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<document id="84E665C166734C8BA40284A8DE647E68" ID-DOI="10.5281/zenodo.5176688" ID-ISSN="0374-1036" ID-Zenodo-Dep="5176688" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628603916739" checkinUser="carolina" docAuthor="Hájek, Jiří" docDate="2006" docId="DE3A87B12019233AFE24FDCD3168FC2A" docLanguage="en" docName="ActaEntMusNatPra.46.43-56.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 46" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Neptosternus circumductus : REGIMBART 1899" docType="treatment" docVersion="3" lastPageNumber="46" masterDocId="2203FFC9201B2339FFA3FF94352AFFBD" masterDocTitle="The westernmost record of Neptosternus circumductus, and a review of Dytiscidae (Coleoptera) of Baluchistan (Iran, Pakistan)" masterLastPageNumber="56" masterPageNumber="43" pageNumber="45" updateTime="1699047380189" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="D98F996CFB19F390B00B14B980A0DF25" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D7EC3FF4447F2D91C2B1D2128B11407F">
<mods:title id="66F6C855AFB699A584CB955994346498">The westernmost record of Neptosternus circumductus, and a review of Dytiscidae (Coleoptera) of Baluchistan (Iran, Pakistan)</mods:title>
</mods:titleInfo>
<mods:name id="4906EFBD9223F34884607B70B7BF5943" type="personal">
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</mods:role>
<mods:namePart id="4CA0B7E8A5A24C9DED4A9C7E3954D323">Hájek, Jiří</mods:namePart>
</mods:name>
<mods:typeOfResource id="E549A9C0C4E4E12D3155C239DB918685">text</mods:typeOfResource>
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<mods:title id="AEB2F9C178235C0470CA83992400BAD7">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="481F838E4E51461307C5653559EC569E">
<mods:date id="19B651811B7D6B8F5177CA24692140EC">2006</mods:date>
<mods:detail id="94461A7F9864D8A58DA79AB630B98CF3" type="pubDate">
<mods:number id="DCD07645FC237441D75A1C7AAA7AC662">2006-11-06</mods:number>
</mods:detail>
<mods:detail id="16AF14D20F60B9A42BBFA50E1406E047" type="volume">
<mods:number id="9E6B71F64E625C54971F0FC9AC1BFD1C">46</mods:number>
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<mods:start id="6054118C4DB4E63433AAC44B8B26D83D">43</mods:start>
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<mods:classification id="C539B7B7ACA8A8149B2F005A8BE1A40C">journal article</mods:classification>
<mods:identifier id="E91D27653EA3904B34624FC3488D8401" type="DOI">10.5281/zenodo.5176688</mods:identifier>
<mods:identifier id="F25A69A31E2B443FEFB683F4C24C26DF" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="0BE591F9D633ED696ED0E730AEDB5865" type="Zenodo-Dep">5176688</mods:identifier>
</mods:mods>
<treatment id="DE3A87B12019233AFE24FDCD3168FC2A" LSID="urn:lsid:plazi:treatment:DE3A87B12019233AFE24FDCD3168FC2A" httpUri="http://treatment.plazi.org/id/DE3A87B12019233AFE24FDCD3168FC2A" lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45">
<subSubSection id="1E89652C2019233BFE24FDCD365DFDCD" box="[391,887,600,625]" pageId="2" pageNumber="45" type="nomenclature">
<paragraph id="562C36A72019233BFE24FDCD365DFDCD" blockId="2.[391,887,600,625]" box="[391,887,600,625]" pageId="2" pageNumber="45">
<heading id="0D6481CB2019233BFE24FDCD365DFDCD" bold="true" box="[391,887,600,625]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="45" reason="3">
<taxonomicName id="91934D242019233BFE24FDCD365DFDCD" ID-CoL="473WX" authority="Regimbart, 1899" authorityName=": REGIMBART" authorityYear="1899" box="[391,887,600,625]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="circumductus">
<emphasis id="64E7EAB52019233BFE24FDCD365DFDCD" bold="true" box="[391,887,600,625]" pageId="2" pageNumber="45">
<emphasis id="64E7EAB52019233BFE24FDCD3798FDCD" bold="true" box="[391,690,600,624]" italics="true" pageId="2" pageNumber="45">Neptosternus circumductus</emphasis>
<bibRefCitation id="32024B562019233BFD19FDCC365DFDCD" author="REGIMBART M." box="[698,887,600,625]" pageId="2" pageNumber="45" pagination="186 - 367" refId="ref8999" refString="REGIMBART M. 1899: Revision des Dytiscidae de la region Indo-sino-malaise. Annales de la Societe Entomologique de France 68: 186 - 367." type="journal article" year="1899">Régimbart, 1899</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1E89652C2019233BFDF3FDED3784FD31" box="[592,686,633,652]" pageId="2" pageNumber="45" type="description">
