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<document id="F8962D56C891E2F5D593FFE22456BA18" ID-CLB-Dataset="20095" ID-DOI="10.11646/phytotaxa.541.2.1" ID-GBIF-Dataset="181ca3af-69ba-42a3-95a7-3654844046f7" ID-ISSN="1179-3163" ID-Zenodo-Dep="6388623" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648450146519" checkinUser="plazi" docAuthor="Ariza-Cortés, William, Cortés-B, Rocio &amp; Fernández-Alonso, José Luis" docDate="2022" docId="DE7D2B43FFE2FFF37DBFA26633AD1830" docLanguage="en" docName="phytotaxa.541.2.1.pdf" docOrigin="Phytotaxa 541 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.541.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Vochysia Aublet 1775" docType="key" docVersion="7" lastPageNumber="108" masterDocId="2244533BFFF1FFE07D37A77836071F47" masterDocTitle="Mahechadendron puntecascarillo (Vochysiaceae), a new genus and forest tree species from Colombia" masterLastPageNumber="112" masterPageNumber="89" pageNumber="108" updateTime="1699318551585" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1728F84B659DE92E382FAC35F45CB63C">Mahechadendron puntecascarillo (Vochysiaceae), a new genus and forest tree species from Colombia</mods:title>
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<mods:namePart id="39AF0862FC8FDEDC051EB392E7C9D091">Ariza-Cortés, William</mods:namePart>
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<mods:affiliation id="1266A11D54DA350A2CF1FDAFC08F22FB">Herbario Forestal, Universidad Distrital Francisco José de Caldas, Carrera 5 Este No. 15 - 82, Bogotá, Colombia &amp; wgarizac @ udistrital. edu. co; https: // orcid. org / 0000 - 0002 - 8423 - 8256</mods:affiliation>
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<mods:namePart id="1CB0F7B9073B49E363CDABA2C8B329AA">Cortés-B, Rocio</mods:namePart>
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<mods:affiliation id="0566CCFA490A3CF0ACD7DFCA28D3226F">Herbario Forestal, Universidad Distrital Francisco José de Caldas, Carrera 5 Este No. 15 - 82, Bogotá, Colombia &amp; rpcortesb @ udistrital. edu. co; https: // orcid. org / 0000 - 0003 - 2781 - 5747</mods:affiliation>
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<mods:namePart id="FFE9F98E98A73EAAC95561493FF697EC">Fernández-Alonso, José Luis</mods:namePart>
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<mods:affiliation id="CE48AB36418E3096DEA2F98AD350B32F">Departamento de Biodiversidad y Conservación, Real Jardín Botánico, CSIC, Claudio Moyano 1, 28014, Madrid, Spain &amp; jlfernandeza @ rjb. csic. es; https: // orcid. org / 0000 - 0002 - 1701 - 480 X</mods:affiliation>
<mods:nameIdentifier id="8147C4E5633E6A7CFE745D4BE8D9D8B4" type="email">jlfernandeza@rjb.csic.es</mods:nameIdentifier>
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<mods:title id="29DB4977323A1629C1C5C1C3C08456EC">Phytotaxa</mods:title>
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<mods:date id="CC8809C1676EFDA6359FCD60D317C6BB">2022</mods:date>
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<mods:number id="F1F44AF6B997258F9A9CBD3C25F0F4F3">541</mods:number>
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<treatment id="DE7D2B43FFE2FFF37DBFA26633AD1830" ID-DOI="http://doi.org/10.5281/zenodo.6391535" ID-GBIF-Taxon="194286105" ID-Zenodo-Dep="6391535" LSID="urn:lsid:plazi:treatment:DE7D2B43FFE2FFF37DBFA26633AD1830" httpUri="http://treatment.plazi.org/id/DE7D2B43FFE2FFF37DBFA26633AD1830" lastPageNumber="108" pageId="19" pageNumber="108">
<subSubSection id="1ECEC9DEFFE2FFF37DBFA26634191A7F" box="[136,542,1310,1336]" pageId="19" pageNumber="108" type="nomenclature">
