan. Maruzen, Tokyo, 330 pp, 100 pls." type="book" year="1965">Sakai 1965: 10</bibRefCitation>
</treatmentCitation>
(Japanese text, in part).—1976: 32 (Japanese text, in part).—
<treatmentCitation id="ED0A10F4397CFFF3FBBB9CDD00DEF93B" author="Ikeda, H." box="[1112,1264,1706,1730]" page="21" pageId="3" pageNumber="49" year="1998">
<bibRefCitation id="083A4B14397CFFF3FBBB9CDD00DEF93B" author="Ikeda, H." box="[1112,1264,1706,1730]" pageId="3" pageNumber="49" refId="ref16530" refString="Ikeda, H. (1998) The Deep-sea Crabs of Sagami Bay. Hayama Shiosai Museum, Hayama, 180 pp. [in Japanese]" type="book" year="1998">Ikeda 1998: 21</bibRefCitation>
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(in part), 59, fig. 3ac.
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</paragraph>
<paragraph id="6C1436E5397CFFF3FF749C90077BF906" blockId="3.[151,1437,1675,2036]" box="[151,853,1767,1791]" pageId="3" pageNumber="49">
<treatmentCitationGroup id="4CBB11CB397CFFF3FF749C90077BF906" box="[151,853,1767,1791]" pageId="3" pageNumber="49">
?
<emphasis id="5EDFEAF7397CFFF3FF4A9C900565F906" box="[169,331,1767,1791]" italics="true" pageId="3" pageNumber="49">
<taxonomicName id="ABAB4D66397CFFF3FF4A9C900569F906" ID-CoL="7DBPQ" authorityName="Tavares" authorityYear="1991" box="[169,327,1767,1791]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="brucei">Tymolus brucei</taxonomicName>
.
</emphasis>
<treatmentCitation id="ED0A10F4397CFFF3FE879C9006BDF906" author="Spiridonov, V. A. &amp; Turkay, M." box="[356,659,1767,1791]" page="7" pageId="3" pageNumber="49" year="2007">
<bibRefCitation id="083A4B14397CFFF3FE879C9006BDF906" author="Spiridonov, V. A. &amp; Turkay, M." box="[356,659,1767,1791]" pageId="3" pageNumber="49" pagination="1 - 9" refId="ref17626" refString="Spiridonov, V. A. &amp; Turkay, M. (2007) Podotreme crabs from the western Indian Ocean: description of a new species of Tymolus Stimpson, 1858 with a review of regional distribution of Cyclodorippidae (Crustacea: Decapoda: Brachyura: Podotremata). Arthropoda Selecta, 16 (1), 1 - 9." type="journal article" year="2007">Spiridonov &amp; Türkay 2007: 7</bibRefCitation>
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. See “Discussion”.
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</paragraph>
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<treatmentCitationGroup id="4CBB11CB397CFFF3FF749D710002F8E7" box="[151,1068,1798,1822]" pageId="3" pageNumber="49">
Not
<taxonomicName id="ABAB4D66397CFFF3FF279D7105B2F8E7" authorityName="Ortmann" authorityYear="1892" box="[196,412,1798,1822]" class="Malacostraca" family="Cyclodorippidae" genus="Cyclodorippe" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF7397CFFF3FF279D7105B2F8E7" box="[196,412,1798,1822]" italics="true" pageId="3" pageNumber="49">Cyclodorippe uncifer</emphasis>
</taxonomicName>
.—
<treatmentCitation id="ED0A10F4397CFFF3FE5A9D71065BF8E7" author="Doflein, F." box="[441,629,1798,1822]" page="653" pageId="3" pageNumber="49" year="1902">
<bibRefCitation id="083A4B14397CFFF3FE5A9D71065BF8E7" author="Doflein, F." box="[441,629,1798,1822]" pageId="3" pageNumber="49" pagination="613 - 670" refId="ref16182" refString="Doflein, F. (1902) Ostasiatische Dekapoden. Abhandlungen der Bayerischen Akademie der Wissenschaften, Munchen, 21, 613 - 670, 6 pls." type="journal article" year="1902">Doflein 1902: 653</bibRefCitation>
</treatmentCitation>
[=
<taxonomicName id="ABAB4D66397CFFF3FD7B9D71000CF8E7" authority="(Doflein, 1904)" baseAuthorityName="Doflein" baseAuthorityYear="1904" box="[664,1058,1798,1822]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="melanommus">
<emphasis id="5EDFEAF7397CFFF3FD7B9D710757F8E7" box="[664,889,1798,1822]" italics="true" pageId="3" pageNumber="49">Tymolus melanommus</emphasis>
(
<bibRefCitation id="083A4B14397CFFF3FC659D710037F8E7" author="Doflein, F." box="[902,1049,1798,1822]" pageId="3" pageNumber="49" pagination="1 - 6" refId="ref16210" refString="Doflein, F. (1904) Brachyura. Wissenschaftliche Ergebnisse der Deutchen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899, 6 (1 - 6), i - xiv + 1 - 314, atlas (pls. 311 - 358)." type="journal article" year="1904">Doflein, 1904</bibRefCitation>
)
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="6C1436E5397CFFF3FF749D520527F8A2" blockId="3.[151,1437,1675,2036]" pageId="3" pageNumber="49">
<treatmentCitationGroup id="4CBB11CB397CFFF3FF749D520527F8A2" pageId="3" pageNumber="49">
Not
<taxonomicName id="ABAB4D66397CFFF3FF219D53067DF8C4" ID-CoL="6BS8X" authorityName="Ortmann" authorityYear="1892" box="[194,595,1828,1853]" class="Malacostraca" family="Cyclodorippidae" genus="Cyclodorippe" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="uncifera" subGenus="Cyclortmannia">
<emphasis id="5EDFEAF7397CFFF3FF219D530562F8C5" box="[194,332,1828,1852]" italics="true" pageId="3" pageNumber="49">Cyclodorippe</emphasis>
(
<emphasis id="5EDFEAF7397CFFF3FEB99D5305DCF8C5" box="[346,498,1828,1852]" italics="true" pageId="3" pageNumber="49">Cyclortmannia</emphasis>
)
<emphasis id="5EDFEAF7397CFFF3FE1C9D52067DF8C4" box="[511,595,1829,1853]" italics="true" pageId="3" pageNumber="49">uncifera</emphasis>
</taxonomicName>
.—
<treatmentCitation id="ED0A10F4397CFFF3FD919D530728F8C4" author="Ihle, J. E. W." box="[626,774,1828,1853]" page="134" pageId="3" pageNumber="49" year="1916">
<bibRefCitation id="083A4B14397CFFF3FD919D530728F8C4" author="Ihle, J. E. W." box="[626,774,1828,1853]" pageId="3" pageNumber="49" pagination="97 - 158" refId="ref16496" refString="Ihle, J. E. W. (1916) Die Decapoda Brachyura der Siboga-Expedition. II. Oxystomata, Dorippidae. Siboga-Expeditie Monographie, 39 B 1, 97 - 158." type="journal article" year="1916">Ihle 1916: 134</bibRefCitation>
</treatmentCitation>
, figs. 40, 43, 49, 50, 52, 57, 62. =?
<taxonomicName id="ABAB4D66397CFFF3FB859D530528F8A2" ID-CoL="7DBPR" authority="(Alcock, 1894)" baseAuthorityName="Alcock" baseAuthorityYear="1894" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="glaucommus">
<emphasis id="5EDFEAF7397CFFF3FB859D530111F8C5" box="[1126,1343,1828,1852]" italics="true" pageId="3" pageNumber="49">Tymolus glaucommus</emphasis>
(
<bibRefCitation id="083A4B14397CFFF3FAA89D5304D0F8A2" author="Alcock, A." pageId="3" pageNumber="49" pagination="225 - 245" refId="ref15590" refString="Alcock, A. (1894) Natural history notes from H. M. Indian Marine Survey Steamer Investigator. Series II, No. 1. On the results of deep-sea dredging during the season of 1890 - 1891. Annals and Magazine of Natural History, Series 6, 13, 225 - 245 + 321 - 334 + 400 - 411 https: // doi. org / 10.1080 / 00222939408677694" type="journal article" year="1894">Alcock, 1894</bibRefCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6C1436E5397CFFF3FF749D1507BDF82F" blockId="3.[151,1437,1675,2036]" pageId="3" pageNumber="49">
<treatmentCitationGroup id="4CBB11CB397CFFF3FF749D1507BDF82F" pageId="3" pageNumber="49">
Not
<taxonomicName id="ABAB4D66397CFFF3FF219D15054AF883" ID-CoL="7DBPK" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[194,356,1890,1914]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF7397CFFF3FF219D15054AF883" box="[194,356,1890,1914]" italics="true" pageId="3" pageNumber="49">Tymolus uncifer</emphasis>
</taxonomicName>
.—Huang &amp; Liu 1979: 1 [=
<taxonomicName id="ABAB4D66397CFFF3FD999D15078AF883" ID-CoL="7DBPQ" authority="Tavares, 1991" authorityName="Tavares" authorityYear="1991" box="[634,932,1890,1914]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="brucei">
<emphasis id="5EDFEAF7397CFFF3FD999D15073AF883" box="[634,788,1890,1914]" italics="true" pageId="3" pageNumber="49">Tymolus brucei</emphasis>
<bibRefCitation id="083A4B14397CFFF3FCFB9D15078AF883" author="Tavares, M. S." box="[792,932,1890,1914]" pageId="3" pageNumber="49" pagination="3 - 4" refId="ref17956" refString="Tavares, M. S. (1991) Revision preliminaire du genre Tymolus Stimpson, avec la description de Tymolus brucei sp. nov. d'Australie occidentale (Crustacea, Brachyura, Cyclodorippoidea). Bulletin du Museum national d'Histoire naturelle, Series 4, 13, Section A (3 - 4), 439 - 456." type="book chapter" year="1991">Tavares, 1991</bibRefCitation>
</taxonomicName>
].—
<treatmentCitation id="ED0A10F4397CFFF3FC299D1500A9F883" author="Dai, A. - Y. &amp; Yang, S. - L. &amp; Song, Y. - Z. &amp; Chen, G. - X." box="[970,1159,1890,1914]" page="31" pageId="3" pageNumber="49" year="1986">
<bibRefCitation id="083A4B14397CFFF3FC299D1500A9F883" author="Dai, A. - Y. &amp; Yang, S. - L. &amp; Song, Y. - Z. &amp; Chen, G. - X." box="[970,1159,1890,1914]" pageId="3" pageNumber="49" refId="ref16080" refString="Dai, A. - Y., Yang, S. - L., Song, Y. - Z. &amp; Chen, G. - X. (1986) Crabs of the China Seas. China Ocean Press, Beijing, xi + 642 pp., 74 pls. [in Chinese]" type="book" year="1986">
Dai
<emphasis id="5EDFEAF7397CFFF3FC149D14000CF883" box="[1015,1058,1890,1914]" italics="true" pageId="3" pageNumber="49">et al</emphasis>
. 1986: 31
</bibRefCitation>
</treatmentCitation>
, pl. 3, fig. 13(2) [=
<taxonomicName id="ABAB4D66397CFFF3FAAB9D150526F861" ID-CoL="7DBPQ" authorityName="Tavares" authorityYear="1991" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="brucei">
<emphasis id="5EDFEAF7397CFFF3FAAB9D150526F861" italics="true" pageId="3" pageNumber="49">Tymolus brucei</emphasis>
</taxonomicName>
].—
<treatmentCitation id="ED0A10F4397CFFF3FEC89DF7066CF861" author="Takeda, M. &amp; Moosa, M. K." box="[299,578,1920,1945]" page="54" pageId="3" pageNumber="49" year="1990">
<bibRefCitation id="083A4B14397CFFF3FEC89DF7066CF861" author="Takeda, M. &amp; Moosa, M. K." box="[299,578,1920,1945]" pageId="3" pageNumber="49" pagination="53 - 71" refId="ref17868" refString="Takeda, M. &amp; Moosa, M. K. (1990) A small collection of deep-sea crabs from the Flores Sea. Indo-Malayan Zoology, 6, 53 - 71." type="journal article" year="1990">Takeda &amp; Moosa 1990: 54</bibRefCitation>
</treatmentCitation>
, 58. [=?]—
<treatmentCitation id="ED0A10F4397CFFF3FD589DF707B4F861" author="Dai, A. - Y. &amp; Yang, S. - L." box="[699,922,1920,1945]" page="36" pageId="3" pageNumber="49" year="1991">
<bibRefCitation id="083A4B14397CFFF3FD589DF707B4F861" author="Dai, A. - Y. &amp; Yang, S. - L." box="[699,922,1920,1945]" pageId="3" pageNumber="49" refId="ref16139" refString="Dai, A. - Y. &amp; Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing and Springer-Verlag, Berlin, 21 + 608 pp., 74 pls." type="book" year="1991">Dai &amp; Yang 1991: 36</bibRefCitation>
</treatmentCitation>
, pl. 3, fig. 13(2) [=
<taxonomicName id="ABAB4D66397CFFF3FB809DF700D0F861" ID-CoL="7DBPQ" authorityName="Tavares" authorityYear="1991" box="[1123,1278,1920,1944]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="brucei">
<emphasis id="5EDFEAF7397CFFF3FB809DF700D0F861" box="[1123,1278,1920,1944]" italics="true" pageId="3" pageNumber="49">Tymolus brucei</emphasis>
</taxonomicName>
].—
<treatmentCitation id="ED0A10F4397CFFF3FAC09DF704C1F84E" author="Chen, H. - L." page="231" pageId="3" pageNumber="49" year="1998">
<bibRefCitation id="083A4B14397CFFF3FAC09DF704C1F84E" author="Chen, H. - L." pageId="3" pageNumber="49" pagination="228 - 264" refId="ref15966" refString="Chen, H. - L. (1998) Further study on the crabs (Crustacea: Brachyura) from Nansha Islands and adjacent waters. In: Studies on Marine Fauna and Flora and Biogeography of the Nansha Islands and Neighbouring Waters. Vol. III. China Ocean Press, Beijing, pp. 228 - 264 [in Chinese with English abstract]" type="book chapter" year="1998">Chen 1998: 231</bibRefCitation>
</treatmentCitation>
, fig. 2 [=
<taxonomicName id="ABAB4D66397CFFF3FEB79DE805C1F84E" ID-CoL="7DBPQ" authorityName="Tavares" authorityYear="1991" box="[340,495,1951,1975]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="brucei">
<emphasis id="5EDFEAF7397CFFF3FEB79DE805C1F84E" box="[340,495,1951,1975]" italics="true" pageId="3" pageNumber="49">Tymolus brucei</emphasis>
</taxonomicName>
].—Tan &amp; Huang 2000: 139.—
<treatmentCitation id="ED0A10F4397CFFF3FCCF9DE807DBF84E" author="Ho, P. - H. &amp; Ng, P. K. L. &amp; Chan, T. - Y. &amp; Lee, D. - A." box="[812,1013,1951,1975]" page="643" pageId="3" pageNumber="49" year="2004">