<paragraph id="562C36A72019233BFDF3FDED3784FD31" blockId="2.[151,1127,633,793]" box="[592,686,633,652]" pageId="2" pageNumber="45">
(
<figureCitation id="CEA82A222019233BFDF5FDED378DFD31" box="[598,679,633,652]" captionStart="Figs" captionStartId="4.[151,190,1461,1480]" captionTargetBox="[189,1083,220,1429]" captionTargetId="figure-10@4.[189,1083,220,1429]" captionTargetPageId="4" captionText="Figs. 1-4. 1, 3: Neptosternus circumductus Régimbart, 1899. 1 – habitus (a – aberrant specimen from Baluchistan; b – lectotype from India); 3 – median lobe of aedeagus (a – apex of median lobe in ventral view; b – median lobe in lateral view). 2, 4: Laccophilus inefficiens (Walker, 1859). 2 – habitus; 4 – median lobe of aedeagus in lateral view. Scale bars: 2 mm (Figs 1-2); 0.25 mm (Fig. 3); 0.5 mm (Fig. 4)." figureDoi="http://doi.org/10.5281/zenodo.5176690" httpUri="https://zenodo.org/record/5176690/files/figure.png" pageId="2" pageNumber="45">Figs. 1, 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1E89652C2019233BFF34FD08314DFD74" pageId="2" pageNumber="45" type="reference_group">
<paragraph id="562C36A72019233BFF34FD08314DFD13" blockId="2.[151,1127,633,793]" box="[151,1127,667,686]" pageId="2" pageNumber="45">
<treatmentCitationGroup id="768311892019233BFF34FD08314DFD13" box="[151,1127,667,686]" pageId="2" pageNumber="45">
<treatmentCitation id="D73210B62019233BFF34FD08376FFD13" author="REGIMBART M." box="[151,581,667,686]" page="83" pageId="2" pageNumber="45" year="1899">
<taxonomicName id="91934D242019233BFF34FD08370CFD13" ID-CoL="473WX" authority=": REGIMBART (1899)" authorityName=": REGIMBART" authorityYear="1899" box="[151,550,667,686]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="circumductus">
<emphasis id="64E7EAB52019233BFF34FD083454FD13" box="[151,382,667,686]" italics="true" pageId="2" pageNumber="45">Neptosternus circumductus</emphasis>
:
<bibRefCitation id="32024B562019233BFE29FD0F370CFD13" author="REGIMBART M." box="[394,550,667,686]" pageId="2" pageNumber="45" pagination="186 - 367" refId="ref8999" refString="REGIMBART M. 1899: Revision des Dytiscidae de la region Indo-sino-malaise. Annales de la Societe Entomologique de France 68: 186 - 367." type="journal article" year="1899">RÉGIMBART (1899)</bibRefCitation>
</taxonomicName>
: 83
</treatmentCitation>
(original description; Genji, South Arkot [
<collectingRegion id="9457F8452019233BFC17FD0F3134FD13" box="[948,1054,667,686]" country="India" name="Tamil Nadu" pageId="2" pageNumber="45">Tamil Nadu</collectingRegion>
,
<collectingCountry id="2E8476372019233BFB84FD0F3172FD13" box="[1063,1112,667,686]" name="India" pageId="2" pageNumber="45">India</collectingCountry>
]);
</treatmentCitationGroup>
</paragraph>
<paragraph id="562C36A72019233BFF15FD22314DFD74" blockId="2.[151,1127,633,793]" box="[182,1127,693,713]" pageId="2" pageNumber="45">
<bibRefCitation id="32024B562019233BFF15FD223469FD75" author="VAZIRANI T. G." box="[182,323,693,713]" pageId="2" pageNumber="45" pagination="423 - 428" refId="ref9090" refString="VAZIRANI T. G. 1953: On a collection of aquatic Coleoptera from the Mettur Dam, Salem District, Madras, with the description of a new species. Records of the Indian Museum (Calcutta) 50 (1952): 423 - 428." type="journal article" year="1953">VAZIRANI (1953)</bibRefCitation>
: 424 (description of variability; Madras [
<collectingRegion id="9457F8452019233BFD03FD223626FD74" box="[672,780,694,713]" country="India" name="Tamil Nadu" pageId="2" pageNumber="45">Tamil Nadu</collectingRegion>
],
<collectingRegion id="9457F8452019233BFCB9FD223686FD74" box="[794,940,694,713]" country="India" name="Madhya Pradesh" pageId="2" pageNumber="45">Madhya Pradesh</collectingRegion>
);
<bibRefCitation id="32024B562019233BFC18FD223162FD74" author="VAZIRANI T. G." box="[955,1096,694,713]" pageId="2" pageNumber="45" pagination="14 - 17" refId="ref9138" refString="VAZIRANI T. G. 1963: On the Indian species of the genus Neptosternus Sharp (Dytiscidae-Coleoptera) with the description of a new species. Bulletin of Entomology, Loyola College (Madras) 4: 14 - 17." type="journal article" year="1963">VAZIRANI (1963)</bibRefCitation>
