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<document id="E6D3199D06530046169BE01061707119" ID-DOI="10.11646/zootaxa.1697.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5107079" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626378524067" checkinUser="felipe" docAuthor="Thandar, Ahmed S." docDate="2008" docId="DE2087C7F52E8749A0ADF66193CA581A" docLanguage="en" docName="zt01697p057.pdf" docOrigin="Zootaxa 1697 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1697.1.1" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Sclerothyone unicolumnus Thandar 2008, sp. nov." docType="treatment" docVersion="3" lastPageNumber="16" masterDocId="2219FFBFF5208746A03AF42E93345C38" masterDocTitle="Additions to the holothuroid fauna of the southern African temperate faunistic provinces, with descriptions of new species" masterLastPageNumber="57" masterPageNumber="1" pageNumber="15" updateTime="1699010251044" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="123C7A1EAD35C37120E18BEE7786A9C4">Additions to the holothuroid fauna of the southern African temperate faunistic provinces, with descriptions of new species</mods:title>
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<mods:namePart id="FC31EB31A72AC399FF845679D30D692C">Thandar, Ahmed S.</mods:namePart>
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<mods:title id="DA5D4D14F9CA5F2043D82B3F6C4A1EFB">Zootaxa</mods:title>
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<mods:date id="EC5E258F5543711FF971F5FBB2421228">2008</mods:date>
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<mods:number id="3C594D11D8A7DFCBA189B7D0C2DB603C">2008-02-05</mods:number>
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<mods:number id="25938DC1066AB7EA3DABD2AB875EBC65">1697</mods:number>
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<mods:identifier id="29990DAE592232677BB1933A8A52DFBE" type="DOI">10.11646/zootaxa.1697.1.1</mods:identifier>
<mods:identifier id="F87EFC743E1A05EC2AD2BF5EC4131324" type="ISSN">1175­5334</mods:identifier>
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<treatment id="DE2087C7F52E8749A0ADF66193CA581A" ID-DOI="http://doi.org/10.5281/zenodo.5107111" ID-Zenodo-Dep="5107111" LSID="urn:lsid:plazi:treatment:DE2087C7F52E8749A0ADF66193CA581A" httpUri="http://treatment.plazi.org/id/DE2087C7F52E8749A0ADF66193CA581A" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="1E93655AF52E8748A0ADF66191745E51" box="[151,576,591,617]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="563636D1F52E8748A0ADF66191745E51" blockId="14.[151,576,591,652]" box="[151,576,591,617]" pageId="14" pageNumber="15">
<heading id="0D7E81BDF52E8748A0ADF66191745E51" bold="true" box="[151,576,591,617]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="64FDEAC3F52E8748A0ADF66191745E51" bold="true" box="[151,576,591,617]" pageId="14" pageNumber="15">
<taxonomicName id="91894D52F52E8748A0ADF66192E85E51" ID-CoL="4VQR2" authority="Thandar, 2008" authorityName="Thandar" authorityYear="2008" box="[151,476,591,617]" class="Holothuroidea" family="Sclerodactylidae" genus="Sclerothyone" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="unicolumnus" status="sp. nov.">
<emphasis id="64FDEAC3F52E8748A0ADF66192E85E51" bold="true" box="[151,476,591,617]" italics="true" pageId="14" pageNumber="15">Sclerothyone unicolumnus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FCE57B8F52E8748A1D9F66191745E51" box="[483,576,591,617]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1E93655AF52E8749A0ADF65C93CA581A" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="563636D1F52E8748A0ADF65C93CE5EB4" blockId="14.[151,576,591,652]" box="[151,250,626,652]" pageId="14" pageNumber="15">