<bibRefCitation id="083A4B14397CFFF3FCCF9DE807DBF84E" author="Ho, P. - H. &amp; Ng, P. K. L. &amp; Chan, T. - Y. &amp; Lee, D. - A." box="[812,1013,1951,1975]" pageId="3" pageNumber="49" pagination="641 - 668" refId="ref16412" refString="Ho, P. - H., Ng, P. K. L., Chan, T. - Y. &amp; Lee, D. - A. (2004) New records of 31 species of brachyuran crabs from the joint Taiwan- France expeditions, &quot; TAIWAN 2000 &quot; and &quot; TAIWAN 2001 &quot;, off deep waters in Taiwan. Crustaceana, 77, 641 - 668. https: // doi. org / 10.1163 / 1568540041958617" type="journal article" year="2004">
Ho
<emphasis id="5EDFEAF7397CFFF3FCB09DD707AEF84E" box="[851,896,1951,1975]" italics="true" pageId="3" pageNumber="49">et al</emphasis>
. 2004: 643
</bibRefCitation>
</treatmentCitation>
, fig. 1C [=?].—
<treatmentCitation id="ED0A10F4397CFFF3FB7A9DE801B6F84E" author="Ahyong, S. T. &amp; Naruse, T. &amp; Tan, S. H. &amp; Ng, P. K. L." box="[1177,1432,1951,1975]" page="170" pageId="3" pageNumber="49" year="2009">
<bibRefCitation id="083A4B14397CFFF3FB7A9DE801B6F84E" author="Ahyong, S. T. &amp; Naruse, T. &amp; Tan, S. H. &amp; Ng, P. K. L." box="[1177,1432,1951,1975]" pageId="3" pageNumber="49" refId="ref15475" refString="Ahyong, S. T., Naruse, T., Tan, S. H. &amp; Ng, P. K. L. (2009) Part II. Infraorder Brachyura: Sections Dromiacea, Raninoida, Cyclodorippoida. In: Chan, T. - Y., Ng, P. K. L., Ahyong, S. T. &amp; Tan, S. H. (Eds.), Crustacean Fauna of Taiwan: Brachyuran Crabs. Vol. 1. Carcinology in Taiwan and Dromiacea, Raninoida, Cyclodorippoida. National Taiwan Ocean University, Keelung, pp. 1 - 198." type="book" year="2009">
Ahyong
<emphasis id="5EDFEAF7397CFFF3FB159DD7010DF84E" box="[1270,1315,1951,1975]" italics="true" pageId="3" pageNumber="49">et al</emphasis>
. 2009: 170
</bibRefCitation>
</treatmentCitation>
, figs. 126128 [=?].—
<treatmentCitation id="ED0A10F4397CFFF3FE459DC9064FF82F" author="Ng, P. K. L. &amp; Shih, H. - T. &amp; Ho, P. - H. &amp; Wang, C. - H." box="[422,609,1982,2006]" page="24" pageId="3" pageNumber="49" year="2017">
<bibRefCitation id="083A4B14397CFFF3FE459DC9064FF82F" author="Ng, P. K. L. &amp; Shih, H. - T. &amp; Ho, P. - H. &amp; Wang, C. - H." box="[422,609,1982,2006]" pageId="3" pageNumber="49" pagination="1 - 185" refId="ref17170" refString="Ng, P. K. L., Shih, H. - T., Ho, P. - H. &amp; Wang, C. - H. (2017) An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum, 70 (3 / 4), 1 - 185. https: // doi. org / 10.6532 / JNTM. 201712 _ 70 (3; 4). 01" type="journal article" year="2017">
Ng
<emphasis id="5EDFEAF7397CFFF3FE2C9DC805D5F82F" box="[463,507,1982,2006]" italics="true" pageId="3" pageNumber="49">et al</emphasis>
. 2017: 24
</bibRefCitation>
</treatmentCitation>
(list) [=?]. See “Discussion”.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6C1436E5397CFFF3FF749DAB05E2F80D" blockId="3.[151,1437,1675,2036]" box="[151,460,2012,2036]" pageId="3" pageNumber="49">Records remain to be verified.</paragraph>
<paragraph id="6C1436E5397BFFF4FF749AE00668FF56" blockId="4.[151,1436,151,297]" box="[151,582,151,175]" pageId="4" pageNumber="50">
<treatmentCitationGroup id="4CBB11CB397BFFF4FF749AE00668FF56" box="[151,582,151,175]" pageId="4" pageNumber="50">
<taxonomicName id="ABAB4D66397BFFF4FF749AE00555FF56" ID-CoL="6BS8X" authorityName="Ortmann" authorityYear="1892" box="[151,379,151,175]" class="Malacostraca" family="Cyclodorippidae" genus="Cyclodorippe" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="uncifera">
<emphasis id="5EDFEAF7397BFFF4FF749AE00555FF56" box="[151,379,151,175]" italics="true" pageId="4" pageNumber="50">Cyclodorippe uncifera</emphasis>
</taxonomicName>
.—
<treatmentCitation id="ED0A10F4397BFFF4FE789AE0066CFF56" author="Balss, H." box="[411,578,151,175]" page="286" pageId="4" pageNumber="50" year="1922">
<bibRefCitation id="083A4B14397BFFF4FE789AE0066CFF56" author="Balss, H." box="[411,578,151,175]" pageId="4" pageNumber="50" pagination="104 - 140" refId="ref15882" refString="Balss, H. (1922) Ostasiatische Decapoden. III. Die Dromiaceen, Oxystomen und Parthenopiden. Archiv fu ¨ r Naturgeschichte, 88 (A), 104 - 140." type="journal article" year="1922">Balss 1922: 286</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6C1436E5397BFFF4FF749AC20097FF34" blockId="4.[151,1436,151,297]" box="[151,1209,181,205]" pageId="4" pageNumber="50">
<treatmentCitationGroup id="4CBB11CB397BFFF4FF749AC20097FF34" box="[151,1209,181,205]" pageId="4" pageNumber="50">
<taxonomicName id="ABAB4D66397BFFF4FF749AC20517FF34" ID-CoL="7DBPK" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,313,181,205]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF7397BFFF4FF749AC20517FF34" box="[151,313,181,205]" italics="true" pageId="4" pageNumber="50">Tymolus uncifer</emphasis>
</taxonomicName>
.—
<treatmentCitation id="ED0A10F4397BFFF4FEB59AC2063EFF34" author="Yokoya, Y." box="[342,528,181,205]" page="103" pageId="4" pageNumber="50" year="1933">
<bibRefCitation id="083A4B14397BFFF4FEB59AC2063EFF34" author="Yokoya, Y." box="[342,528,181,205]" pageId="4" pageNumber="50" pagination="1 - 226" refId="ref18152" refString="Yokoya, Y. (1933) On the distribution of decapod crustaceans inhabiting the continental shelf around Japan, based upon the materials collected by S. S. Soyo-maru, during the years 1923 - 1930. Journal of the College of Agriculture, University of Tokyo, 12 (1), 1 - 226." type="journal article" year="1933">Yokoya 1933: 103</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="ED0A10F4397BFFF4FDCD9AC206C5FF34" author="Takeda, M." box="[558,747,181,205]" page="232" pageId="4" pageNumber="50" year="1997">
<bibRefCitation id="083A4B14397BFFF4FDCD9AC206C5FF34" author="Takeda, M." box="[558,747,181,205]" pageId="4" pageNumber="50" pagination="229 - 255" refId="ref17762" refString="Takeda, M. (1997) Deep-sea decapod crustacean fauna of Suruga Bay, central Japan. National Science Museum Monographs, 12, pp. 229 - 255." type="journal article" year="1997">Takeda 1997: 232</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="ED0A10F4397BFFF4FD169AC20774FF34" author="Takeda, M." box="[757,858,181,205]" page="223" pageId="4" pageNumber="50" year="2001">
<bibRefCitation id="083A4B14397BFFF4FD169AC20774FF34" author="Takeda, M." box="[757,858,181,205]" pageId="4" pageNumber="50" pagination="217 - 262" refId="ref17793" refString="Takeda, M. (2001) Annotated list of crabs from Tosa Bay, Southwest Japan, collected by the R / V Kotaka Maru during the years 1997 - 2000. In: Fujita, T., Saito, T. &amp; Takeda, M. (Eds.), Deep-Sea Fauna and Pollutants in Tosa Bay. National Science Museum Monographs, 20, pp. 217 - 262." type="journal article" year="2001">2001: 223</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="ED0A10F4397BFFF4FC9B9AC2009BFF34" author="Marumura, M. &amp; Kosaka, A." box="[888,1205,181,205]" page="21" pageId="4" pageNumber="50" year="2003">
<bibRefCitation id="083A4B14397BFFF4FC9B9AC2009BFF34" author="Marumura, M. &amp; Kosaka, A." box="[888,1205,181,205]" pageId="4" pageNumber="50" refId="ref16939" refString="Marumura, M. &amp; Kosaka, A. (2003) Catalogue of the Brachyuran and Anomuran Crabs Donated by the Late Mr. Seiji Nagai to the Wakayama Prefectural Museum of Natural History. Wakayama Prefectural Museum of Natural History, Kainan, 73 pp. [in Japanese]" type="book" year="2003">Marumura &amp; Kosaka 2003: 21</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6C1436E5397BFFF4FF749AA3078AFF15" blockId="4.[151,1436,151,297]" box="[151,932,212,236]" pageId="4" pageNumber="50">Fossil records remain to be verified or in uncertain status (see “Discussion”).</paragraph>
<paragraph id="6C1436E5397BFFF4FF749A840582FED0" blockId="4.[151,1436,151,297]" pageId="4" pageNumber="50">
<treatmentCitationGroup id="4CBB11CB397BFFF4FF749A840582FED0" pageId="4" pageNumber="50">
<taxonomicName id="ABAB4D66397BFFF4FF749A840515FEF2" ID-CoL="7DBPK" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,315,243,267]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF7397BFFF4FF749A840515FEF2" box="[151,315,243,267]" italics="true" pageId="4" pageNumber="50">Tymolus uncifer</emphasis>
</taxonomicName>
.—
<treatmentCitation id="ED0A10F4397BFFF4FEBB9A84060DFEF2" author="Karasawa, H." box="[344,547,243,267]" page="43" pageId="4" pageNumber="50" year="1993">
<bibRefCitation id="083A4B14397BFFF4FEBB9A84060DFEF2" author="Karasawa, H." box="[344,547,243,267]" pageId="4" pageNumber="50" pagination="1 - 92" refId="ref16586" refString="Karasawa, H. (1993) Cenozoic Decapod Crustacea from Southwest Japan. Bulletin of the Mizunami Fossil Museum, 20, 1 - 92." type="journal article" year="1993">Karasawa 1993: 43</bibRefCitation>
</treatmentCitation>
, pl. 6, fig. 4.—
<treatmentCitation id="ED0A10F4397BFFF4FD5C9A8407A4FEF2" author="Karasawa, H." box="[703,906,243,267]" page="16" pageId="4" pageNumber="50" year="1997">
<bibRefCitation id="083A4B14397BFFF4FD5C9A8407A4FEF2" author="Karasawa, H." box="[703,906,243,267]" pageId="4" pageNumber="50" pagination="1 - 81" refId="ref16613" refString="Karasawa, H. (1997) A monograph of Cenozoic stomatopod, decapod, isopod and amphipod Crustacea from West Japan. Monograph of the Mizunami Fossil Museum, 8, 1 - 81." type="journal article" year="1997">Karasawa 1997: 16</bibRefCitation>
</treatmentCitation>
, 41, pl. 8, fig. 10.—
<treatmentCitation id="ED0A10F4397BFFF4FBBA9A840155FEF3" author="Kitamura, K. &amp; Shiba, M." box="[1113,1403,242,267]" page="15" pageId="4" pageNumber="50" year="2008">
<bibRefCitation id="083A4B14397BFFF4FBBA9A840155FEF3" author="Kitamura, K. &amp; Shiba, M." box="[1113,1403,242,267]" pageId="4" pageNumber="50" pagination="11 - 18" refId="ref16649" refString="Kitamura, K. &amp; Shiba, M. (2008) Fossil decapod crustaceans from the Hijikata Formation of the Kakegawa Group, distributed in Nagaya, Kakegawa City, Shizuoka Prefecture, Central Japan. Science Reports of the Museum, Tokai University, 9, 11 - 18." type="journal article" year="2008">Kitamura &amp; Shiba 2008: 15</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="ED0A10F4397BFFF4FF249B660586FED0" ID-DOI="http://doi.org/10.5281/zenodo.4426214" ID-Zenodo-Dep="4426214" author="Ando, Y. &amp; Kawano, S. &amp; Muramiya, Y. &amp; Niiyama, S. &amp; Kameyama, S. &amp; Shimoyama, S." box="[199,424,273,297]" httpUri="http://treatment.plazi.org/id/03C987D66A0EB85D83E9FDD95C58FC76" page="532" pageId="4" pageNumber="50" year="2020">
<bibRefCitation id="083A4B14397BFFF4FF249B660586FED0" author="Ando, Y. &amp; Kawano, S. &amp; Muramiya, Y. &amp; Niiyama, S. &amp; Kameyama, S. &amp; Shimoyama, S." box="[199,424,273,297]" pageId="4" pageNumber="50" pagination="523 - 541" refId="ref15796" refString="Ando, Y., Kawano, S., Muramiya, Y., Niiyama, S., Kameyama, S. &amp; Shimoyama, S. (2020) Fossil decapods from the Upper Quaternary in Shinjima Island in Kagoshima, Kyushu, Japan, and description of a new species of ghost shrimp (Axiidea, Eucalliacidae). Zootaxa, 4878 (3), 523 - 541. https: // doi. org / 10.11646 / zootaxa. 4878.3.5" type="journal article" year="2020">
Ando
<emphasis id="5EDFEAF7397BFFF4FEE59B65051CFED0" box="[262,306,273,297]" italics="true" pageId="4" pageNumber="50">et al</emphasis>
. 2020: 532
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="38D4666D397BFFF4FF749E6B05D2FBA1" ID-DOI="http://doi.org/10.5281/zenodo.8406962" ID-Zenodo-Dep="8406962" httpUri="https://zenodo.org/record/8406962/files/figure.png" pageId="4" pageNumber="50" startId="4.[151,250,1052,1076]" targetBox="[151,1436,322,1029]" targetPageId="4" targetType="figure">
<paragraph id="6C1436E5397BFFF4FF749E6B05D2FBA1" blockId="4.[151,1436,1052,1112]" pageId="4" pageNumber="50">
<emphasis id="5EDFEAF7397BFFF4FF749E6B053BFBCD" bold="true" box="[151,277,1052,1076]" pageId="4" pageNumber="50">FIGURE 2.</emphasis>
<taxonomicName id="ABAB4D66397BFFF4FEFD9E6B0654FBCD" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[286,634,1052,1076]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF7397BFFF4FEFD9E6B05EDFBCD" box="[286,451,1052,1076]" italics="true" pageId="4" pageNumber="50">Tymolus uncifer</emphasis>
(
<bibRefCitation id="083A4B14397BFFF4FE319E6B065DFBCD" author="Ortmann, A. E." box="[466,627,1052,1076]" pageId="4" pageNumber="50" pagination="532 - 588" refId="ref17259" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit bedonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit Strassburger Museum aufwbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
)
</taxonomicName>
, lectotype, male (5.0×5.0 mm), MZS Cru 3187. A, cephalothorax and pleon, dorsal view; B, same, ventral view.