: 16
</paragraph>
</subSubSection>
<subSubSection id="1E89652C2019233BFF15FD443459FCA4" pageId="2" pageNumber="45" type="description">
<paragraph id="562C36A72019233BFF15FD44314DFD5F" blockId="2.[151,1127,633,793]" box="[182,1127,720,740]" pageId="2" pageNumber="45">
(
<collectingRegion id="9457F8452019233BFF1FFD4435C5FD5E" box="[188,239,720,739]" country="India" name="Bihar" pageId="2" pageNumber="45">Bihar</collectingRegion>
); (1969a): 251 (Maharahstra,
<collectingRegion id="9457F8452019233BFE53FD443702FD5E" box="[496,552,720,739]" country="India" name="Odisha" pageId="2" pageNumber="45">Orissa</collectingRegion>
,
<collectingRegion id="9457F8452019233BFD91FD453781FD5E" box="[562,683,720,740]" country="India" name="Uttar Pradesh" pageId="2" pageNumber="45">Uttar Pradesh</collectingRegion>
);
<bibRefCitation id="32024B562019233BFD19FD453680FD5E" author="HOLMEN M. &amp; VAZIRANI T. G." box="[698,938,720,740]" pageId="2" pageNumber="45" pagination="19 - 31" refId="ref8718" refString="HOLMEN M. &amp; VAZIRANI T. G. 1990: Notes on the genera Neptosternus Sharp and Copelatus Erichson from Sri Lanka and India with the description of new species (Coleoptera: Dytiscidae). Koleopterologische Rundschau 60: 19 - 31." type="journal article" year="1990">HOLMEN &amp; VAZIRANI (1990)</bibRefCitation>
: 20 (
<collectingCountry id="2E8476372019233BFC75FD44312CFD5E" box="[982,1030,720,739]" name="India" pageId="2" pageNumber="45">India</collectingCountry>
);
<bibRefCitation id="32024B562019233BFBB5FD45314DFD5F" author="HENDRICH L. &amp; BALKE M." box="[1046,1127,721,740]" pageId="2" pageNumber="45" pagination="57 - 62" refId="ref8424" refString="HENDRICH L. &amp; BALKE M. 1999: Two new species of Neptosternus Sharp 1882 from Southern India (Coleoptera: Dytiscidae). Linzer Biolologische Beitrage 31: 57 - 62." type="journal article" year="1999">HENDRICH</bibRefCitation>
</paragraph>
<paragraph id="562C36A72019233BFF15FD7F314CFD43" blockId="2.[151,1127,633,793]" box="[182,1126,747,766]" pageId="2" pageNumber="45">
&amp; BALKE (1999): 60 (
<collectingCountry id="2E8476372019233BFED5FD7F348CFD43" box="[374,422,747,766]" name="India" pageId="2" pageNumber="45">India</collectingCountry>
); HENDRICH &amp; BALKE (2000): 1286 (
<collectingRegion id="9457F8452019233BFD45FD7F3650FD43" box="[742,890,747,766]" country="India" name="Madhya Pradesh" pageId="2" pageNumber="45">Madhya Pradesh</collectingRegion>
,
<collectingRegion id="9457F8452019233BFC26FD7F36CDFD43" box="[901,999,747,766]" country="India" name="Maharashtra" pageId="2" pageNumber="45">Maharastra</collectingRegion>
,
<collectingRegion id="9457F8452019233BFC51FD7F3100FD43" box="[1010,1066,747,766]" country="India" name="Odisha" pageId="2" pageNumber="45">Orissa</collectingRegion>
,
<collectingRegion id="9457F8452019233BFB96FD7F314CFD43" box="[1077,1126,747,766]" country="India" name="Tamil Nadu" pageId="2" pageNumber="45">Tamil</collectingRegion>
</paragraph>
<paragraph id="562C36A72019233BFF15FC923459FCA4" blockId="2.[151,1127,633,793]" box="[182,371,774,793]" pageId="2" pageNumber="45">
Nadu,
<collectingRegion id="9457F8452019233BFF53FC923443FCA4" box="[240,361,774,793]" country="India" name="Uttar Pradesh" pageId="2" pageNumber="45">Uttar Pradesh</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="1E89652C2019233BFF34FCD73750FB0D" pageId="2" pageNumber="45" type="materials_examined">
<paragraph id="562C36A72019233BFF34FCD7369BFCEB" blockId="2.[151,1127,834,1201]" box="[151,945,834,854]" pageId="2" pageNumber="45">
<emphasis id="64E7EAB52019233BFF34FCD7343AFCE8" bold="true" box="[151,272,834,854]" pageId="2" pageNumber="45">
<typeStatus id="892888052019233BFF34FCD735E9FCEB" box="[151,195,835,854]" pageId="2" pageNumber="45">Type</typeStatus>
locality.
</emphasis>
‘l’Inde anglaise. Genji (South Arkot)’ [Province
<collectingRegion id="9457F8452019233BFD14FCD73635FCEB" box="[695,799,835,854]" country="India" name="Tamil Nadu" pageId="2" pageNumber="45">Tamil Nadu</collectingRegion>
, southern
<collectingCountry id="2E8476372019233BFCDBFCD73682FCEB" box="[888,936,835,854]" name="India" pageId="2" pageNumber="45">India</collectingCountry>
].