<figureCitation id="CEB22A54F52E8748A0ADF65C93CE5EB4" box="[151,250,626,652]" captionStart="FIGURE 7" captionStartId="16.[151,255,1937,1961]" captionTargetBox="[179,1416,195,1917]" captionTargetId="figure-77@16.[160,1436,180,1932]" captionTargetPageId="16" captionText="FIGURE 7. Sclerothyone unicolumnus sp. nov. Holotype, SAM-A27987. A. Tables from body wall; B. tables from anal region; C. plates from anal papillae; D. end-plate from anal podium; E. end-plate from ventral podium; F. reduced tables from tentacles; G. rods from tentacles; H. plates from tentacles; I. calcareous ring. C scale a; A, B &amp; D-H scale b." figureDoi="http://doi.org/10.5281/zenodo.5107101" httpUri="https://zenodo.org/record/5107101/files/figure.png" pageId="14" pageNumber="15">Figure 7</figureCitation>
</paragraph>
<paragraph id="563636D1F52E8748A0ADF6EC92385EE4" blockId="14.[151,1436,706,932]" box="[151,268,706,732]" pageId="14" pageNumber="15">Diagnosis</paragraph>
<paragraph id="563636D1F52E8748A084F6C497835F9C" blockId="14.[151,1436,706,932]" pageId="14" pageNumber="15">
Perhaps a small species,
<typeStatus id="89328873F52E8748A1D8F6C4917C5F3C" box="[482,584,746,772]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(juvenile)
<quantity id="91719B34F52E8748A2F0F6C4903A5F3C" box="[714,782,746,772]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="mm" value="6.0">6 mm</quantity>
long. Podia mostly in double rows. Posterior processes of calcareous ring about five times length of radial plates, fragmented. Polian vesicle single. Body wall tables with smooth disc,
<quantity id="91719B34F52E8748A14AF71492D25F6C" box="[368,486,826,852]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.3" metricValueMin="4.2" pageId="14" pageNumber="15" unit="mm" value="47.5" valueMax="53.0" valueMin="42.0">42-53 µm</quantity>
, with undulating margin and perforated by four large central and usually four or more peripheral holes; spire of fused pillars, terminating in usually a tri-forked crown. Podia without spicules except for reduced end-plates. Tentacles with rods and usually smooth multilocular plates.
</paragraph>
<paragraph id="563636D1F52E8748A0ADF7F490E35824" blockId="14.[151,983,986,1052]" box="[151,983,986,1052]" pageId="14" pageNumber="15">Etymology The specific name refers to the fused pillars of the body wall tables.</paragraph>
<paragraph id="563636D1F52E8748A0ADF07C92475854" blockId="14.[151,1436,1106,1215]" box="[151,371,1106,1132]" pageId="14" pageNumber="15">Material examined</paragraph>
<paragraph id="563636D1F52E8748A086F05090B35887" blockId="14.[151,1436,1106,1215]" pageId="14" pageNumber="15">
<materialsCitation id="E6E13C8CF52E8748A086F05090B35887" accessionNumber="A27987" collectingDate="1960-08-28" collectionCode="UCT, SCD" location="Mossel Bay" longitude="22.3" pageId="14" pageNumber="15" specimenCount="2" typeStatus="holotype">
<emphasis id="64FDEAC3F52E8748A086F050921358AF" box="[188,295,1150,1175]" italics="true" pageId="14" pageNumber="15">
<typeStatus id="89328873F52E8748A086F050921358AF" box="[188,295,1150,1175]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
SAM A27987,
<collectionCode id="3098AE14F52E8748A1D9F053912958AF" box="[483,541,1149,1175]" pageId="14" pageNumber="15">UCT</collectionCode>
Ecol. Surv., off
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, St.
<collectionCode id="3098AE14F52E8748A3A4F05390E358AF" box="[926,983,1149,1175]" pageId="14" pageNumber="15">SCD</collectionCode>
<geoCoordinate id="33BD5016F52E8748A3E4F053970A58B0" box="[990,1086,1146,1175]" degrees="150, 34" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="555" value="-150.34">
150, 34
<superScript id="A1FC9B99F52E8748A40CF054970A58B0" attach="left" box="[1078,1086,1146,1160]" fontSize="6" pageId="14" pageNumber="15">o</superScript>
</geoCoordinate>
59' S,
<geoCoordinate id="33BD5016F52E8748A4B5F05397CE58AF" box="[1167,1274,1146,1175]" degrees="22" direction="east" minutes="18" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="22.3">
22
<superScript id="A1FC9B99F52E8748A496F054978058B0" attach="left" box="[1196,1204,1146,1160]" fontSize="6" pageId="14" pageNumber="15">o</superScript>
18' E
</geoCoordinate>
,
<date id="22371011F52E8748A53DF05396A358AF" box="[1287,1431,1149,1175]" pageId="14" pageNumber="15" value="1960-08-28">
<collectingDate id="3273E9F9F52E8748A53DF05396A358AF" box="[1287,1431,1149,1175]" pageId="14" pageNumber="15" value="1960-08-28">
28.viii.