</paragraph>
</caption>
<caption id="38D4666D397BFFF4FF749CE60728F937" ID-DOI="http://doi.org/10.5281/zenodo.8406964" ID-Zenodo-Dep="8406964" httpUri="https://zenodo.org/record/8406964/files/figure.png" pageId="4" pageNumber="50" startId="4.[151,250,1681,1705]" targetBox="[151,1435,1136,1658]" targetPageId="4" targetType="figure">
<paragraph id="6C1436E5397BFFF4FF749CE60728F937" blockId="4.[151,1436,1681,1742]" pageId="4" pageNumber="50">
<emphasis id="5EDFEAF7397BFFF4FF749CE6053BF950" bold="true" box="[151,277,1681,1705]" pageId="4" pageNumber="50">FIGURE 3.</emphasis>
<taxonomicName id="ABAB4D66397BFFF4FEFE9CE60686F953" authority="(Doflein, 1904)" baseAuthorityName="Doflein" baseAuthorityYear="1904" box="[285,680,1681,1706]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="melanommus">
<emphasis id="5EDFEAF7397BFFF4FEFE9CE605D1F950" box="[285,511,1681,1705]" italics="true" pageId="4" pageNumber="50">Tymolus melanommus</emphasis>
(
<bibRefCitation id="083A4B14397BFFF4FDED9CE6068FF953" author="Doflein, F." box="[526,673,1681,1706]" pageId="4" pageNumber="50" pagination="1 - 6" refId="ref16210" refString="Doflein, F. (1904) Brachyura. Wissenschaftliche Ergebnisse der Deutchen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899, 6 (1 - 6), i - xiv + 1 - 314, atlas (pls. 311 - 358)." type="journal article" year="1904">Doflein, 1904</bibRefCitation>
)
</taxonomicName>
, lectotype, male (7.5×7.0 mm), ZSM 393/1. A, cephalothorax and pleon, dorsal view; B, detached left cheliped and pereopods 2 and 3.
</paragraph>
</caption>
<subSubSection id="24B1656E397BFFF6FF749D68016FFE70" lastPageId="6" lastPageNumber="52" pageId="4" pageNumber="50" type="materials_examined">
<paragraph id="6C1436E5397BFFF4FF749D6806ABF8A7" blockId="4.[151,1437,1823,2030]" pageId="4" pageNumber="50">
<emphasis id="5EDFEAF7397BFFF4FF749D6805AEF8C0" bold="true" box="[151,384,1823,1849]" pageId="4" pageNumber="50">Material examined.</emphasis>
<materialsCitation id="DCC33CB8397BFFF4FE649D5706ABF8A7" ID-GBIF-Occurrence="4421509322" collectionCode="MZS" collectorName="L. Doderlein" elevation="180" location="Cru" municipality="Sagami Bay" pageId="4" pageNumber="50" specimenCount="1" typeStatus="lectotype">
<typeStatus id="B3108847397BFFF4FE649D5705D5F8C3" box="[391,507,1824,1850]" pageId="4" pageNumber="50" type="lectotype">Lectotype</typeStatus>
.
<collectionCode id="0ABAAE20397BFFF4FDE79D570610F8C3" box="[516,574,1824,1850]" country="France" httpUri="http://grbio.org/cool/6hz5-fzvb" name="Universite de Strasbourg, Musee de Zoologie" pageId="4" pageNumber="50">MZS</collectionCode>
<location id="6974603E397BFFF4FDA69D57065FF8C3" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6:6974603E397BFFF4FDA69D57065FF8C3" box="[581,625,1824,1850]" municipality="Sagami Bay" name="Cru" pageId="4" pageNumber="50">Cru</location>
3187, male (5.0×5.0 mm),
<collectingMunicipality id="8C70AC9F397BFFF4FC799D57000CF8C3" box="[922,1058,1824,1850]" pageId="4" pageNumber="50">Sagami Bay</collectingMunicipality>
, 100 fathoms (
<quantity id="AB539B00397BFFF4FB249D57013CF8C3" box="[1223,1298,1824,1850]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="50" unit="m" value="180.0">
<elevation id="E786D1D6397BFFF4FB249D57013CF8C3" box="[1223,1298,1824,1850]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="50" unit="m" value="180.0">180 m</elevation>
</quantity>
), 1882, coll.
<collectorName id="C15E5333397BFFF4FF749D330507F8A7" box="[151,297,1860,1886]" pageId="4" pageNumber="50">L. Döderlein</collectorName>
, designated by
<bibRefCitation id="083A4B14397BFFF4FE369D3306AFF8A7" author="Tavares, M. S." box="[469,641,1860,1886]" pageId="4" pageNumber="50" pagination="201 - 207" refId="ref18016" refString="Tavares, M. S. (1992) Revalidation de Tymolus dromioides (Ortmann, 1892) (Crustacea, Decapoda, Brachyura, Cyclodorippidae). Bulletin du Museum national d'Histoire naturelle, Paris, Series 4, 14, section A, (1), 201 - 207. https: // doi. org / 10.5962 / p. 289871" type="journal article" year="1992">Tavares (1992)</bibRefCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="6C1436E5397BFFF4FF249D1F050AF817" blockId="4.[151,1437,1823,2030]" pageId="4" pageNumber="50">
<materialsCitation id="DCC33CB8397BFFF4FF249D1F0505F85F" ID-GBIF-Occurrence="4421509331" collectionCode="MZS" collectorName="L. Doderlein" elevation="225" location="Cru" municipality="Sagami Bay" pageId="4" pageNumber="50" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<typeStatus id="B3108847397BFFF4FF249D1F0540F87B" box="[199,366,1896,1922]" pageId="4" pageNumber="50" type="paralectotype">Paralectotypes</typeStatus>
.
<collectionCode id="0ABAAE20397BFFF4FE999D1F059AF87B" box="[378,436,1896,1922]" country="France" httpUri="http://grbio.org/cool/6hz5-fzvb" name="Universite de Strasbourg, Musee de Zoologie" pageId="4" pageNumber="50">MZS</collectionCode>
<location id="6974603E397BFFF4FE5F9D1005C6F87B" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6:6974603E397BFFF4FE5F9D1005C6F87B" box="[444,488,1895,1922]" municipality="Sagami Bay" name="Cru" pageId="4" pageNumber="50">Cru</location>
569,
<specimenCount id="7AADFD6C397BFFF4FDC39D1F0643F87B" box="[544,621,1896,1922]" pageId="4" pageNumber="50" type="male">1 male</specimenCount>
(8.6×8.2 mmm),
<collectingMunicipality id="8C70AC9F397BFFF4FCDB9D1F07ECF87B" box="[824,962,1896,1922]" pageId="4" pageNumber="50">Sagami Bay</collectingMunicipality>
, 50200 fathoms (
<quantity id="AB539B00397BFFF4FB759D100120F87B" box="[1174,1294,1895,1922]" metricMagnitude="2" metricUnit="m" metricValue="2.25" metricValueMax="3.6" metricValueMin="0.9" pageId="4" pageNumber="50" unit="m" value="225.0" valueMax="360.0" valueMin="90.0">
<elevation id="E786D1D6397BFFF4FB759D100120F87B" box="[1174,1294,1895,1922]" metricMagnitude="2" metricUnit="m" metricValue="2.25" metricValueMax="3.6" metricValueMin="0.9" pageId="4" pageNumber="50" unit="m" value="225.0" valueMax="360.0" valueMin="90.0">90360 m</elevation>
</quantity>
), 1882, coll.
<collectorName id="C15E5333397BFFF4FF749DFB0505F85F" box="[151,299,1932,1958]" pageId="4" pageNumber="50">L. Döderlein</collectorName>
</materialsCitation>
;
<materialsCitation id="DCC33CB8397BFFF4FED49DFB075EF85F" ID-GBIF-Occurrence="4421509307" box="[311,880,1932,1958]" collectionCode="MZS" collectorName="L. Doderlein" elevation="225" location="Cru" municipality="Sagami Bay" pageId="4" pageNumber="50" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<collectionCode id="0ABAAE20397BFFF4FED49DFB055FF85F" box="[311,369,1932,1958]" country="France" httpUri="http://grbio.org/cool/6hz5-fzvb" name="Universite de Strasbourg, Musee de Zoologie" pageId="4" pageNumber="50">MZS</collectionCode>
<location id="6974603E397BFFF4FE9B9DFB058DF85F" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6:6974603E397BFFF4FE9B9DFB058DF85F" box="[376,419,1932,1958]" name="Cru" pageId="4" pageNumber="50">Cru</location>
4224,
<specimenCount id="7AADFD6C397BFFF4FE0A9DFB0664F85F" box="[489,586,1932,1958]" pageId="4" pageNumber="50" type="female">1 female</specimenCount>
(7.8×
<quantity id="AB539B00397BFFF4FD6E9DFB06C7F85F" box="[653,745,1932,1958]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="4" pageNumber="50" unit="mm" value="7.8">7.8 mm</quantity>
), same data
</materialsCitation>
;
<materialsCitation id="DCC33CB8397BFFF4FC9F9DFB078AF833" ID-GBIF-Occurrence="4421509330" collectionCode="MZS" collectorName="L. Doderlein" elevation="180" location="Cru" municipality="Cru" pageId="4" pageNumber="50" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paralectotype">
<collectionCode id="0ABAAE20397BFFF4FC9F9DFB0798F85F" box="[892,950,1932,1958]" country="France" httpUri="http://grbio.org/cool/6hz5-fzvb" name="Universite de Strasbourg, Musee de Zoologie" pageId="4" pageNumber="50">MZS</collectionCode>
<location id="6974603E397BFFF4FC5D9DFC07C4F85F" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6:6974603E397BFFF4FC5D9DFC07C4F85F" box="[958,1002,1931,1958]" municipality="Cru" name="Cru" pageId="4" pageNumber="50">Cru</location>
570,
<collectingMunicipality id="8C70AC9F397BFFF4FBC39DFC0065F85F" box="[1056,1099,1931,1958]" pageId="4" pageNumber="50">Cru</collectingMunicipality>
42254228,
<specimenCount id="7AADFD6C397BFFF4FB349DFB0100F85F" box="[1239,1326,1932,1958]" pageId="4" pageNumber="50" type="male">4 males</specimenCount>
(4.0×4.0, 4.2×4.2, 5.0×
<quantity id="AB539B00397BFFF4FECE9DD805A9F833" box="[301,391,1967,1994]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="4" pageNumber="50" unit="mm" value="5.2">5.2 mm</quantity>
),
<specimenCount id="7AADFD6C397BFFF4FE7A9DC705D7F833" box="[409,505,1968,1994]" pageId="4" pageNumber="50" type="female">1 female</specimenCount>
(4.0×
<quantity id="AB539B00397BFFF4FDD89DC706BBF833" box="[571,661,1968,1994]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="50" unit="mm" value="4.2">4.2 mm</quantity>
), same data as lectotype
</materialsCitation>
;
<materialsCitation id="DCC33CB8397BFFF4FC4C9DC7050EF817" ID-GBIF-Occurrence="4421509319" collectionCode="MZS" collectorName="L. Doderlein" elevation="180" location="Cru" municipality="Sagami Bay" pageId="4" pageNumber="50" specimenCode="MZS 571" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCode id="3C0D9E9E397BFFF4FC4C9DC70035F833" box="[943,1051,1967,1994]" collectionCode="MZS" country="France" httpUri="http://grbio.org/cool/6hz5-fzvb" name="Universite de Strasbourg, Musee de Zoologie" pageId="4" pageNumber="50">MZS 571</specimenCode>
,
<specimenCount id="7AADFD6C397BFFF4FBC49DC7005FF833" box="[1063,1137,1968,1994]" pageId="4" pageNumber="50" type="male">1 male</specimenCount>
(not measured), same data as lectotype
</materialsCitation>
.