</paragraph>
<paragraph id="562C36A72019233BFF34FCC934A2FB96" blockId="2.[151,1127,834,1201]" pageId="2" pageNumber="45">
<emphasis id="64E7EAB52019233BFF34FCC93435FCCD" bold="true" box="[151,287,861,880]" pageId="2" pageNumber="45">Type material.</emphasis>
<materialsCitation id="E6FB3CFA2019233BFE85FCCA37ECFC1B" collectionCode="MNHN" country="India" determinerName="Regimbart" location="INDE" pageId="2" pageNumber="45" specimenCount="1" typeStatus="lectotype">
<typeStatus id="892888052019233BFE85FCCA3455FCD2" box="[294,383,862,881]" pageId="2" pageNumber="45" type="lectotype">LECTOTYPE</typeStatus>
: 3, by present designation (
<collectionCode id="3082AE622019233BFDDDFCC937EEFCCD" box="[638,708,861,880]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="45">MNHN</collectionCode>
), labelled: ‘
<collectingCountry id="2E8476372019233BFC8AFCC9364AFCCD" box="[809,864,861,880]" name="India" pageId="2" pageNumber="45">INDE</collectingCountry>
ANGLAIS, Maindron 1881 / MUSEUM PARIS, COLL.
<collectingCountry id="2E8476372019233BFE26FCEC34CCFC36" box="[389,486,888,907]" name="Mauritius" pageId="2" pageNumber="45">MAURICE</collectingCountry>
REGIMBART, 1908 [printed] / TYPE [red label, printed] /
<taxonomicName id="91934D242019233BFC54FCEC3412FC1B" authority="Reg." authorityName="Reg." authorityYear="2006" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="circumductus" status="sp. nov.">Neptosternus circumductus Rég.</taxonomicName>
<taxonomicNameLabel id="7FD457CE2019233BFE9FFC073441FC1B" box="[316,363,915,934]" pageId="2" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
typ. [hw],
<determinerName id="A3F0DA632019233BFE6BFC07370BFC1B" box="[456,545,915,934]" pageId="2" pageNumber="45">Régimbart</determinerName>
det. 1896 [printed]
</materialsCitation>
/
<typeStatus id="892888052019233BFD75FC073677FC1B" box="[726,861,915,934]" pageId="2" pageNumber="45" type="lectotype">LECTOTYPUS</typeStatus>
3,
<taxonomicName id="91934D242019233BFCDDFC07344EFC7D" authority="Regimbart, 1899" authorityName=": REGIMBART" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="circumductus">
NEPTOSTERNUS circumductus
<bibRefCitation id="32024B562019233BFF77FC39344EFC7D" author="REGIMBART M." box="[212,356,941,960]" pageId="2" pageNumber="45" pagination="186 - 367" refId="ref8999" refString="REGIMBART M. 1899: Revision des Dytiscidae de la region Indo-sino-malaise. Annales de la Societe Entomologique de France 68: 186 - 367." type="journal article" year="1899">Régimbart, 1899</bibRefCitation>
</taxonomicName>
, Jiří Hájek des. 2002 [red label, printed]’.
<specimenCount id="4095FD2E2019233BFD68FC39367FFC7D" box="[715,853,941,960]" pageId="2" pageNumber="45" type="generic" typeStatus="paralectotypes">2 paralectotypes</specimenCount>
(
<collectionCode id="3082AE622019233BFCC2FC39368CFC7D" box="[865,934,941,960]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="45">MNHN</collectionCode>
), labelled: ‘MUSEUM PARIS,
<collectingCountry id="2E8476372019233BFF7EFC5C343AFC66" box="[221,272,968,987]" name="India" pageId="2" pageNumber="45">INDE</collectingCountry>
ANGLAISE, ARKOT NORD, GENJI, M. MAINDRON 1881 / Genji (South Arkot),
<collectingCountry id="2E8476372019233BFC53FC5C313CFC66" box="[1008,1046,968,987]" name="India" pageId="2" pageNumber="45">Inde</collectingCountry>
anglaise,
<date id="222D10672019233BFF34FC773423FC4B" box="[151,265,995,1014]" pageId="2" pageNumber="45" value="1881-01">Janvier 1881</date>
/
<taxonomicName id="91934D242019233BFEBDFC7734EFFC4B" authority="Reg." authorityName=": REGIMBART" authorityYear="1899" box="[286,453,995,1014]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="circumductus">circumductus Rég.</taxonomicName>
,
<collectingCountry id="2E8476372019233BFE73FC7734DCFC4B" box="[464,502,995,1014]" name="India" pageId="2" pageNumber="45">Inde</collectingCountry>
(typ. museum) [hw]’ and the respective
<typeStatus id="892888052019233BFCC5FC7736F0FC4B" box="[870,986,995,1014]" pageId="2" pageNumber="45" type="paralectotype">paralectotype</typeStatus>
labels. I herein designate the
<typeStatus id="892888052019233BFEAEFC693470FBAD" box="[269,346,1021,1040]" pageId="2" pageNumber="45" type="lectotype">lectotype</typeStatus>
to fix the identity of this species, as available taxonomic works do not allow unambiguous identification of specimens.