<paragraph id="566B9A55FFE2FFF37DBFA26634191A7F" blockId="19.[136,1452,1310,1912]" box="[136,542,1310,1336]" pageId="19" pageNumber="108">
<heading id="0D232D39FFE2FFF37DBFA26634191A7F" bold="true" box="[136,542,1310,1336]" fontSize="11" level="1" pageId="19" pageNumber="108" reason="1">
<emphasis id="64A04647FFE2FFF37DBFA26634191A7F" bold="true" box="[136,542,1310,1336]" pageId="19" pageNumber="108">
Key to the genera of
<taxonomicName id="91D4E1D6FFE2FFF37CB7A26634191A7F" ID-CoL="HV2" authority="Aublet (1775 a: 18)" authorityName="Aublet" authorityPageNumber="18" authorityYear="1775" box="[384,542,1310,1336]" class="Magnoliopsida" family="Vochysiaceae" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="108" phylum="Tracheophyta" rank="family">Vochysiaceae</taxonomicName>
</emphasis>
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<subSubSection id="1ECEC9DEFFE2FFF37DBFA24733AD1830" pageId="19" pageNumber="108" type="key">
<keyStep id="ED2082F0FFE2FFF37DBFA24733AC1A36" pageId="19" pageNumber="108">
<paragraph id="566B9A55FFE2FFF37DBFA24733AC1A13" blockId="19.[136,1452,1310,1912]" box="[136,1451,1343,1365]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA24733AC1A13" box="[136,1451,1343,1365]" pageId="19" pageNumber="108">1. Trees with unilocular inferior ovary, samaroid fruits .........................................................................................................................2</keyLead>
</paragraph>
<paragraph id="566B9A55FFE2FFF37DBFA22433AC1A36" blockId="19.[136,1452,1310,1912]" box="[136,1451,1371,1393]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA22433AC1A36" box="[136,1451,1371,1393]" pageId="19" pageNumber="108">- Trees, shrubs or subshrubs with trilocular superior ovary, capsular fruits (Neotropics)...................................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep id="ED2082F0FFE2FFF37DBFA20033AD1AEC" pageId="19" pageNumber="108">
<paragraph id="566B9A55FFE2FFF37DBFA20033AC1ACA" blockId="19.[136,1452,1310,1912]" box="[136,1451,1400,1422]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA20033AC1ACA" box="[136,1451,1400,1422]" pageId="19" pageNumber="108">2. Flowers with five petals, fruits with five wings (West Africa)...........................................................................................................3</keyLead>
</paragraph>
<paragraph id="566B9A55FFE2FFF37DBFA2ED33AD1AEC" blockId="19.[136,1452,1310,1912]" box="[136,1450,1429,1451]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA2ED33AD1AEC" box="[136,1450,1429,1451]" pageId="19" pageNumber="108">
- Flowers with a single petal, fruits with four wings (Neotropics) .............................................................................................
<taxonomicName id="91D4E1D6FFE2FFF37852A2EE33AD1AEC" ID-CoL="62S22" authorityName="Rudge" authorityYear="1805" box="[1381,1450,1430,1451]" class="Magnoliopsida" family="Vochysiaceae" genus="Erisma" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="108" phylum="Tracheophyta" rank="genus">
<emphasis id="64A04647FFE2FFF37852A2EE33AD1AEC" box="[1381,1450,1430,1451]" italics="true" pageId="19" pageNumber="108">Erisma</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="ED2082F0FFE2FFF37DBFA2CA33AC1AA3" pageId="19" pageNumber="108">
<paragraph id="566B9A55FFE2FFF37DBFA2CA33AD1A80" blockId="19.[136,1452,1310,1912]" box="[136,1450,1458,1480]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA2CA33AD1A80" box="[136,1450,1458,1480]" pageId="19" pageNumber="108">
3. Calyx with three petaloid sepals and two non-petaloid..............................................................................................