</paragraph>
<caption id="38D4666D397AFFF5FF749CD706E3F925" ID-DOI="http://doi.org/10.5281/zenodo.8406966" ID-Zenodo-Dep="8406966" httpUri="https://zenodo.org/record/8406966/files/figure.png" pageId="5" pageNumber="51" startId="5.[151,250,1696,1720]" targetBox="[156,1431,181,1672]" targetPageId="5" targetType="figure">
<paragraph id="6C1436E5397AFFF5FF749CD706E3F925" blockId="5.[151,1436,1696,1756]" pageId="5" pageNumber="51">
<emphasis id="5EDFEAF7397AFFF5FF749CD7053AF941" bold="true" box="[151,276,1696,1720]" pageId="5" pageNumber="51">FIGURE 4.</emphasis>
<taxonomicName id="ABAB4D66397AFFF5FEFF9CD7065AF941" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[284,628,1696,1720]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF7397AFFF5FEFF9CD705EEF941" box="[284,448,1696,1720]" italics="true" pageId="5" pageNumber="51">Tymolus uncifer</emphasis>
(
<bibRefCitation id="083A4B14397AFFF5FE2D9CD70643F941" author="Ortmann, A. E." box="[462,621,1696,1720]" pageId="5" pageNumber="51" pagination="532 - 588" refId="ref17259" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit bedonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit Strassburger Museum aufwbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
)
</taxonomicName>
, habitus in dorsal view. A, male (9.8×9.8 mm), CBM-ZC 16951; B, ovigerous female (7.6×7.6 mm), CBM-ZC 16943. In preservative.
</paragraph>
</caption>
<paragraph id="6C1436E5397AFFF6FF249D59016FFE70" blockId="5.[151,1436,1838,2009]" lastBlockId="6.[151,1437,151,393]" lastPageId="6" lastPageNumber="52" pageId="5" pageNumber="51">
Other material.
<materialsCitation id="DCC33CB8397AFFF5FE9B9D580691F869" ID-GBIF-Occurrence="4421509312" collectingDate="1996-10-24" collectionCode="CBM-ZC" elevation="325" latitude="34.998165" location="Tateyama Bay" longLatPrecision="11" longitude="139.76917" pageId="5" pageNumber="51" specimenCode="CBM-ZC 4507" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" stateProvince="Chiba">
<specimenCode id="3C0D9E9E397AFFF5FE9B9D58060BF8B0" box="[376,549,1838,1865]" collectionCode="CBM-ZC" pageId="5" pageNumber="51">CBM-ZC 4507</specimenCode>
,
<specimenCount id="7AADFD6C397AFFF5FDD39D5806BFF8B1" box="[560,657,1838,1865]" pageId="5" pageNumber="51" type="female">1 female</specimenCount>
(4.9×
<quantity id="AB539B00397AFFF5FD379D590700F8B0" box="[724,814,1838,1865]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="51" unit="mm" value="5.1">5.1 mm</quantity>
), 1 ovigerous female (6.9×
<quantity id="AB539B00397AFFF5FBBA9D59009AF8B0" box="[1113,1204,1838,1865]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="5" pageNumber="51" unit="mm" value="7.2">7.2 mm</quantity>
),
<specimenCount id="7AADFD6C397AFFF5FB249D580118F8B1" box="[1223,1334,1838,1865]" pageId="5" pageNumber="51" type="juvenile">1 juvenile</specimenCount>
(3.5×
<quantity id="AB539B00397AFFF5FA9B9D5904E9F894" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="51" unit="mm" value="3.6">3.6 mm</quantity>
),
<location id="6974603E397AFFF5FF3A9D240554F894" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6:6974603E397AFFF5FF3A9D240554F894" box="[217,378,1875,1901]" latitude="34.998165" longLatPrecision="11" longitude="139.76917" name="Tateyama Bay" pageId="5" pageNumber="51" stateProvince="Chiba">Tateyama Bay</location>
,
<collectingRegion id="AE6FF807397AFFF5FE669D25066AF895" box="[389,580,1874,1900]" country="Japan" name="Chiba" pageId="5" pageNumber="51">Chiba Prefecture</collectingRegion>
, TRV
<emphasis id="5EDFEAF7397AFFF5FD699D25070AF895" box="[650,804,1874,1900]" italics="true" pageId="5" pageNumber="51">Shinyo-maru</emphasis>
, 1996 research cruise, stn 14,
<geoCoordinate id="099F5022397AFFF5FB969D2500D2F894" box="[1141,1276,1874,1901]" degrees="34" direction="north" minutes="59.89" orientation="latitude" pageId="5" pageNumber="51" precision="9" value="34.998165">34°59.89N</geoCoordinate>
,
<geoCoordinate id="099F5022397AFFF5FAE49D2501B6F894" box="[1287,1432,1874,1901]" degrees="139" direction="east" minutes="46.15" orientation="longitude" pageId="5" pageNumber="51" precision="9" value="139.76917">139°46.15E</geoCoordinate>
,
<quantity id="AB539B00397AFFF5FF749D010534F868" box="[151,282,1910,1937]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.9" metricValueMin="2.6" pageId="5" pageNumber="51" unit="m" value="325.0" valueMax="390.0" valueMin="260.0">
<elevation id="E786D1D6397AFFF5FF749D010534F868" box="[151,282,1910,1937]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.9" metricValueMin="2.6" pageId="5" pageNumber="51" unit="m" value="325.0" valueMax="390.0" valueMin="260.0">260390 m</elevation>
</quantity>
, mud,
<date id="18151025397AFFF5FE8B9D000604F869" box="[360,554,1910,1937]" pageId="5" pageNumber="51" value="1996-10-24">
<collectingDate id="0851E9CD397AFFF5FE8B9D000604F869" box="[360,554,1910,1937]" pageId="5" pageNumber="51" value="1996-10-24">24 October 1996</collectingDate>
</date>
, dredge, coll
</materialsCitation>
.
<materialsCitation id="DCC33CB8397AFFF5FD289D0000E1F868" ID-GBIF-Occurrence="4421509335" box="[715,1231,1910,1937]" collectionCode="T" collectorName="Tan &amp; Huang" location="Komai" pageId="5" pageNumber="51" specimenCount="1" stateProvince="Chiba">
<collectionCode id="0ABAAE20397AFFF5FD289D0006F3F868" box="[715,733,1911,1937]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="51">T</collectionCode>
.
<location id="6974603E397AFFF5FD0A9D010716F869" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6:6974603E397AFFF5FD0A9D010716F869" box="[745,824,1910,1936]" name="Komai" pageId="5" pageNumber="51" stateProvince="Chiba">Komai</location>
, examined by
<collectorName id="C15E5333397AFFF5FC3D9D000026F868" box="[990,1032,1911,1937]" pageId="5" pageNumber="51">Tan</collectorName>
&amp;
<collectorName id="C15E5333397AFFF5FBCD9D000057F868" box="[1070,1145,1911,1937]" pageId="5" pageNumber="51">Huang</collectorName>
(2000)
</materialsCitation>
;
<materialsCitation id="DCC33CB8397AFFF5FB3F9D000000F821" ID-GBIF-Occurrence="4421509310" collectingDate="2017-02-15" collectionCode="RV" collectorName="Rinkai-maru" elevation="240" latitude="35.116665" location="Sagami Bay" longLatPrecision="11" longitude="139.57" pageId="5" pageNumber="51" specimenCode="CBM-ZC 16734" specimenCount="1" specimenCount-male="1" stateProvince="Kanagawa">
<specimenCode id="3C0D9E9E397AFFF5FB3F9D0001B6F869" box="[1244,1432,1910,1937]" collectionCode="CBM-ZC" pageId="5" pageNumber="51">CBM-ZC 16734</specimenCode>
,
<specimenCount id="7AADFD6C397AFFF5FF749DEC04CBF84D" box="[151,229,1946,1973]" pageId="5" pageNumber="51" type="male">1 male</specimenCount>
(7.1×
<quantity id="AB539B00397AFFF5FEC89DED05A7F84C" box="[299,393,1946,1973]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="5" pageNumber="51" unit="mm" value="6.7">6.7 mm</quantity>
),
<location id="6974603E397AFFF5FE439DED0602F84C" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6:6974603E397AFFF5FE439DED0602F84C" box="[416,556,1946,1973]" latitude="35.116665" longLatPrecision="11" longitude="139.57" name="Sagami Bay" pageId="5" pageNumber="51" stateProvince="Kanagawa">Sagami Bay</location>
, SW of
<locationDeviation id="7DA62FC7397AFFF5FD739DED0727F84D" box="[656,777,1946,1972]" pageId="5" pageNumber="51">Jogashima</locationDeviation>
,
<locationDeviation id="7DA62FC7397AFFF5FCF49DED0770F84D" box="[791,862,1946,1972]" pageId="5" pageNumber="51">Miura</locationDeviation>
,
<collectingRegion id="AE6FF807397AFFF5FC8E9DEC004DF84D" box="[877,1123,1946,1973]" country="Japan" name="Kanagawa" pageId="5" pageNumber="51">Kanagawa Prefecture</collectingRegion>
,
<geoCoordinate id="099F5022397AFFF5FB919DED00D6F84C" box="[1138,1272,1946,1973]" degrees="35" direction="north" minutes="07.00" orientation="latitude" pageId="5" pageNumber="51" precision="9" value="35.116665">35°07.00N</geoCoordinate>
,
<geoCoordinate id="099F5022397AFFF5FAE49DEC01B6F84C" box="[1287,1432,1947,1973]" degrees="139" direction="east" minutes="34.20" orientation="longitude" pageId="5" pageNumber="51" precision="9" value="139.57">139°34.20E</geoCoordinate>
,
<quantity id="AB539B00397AFFF5FF749DC90533F820" box="[151,285,1982,2009]" metricMagnitude="2" metricUnit="m" metricValue="2.4" metricValueMax="3.2299999999999995" metricValueMin="1.5699999999999998" pageId="5" pageNumber="51" unit="m" value="240.0" valueMax="323.0" valueMin="157.0">
<elevation id="E786D1D6397AFFF5FF749DC90533F820" box="[151,285,1982,2009]" metricMagnitude="2" metricUnit="m" metricValue="2.4" metricValueMax="3.2299999999999995" metricValueMin="1.5699999999999998" pageId="5" pageNumber="51" unit="m" value="240.0" valueMax="323.0" valueMin="157.0">157323 m</elevation>
</quantity>
, mud and sand,
<date id="18151025397AFFF5FE079DC90696F820" box="[484,696,1982,2009]" pageId="5" pageNumber="51" value="2017-02-15">
<collectingDate id="0851E9CD397AFFF5FE079DC90696F820" box="[484,696,1982,2009]" pageId="5" pageNumber="51" value="2017-02-15">15 February 2017</collectingDate>
</date>
, dredge (
<collectionCode id="0ABAAE20397AFFF5FCC59DC80762F820" box="[806,844,1983,2009]" country="Italy" name="Collection of Leptospira Strains" pageId="5" pageNumber="51">RV</collectionCode>
<emphasis id="5EDFEAF7397AFFF5FCBB9DC907C8F821" box="[856,998,1982,2008]" italics="true" pageId="5" pageNumber="51">
<collectorName id="C15E5333397AFFF5FCBB9DC907C8F821" box="[856,998,1982,2008]" pageId="5" pageNumber="51">Rinkai-maru</collectorName>
</emphasis>
), coll
</materialsCitation>
.
<materialsCitation id="DCC33CB8397AFFF5FBDD9DC80081F821" ID-GBIF-Occurrence="4421509320" box="[1086,1199,1982,2009]" collectionCode="T" collectorName="Komai" pageId="5" pageNumber="51" specimenCount="1" stateProvince="Kanagawa">
<collectionCode id="0ABAAE20397AFFF5FBDD9DC8007EF820" box="[1086,1104,1983,2009]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="51">T</collectionCode>
.
<collectorName id="C15E5333397AFFF5FBBC9DC90081F821" box="[1119,1199,1982,2008]" pageId="5" pageNumber="51">Komai</collectorName>
</materialsCitation>
;
<materialsCitation id="DCC33CB8397AFFF6FB5C9DC804EAFF00" ID-GBIF-Occurrence="4421509316" collectingDate="2016-02-18" collectionCode="RV" collectorName="Rinkai-maru" elevation="532" lastPageId="6" lastPageNumber="52" latitude="35.116665" location="Sagami Bay" longLatPrecision="11" longitude="139.55867" pageId="5" pageNumber="51" specimenCode="CBM-ZC 16943" specimenCount="1" specimenCount-female="1" stateProvince="Kanagawa">
<specimenCode id="3C0D9E9E397AFFF5FB5C9DC80151F821" box="[1215,1407,1982,2009]" collectionCode="CBM-ZC" pageId="5" pageNumber="51">CBM-ZC 16943</specimenCode>
,
<specimenCount id="7AADFD6C397AFFF6FA6D9DC804CDFF48" lastPageId="6" lastPageNumber="52" pageId="5" pageNumber="51" type="female">1 female</specimenCount>
(cl
<quantity id="AB539B003979FFF6FEEC9AE00546FF48" box="[271,360,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="6" pageNumber="52" unit="mm" value="8.1">8.1 mm</quantity>
; infested by bopyrid), 1 ovigerous female (7.6×
<quantity id="AB539B003979FFF6FC9F9AE007F9FF48" box="[892,983,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="6" pageNumber="52" unit="mm" value="7.6">7.6 mm</quantity>
),
<location id="6974603E3979FFF6FC099AE0005DFF48" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6:6974603E3979FFF6FC099AE0005DFF48" box="[1002,1139,151,177]" latitude="35.116665" longLatPrecision="11" longitude="139.55867" name="Sagami Bay" pageId="6" pageNumber="52" stateProvince="Kanagawa">Sagami Bay</location>
, SW of
<locationDeviation id="7DA62FC73979FFF6FB2E9AE00168FF48" box="[1229,1350,151,177]" pageId="6" pageNumber="52">Jogashima</locationDeviation>
,
<locationDeviation id="7DA62FC73979FFF6FAB29AE001B6FF48" box="[1361,1432,151,177]" pageId="6" pageNumber="52">Miura</locationDeviation>
,
<collectingRegion id="AE6FF8073979FFF6FF749ACC05A2FF2C" box="[151,396,187,213]" country="Japan" name="Kanagawa" pageId="6" pageNumber="52">Kanagawa Prefecture</collectingRegion>
,
<geoCoordinate id="099F50223979FFF6FE7B9ACD0630FF2C" box="[408,542,186,213]" degrees="35" direction="north" minutes="07.00" orientation="latitude" pageId="6" pageNumber="52" precision="9" value="35.116665">35°07.00N</geoCoordinate>
,
<geoCoordinate id="099F50223979FFF6FDC89ACD0693FF2C" box="[555,701,186,213]" degrees="139" direction="east" minutes="33.52" orientation="longitude" pageId="6" pageNumber="52" precision="9" value="139.55867">139°33.52E</geoCoordinate>
,
<quantity id="AB539B003979FFF6FD299ACD0762FF2C" box="[714,844,186,213]" metricMagnitude="2" metricUnit="m" metricValue="5.32" metricValueMax="5.55" metricValueMin="5.09" pageId="6" pageNumber="52" unit="m" value="532.0" valueMax="555.0" valueMin="509.0">
<elevation id="E786D1D63979FFF6FD299ACD0762FF2C" box="[714,844,186,213]" metricMagnitude="2" metricUnit="m" metricValue="5.32" metricValueMax="5.55" metricValueMin="5.09" pageId="6" pageNumber="52" unit="m" value="532.0" valueMax="555.0" valueMin="509.0">509555 m</elevation>
</quantity>
, mud,
<date id="181510253979FFF6FC799ACC0049FF2C" box="[922,1127,187,213]" pageId="6" pageNumber="52" value="2016-02-18">
<collectingDate id="0851E9CD3979FFF6FC799ACC0049FF2C" box="[922,1127,187,213]" pageId="6" pageNumber="52" value="2016-02-18">18 February 2016</collectingDate>
</date>
, dredge (
<collectionCode id="0ABAAE203979FFF6FB339ACC00D8FF2C" box="[1232,1270,187,213]" country="Italy" name="Collection of Leptospira Strains" pageId="6" pageNumber="52">RV</collectionCode>
<emphasis id="5EDFEAF73979FFF6FB1D9ACC01A2FF2C" box="[1278,1420,187,213]" italics="true" pageId="6" pageNumber="52">
<collectorName id="C15E53333979FFF6FB1D9ACC01A2FF2C" box="[1278,1420,187,213]" pageId="6" pageNumber="52">Rinkai-maru</collectorName>
</emphasis>
), coll
</materialsCitation>
.