</paragraph>
<paragraph id="562C36A72019233BFF34FBA63750FB0D" blockId="2.[151,1127,834,1201]" pageId="2" pageNumber="45">
<emphasis id="64E7EAB52019233BFF34FBA63485FBF8" bold="true" box="[151,431,1074,1093]" pageId="2" pageNumber="45">Additional material examined.</emphasis>
<materialsCitation id="E6FB3CFA2019233BFE14FBA63642FBDD" collectionCode="NMW" collectorName="Boukal &amp; Kejval'" country="India" elevation="900" location="Kunchappanai" longitude="11.366667" municipality="Nilgiri Hills" pageId="2" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu">
<emphasis id="64E7EAB52019233BFE14FBA634D6FBF8" bold="true" box="[439,508,1074,1093]" pageId="2" pageNumber="45">
<collectingCountry id="2E8476372019233BFE14FBA634D3FBF8" box="[439,505,1074,1093]" name="India" pageId="2" pageNumber="45">INDIA</collectingCountry>
:
</emphasis>
13
<specimenCount id="4095FD2E2019233BFD86FBA73716FBFA" box="[549,572,1071,1095]" pageId="2" pageNumber="45" type="female">1♀</specimenCount>
, ‘S-India,
<collectingRegion id="9457F8452019233BFD3DFBA7362DFBFB" box="[670,775,1075,1094]" country="India" name="Tamil Nadu" pageId="2" pageNumber="45">Tamil Nadu</collectingRegion>
,
<collectingMunicipality id="B648ACDD2019233BFCB1FBA73657FBFB" box="[786,893,1075,1094]" pageId="2" pageNumber="45">Nilgiri Hills</collectingMunicipality>
,
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SE Kotagiri
</locationDeviation>
,
<location id="534C607C2019233BFB98FBA735C5FBDD" LSID="urn:lsid:plazi:treatment:DE3A87B12019233AFE24FDCD3168FC2A:534C607C2019233BFB98FBA735C5FBDD" country="India" longitude="11.366667" municipality="Nilgiri Hills" name="Kunchappanai" pageId="2" pageNumber="45" stateProvince="Tamil Nadu">Kunchappanai</location>
,
<quantity id="916B9B422019233BFF5BFBD9341BFBDC" box="[248,305,1101,1121]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="45" unit="m" value="900.0">
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</quantity>
,
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, 17-28.11.1993, leg.
<collectorName id="FB6653712019233BFD21FBD93795FBDD" box="[642,703,1101,1120]" pageId="2" pageNumber="45">Boukal</collectorName>
&amp;
<collectorName id="FB6653712019233BFD7EFBD93631FBDD" box="[733,795,1101,1120]" pageId="2" pageNumber="45">Kejval’</collectorName>
(
<collectionCode id="3082AE622019233BFC84FBD93648FBDD" box="[807,866,1101,1120]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="45">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E6FB3CFA2019233BFCD1FBD93641FBC6" collectingDate="1988-08" collectionCode="LHCB, NHMB, NMPC" collectorName="C India, MP &amp; C. Werner" country="India" location="Panna National Park" pageId="2" pageNumber="45" specimenCount="1" stateProvince="Madhya Pradesh">
4 ex, ‘
<collectorName id="FB6653712019233BFC04FBD9313EFBDC" box="[935,1044,1101,1121]" pageId="2" pageNumber="45">C India, MP</collectorName>
[
<collectingRegion id="9457F8452019233BFBBCFBD935F4FBC6" country="India" name="Madhya Pradesh" pageId="2" pageNumber="45">Madhya Pradesh</collectingRegion>
],
<location id="534C607C2019233BFF4EFBFC34B5FBC6" LSID="urn:lsid:plazi:treatment:DE3A87B12019233AFE24FDCD3168FC2A:534C607C2019233BFF4EFBFC34B5FBC6" box="[237,415,1128,1147]" country="India" name="Panna National Park" pageId="2" pageNumber="45" stateProvince="Madhya Pradesh">Panna National Park</location>
,
<date id="222D10672019233BFE0BFBFC34D4FBC6" box="[424,510,1128,1147]" pageId="2" pageNumber="45" value="1988-08">
<collectingDate id="3269E98F2019233BFE0BFBFC34D4FBC6" box="[424,510,1128,1147]" pageId="2" pageNumber="45" value="1988-08">VIII.1988</collectingDate>
</date>
,
<collectorName id="FB6653712019233BFDA4FBFC3774FBC6" box="[519,606,1128,1147]" pageId="2" pageNumber="45">C. Werner</collectorName>
leg.’ (
<collectionCode id="3082AE622019233BFD36FBFC37FBFBC6" box="[661,721,1128,1147]" pageId="2" pageNumber="45">LHCB</collectionCode>
,
<collectionCode id="3082AE622019233BFD79FBFC3636FBC6" box="[730,796,1128,1147]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="2" pageNumber="45">NHMB</collectionCode>
,