<taxonomicName id="91D4E1D6FFE2FFF37825A2CA33AD1A80" ID-CoL="58Y9" authorityName="Litt &amp; Cheek" authorityYear="2002" box="[1298,1450,1458,1479]" class="Magnoliopsida" family="Vochysiaceae" genus="Korupodendron" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="108" phylum="Tracheophyta" rank="genus">
<emphasis id="64A04647FFE2FFF37825A2CA33AD1A80" box="[1298,1450,1458,1479]" italics="true" pageId="19" pageNumber="108">Korupodendron</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="566B9A55FFE2FFF37DBFA2B733AC1AA3" blockId="19.[136,1452,1310,1912]" box="[136,1451,1487,1509]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA2B733AC1AA3" box="[136,1451,1487,1509]" pageId="19" pageNumber="108">
- Calyx with five non-petaloid sepals .............................................................................................................................
<taxonomicName id="91D4E1D6FFE2FFF3782CA2B733AC1AA3" ID-CoL="633PQ" authorityName="Mildbraed" authorityYear="1913" box="[1307,1451,1487,1508]" class="Magnoliopsida" family="Vochysiaceae" genus="Erismadelphus" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="108" phylum="Tracheophyta" rank="genus">
<emphasis id="64A04647FFE2FFF3782CA2B733AC1AA3" box="[1307,1451,1487,1508]" italics="true" pageId="19" pageNumber="108">Erismadelphus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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<keyStep id="ED2082F0FFE2FFF37DBFA29433AC1959" pageId="19" pageNumber="108">
<paragraph id="566B9A55FFE2FFF37DBFA29433AC1947" blockId="19.[136,1452,1310,1912]" box="[136,1451,1515,1537]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA29433AC1947" box="[136,1451,1515,1537]" pageId="19" pageNumber="108">
4. Petals 5, branches with 59 leaves per node .........................................................................................................................
<taxonomicName id="91D4E1D6FFE2FFF37863A29333AC1947" ID-CoL="7C26" authorityName="Saint-Hilaire" authorityYear="1820" box="[1364,1451,1515,1536]" class="Magnoliopsida" family="Vochysiaceae" genus="Salvertia" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="108" phylum="Tracheophyta" rank="genus">
<emphasis id="64A04647FFE2FFF37863A29333AC1947" box="[1364,1451,1515,1536]" italics="true" pageId="19" pageNumber="108">Salvertia</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="566B9A55FFE2FFF37DBFA17133AC1959" blockId="19.[136,1452,1310,1912]" box="[136,1451,1544,1566]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA17133AC1959" box="[136,1451,1544,1566]" pageId="19" pageNumber="108">- Petals absent, 1 or 3, branches with 25(8) leaves per node ..............................................................................................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="ED2082F0FFE2FFF37DBFA15D33AC191F" pageId="19" pageNumber="108">
<paragraph id="566B9A55FFE2FFF37DBFA15D33AD197D" blockId="19.[136,1452,1310,1912]" box="[136,1450,1573,1595]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA15D33AD197D" box="[136,1450,1573,1595]" pageId="19" pageNumber="108">
5. Petals usually 3, less often absent or 1 ..................................................................................................................................
<taxonomicName id="91D4E1D6FFE2FFF37862A15D33AD197D" ID-CoL="875R" authorityName="Aublet" authorityYear="1775" box="[1365,1450,1573,1594]" class="Magnoliopsida" family="Vochysiaceae" genus="Vochysia" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="108" phylum="Tracheophyta" rank="genus">
<emphasis id="64A04647FFE2FFF37862A15D33AD197D" box="[1365,1450,1573,1594]" italics="true" pageId="19" pageNumber="108">Vochysia</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="566B9A55FFE2FFF37DBFA13A33AC191F" blockId="19.[136,1452,1310,1912]" box="[136,1451,1602,1624]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA13A33AC191F" box="[136,1451,1602,1624]" pageId="19" pageNumber="108">- Petals 1................................................................................................................................................................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="ED2082F0FFE2FFF37DBFA12733AC19D6" pageId="19" pageNumber="108">
<paragraph id="566B9A55FFE2FFF37DBFA12733AC1933" blockId="19.[136,1452,1310,1912]" box="[136,1451,1631,1653]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA12733AC1933" box="[136,1451,1631,1653]" pageId="19" pageNumber="108">6. Stipules transformed into glands, fruit without a central column ......................................................................................................7</keyLead>
</paragraph>
<paragraph id="566B9A55FFE2FFF37DBFA10433AC19D6" blockId="19.[136,1452,1310,1912]" box="[136,1451,1659,1681]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA10433AC19D6" box="[136,1451,1659,1681]" pageId="19" pageNumber="108">- Stipules obsolete, not transformed into glands, fruit with a thick and persistent central column ......................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="ED2082F0FFE2FFF37DBFA1E133AC1843" pageId="19" pageNumber="108">
<paragraph id="566B9A55FFE2FFF37DBFA1E133AC198C" blockId="19.[136,1452,1310,1912]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA1E133AC198C" pageId="19" pageNumber="108">
7. Unspurred sepal lobes appressed to the spurred sepal and much shorter than it; anther with unilaterally ciliated theca.................... ...........................................................................................................................................................................................