<materialsCitation id="DCC33CB83979FFF6FF2C9AA80512FF00" ID-GBIF-Occurrence="4421509336" box="[207,316,223,249]" collectionCode="T" collectorName="Komai" pageId="6" pageNumber="52" specimenCount="1" stateProvince="Kanagawa">
<collectionCode id="0ABAAE203979FFF6FF2C9AA804CFFF00" box="[207,225,223,249]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="52">T</collectionCode>
.
<collectorName id="C15E53333979FFF6FF0F9AA80512FF00" box="[236,316,223,249]" pageId="6" pageNumber="52">Komai</collectorName>
</materialsCitation>
;
<materialsCitation id="DCC33CB83979FFF6FEA49AA8005AFF00" ID-GBIF-Occurrence="4421509303" box="[327,1140,222,249]" collectionCode="T" collectorName="Komai" pageId="6" pageNumber="52" specimenCode="CBM-ZC 16944, 2" specimenCount="1" stateProvince="Kanagawa">
<specimenCode id="3C0D9E9E3979FFF6FEA49AA8062CFF00" box="[327,514,223,249]" collectionCode="CBM-ZC" pageId="6" pageNumber="52">CBM-ZC 16944</specimenCode>
,
<specimenCode id="3C0D9E9E3979FFF6FDEE9AA80635FF00" box="[525,539,223,249]" collectionCode="CBM-ZC" pageId="6" pageNumber="52">2</specimenCode>
ovigerous females (7.2×7.6, 7.4×
<quantity id="AB539B003979FFF6FC769AA907C1FF00" box="[917,1007,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="52" unit="mm" value="7.5">7.5 mm</quantity>
), same data
</materialsCitation>
;
<materialsCitation id="DCC33CB83979FFF6FB639AA80681FE9C" ID-GBIF-Occurrence="4421509329" collectingDate="2002-10-23" collectionCode="CBM-ZC" elevation="486" latitude="35.0085" location="Izu Islands" longLatPrecision="11" longitude="139.205" municipality="Izu-ohshima Island" pageId="6" pageNumber="52" specimenCode="CBM-ZC 16951" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Kanagawa">
<specimenCode id="3C0D9E9E3979FFF6FB639AA80115FF00" box="[1152,1339,222,249]" collectionCode="CBM-ZC" pageId="6" pageNumber="52">CBM-ZC 16951</specimenCode>
,
<specimenCount id="7AADFD6C3979FFF6FAA59AA901B2FF00" box="[1350,1436,222,249]" pageId="6" pageNumber="52" type="male">5 males</specimenCount>
(5.0×5.0, 7.1×6.7, 7.3×7.3, 8.5×8.6, 9.8×
<quantity id="AB539B003979FFF6FD819B740693FEE4" box="[610,701,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="6" pageNumber="52" unit="mm" value="9.8">9.8 mm</quantity>
),
<specimenCount id="7AADFD6C3979FFF6FD339B74071FFEE4" box="[720,817,259,285]" pageId="6" pageNumber="52" type="female">1 female</specimenCount>
(6.8×
<quantity id="AB539B003979FFF6FC919B7407E0FEE4" box="[882,974,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="52" unit="mm" value="7.4">7.4 mm</quantity>
), 1 ovigerous female (6.7×
<quantity id="AB539B003979FFF6FB1A9B74017BFEE4" box="[1273,1365,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="6" pageNumber="52" unit="mm" value="6.9">6.9 mm</quantity>
), TRV
<emphasis id="5EDFEAF73979FFF6FF749B50051FFEB8" box="[151,305,295,321]" italics="true" pageId="6" pageNumber="52">Shinyo-maru</emphasis>
, 2002 research cruise, stn 21, SW of
<collectingMunicipality id="8C70AC9F3979FFF6FD179B5007FFFEB8" box="[756,977,295,321]" pageId="6" pageNumber="52">Izu-ohshima Island</collectingMunicipality>
,
<location id="6974603E3979FFF6FC029B50004CFEB8" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6:6974603E3979FFF6FC029B50004CFEB8" box="[993,1122,295,321]" latitude="35.0085" longLatPrecision="11" longitude="139.205" municipality="Izu-ohshima Island" name="Izu Islands" pageId="6" pageNumber="52" stateProvince="Kanagawa">Izu Islands</location>
,
<geoCoordinate id="099F50223979FFF6FB929B5100D6FEB8" box="[1137,1272,294,321]" degrees="35" direction="north" minutes="00.51" orientation="latitude" pageId="6" pageNumber="52" precision="9" value="35.0085">35°00.51N</geoCoordinate>
,
<geoCoordinate id="099F50223979FFF6FAE49B5001B6FEB8" box="[1287,1432,295,321]" degrees="139" direction="east" minutes="12.30" orientation="longitude" pageId="6" pageNumber="52" precision="9" value="139.205">139°12.30E</geoCoordinate>
, 549
<quantity id="AB539B003979FFF6FF2C9B3D0539FE9C" box="[207,279,330,357]" metricMagnitude="2" metricUnit="m" metricValue="4.86" pageId="6" pageNumber="52" unit="m" value="486.0">
<elevation id="E786D1D63979FFF6FF2C9B3D0539FE9C" box="[207,279,330,357]" metricMagnitude="2" metricUnit="m" metricValue="4.86" pageId="6" pageNumber="52" unit="m" value="486.0">486 m</elevation>
</quantity>
, mud,
<date id="181510253979FFF6FE829B3C0631FE9C" box="[353,543,331,357]" pageId="6" pageNumber="52" value="2002-10-23">
<collectingDate id="0851E9CD3979FFF6FE829B3C0631FE9C" box="[353,543,331,357]" pageId="6" pageNumber="52" value="2002-10-23">23 October 2002</collectingDate>
</date>
, dredge, coll
</materialsCitation>
.
<materialsCitation id="DCC33CB83979FFF6FD5A9B3C070BFE9C" ID-GBIF-Occurrence="4421509305" box="[697,805,331,357]" collectionCode="T" collectorName="Komai" pageId="6" pageNumber="52" specimenCount="1" stateProvince="Kanagawa">
<collectionCode id="0ABAAE203979FFF6FD5A9B3C06E5FE9C" box="[697,715,331,357]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="52">T</collectionCode>
.
<collectorName id="C15E53333979FFF6FD369B3C070BFE9C" box="[725,805,331,357]" pageId="6" pageNumber="52">Komai</collectorName>
</materialsCitation>
;
<materialsCitation id="DCC33CB83979FFF6FCCC9B3C0112FE70" ID-GBIF-Occurrence="4421509317" collectingDate="2021-10-24" collectionCode="CBM-ZC" collectorName="Ukusu &amp; Nishi-izu" elevation="350" location="Suruga Bay" pageId="6" pageNumber="52" specimenCode="CBM-ZC 16841, 2" specimenCount="1" stateProvince="Kanagawa">
<specimenCode id="3C0D9E9E3979FFF6FCCC9B3C07C4FE9C" box="[815,1002,331,357]" collectionCode="CBM-ZC" pageId="6" pageNumber="52">CBM-ZC 16841</specimenCode>
,
<specimenCode id="3C0D9E9E3979FFF6FC179B3C002CFE9C" box="[1012,1026,331,357]" collectionCode="CBM-ZC" pageId="6" pageNumber="52">2</specimenCode>
ovigerous females (6.6×7.0, 7.8×
<quantity id="AB539B003979FFF6FA999B3C04E8FE70" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="6" pageNumber="52" unit="mm" value="8.1">8.1 mm</quantity>
; DNA voucher),
<location id="6974603E3979FFF6FE669B180627FE70" LSID="urn:lsid:plazi:treatment:E40287F3397CFFFBFF749F8907D9FCD6:6974603E3979FFF6FE669B180627FE70" box="[389,521,367,393]" name="Suruga Bay" pageId="6" pageNumber="52" stateProvince="Kanagawa">Suruga Bay</location>
, off
<collectorName id="C15E53333979FFF6FDDF9B1806A6FE70" box="[572,648,367,393]" pageId="6" pageNumber="52">Ukusu</collectorName>
,
<collectorName id="C15E53333979FFF6FD709B1806D0FE70" box="[659,766,367,393]" pageId="6" pageNumber="52">Nishi-izu</collectorName>
,
<quantity id="AB539B003979FFF6FCEA9B1807A4FE70" box="[777,906,367,393]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="6" pageNumber="52" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">
<elevation id="E786D1D63979FFF6FCEA9B1807A4FE70" box="[777,906,367,393]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="6" pageNumber="52" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">300400 m</elevation>
</quantity>
,
<date id="181510253979FFF6FC759B18007BFE70" box="[918,1109,367,393]" pageId="6" pageNumber="52" value="2021-10-24">
<collectingDate id="0851E9CD3979FFF6FC759B18007BFE70" box="[918,1109,367,393]" pageId="6" pageNumber="52" value="2021-10-24">24 October 2021</collectingDate>
</date>
, commercial trawler
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="38D4666D3979FFF6FF749F7D073BFABF" ID-DOI="http://doi.org/10.5281/zenodo.8406969" ID-Zenodo-Dep="8406969" httpUri="https://zenodo.org/record/8406969/files/figure.png" pageId="6" pageNumber="52" startId="6.[151,250,1290,1314]" targetBox="[151,1436,417,1267]" targetPageId="6" targetType="figure">
<paragraph id="6C1436E53979FFF6FF749F7D073BFABF" blockId="6.[151,1436,1290,1350]" pageId="6" pageNumber="52">
<emphasis id="5EDFEAF73979FFF6FF749F7D053BFADB" bold="true" box="[151,277,1290,1314]" pageId="6" pageNumber="52">FIGURE 5.</emphasis>
<taxonomicName id="ABAB4D663979FFF6FEFD9F7D0657FADB" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[286,633,1290,1314]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF73979FFF6FEFD9F7D05ECFADB" box="[286,450,1290,1314]" italics="true" pageId="6" pageNumber="52">Tymolus uncifer</emphasis>
(
<bibRefCitation id="083A4B143979FFF6FE329F7D065CFADB" author="Ortmann, A. E." box="[465,626,1290,1314]" pageId="6" pageNumber="52" pagination="532 - 588" refId="ref17259" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit bedonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit Strassburger Museum aufwbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
)
</taxonomicName>
, male (9.8×9.8 mm), CBM-ZC 16951. A, carapace and cephalic appendages, dorsal view; B, thoracic sternum, ventral view. In preservative.
</paragraph>
</caption>
<subSubSection id="24B1656E3979FFF6FF249FEF01ABF80A" pageId="6" pageNumber="52" type="diagnosis">
<paragraph id="6C1436E53979FFF6FF249FEF01ABF80A" blockId="6.[151,1437,1432,2035]" pageId="6" pageNumber="52">
<emphasis id="5EDFEAF73979FFF6FF249FEF056CFA4B" bold="true" box="[199,322,1432,1458]" pageId="6" pageNumber="52">Diagnosis.</emphasis>
Carapace subcircular in outline with subparallel lateral margins, approximately as long as wide; dorsal surface with short setae (sparse on dorsal surface, more numerous on lateral sides), devoid of conspicuous tubercles or nodules on protogastric and metagastric regions; anterolateral margin with 1 small tubercle (laterobranchial tubercle); pterygostomial region with minute granules, none spiniform; front with median projection distally bifurcate by V-shaped median notch and triangular frontal lateral teeth falling far short of tips of frontal submedian teeth; frontal width approximately 0.4 of carapace width; exorbital teeth slightly produced; orbital notch deeply incised, V-shaped, relatively narrow. Ocular peduncle relatively stout, short, overreaching exorbital tooth only by length of cornea, not constricted; cornea large (width more than half distance between frontal submedian and lateral teeth), poorly pigmented even in life. Maxilliped 3 ischium-merus external surface microscopically granular, none of granules spinulose; merus tapering to rounded distal margin, subovate in outline. Pereopod 2 merus 1.72.3 times as long as carapace in males, 1.52.0 times as long in females, 14.317.0 times as long as distal width in males, 10.116.3 time as long in females, armed with cluster of 37 small, hook-like spines at anterodistal angle; carpus and propodus combined 0.40.6 times as long as merus in males, 0.50.8 times as long as merus in females. Pereopod 3 merus 1.62.0 times as long as carapace in males, 1.21.8 times as long as carapace in females, 12.915.8 times as long as distal width in males, 10.014.5 time as long as distal width in females, armed with single or cluster of 24 hook-like spines at anterodistal angle; carpus and propodus combined 0.60.7 times as long as merus in males, 0.60.9 times as long as merus in females. No fringe of setae on flexor margins of propodi of pereopods 2 and 3.