<collectionCode id="3082AE622019233BFC86FBFC364FFBC6" box="[805,869,1128,1147]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="45">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E6FB3CFA2019233BFCD7FBFC375CFB0D" collectingDate="2000-04-11" collectingDateMax="2000-04-12" collectingDateMin="2000-04-11" collectionCode="LHBC, NMPC" collectorName="Iran &amp; Czech Biological Expedition, J &amp; M. Mikat" country="Iran" elevation="100" latitude="25.733334" location="Pir Sohrab" longLatPrecision="1245" longitude="60.833332" pageId="2" pageNumber="45" specimenCount="84" stateProvince="Sistan va Baluchestan">
<emphasis id="64E7EAB52019233BFCD7FBFC369BFBC6" bold="true" box="[884,945,1128,1147]" pageId="2" pageNumber="45">
<collectingCountry id="2E8476372019233BFCD7FBFC3687FBC6" box="[884,941,1128,1147]" name="Iran" pageId="2" pageNumber="45">IRAN</collectingCountry>
:
</emphasis>
<specimenCount id="4095FD2E2019233BFC14FBFC36DDFBC6" box="[951,1015,1128,1147]" pageId="2" pageNumber="45" type="generic">84 exs.</specimenCount>
, ‘IRAN,
<date id="222D10672019233BFBE5FBFC35DDFB2B" pageId="2" pageNumber="45" value="2000-04-11" valueMax="2000-04-12" valueMin="2000-04-11">
<collectingDate id="3269E98F2019233BFBE5FBFC35DDFB2B" pageId="2" pageNumber="45" value="2000-04-11" valueMax="2000-04-12" valueMin="2000-04-11">11.- 12.IV.2000</collectingDate>
</date>
,
<collectingRegion id="9457F8452019233BFF5CFB1734C7FB2B" box="[255,493,1155,1174]" country="Iran" name="Sistan va Baluchestan" pageId="2" pageNumber="45">Sistan va Baluchestan Prov.</collectingRegion>
,
<location id="534C607C2019233BFE57FB173764FB2B" LSID="urn:lsid:plazi:treatment:DE3A87B12019233AFE24FDCD3168FC2A:534C607C2019233BFE57FB173764FB2B" box="[500,590,1155,1174]" country="Iran" latitude="25.733334" longLatPrecision="1245" longitude="60.833332" name="Pir Sohrab" pageId="2" pageNumber="45" stateProvince="Sistan va Baluchestan">Pir Sohrab</location>
env.,
<quantity id="916B9B422019233BFD23FB17379DFB2B" box="[640,695,1155,1174]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="45" unit="m" value="100.0">
<elevation id="DDBED1942019233BFD23FB17379DFB2B" box="[640,695,1155,1174]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="45" unit="m" value="100.0">100 m</elevation>
</quantity>
, (
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,
<geoCoordinate id="33A750602019233BFCBAFB163648FB2B" box="[793,866,1153,1174]" degrees="60" direction="east" minutes="50" orientation="longitude" pageId="2" pageNumber="45" precision="925" value="60.833332">60°50′E</geoCoordinate>
)’, ‘
<collectorName id="FB6653712019233BFCD9FB17368AFB2B" box="[890,928,1155,1174]" pageId="2" pageNumber="45">Iran</collectorName>
2000
<collectorName id="FB6653712019233BFC77FB173423FB0C" pageId="2" pageNumber="45">Czech Biological Expedition, J</collectorName>
. Hájek &amp;
<collectorName id="FB6653712019233BFEC1FB0A3499FB0D" box="[354,435,1181,1201]" pageId="2" pageNumber="45">M. Mikát</collectorName>
leg.’ (
<collectionCode id="3082AE622019233BFE48FB09370DFB0D" box="[491,551,1181,1200]" pageId="2" pageNumber="45">LHBC</collectionCode>
,
<collectionCode id="3082AE622019233BFD93FB09375AFB0D" box="[560,624,1181,1200]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="45">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1E89652C2019233AFF34FB533585FCCA" lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45" type="description">
<paragraph id="562C36A72019233BFF34FB53341CFAA3" blockId="2.[151,1127,1223,1598]" pageId="2" pageNumber="45">
<emphasis id="64E7EAB52019233BFF34FB533411FB62" bold="true" box="[151,315,1223,1247]" pageId="2" pageNumber="45">Redescription.</emphasis>
Large, elongate species; body arched in lateral view. Head and pronotum testaceous, elytra brown with variable yellowish pattern (
<figureCitation id="CEA82A222019233BFD54FB73361FFB42" box="[759,821,1255,1279]" captionStart="Figs" captionStartId="4.[151,190,1461,1480]" captionTargetBox="[189,1083,220,1429]" captionTargetId="figure-10@4.[189,1083,220,1429]" captionTargetPageId="4" captionText="Figs. 1-4. 1, 3: Neptosternus circumductus Régimbart, 1899. 1 – habitus (a – aberrant specimen from Baluchistan; b – lectotype from India); 3 – median lobe of aedeagus (a – apex of median lobe in ventral view; b – median lobe in lateral view). 2, 4: Laccophilus inefficiens (Walker, 1859). 2 – habitus; 4 – median lobe of aedeagus in lateral view. Scale bars: 2 mm (Figs 1-2); 0.25 mm (Fig. 3); 0.5 mm (Fig. 4)." figureDoi="http://doi.org/10.5281/zenodo.5176690" httpUri="https://zenodo.org/record/5176690/files/figure.png" pageId="2" pageNumber="45">Fig. 1</figureCitation>
). Ventral parts and appendages testaceous.