<taxonomicName id="91D4E1D6FFE2FFF3780CA1CE33AC198C" ID-CoL="5TFWP" authorityName="Marcano-Berti" authorityYear="1969" box="[1339,1451,1718,1739]" class="Magnoliopsida" family="Vochysiaceae" genus="Ruizterania" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="108" phylum="Tracheophyta" rank="genus">
<emphasis id="64A04647FFE2FFF3780CA1CE33AC198C" box="[1339,1451,1718,1739]" italics="true" pageId="19" pageNumber="108">Ruizterania</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="566B9A55FFE2FFF37DBFA1AA33AC1843" blockId="19.[136,1452,1310,1912]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA1AA33AC1843" pageId="19" pageNumber="108">
- Unspurred sepal lobes not appressed to the spurred sepal, generally of similar length; anther glabrous or hairy only on back ......... ..................................................................................................................................................................................................
<taxonomicName id="91D4E1D6FFE2FFF37853A19733AC1843" ID-CoL="76DR" authorityName="Aublet" authorityYear="1775" box="[1380,1451,1775,1796]" class="Magnoliopsida" family="Vochysiaceae" genus="Qualea" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="108" phylum="Tracheophyta" rank="genus">
<emphasis id="64A04647FFE2FFF37853A19733AC1843" box="[1380,1451,1775,1796]" italics="true" pageId="19" pageNumber="108">Qualea</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="ED2082F0FFE2FFF37DBFA07433AD1830" pageId="19" pageNumber="108">
<paragraph id="566B9A55FFE2FFF37DBFA07433AD187A" blockId="19.[136,1452,1310,1912]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA07433AD187A" pageId="19" pageNumber="108">
8. Inflorescences in cincinnus with 15 flowers, unguiculate petal, fruits with brittle exocarp that separates unevenly from the endocarp, circularly winged seeds.....................................................................................................................................
<taxonomicName id="91D4E1D6FFE2FFF3780AA05033AD187A" ID-CoL="3FKK" baseAuthorityName="Stafleu" baseAuthorityYear="1952" box="[1341,1450,1832,1853]" class="Magnoliopsida" family="Vochysiaceae" genus="Callisthene" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="108" phylum="Tracheophyta" rank="genus">
<emphasis id="64A04647FFE2FFF3780AA05033AD187A" box="[1341,1450,1832,1853]" italics="true" pageId="19" pageNumber="108">Callisthene</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="566B9A55FFE2FFF37DBFA03D33AD1830" blockId="19.[136,1452,1310,1912]" pageId="19" pageNumber="108">
<keyLead id="ED253960FFE2FFF37DBFA03D33AD1830" pageId="19" pageNumber="108">
- Inflorescences in compound dichasia with more than 16 flowers, sessile petal, fruits with exocarp soft, not brittle, not separated from the endocarp, unilaterally winged seeds ..........................................................................................................
<taxonomicName id="91D4E1D6FFE2FFF37835A01A33AD1830" ID-CoL="98YFN" box="[1282,1450,1890,1911]" class="Magnoliopsida" family="Vochysiaceae" genus="Mahechadendron" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="108" phylum="Tracheophyta" rank="genus">
<emphasis id="64A04647FFE2FFF37835A01A33AD1830" box="[1282,1450,1890,1911]" italics="true" pageId="19" pageNumber="108">Mahechadendron</emphasis>
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