</paragraph>
</subSubSection>
<caption id="38D4666D3978FFF7FF749FE80798F9DA" ID-DOI="http://doi.org/10.5281/zenodo.8406971" ID-Zenodo-Dep="8406971" httpUri="https://zenodo.org/record/8406971/files/figure.png" pageId="7" pageNumber="53" startId="7.[151,250,1439,1463]" targetBox="[159,1428,189,1407]" targetPageId="7" targetType="figure">
<paragraph id="6C1436E53978FFF7FF749FE80798F9DA" blockId="7.[151,1436,1439,1572]" pageId="7" pageNumber="53">
<emphasis id="5EDFEAF73978FFF7FF749FE80539FA4E" bold="true" box="[151,279,1439,1463]" pageId="7" pageNumber="53">FIGURE 6.</emphasis>
<taxonomicName id="ABAB4D663978FFF7FEC29FE806ACFA41" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[289,642,1439,1464]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF73978FFF7FEC29FE805E6FA4E" box="[289,456,1439,1463]" italics="true" pageId="7" pageNumber="53">Tymolus uncifer</emphasis>
(
<bibRefCitation id="083A4B143978FFF7FE3B9FD70655FA41" author="Ortmann, A. E." box="[472,635,1440,1464]" pageId="7" pageNumber="53" pagination="532 - 588" refId="ref17259" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit bedonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit Strassburger Museum aufwbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
)
</taxonomicName>
, male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted).
</paragraph>
</caption>
<subSubSection id="24B1656E3978FFFBFF249C02004BFDCE" lastPageId="11" lastPageNumber="57" pageId="7" pageNumber="53" type="description">
<paragraph id="6C1436E53978FFF8FF249C0200E8FEB8" blockId="7.[151,1437,1653,2040]" lastBlockId="8.[151,1437,151,1977]" lastPageId="8" lastPageNumber="54" pageId="7" pageNumber="53">
<emphasis id="5EDFEAF73978FFF7FF249C02055BF976" bold="true" box="[199,373,1653,1679]" pageId="7" pageNumber="53">Redescription.</emphasis>
<emphasis id="5EDFEAF73978FFF7FE9A9C010594F969" box="[377,442,1654,1680]" italics="true" pageId="7" pageNumber="53">Male.</emphasis>
Carapace (
<figureCitation id="F4902A603978FFF7FDD19C0106BDF969" box="[562,659,1654,1680]" captionStart="FIGURE 4" captionStartId="5.[151,250,1696,1720]" captionTargetBox="[156,1431,181,1672]" captionTargetId="figure-21@5.[156,1431,181,1672]" captionTargetPageId="5" captionText="FIGURE 4. Tymolus uncifer (Ortmann, 1892), habitus in dorsal view. A, male (9.8×9.8 mm), CBM-ZC 16951; B, ovigerous female (7.6×7.6 mm), CBM-ZC 16943. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406966" httpUri="https://zenodo.org/record/8406966/files/figure.png" pageId="7" pageNumber="53">Figs.4A</figureCitation>
,
<figureCitation id="F4902A603978FFF7FD7F9C0206EEF969" box="[668,704,1653,1680]" captionStart="FIGURE 5" captionStartId="6.[151,250,1290,1314]" captionTargetBox="[151,1436,417,1267]" captionTargetId="figure-127@6.[151,1436,417,1267]" captionTargetPageId="6" captionText="FIGURE 5. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, carapace and cephalic appendages, dorsal view; B, thoracic sternum, ventral view. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406969" httpUri="https://zenodo.org/record/8406969/files/figure.png" pageId="7" pageNumber="53">5A</figureCitation>
,
<figureCitation id="F4902A603978FFF7FD2A9C020727F969" box="[713,777,1653,1680]" captionStart="FIGURE 6" captionStartId="7.[151,250,1439,1463]" captionTargetBox="[159,1428,189,1407]" captionTargetId="figure-21@7.[151,1436,181,1415]" captionTargetPageId="7" captionText="FIGURE 6. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.8406971" httpUri="https://zenodo.org/record/8406971/files/figure.png" pageId="7" pageNumber="53">6A,B</figureCitation>
)subcircular with lateral margins subparallel, approximately as long as wide, greatest width at 0.60.7 length of carapace; surface finely granular, with short setae (sparse on dorsal surface, more numerous on lateral sides). Regions on dorsal surface rather clearly indicated by shallow grooves, without conspicuous tubercles or nodules on metagastric and protogastric regions; metagastric region widened posteriorly; protogastric regions each divided into two parts by shallow longitudinal grooves; epibranchial regions demarcated; cardiac region elevated, dome-like, clearly demarcated by deep gastric fossae and connected deep longitudinal grooves; intestinal region extremely narrow. Anterolateral margins not clearly delimited, with 2 obsolescent tubercles (anterolateral and laterobranchial tubercles) on each side; posterolateral margins each with row of minute, sharp granules. Front with median projection distally bifurcate by wide V-shaped median notch and triangular frontal lateral teeth falling far short of tips of submedian teeth on median projection, tips of frontal lateral teeth blunt or subacute; dorsal surface with shallow median groove, demarcated from carapace mesogastric region by shallow transverse groove extending to anteroventral margins. Frontal width approximately 0.4 of carapace width. Exorbital teeth laterally compressed, falling far short of frontal lateral teeth, separated from frontal lateral teeth by deeply incised, V-shaped orbital notch, each tip bluntly pointed. Inferior orbital tooth small, blunt, flanked by anteroventral angle of exorbital tooth and article 2 of antennal peduncle (
<figureCitation id="F4902A603977FFF8FC089B74006DFEE4" box="[1003,1091,259,285]" captionStart="FIGURE 6" captionStartId="7.[151,250,1439,1463]" captionTargetBox="[159,1428,189,1407]" captionTargetId="figure-21@7.[151,1436,181,1415]" captionTargetPageId="7" captionText="FIGURE 6. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.8406971" httpUri="https://zenodo.org/record/8406971/files/figure.png" pageId="8" pageNumber="54">Fig. 6C</figureCitation>
). Pterygostomial region gently inflated, divided into 2 unequal parts by shallow oblique groove, covered with minute granules.
</paragraph>
<paragraph id="6C1436E53977FFF8FF249B3C0592FE70" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="54">
Endostome (
<figureCitation id="F4902A603977FFF8FEB59B3C0583FE9C" box="[342,429,331,357]" captionStart="FIGURE 6" captionStartId="7.[151,250,1439,1463]" captionTargetBox="[159,1428,189,1407]" captionTargetId="figure-21@7.[151,1436,181,1415]" captionTargetPageId="7" captionText="FIGURE 6. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.8406971" httpUri="https://zenodo.org/record/8406971/files/figure.png" pageId="8" pageNumber="54">Fig. 6C</figureCitation>
) narrowing anteriorly to form efferent channel, anterior end reaching to median notch of frontal median projection.
</paragraph>
<paragraph id="6C1436E53977FFF8FF249BE405F7FD34" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="54">
Thoracic sternum (
<figureCitation id="F4902A603977FFF8FE439BE405D7FE54" box="[416,505,402,429]" captionStart="FIGURE 5" captionStartId="6.[151,250,1290,1314]" captionTargetBox="[151,1436,417,1267]" captionTargetId="figure-127@6.[151,1436,417,1267]" captionTargetPageId="6" captionText="FIGURE 5. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, carapace and cephalic appendages, dorsal view; B, thoracic sternum, ventral view. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406969" httpUri="https://zenodo.org/record/8406969/files/figure.png" pageId="8" pageNumber="54">Fig. 5B</figureCitation>
) narrowing anteriorly, widest at sternite 5; surface granular, with median groove almost over entire length of sternites 35. Anteriormost sternite subtriangular with blunt or subacute anterior end. Sternites 35 partially fused medially; sternite 3 subhexagonal; sternites 4 elevated along lateral margins, forming protuberance-like prominences posteriorly. Sternites 5 and 6 partially fused mesially, lateral parts of former narrowing toward midline; groove separating sternites 5 and 6 deep, oblique, ending mesially in deep depression far remote from midline. Sternite 6 with posterolateral parts perpendicular against anterolateral parts (either side of sterno-pleonal cavity), extending dorsally to surround coxae of pereopods 3. Sternite 7 with lateral parts small, subtriangular in outline, weakly elevated. Sterno-pleonal cavity very deep, reaching nearly to anterior end of sternite 6, margins clearly delimited.
</paragraph>
<paragraph id="6C1436E53977FFF8FF2498A0054AFCA4" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="54">
Ocular peduncles (
<figureCitation id="F4902A603977FFF8FE7998A005D0FD08" box="[410,510,726,753]" captionStart="FIGURE 5" captionStartId="6.[151,250,1290,1314]" captionTargetBox="[151,1436,417,1267]" captionTargetId="figure-127@6.[151,1436,417,1267]" captionTargetPageId="6" captionText="FIGURE 5. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, carapace and cephalic appendages, dorsal view; B, thoracic sternum, ventral view. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406969" httpUri="https://zenodo.org/record/8406969/files/figure.png" pageId="8" pageNumber="54">Figs. 5A</figureCitation>
,
<figureCitation id="F4902A603977FFF8FDE998A00661FD08" box="[522,591,727,753]" captionStart="FIGURE 6" captionStartId="7.[151,250,1439,1463]" captionTargetBox="[159,1428,189,1407]" captionTargetId="figure-21@7.[151,1436,181,1415]" captionTargetPageId="7" captionText="FIGURE 6. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.8406971" httpUri="https://zenodo.org/record/8406971/files/figure.png" pageId="8" pageNumber="54">6A, C</figureCitation>
) slightly divergent, subcylindrical, relatively stout, not constricted, slightly inflated basally, overreaching exorbital teeth only by lengths of corneas. Cornea large (width more than half distance between tips of frontal submedian tooth and frontal lateral tooth), poorly pigmented even in life; ommatidia apparently absent.
</paragraph>
<paragraph id="6C1436E53977FFF8FF249910000CFC5C" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="54">
Antennular peduncle (
<figureCitation id="F4902A603977FFF8FE2B99100666FC78" box="[456,584,871,897]" captionStart="FIGURE 6" captionStartId="7.[151,250,1439,1463]" captionTargetBox="[159,1428,189,1407]" captionTargetId="figure-21@7.[151,1436,181,1415]" captionTargetPageId="7" captionText="FIGURE 6. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.8406971" httpUri="https://zenodo.org/record/8406971/files/figure.png" pageId="8" pageNumber="54">Fig. 6A, C</figureCitation>
) about half-length of carapace. Basal articles stout, separated by anterior extension of endostome. Article 3 subequal in length and slenderer than article 2.
</paragraph>
<paragraph id="6C1436E53977FFF8FF2499D80104FBE8" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="54">
Antennal peduncle (
<figureCitation id="F4902A603977FFF8FE4F99D80609FC30" box="[428,551,943,969]" captionStart="FIGURE 6" captionStartId="7.[151,250,1439,1463]" captionTargetBox="[159,1428,189,1407]" captionTargetId="figure-21@7.[151,1436,181,1415]" captionTargetPageId="7" captionText="FIGURE 6. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.8406971" httpUri="https://zenodo.org/record/8406971/files/figure.png" pageId="8" pageNumber="54">Fig. 6A, C</figureCitation>
) much shorter than antennular peduncle. Article
<quantity id="AB539B003977FFF8FBB799D800AEFC30" box="[1108,1152,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="54" unit="in" value="1.0">1 in</quantity>
line with basal article of antennular peduncle. Articles 2 subcylindrical, slightly widened distally. Article 3 also subcylindrical, subequal to and slightly narrower than article 2. Terminal article very short, conical, bearing sensilia-like apical seta.
</paragraph>
<paragraph id="6C1436E53977FFF8FF249E6C07BFFB10" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="54">
Maxilliped 3 (
<figureCitation id="F4902A603977FFF8FE8C9E6C05E4FBCC" box="[367,458,1051,1077]" captionStart="FIGURE 6" captionStartId="7.[151,250,1439,1463]" captionTargetBox="[159,1428,189,1407]" captionTargetId="figure-21@7.[151,1436,181,1415]" captionTargetPageId="7" captionText="FIGURE 6. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.8406971" httpUri="https://zenodo.org/record/8406971/files/figure.png" pageId="8" pageNumber="54">Fig. 6D</figureCitation>
) elongate, reaching distally to anterior end of endostome, upturned to fit anteriorly located buccal cavern; external surface of ischium and merus minutely granular, none of granules spinulose. Ischium elongate subrectangular. Merus tapering distally to rounded tip, elongate subovate; lateral margin gently convex, mesial margin nearly straight. Palp (carpus- propodus-dactylus) partially hidden by merus; dactylus small, tapering distally, with brush-like setae. Exopod narrow, closely fit endopod ischium along entire length, reaching distal margin of ischium, lateral margin slightly convex; flagellum absent.