</paragraph>
<paragraph id="562C36A72019233BFF15FAB33170FAE2" blockId="2.[151,1127,1223,1598]" pageId="2" pageNumber="45">
Measurements. Total length of beetle
<quantity id="916B9B422019233BFDE2FAB33790FA83" box="[577,698,1319,1343]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.6" pageId="2" pageNumber="45" unit="mm" value="3.75" valueMax="3.9" valueMin="3.6">3.6-3.9 mm</quantity>
(
<typeStatus id="892888052019233BFD64FAB33600FA82" box="[711,810,1319,1343]" pageId="2" pageNumber="45" type="lectotype">lectotype</typeStatus>
<quantity id="916B9B422019233BFC93FAB336AFFA82" box="[816,901,1319,1343]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="45" unit="mm" value="3.7">3.7 mm</quantity>
); length without head
<quantity id="916B9B422019233BFF34FAD33438FAE3" box="[151,274,1351,1375]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="2" pageNumber="45" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3-3.5 mm</quantity>
(
<typeStatus id="892888052019233BFE83FAD334A9FAE2" box="[288,387,1351,1375]" pageId="2" pageNumber="45" type="lectotype">lectotype</typeStatus>
<quantity id="916B9B422019233BFE2AFAD334F5FAE2" box="[393,479,1351,1375]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="45" unit="mm" value="3.3">3.3 mm</quantity>
); maximum width of beetle 1.9-2.1 mm (
<typeStatus id="892888052019233BFC33FAD336D9FAE2" box="[912,1011,1351,1375]" pageId="2" pageNumber="45" type="lectotype">lectotype</typeStatus>
2.0 mm).
</paragraph>
<paragraph id="562C36A72019233BFF15FAF337FDFA23" blockId="2.[151,1127,1223,1598]" pageId="2" pageNumber="45">Head. Coloration testaceous. Reticulation composed of slightly impressed polygonal meshes, but medially densely punctate. Mouthparts testaceous.</paragraph>
<paragraph id="562C36A72019233BFF15FA3335F8F983" blockId="2.[151,1127,1223,1598]" pageId="2" pageNumber="45">Pronotum. Coloration testaceous, with thin brown stripe along posterior margin, darker anteriorly along the row of large punctures, and in some specimens also with median brown spot along anterior margin. Reticulation similar to that on head but more impressed. Surface densely punctate; basomedially, along anterior margin and laterally with some larger punctures.</paragraph>
<paragraph id="562C36A72018233AFF33FF4B3618FE2A" blockId="3.[113,1090,223,919]" pageId="3" pageNumber="46">
Elytra. Blackish brown, with uninterrupted lateral yellow margin and two variable, sometimes confluent yellowish spots on disc of each elytron; lateral margin dilated apically towards suture in form of a triangle (
<figureCitation id="CEA82A222018233AFE7CFE8B3737FE8A" box="[479,541,287,311]" captionStart="Figs" captionStartId="4.[151,190,1461,1480]" captionTargetBox="[189,1083,220,1429]" captionTargetId="figure-10@4.[189,1083,220,1429]" captionTargetPageId="4" captionText="Figs. 1-4. 1, 3: Neptosternus circumductus Régimbart, 1899. 1 – habitus (a – aberrant specimen from Baluchistan; b – lectotype from India); 3 – median lobe of aedeagus (a – apex of median lobe in ventral view; b – median lobe in lateral view). 2, 4: Laccophilus inefficiens (Walker, 1859). 2 – habitus; 4 – median lobe of aedeagus in lateral view. Scale bars: 2 mm (Figs 1-2); 0.25 mm (Fig. 3); 0.5 mm (Fig. 4)." figureDoi="http://doi.org/10.5281/zenodo.5176690" httpUri="https://zenodo.org/record/5176690/files/figure.png" pageId="3" pageNumber="46">Fig. 1</figureCitation>
). Microreticulation consisting of slightly transverse polygonal meshes. Large punctures sparse on disc. Three rows of serial punctures perceptible (cf.
<bibRefCitation id="32024B562018233AFF3FFEF4348EFECA" author="HENDRICH L. &amp; BALKE M." box="[156,420,351,375]" pageId="3" pageNumber="46" pagination="53 - 97" refId="ref8388" refString="HENDRICH L. &amp; BALKE M. 1997: Taxonomische Revision der sudostasiatischen Arten der Gattung Neptosternus Sharp, 1882 (Coleoptera: Dytiscidae: Laccophilinae). Koleopterologische Rundschau 67: 53 - 97." type="journal article" year="1997">HENDRICH &amp; BALKE 1997</bibRefCitation>
): discal and first lateral row distinct, second lateral row perceptible only in second third of elytral length, sutural row not present.