<quantity id="91719B34F52E8748A562F05396A358AF" box="[1368,1431,1149,1175]" metricMagnitude="3" metricUnit="m" metricValue="1.960106" pageId="14" pageNumber="15" unit="m" value="1960.106">1960</quantity>
</collectingDate>
</date>
, 106 m;
<typeStatus id="89328873F52E8748A0CAF08B92575887" box="[240,355,1189,1215]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
, SAM-A27988, same data as holotype,
<specimenCount id="408FFD58F52E8748A30FF08B90B35887" box="[821,903,1189,1215]" pageId="14" pageNumber="15" type="generic">2 spec.</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="563636D1F52E8748A0ADF0DB92995937" blockId="14.[151,1437,1269,2015]" box="[151,429,1269,1295]" pageId="14" pageNumber="15">
Description of
<typeStatus id="89328873F52E8748A17CF0DB92995937" box="[326,429,1269,1295]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
</paragraph>
<paragraph id="563636D1F52E8748A0F5F133911F59EF" blockId="14.[151,1437,1269,2015]" pageId="14" pageNumber="15">
Specimen minute, previously dissected, calcareous ring partially damaged and few tentacles missing. Length
<quantity id="91719B34F52E8748A0C8F16B920E5967" box="[242,314,1349,1375]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="mm" value="6.0">6 mm</quantity>
, width in mid-body 1.25 mm. Colour in alcohol yellowish-white. Body form straight, cylindrical, mouth and anus terminal. Tentacles perhaps 10, only six intact, large. Anus encircled by large anal papillae, anal teeth not observed. Podia restricted to ambulacra in double rows, better developed ventrally, few also scattered in dorsal interambulacra.
</paragraph>
<paragraph id="563636D1F52E8748A0F9F1CB91835B2F" blockId="14.[151,1437,1269,2015]" pageId="14" pageNumber="15">
Calcareous ring (
<figureCitation id="CEB22A54F52E8748A1B4F1CB92CA59C7" box="[398,510,1509,1535]" captionStart="FIGURE 7" captionStartId="16.[151,255,1937,1961]" captionTargetBox="[179,1416,195,1917]" captionTargetId="figure-77@16.[160,1436,180,1932]" captionTargetPageId="16" captionText="FIGURE 7. Sclerothyone unicolumnus sp. nov. Holotype, SAM-A27987. A. Tables from body wall; B. tables from anal region; C. plates from anal papillae; D. end-plate from anal podium; E. end-plate from ventral podium; F. reduced tables from tentacles; G. rods from tentacles; H. plates from tentacles; I. calcareous ring. C scale a; A, B &amp; D-H scale b." figureDoi="http://doi.org/10.5281/zenodo.5107101" httpUri="https://zenodo.org/record/5107101/files/figure.png" pageId="14" pageNumber="15">Figure 7I</figureCitation>
) delicate, not tube-like, radial and interradial plates compact, meeting only at base, radial plates triangular with slightly concave posterior margin and anterior depression for radial muscle; interradial plates also triangular with deeper posterior concavity and a slight depression at anterior end; posterior processes of radial plates extremely long, about five times the length of radial plates, reaching half body length, each fragmented into a single series of elements. Polian vesicle single, elongate; stone canal not detected. Respiratory trees reaching two thirds body length poorly branched, branches few (ca. 8-10), small, sac-like, right tree slightly longer than left. Gonad absent. Retractor muscles arising from anterior third of longitudinal bands, inserting without subdivision.
</paragraph>
<paragraph id="563636D1F52E8749A081F30B91605D6A" blockId="14.[151,1437,1269,2015]" lastBlockId="15.[151,1436,152,338]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Body wall spicules scarce tables (
<figureCitation id="CEB22A54F52E8748A27FF30B91D75B07" box="[581,739,1829,1855]" captionStart="FIGURE 7" captionStartId="16.[151,255,1937,1961]" captionTargetBox="[179,1416,195,1917]" captionTargetId="figure-77@16.[160,1436,180,1932]" captionTargetPageId="16" captionText="FIGURE 7. Sclerothyone unicolumnus sp. nov. Holotype, SAM-A27987. A. Tables from body wall; B. tables from anal region; C. plates from anal papillae; D. end-plate from anal podium; E. end-plate from ventral podium; F. reduced tables from tentacles; G. rods from tentacles; H. plates from tentacles; I. calcareous ring. C scale a; A, B &amp; D-H scale b." figureDoi="http://doi.org/10.5281/zenodo.5107101" httpUri="https://zenodo.org/record/5107101/files/figure.png" pageId="14" pageNumber="15">Figure 7A, B</figureCitation>