</paragraph>
<paragraph id="6C1436E53977FFF8FF249E8407AFFA1C" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="54">
Chelipeds (
<figureCitation id="F4902A603977FFF8FEA99E84059CFAF4" box="[330,434,1267,1293]" captionStart="FIGURE 4" captionStartId="5.[151,250,1696,1720]" captionTargetBox="[156,1431,181,1672]" captionTargetId="figure-21@5.[156,1431,181,1672]" captionTargetPageId="5" captionText="FIGURE 4. Tymolus uncifer (Ortmann, 1892), habitus in dorsal view. A, male (9.8×9.8 mm), CBM-ZC 16951; B, ovigerous female (7.6×7.6 mm), CBM-ZC 16943. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406966" httpUri="https://zenodo.org/record/8406966/files/figure.png" pageId="8" pageNumber="54">Figs. 4A</figureCitation>
,
<figureCitation id="F4902A603977FFF8FE229E84062DFAF4" box="[449,515,1267,1293]" captionStart="FIGURE 6" captionStartId="7.[151,250,1439,1463]" captionTargetBox="[159,1428,189,1407]" captionTargetId="figure-21@7.[151,1436,181,1415]" captionTargetPageId="7" captionText="FIGURE 6. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.8406971" httpUri="https://zenodo.org/record/8406971/files/figure.png" pageId="8" pageNumber="54">6E, F</figureCitation>
) equal and similar, surfaces of articles minutely granular. Merus upper surface rounded; lower surface flattened. Carpus upper surface gently convex; inner surface weakly produced with few spinules. Palm widened distally, 1.1 times as high as long in adults; upper inner margin bluntly ridged; outer surface feebly convex; inner surface nearly flat; lower surface rounded. Fixed finger slightly deflexed, well compressed, terminating in subacute tip, occlusal margin irregularly denticulate; dactylus 1.1 times as long as palm, slightly curved, terminating in blunt or subacute tip, upper surface rounded with short setae along inner side, occlusal margin irregularly denticulate like opposite margin of fixed finger.
</paragraph>
<paragraph id="6C1436E53977FFF8FF249F98062FF918" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="54">
Pereopod 2 (
<figureCitation id="F4902A603977FFF8FEB89F9805EFF9F0" box="[347,449,1518,1545]" captionStart="FIGURE 5" captionStartId="6.[151,250,1290,1314]" captionTargetBox="[151,1436,417,1267]" captionTargetId="figure-127@6.[151,1436,417,1267]" captionTargetPageId="6" captionText="FIGURE 5. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, carapace and cephalic appendages, dorsal view; B, thoracic sternum, ventral view. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406969" httpUri="https://zenodo.org/record/8406969/files/figure.png" pageId="8" pageNumber="54">Figs. 5A</figureCitation>
,
<figureCitation id="F4902A603977FFF8FE2C9F98063BF9F0" box="[463,533,1519,1545]" captionStart="FIGURE 7" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[206,1380,189,1880]" captionTargetId="figure-21@9.[198,1388,181,1885]" captionTargetPageId="9" captionText="FIGURE 7. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951.A, right pereopod 2, dorsal view; B, same, distal part of merus, dorsal view; C, right pereopod 3, dorsal view (setae omitted); D, same, distal part of merus, dorsal view; E, right pereopod 4, dorsal view; F, right pereopod 5, dorsal view; G, left gonopod 1, mesial view; H, same, ventral view; I, same, lateral view; J, same, dorsal view; K, left gonopod 2, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8406973" httpUri="https://zenodo.org/record/8406973/files/figure.png" pageId="8" pageNumber="54">7A, B</figureCitation>
) markedly elongate, surfaces of articles covered with minute granules (except for dactylus) and short setae, but not particularly spinulose. Coxa large, with condyle at anteroventral distal angle. Ischium articulation to merus strongly oblique. Merus 1.72.3 times as long as carapace, 14.317.0 times as long as distal width, nearly straight, rod-like, slightly widened distally, with cluster of 37 small, mesially curved, hook-like spines at anterodistal angle. Carpus and propodus combined 0.40.6 times as long as merus, both articles unarmed; carpus slightly widened distally. Dactylus slender, slightly flattened, gently arcuate, subequal in length to propodus; upper and lower margins setose.
</paragraph>
<paragraph id="6C1436E53977FFF8FF249C9C0129F8B4" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="54">
Pereopod 3 (
<figureCitation id="F4902A603977FFF8FEB69C9C0596F8FC" box="[341,440,1770,1797]" captionStart="FIGURE 5" captionStartId="6.[151,250,1290,1314]" captionTargetBox="[151,1436,417,1267]" captionTargetId="figure-127@6.[151,1436,417,1267]" captionTargetPageId="6" captionText="FIGURE 5. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, carapace and cephalic appendages, dorsal view; B, thoracic sternum, ventral view. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406969" httpUri="https://zenodo.org/record/8406969/files/figure.png" pageId="8" pageNumber="54">Figs. 5A</figureCitation>
,
<figureCitation id="F4902A603977FFF8FE279C9C0629F8FC" box="[452,519,1771,1797]" captionStart="FIGURE 7" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[206,1380,189,1880]" captionTargetId="figure-21@9.[198,1388,181,1885]" captionTargetPageId="9" captionText="FIGURE 7. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951.A, right pereopod 2, dorsal view; B, same, distal part of merus, dorsal view; C, right pereopod 3, dorsal view (setae omitted); D, same, distal part of merus, dorsal view; E, right pereopod 4, dorsal view; F, right pereopod 5, dorsal view; G, left gonopod 1, mesial view; H, same, ventral view; I, same, lateral view; J, same, dorsal view; K, left gonopod 2, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8406973" httpUri="https://zenodo.org/record/8406973/files/figure.png" pageId="8" pageNumber="54">7C, D</figureCitation>
) generally similar to pereopod 2, but slightly shorter. Merus 1.62.0 times as long as carapace, 12.915.8 times as long as distal width, anterodistal angle with cluster of 3 or 4 small, hook-like spines. Carpus-propodus combined 0.60.7 times as long as merus. Dactylus subequal in length to propodus.
</paragraph>
<paragraph id="6C1436E53977FFF8FF249D200605F840" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="54">
Pereopods 4 and 5 (
<figureCitation id="F4902A603977FFF8FE479D200626F888" box="[420,520,1878,1905]" captionStart="FIGURE 5" captionStartId="6.[151,250,1290,1314]" captionTargetBox="[151,1436,417,1267]" captionTargetId="figure-127@6.[151,1436,417,1267]" captionTargetPageId="6" captionText="FIGURE 5. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, carapace and cephalic appendages, dorsal view; B, thoracic sternum, ventral view. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406969" httpUri="https://zenodo.org/record/8406969/files/figure.png" pageId="8" pageNumber="54">Figs. 5A</figureCitation>
,
<figureCitation id="F4902A603977FFF8FDF09D20067FF888" box="[531,593,1879,1905]" captionStart="FIGURE 7" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[206,1380,189,1880]" captionTargetId="figure-21@9.[198,1388,181,1885]" captionTargetPageId="9" captionText="FIGURE 7. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951.A, right pereopod 2, dorsal view; B, same, distal part of merus, dorsal view; C, right pereopod 3, dorsal view (setae omitted); D, same, distal part of merus, dorsal view; E, right pereopod 4, dorsal view; F, right pereopod 5, dorsal view; G, left gonopod 1, mesial view; H, same, ventral view; I, same, lateral view; J, same, dorsal view; K, left gonopod 2, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8406973" httpUri="https://zenodo.org/record/8406973/files/figure.png" pageId="8" pageNumber="54">7E, F</figureCitation>
) located dorsally, much shorter and slenderer than pereopods 2 and 3. Meri extending well beyond posterolateral margin of carapace. Dactyli slender, sickle-shaped, shorter than propodi, fully folded against propodi when flexed.
</paragraph>
<caption id="38D4666D3976FFF9FF749D02071FF800" ID-DOI="http://doi.org/10.5281/zenodo.8406973" ID-Zenodo-Dep="8406973" httpUri="https://zenodo.org/record/8406973/files/figure.png" pageId="9" pageNumber="55" startId="9.[151,250,1909,1933]" targetBox="[206,1380,189,1880]" targetPageId="9" targetType="figure">
<paragraph id="6C1436E53976FFF9FF749D02071FF800" blockId="9.[151,1436,1909,2041]" pageId="9" pageNumber="55">
<emphasis id="5EDFEAF73976FFF9FF749D02053CF874" bold="true" box="[151,274,1909,1933]" pageId="9" pageNumber="55">FIGURE 7.</emphasis>
<taxonomicName id="ABAB4D663976FFF9FEFB9D020644F874" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[280,618,1909,1933]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF73976FFF9FEFB9D020594F874" box="[280,442,1909,1933]" italics="true" pageId="9" pageNumber="55">Tymolus uncifer</emphasis>
(
<bibRefCitation id="083A4B143976FFF9FE259D02064DF874" author="Ortmann, A. E." box="[454,611,1909,1933]" pageId="9" pageNumber="55" pagination="532 - 588" refId="ref17259" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit bedonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit Strassburger Museum aufwbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
)
</taxonomicName>
, male (9.8×9.8 mm), CBM-ZC 16951. A, right pereopod 2, dorsal view; B, same, distal part of merus, dorsal view; C, right pereopod 3, dorsal view (setae omitted); D, same, distal part of merus, dorsal view; E, right pereopod 4, dorsal view; F, right pereopod 5, dorsal view; G, left gonopod 1, mesial view; H, same, ventral view; I, same, lateral view; J, same, dorsal view; K, left gonopod 2, lateral view.
</paragraph>
</caption>
<paragraph id="6C1436E53975FFFAFF249AE00759FEE4" blockId="10.[151,1437,150,825]" pageId="10" pageNumber="56">
Pleon (
<figureCitation id="F4902A603975FFFAFEFB9AE00552FF48" box="[280,380,151,177]" captionStart="FIGURE 6" captionStartId="7.[151,250,1439,1463]" captionTargetBox="[159,1428,189,1407]" captionTargetId="figure-21@7.[151,1436,181,1415]" captionTargetPageId="7" captionText="FIGURE 6. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951. A, anterior part of carapace, ocular peduncles, antennules and antennae, dorsal view; B, left anterolateral part of carapace (setae omitted); C, left anterior part of carapace and cephalic appendages, ventral view (setae omitted); D, right maxilliped 3, external view; E, carpus of right cheliped, dorsal view; F, right chela, outer view; G, pleon, external view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.8406971" httpUri="https://zenodo.org/record/8406971/files/figure.png" pageId="10" pageNumber="56">Figs. 6G</figureCitation>
) consisting of 5 somites, pleomere 1 narrowest, pleomere 4 widest; pleomeres 13 visible in dorsal view; midline of pleomeres 14 weakly elevated in blunt ridge, lateral margins also weakly elevated, separated from median ridge by shallow sulci. Pleotelson formed by fusion of pleomeres 5, 6 and telson, subtriangular with rounded apex; outer surface with low median elevation near base.
</paragraph>
<paragraph id="6C1436E53975FFFAFF249B500705FE0C" blockId="10.[151,1437,150,825]" pageId="10" pageNumber="56">
Gonopod 1 (
<figureCitation id="F4902A603975FFFAFEB49B5005E7FEB8" box="[343,457,295,321]" captionStart="FIGURE 7" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[206,1380,189,1880]" captionTargetId="figure-21@9.[198,1388,181,1885]" captionTargetPageId="9" captionText="FIGURE 7. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951.A, right pereopod 2, dorsal view; B, same, distal part of merus, dorsal view; C, right pereopod 3, dorsal view (setae omitted); D, same, distal part of merus, dorsal view; E, right pereopod 4, dorsal view; F, right pereopod 5, dorsal view; G, left gonopod 1, mesial view; H, same, ventral view; I, same, lateral view; J, same, dorsal view; K, left gonopod 2, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8406973" httpUri="https://zenodo.org/record/8406973/files/figure.png" pageId="10" pageNumber="56">Fig. 7GJ</figureCitation>
) consisting of 3 articles; angle between ultimate and penultimate article 30° in lateral view; ultimate article bean-shaped in general outline, cannulate, forming copulatory tube embracing gonopod
<quantity id="AB539B003975FFFAFA929B3C01B2FE9C" box="[1393,1436,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="56" unit="in" value="2.0">2 in</quantity>
situ, distal margin with setae extending onto proximal margins, ventral surface with deep longitudinal groove along over entire length; inner margin of penultimate article slightly expanded. G2 (
<figureCitation id="F4902A603975FFFAFBFD9BE40056FE54" box="[1054,1144,403,429]" captionStart="FIGURE 7" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[206,1380,189,1880]" captionTargetId="figure-21@9.[198,1388,181,1885]" captionTargetPageId="9" captionText="FIGURE 7. Tymolus uncifer (Ortmann, 1892), male (9.8×9.8 mm), CBM-ZC 16951.A, right pereopod 2, dorsal view; B, same, distal part of merus, dorsal view; C, right pereopod 3, dorsal view (setae omitted); D, same, distal part of merus, dorsal view; E, right pereopod 4, dorsal view; F, right pereopod 5, dorsal view; G, left gonopod 1, mesial view; H, same, ventral view; I, same, lateral view; J, same, dorsal view; K, left gonopod 2, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8406973" httpUri="https://zenodo.org/record/8406973/files/figure.png" pageId="10" pageNumber="56">Fig. 7K</figureCitation>
) stout, fairly compressed laterally, lower distal end extending into slender, chitinous projection, upper distal margin truncate, hollowed; upper margin gently convex, while lower margin gently concave.