</paragraph>
<paragraph id="562C36A72018233AFF33FE0B3478FE6A" blockId="3.[113,1090,223,919]" pageId="3" pageNumber="46">Ventral side. Venter testaceous and microreticulated. Abdominal sternites with several large setigerous punctures.</paragraph>
<paragraph id="562C36A72018233AFF33FE743407FDAA" blockId="3.[113,1090,223,919]" pageId="3" pageNumber="46">Legs. Completely testaceous. Male tarsomeres of fore and middle legs not dilated. Claws simple and equal.</paragraph>
<paragraph id="562C36A72018233AFF33FD8B37B5FD8A" blockId="3.[113,1090,223,919]" box="[144,671,543,567]" pageId="3" pageNumber="46">
Male genitalia. Shape of median lobe as in
<figureCitation id="CEA82A222018233AFDF8FD8B37B1FD8A" box="[603,667,543,567]" captionStart="Figs" captionStartId="4.[151,190,1461,1480]" captionTargetBox="[189,1083,220,1429]" captionTargetId="figure-10@4.[189,1083,220,1429]" captionTargetPageId="4" captionText="Figs. 1-4. 1, 3: Neptosternus circumductus Régimbart, 1899. 1 – habitus (a – aberrant specimen from Baluchistan; b – lectotype from India); 3 – median lobe of aedeagus (a – apex of median lobe in ventral view; b – median lobe in lateral view). 2, 4: Laccophilus inefficiens (Walker, 1859). 2 – habitus; 4 – median lobe of aedeagus in lateral view. Scale bars: 2 mm (Figs 1-2); 0.25 mm (Fig. 3); 0.5 mm (Fig. 4)." figureDoi="http://doi.org/10.5281/zenodo.5176690" httpUri="https://zenodo.org/record/5176690/files/figure.png" pageId="3" pageNumber="46">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="562C36A72018233AFF33FDAB3675FDEA" blockId="3.[113,1090,223,919]" box="[144,863,575,599]" pageId="3" pageNumber="46">Female. Similar to male. Last abdominal sternite narrowed apically.</paragraph>
<paragraph id="562C36A72018233AFFD2FDCB3497FD4A" blockId="3.[113,1090,223,919]" pageId="3" pageNumber="46">
<emphasis id="64E7EAB52018233AFFD2FDCB35CAFDCA" bold="true" box="[113,224,607,631]" pageId="3" pageNumber="46">Affinities.</emphasis>
According to the elongate shape of body and elytral coloration,
<taxonomicName id="91934D242018233AFC30FDF4316BFDCA" authorityName=": REGIMBART" authorityYear="1899" box="[915,1089,607,631]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="circumductus">
<emphasis id="64E7EAB52018233AFC30FDF4316BFDCA" box="[915,1089,607,631]" italics="true" pageId="3" pageNumber="46">N. circumductus</emphasis>
</taxonomicName>
stands relatively solitary within the Indian and Oriental species of the genus. Habitually, it is more similar to the African species arranged by OMER- COOPER (1970) in species group I (
<taxonomicName id="91934D242018233AFB8BFD3435E8FD6A" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="64E7EAB52018233AFB8BFD3435E8FD6A" italics="true" pageId="3" pageNumber="46">N. ornatus</emphasis>
</taxonomicName>
-species group). At present, this group contains 14 species occurring in southern and central Africa and
<collectingCountry id="2E8476372018233AFE94FD4B3493FD4A" box="[311,441,735,759]" name="Madagascar" pageId="3" pageNumber="46">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="562C36A72018233AFFD2FD6B3585FCCA" blockId="3.[113,1090,223,919]" pageId="3" pageNumber="46">
<emphasis id="64E7EAB52018233AFFD2FD6B35E5FCAA" bold="true" box="[113,207,767,791]" pageId="3" pageNumber="46">Habitat.</emphasis>
The Iranian specimens of
<taxonomicName id="91934D242018233AFE5BFC943782FCAA" authorityName=": REGIMBART" authorityYear="1899" box="[504,680,767,791]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="circumductus">
<emphasis id="64E7EAB52018233AFE5BFC943782FCAA" box="[504,680,767,791]" italics="true" pageId="3" pageNumber="46">N. circumductus</emphasis>
</taxonomicName>
were collected in a small pool of an almost dry river bed (around 5 metres in diameter and
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in depth). The bottom consist- ed of sand and few larger stones (
<figureCitation id="CEA82A222018233AFE78FCAB3714FCEA" box="[475,574,831,855]" captionStart="Figs" captionStartId="5.[113,152,1576,1595]" captionTargetBox="[114,1089,220,1538]" captionTargetPageId="5" captionText="Figs. 5-6. Locality ‘Pir Sohrab’, 100 m a.s.l., 12.iv.2000, Sistan va Baluchestan Province, Iran. Photo author." figureDoi="http://doi.org/10.5281/zenodo.5176692" httpUri="https://zenodo.org/record/5176692/files/figure.png" pageId="3" pageNumber="46">Figs. 5-6</figureCitation>
). All specimens were collected with a dip net at night.
</paragraph>
</subSubSection>
<subSubSection id="1E89652C2018233AFFD2FCEB3168FC2A" box="[113,1090,895,919]" pageId="3" pageNumber="46" type="distribution">
<paragraph id="562C36A72018233AFFD2FCEB3168FC2A" blockId="3.[113,1090,223,919]" box="[113,1090,895,919]" pageId="3" pageNumber="46">
<emphasis id="64E7EAB52018233AFFD2FCEB3428FC2A" bold="true" box="[113,258,895,919]" pageId="3" pageNumber="46">Distribution.</emphasis>
<collectingCountry id="2E8476372018233AFEABFCEB3414FC2A" box="[264,318,895,919]" name="India" pageId="3" pageNumber="46">India</collectingCountry>
and
<collectingCountry id="2E8476372018233AFED2FC1434B6FC2A" box="[369,412,896,919]" name="Iran" pageId="3" pageNumber="46">Iran</collectingCountry>
(
<collectingRegion id="9457F8452018233AFE0AFCEB37D3FC2A" box="[425,761,895,919]" country="Iran" name="Sistan va Baluchestan" pageId="3" pageNumber="46">Sistan va Baluchestan province</collectingRegion>
). Presence in
<collectingCountry id="2E8476372018233AFC2BFCEB36C8FC2A" box="[904,994,895,919]" name="Pakistan" pageId="3" pageNumber="46">Pakistan</collectingCountry>
is likely.
</paragraph>
</subSubSection>
</treatment>
</document>
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