), restricted to posterior end, with quadrangular to subcircular, smooth disc,
<quantity id="91719B34F52E8748A165F36392E15B5F" box="[351,469,1869,1895]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.3" metricValueMin="4.2" pageId="14" pageNumber="15" unit="mm" value="47.5" valueMax="53.0" valueMin="42.0">42-53 µm</quantity>
, with undulating margin, perforated by four large central holes and usually four or more peripheral ones; spire of moderate height, comprising fused pillars without perforation, terminating in usually a tri-forked, non-perforate crown, each fork often terminally bifid. Large periproctal plates also present. Podia without spicules except for reduced end-plates (ca
<quantity id="91719B34F52E8748A38CF3EB973C5BE7" box="[950,1032,1989,2015]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="14" pageNumber="15" unit="mm" value="57.0">57 µm</quantity>
), with large peripheral and much smaller central holes (
<figureCitation id="CEB22A54F52F8749A19AF4B691775C8A" box="[416,579,152,178]" captionStart="FIGURE 7" captionStartId="16.[151,255,1937,1961]" captionTargetBox="[179,1416,195,1917]" captionTargetId="figure-77@16.[160,1436,180,1932]" captionTargetPageId="16" captionText="FIGURE 7. Sclerothyone unicolumnus sp. nov. Holotype, SAM-A27987. A. Tables from body wall; B. tables from anal region; C. plates from anal papillae; D. end-plate from anal podium; E. end-plate from ventral podium; F. reduced tables from tentacles; G. rods from tentacles; H. plates from tentacles; I. calcareous ring. C scale a; A, B &amp; D-H scale b." figureDoi="http://doi.org/10.5281/zenodo.5107101" httpUri="https://zenodo.org/record/5107101/files/figure.png" pageId="15" pageNumber="16">Figure 7D, E</figureCitation>
). Anal papillae with irregular, smooth, multilocular plates of variety of shapes and sizes (
<figureCitation id="CEB22A54F52F8749A151F4EE92DC5CE2" box="[363,488,192,218]" captionStart="FIGURE 7" captionStartId="16.[151,255,1937,1961]" captionTargetBox="[179,1416,195,1917]" captionTargetId="figure-77@16.[160,1436,180,1932]" captionTargetPageId="16" captionText="FIGURE 7. Sclerothyone unicolumnus sp. nov. Holotype, SAM-A27987. A. Tables from body wall; B. tables from anal region; C. plates from anal papillae; D. end-plate from anal podium; E. end-plate from ventral podium; F. reduced tables from tentacles; G. rods from tentacles; H. plates from tentacles; I. calcareous ring. C scale a; A, B &amp; D-H scale b." figureDoi="http://doi.org/10.5281/zenodo.5107101" httpUri="https://zenodo.org/record/5107101/files/figure.png" pageId="15" pageNumber="16">Figure 7C</figureCitation>
). Tentacles with
<specimenCount id="408FFD58F52F8749A289F4EE90005CE2" box="[691,820,192,218]" pageId="15" pageNumber="16" type="generic" typeStatus="types">three types</specimenCount>
of deposits: rods (
<quantity id="91719B34F52F8749A426F4EE97975CE2" box="[1052,1187,191,218]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" metricValueMax="11.8" metricValueMin="5.8" pageId="15" pageNumber="16" unit="mm" value="88.0" valueMax="118.0" valueMin="58.0">58-118 µm</quantity>
), usually with perforated and/or digitated ends (
<figureCitation id="CEB22A54F52F8749A1E5F4C691695D3A" box="[479,605,232,258]" captionStart="FIGURE 7" captionStartId="16.[151,255,1937,1961]" captionTargetBox="[179,1416,195,1917]" captionTargetId="figure-77@16.[160,1436,180,1932]" captionTargetPageId="16" captionText="FIGURE 7. Sclerothyone unicolumnus sp. nov. Holotype, SAM-A27987. A. Tables from body wall; B. tables from anal region; C. plates from anal papillae; D. end-plate from anal podium; E. end-plate from ventral podium; F. reduced tables from tentacles; G. rods from tentacles; H. plates from tentacles; I. calcareous ring. C scale a; A, B &amp; D-H scale b." figureDoi="http://doi.org/10.5281/zenodo.5107101" httpUri="https://zenodo.org/record/5107101/files/figure.png" pageId="15" pageNumber="16">Figure 7G</figureCitation>
); small (
<quantity id="91719B34F52F8749A2F8F4C6900F5D3A" box="[706,827,231,258]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" metricValueMax="5.6" metricValueMin="4.2" pageId="15" pageNumber="16" unit="mm" value="49.0" valueMax="56.0" valueMin="42.0">42-56 µm</quantity>
), smooth, multilocular plates with small holes (