</paragraph>
<paragraph id="6C1436E53975FFFAFF249B8807B0FCC0" blockId="10.[151,1437,150,825]" pageId="10" pageNumber="56">
<emphasis id="5EDFEAF73975FFFAFF249B880500FDE0" box="[199,302,511,537]" italics="true" pageId="10" pageNumber="56">Females.</emphasis>
Carapace generally similar to that of male, but pereopods 2 and 3 proportionally shorter (
<figureCitation id="F4902A603975FFFAFAD69B8801A1FDE0" box="[1333,1423,511,537]" captionStart="FIGURE 4" captionStartId="5.[151,250,1696,1720]" captionTargetBox="[156,1431,181,1672]" captionTargetId="figure-21@5.[156,1431,181,1672]" captionTargetPageId="5" captionText="FIGURE 4. Tymolus uncifer (Ortmann, 1892), habitus in dorsal view. A, male (9.8×9.8 mm), CBM-ZC 16951; B, ovigerous female (7.6×7.6 mm), CBM-ZC 16943. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406966" httpUri="https://zenodo.org/record/8406966/files/figure.png" pageId="10" pageNumber="56">Fig. 4B</figureCitation>
). Thoracic sternum (
<figureCitation id="F4902A603975FFFAFE8C985405E7FDC4" box="[367,457,547,573]" captionStart="FIGURE 8" captionStartId="10.[151,250,1808,1832]" captionTargetBox="[159,1428,857,1777]" captionTargetId="figure-298@10.[151,1436,849,1785]" captionTargetPageId="10" captionText="FIGURE 8. Tymolus uncifer (Ortmann, 1892), ovigerous female 7.6×7.6 mm), CBM-ZC 16943. A, thoracic sternum, ventral view; B, right chela, outer view; C, pleon, external view. Setae omitted." figureDoi="http://doi.org/10.5281/zenodo.8406975" httpUri="https://zenodo.org/record/8406975/files/figure.png" pageId="10" pageNumber="56">Fig. 8A</figureCitation>
) narrowing anteriorly, widest at sternite 5; surface microscopically granular, medially sulcate on sternites 35. Anteriormost sternite subtriangular with blunt anterior margin. Sternites 35 demarcated by transverse furrows; sternite 3 subhexagonal; sternites 4 with paired rounded elevations, lateral margins accommodating coxae of chelipeds substantially elevated, flange-like; small deep pit on either side just proximal to base of maxilliped 3. Sternite 5 posteriorly with transverse ridge forming anterior boundary of sterno-pleonal cavity. Entire sternite 6 forming deep sterno-pleonal cavity, steeply sloping, medially with blunt transverse ridge adjacent to sternite 5; suture demarcating sternites 5 and 6 distinct laterally, faded away medially; suture demarcating sternites 6 and 7 also distinct laterally, fairly oblique against longitudinal axis.
</paragraph>
<caption id="38D4666D3975FFFAFF749D670743F8B5" ID-DOI="http://doi.org/10.5281/zenodo.8406975" ID-Zenodo-Dep="8406975" httpUri="https://zenodo.org/record/8406975/files/figure.png" pageId="10" pageNumber="56" startId="10.[151,250,1808,1832]" targetBox="[159,1428,857,1777]" targetPageId="10" targetType="figure">
<paragraph id="6C1436E53975FFFAFF749D670743F8B5" blockId="10.[151,1436,1808,1868]" pageId="10" pageNumber="56">
<emphasis id="5EDFEAF73975FFFAFF749D67053AF8D1" bold="true" box="[151,276,1808,1832]" pageId="10" pageNumber="56">FIGURE 8.</emphasis>
<taxonomicName id="ABAB4D663975FFFAFEFF9D67065DF8D1" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[284,627,1808,1832]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="56" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF73975FFFAFEFF9D670591F8D1" box="[284,447,1808,1832]" italics="true" pageId="10" pageNumber="56">Tymolus uncifer</emphasis>
(
<bibRefCitation id="083A4B143975FFFAFE2E9D670642F8D1" author="Ortmann, A. E." box="[461,620,1808,1832]" pageId="10" pageNumber="56" pagination="532 - 588" refId="ref17259" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit bedonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit Strassburger Museum aufwbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
)
</taxonomicName>
, ovigerous female 7.6×7.6 mm), CBM-ZC 16943. A, thoracic sternum, ventral view; B, right chela, outer view; C, pleon, external view. Setae omitted.
</paragraph>
</caption>
<paragraph id="6C1436E53974FFFBFF249AE0076BFF0E" blockId="11.[151,1437,150,816]" pageId="11" pageNumber="57">
Chelipeds (
<figureCitation id="F4902A603974FFFBFEA59AE00589FF48" box="[326,423,151,177]" captionStart="FIGURE 4" captionStartId="5.[151,250,1696,1720]" captionTargetBox="[156,1431,181,1672]" captionTargetId="figure-21@5.[156,1431,181,1672]" captionTargetPageId="5" captionText="FIGURE 4. Tymolus uncifer (Ortmann, 1892), habitus in dorsal view. A, male (9.8×9.8 mm), CBM-ZC 16951; B, ovigerous female (7.6×7.6 mm), CBM-ZC 16943. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406966" httpUri="https://zenodo.org/record/8406966/files/figure.png" pageId="11" pageNumber="57">Figs. 4B</figureCitation>
,
<figureCitation id="F4902A603974FFFBFE509AE005FAFF48" box="[435,468,151,177]" captionStart="FIGURE 8" captionStartId="10.[151,250,1808,1832]" captionTargetBox="[159,1428,857,1777]" captionTargetId="figure-298@10.[151,1436,849,1785]" captionTargetPageId="10" captionText="FIGURE 8. Tymolus uncifer (Ortmann, 1892), ovigerous female 7.6×7.6 mm), CBM-ZC 16943. A, thoracic sternum, ventral view; B, right chela, outer view; C, pleon, external view. Setae omitted." figureDoi="http://doi.org/10.5281/zenodo.8406975" httpUri="https://zenodo.org/record/8406975/files/figure.png" pageId="11" pageNumber="57">8B</figureCitation>
) slenderer than those of males. Chela 2.5 times as long as wide; palm widened distally, 1.1 times as wide as long; fingers terminating in acute tips, fixed finger somewhat deflexed; denticles on occlusal margins obsolescent or blunt; dactylus twice as long as palm.
</paragraph>
<paragraph id="6C1436E53974FFFBFF249B76058CFE7F" blockId="11.[151,1437,150,816]" pageId="11" pageNumber="57">
Pereopod 2 (
<figureCitation id="F4902A603974FFFBFEB59B760583FEE2" box="[342,429,257,283]" captionStart="FIGURE 4" captionStartId="5.[151,250,1696,1720]" captionTargetBox="[156,1431,181,1672]" captionTargetId="figure-21@5.[156,1431,181,1672]" captionTargetPageId="5" captionText="FIGURE 4. Tymolus uncifer (Ortmann, 1892), habitus in dorsal view. A, male (9.8×9.8 mm), CBM-ZC 16951; B, ovigerous female (7.6×7.6 mm), CBM-ZC 16943. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406966" httpUri="https://zenodo.org/record/8406966/files/figure.png" pageId="11" pageNumber="57">Fig. 4B</figureCitation>
) merus 1.52.0 times as long as carapace, 10.116.3 times as long as distal width; carpus and propodus combined 0.50.8 times as long as merus. Pereopod 3 (
<figureCitation id="F4902A603974FFFBFC259B53000EFEC6" box="[966,1056,292,319]" captionStart="FIGURE 4" captionStartId="5.[151,250,1696,1720]" captionTargetBox="[156,1431,181,1672]" captionTargetId="figure-21@5.[156,1431,181,1672]" captionTargetPageId="5" captionText="FIGURE 4. Tymolus uncifer (Ortmann, 1892), habitus in dorsal view. A, male (9.8×9.8 mm), CBM-ZC 16951; B, ovigerous female (7.6×7.6 mm), CBM-ZC 16943. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406966" httpUri="https://zenodo.org/record/8406966/files/figure.png" pageId="11" pageNumber="57">Fig. 4B</figureCitation>
) merus 1.21.8 times as long as carapace, 1014.5 times as long as distal width; carpus and propodus combined 0.50.8 times as long as merus; gonopore on each coxa.
</paragraph>
<paragraph id="6C1436E53974FFFBFF249BF80086FE09" blockId="11.[151,1437,150,816]" pageId="11" pageNumber="57">
Pleon (
<figureCitation id="F4902A603974FFFBFEF99BF805AEFE50" box="[282,384,399,425]" captionStart="FIGURE 4" captionStartId="5.[151,250,1696,1720]" captionTargetBox="[156,1431,181,1672]" captionTargetId="figure-21@5.[156,1431,181,1672]" captionTargetPageId="5" captionText="FIGURE 4. Tymolus uncifer (Ortmann, 1892), habitus in dorsal view. A, male (9.8×9.8 mm), CBM-ZC 16951; B, ovigerous female (7.6×7.6 mm), CBM-ZC 16943. In preservative." figureDoi="http://doi.org/10.5281/zenodo.8406966" httpUri="https://zenodo.org/record/8406966/files/figure.png" pageId="11" pageNumber="57">Figs. 4B</figureCitation>
,
<figureCitation id="F4902A603974FFFBFE739BF8059FFE50" box="[400,433,399,425]" captionStart="FIGURE 8" captionStartId="10.[151,250,1808,1832]" captionTargetBox="[159,1428,857,1777]" captionTargetId="figure-298@10.[151,1436,849,1785]" captionTargetPageId="10" captionText="FIGURE 8. Tymolus uncifer (Ortmann, 1892), ovigerous female 7.6×7.6 mm), CBM-ZC 16943. A, thoracic sternum, ventral view; B, right chela, outer view; C, pleon, external view. Setae omitted." figureDoi="http://doi.org/10.5281/zenodo.8406975" httpUri="https://zenodo.org/record/8406975/files/figure.png" pageId="11" pageNumber="57">8C</figureCitation>
) consisting of 5 somites, pleomere 6 fused with telson (= pleotelson). Pleomeres 15 each with low, rounded protuberance on midline; pleomeres 14 visible in dorsal view; pleomere 1 narrowest, subtrapezoidal; pleomeres 4 and 5 widest. Pleotelson semicircular, 1.6 times as long as wide.
</paragraph>
<paragraph id="6C1436E53974FFFBFF249B8D0611FDED" blockId="11.[151,1437,150,816]" box="[199,575,505,532]" pageId="11" pageNumber="57">
Eggs spherical, diameter
<quantity id="AB539B003974FFFBFE019B8E0615FDED" box="[482,571,505,532]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="57" unit="mm" value="0.7">0.7 mm</quantity>
.
</paragraph>
<paragraph id="6C1436E53974FFFBFF24986B004BFDCE" blockId="11.[151,1437,150,816]" box="[199,1125,540,567]" pageId="11" pageNumber="57">
<emphasis id="5EDFEAF73974FFFBFF24986B0584FDCF" bold="true" box="[199,426,540,566]" pageId="11" pageNumber="57">Colouration in life.</emphasis>
Entirely white; corneas opaque, poorly pigmented (
<figureCitation id="F4902A603974FFFBFC13986A0076FDCE" box="[1008,1112,541,567]" captionStart="FIGURE 14" captionStartId="18.[151,250,1691,1715]" captionTargetBox="[151,1436,181,1668]" captionTargetId="figure-15@18.[151,1436,181,1668]" captionTargetPageId="18" captionText="FIGURE 14. Habitus showing living colouration.A, Tymolus uncifer (Ortmann, 1892), ovigerous female (7.8×8.1 mm), CBM- ZC 16841; B, Tymolus melanommus (Doflein, 1904), male (13.0×13.0 mm), CBM-ZC 17323." figureDoi="http://doi.org/10.5281/zenodo.8406990" httpUri="https://zenodo.org/record/8406990/files/figure.png" pageId="11" pageNumber="57">Fig. 14A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="24B1656E3974FFFBFF2498370671FD87" pageId="11" pageNumber="57" type="distribution">
<paragraph id="6C1436E53974FFFBFF2498370671FD87" blockId="11.[151,1437,150,816]" pageId="11" pageNumber="57">
<emphasis id="5EDFEAF73974FFFBFF249837054FFDA3" bold="true" box="[199,353,576,602]" pageId="11" pageNumber="57">Distribution.</emphasis>
Known with certainty only from the Pacific coast of
<collectingCountry id="14BC76753974FFFBFC57983607D9FDA2" box="[948,1015,577,603]" name="Japan" pageId="11" pageNumber="57">Japan</collectingCountry>
, Sagami Sea to Suruga Bay, at depths of
<quantity id="AB539B003974FFFBFF559814051BFD87" box="[182,309,611,638]" metricMagnitude="2" metricUnit="m" metricValue="3.56" metricValueMax="5.55" metricValueMin="1.5699999999999998" pageId="11" pageNumber="57" unit="m" value="356.0" valueMax="555.0" valueMin="157.0">157555 m</quantity>
(see below “Discussion”).
</paragraph>
</subSubSection>
<subSubSection id="24B1656E3974FFFBFF2498F007D9FCD6" pageId="11" pageNumber="57" type="discussion">
<paragraph id="6C1436E53974FFFBFF2498F007D9FCD6" blockId="11.[151,1437,150,816]" pageId="11" pageNumber="57">
<emphasis id="5EDFEAF73974FFFBFF2498F00515FD58" bold="true" box="[199,315,647,673]" pageId="11" pageNumber="57">Remarks.</emphasis>
One juvenile (CBM-ZC 4507, 3.5×
<quantity id="AB539B003974FFFBFD3498F0071AFD58" box="[727,820,647,674]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="11" pageNumber="57" unit="mm" value="3.6">3.6 mm</quantity>
), the smallest among the examined specimens, differs from adult specimens in having proportionally shorter carpus+propodus (0.9 times as long as merus in the pereopod 2, 0.7 times in the pereopod 3), approaching the condition seen in
<taxonomicName id="ABAB4D663974FFFBFC8098B8003FFD11" baseAuthorityName="Doflein" baseAuthorityYear="1904" box="[867,1041,718,744]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="57" phylum="Arthropoda" rank="species" species="melanommus">
<emphasis id="5EDFEAF73974FFFBFC8098B8003FFD11" box="[867,1041,718,744]" italics="true" pageId="11" pageNumber="57">T. melanommus</emphasis>
</taxonomicName>
. As discussed below, morphometric characters of the pereopods 2 and 3 are useful in differentiating
<taxonomicName id="ABAB4D663974FFFBFC84988507FFFCF5" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[871,977,754,780]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="57" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="5EDFEAF73974FFFBFC84988507FFFCF5" box="[871,977,754,780]" italics="true" pageId="11" pageNumber="57">T. uncifer</emphasis>
</taxonomicName>
and
<taxonomicName id="ABAB4D663974FFFBFBEA98850080FCF2" authorityName="Doflein" authorityYear="1904" box="[1033,1198,753,779]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="57" phylum="Arthropoda" rank="species" species="melanomma">
<emphasis id="5EDFEAF73974FFFBFBEA98850080FCF2" box="[1033,1198,753,779]" italics="true" pageId="11" pageNumber="57">T. melanomma</emphasis>
</taxonomicName>
in adults, but careful observation will be necessary when identifying juvenile or young specimens.
</paragraph>
</subSubSection>
</treatment>
</document>