<figureCitation id="CEB22A54F52F8749A553F4C693D85D12" captionStart="FIGURE 7" captionStartId="16.[151,255,1937,1961]" captionTargetBox="[179,1416,195,1917]" captionTargetId="figure-77@16.[160,1436,180,1932]" captionTargetPageId="16" captionText="FIGURE 7. Sclerothyone unicolumnus sp. nov. Holotype, SAM-A27987. A. Tables from body wall; B. tables from anal region; C. plates from anal papillae; D. end-plate from anal podium; E. end-plate from ventral podium; F. reduced tables from tentacles; G. rods from tentacles; H. plates from tentacles; I. calcareous ring. C scale a; A, B &amp; D-H scale b." figureDoi="http://doi.org/10.5281/zenodo.5107101" httpUri="https://zenodo.org/record/5107101/files/figure.png" pageId="15" pageNumber="16">Figure 7H</figureCitation>
); and other, slightly larger (
<quantity id="91719B34F52F8749A20DF53E91845D12" box="[567,688,271,298]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" metricValueMax="5.8" metricValueMin="4.6" pageId="15" pageNumber="16" unit="mm" value="52.0" valueMax="58.0" valueMin="46.0">46-58 µm</quantity>
), multilocular, plates with large holes and central knob (
<figureCitation id="CEB22A54F52F8749A577F53E938D5D6A" captionStart="FIGURE 7" captionStartId="16.[151,255,1937,1961]" captionTargetBox="[179,1416,195,1917]" captionTargetId="figure-77@16.[160,1436,180,1932]" captionTargetPageId="16" captionText="FIGURE 7. Sclerothyone unicolumnus sp. nov. Holotype, SAM-A27987. A. Tables from body wall; B. tables from anal region; C. plates from anal papillae; D. end-plate from anal podium; E. end-plate from ventral podium; F. reduced tables from tentacles; G. rods from tentacles; H. plates from tentacles; I. calcareous ring. C scale a; A, B &amp; D-H scale b." figureDoi="http://doi.org/10.5281/zenodo.5107101" httpUri="https://zenodo.org/record/5107101/files/figure.png" pageId="15" pageNumber="16">Figure 7F</figureCitation>
). No deposits detected in introvert.
</paragraph>
<paragraph id="563636D1F52F8749A0ADF5A6923D5D9A" blockId="15.[151,1436,392,618]" box="[151,265,392,418]" pageId="15" pageNumber="16">
<typeStatus id="89328873F52F8749A0ADF5A6923D5D9A" box="[151,265,392,418]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</paragraph>
<paragraph id="563636D1F52F8749A0FBF59E90135E52" blockId="15.[151,1436,392,618]" pageId="15" pageNumber="16">
Both specimens minute,
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x 1.5 mm and
<quantity id="91719B34F52F8749A2DAF59E90115DF2" box="[736,805,432,458]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="mm" value="2.0">2 mm</quantity>
x
<quantity id="91719B34F52F8749A37EF59E90B85DF2" box="[836,908,432,458]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="mm" value="1.0">1 mm</quantity>
. Colour and distribution of podia as in
<typeStatus id="89328873F52F8749A564F59E93F95DCA" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
. Form slightly U-shaped. Tentacles retracted, anal teeth and papillae present. Body wall spicules as wellformed tables, present throughout integument, disc
<quantity id="91719B34F52F8749A2C9F62E90525E22" box="[755,870,511,538]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" metricValueMax="6.5" metricValueMin="4.3" pageId="15" pageNumber="16" unit="mm" value="54.0" valueMax="65.0" valueMin="43.0">43-65 µm</quantity>
(mean
<quantity id="91719B34F52F8749A384F62E973F5E22" box="[958,1035,511,538]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="15" pageNumber="16" unit="mm" value="55.0">55 µm</quantity>
) with four central and 1-5 smaller peripheral holes; spire of same form as that of the
<typeStatus id="89328873F52F8749A2D1F60690605E7A" box="[747,852,552,578]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
,
<quantity id="91719B34F52F8749A35AF60690E05E7A" box="[864,980,551,578]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="15" pageNumber="16" unit="mm" value="19.0" valueMax="23.0" valueMin="15.0">15-23 µm</quantity>
(mean
<quantity id="91719B34F52F8749A416F60697495E7A" box="[1068,1149,551,578]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="mm" value="20.0">20 µm</quantity>
). Podia without deposits except for reduced end-plates,
<quantity id="91719B34F52F8749A1C5F67E91475E52" box="[511,627,591,618]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="6.4" metricValueMin="5.6" pageId="15" pageNumber="16" unit="mm" value="60.0" valueMax="64.0" valueMin="56.0">56-64 µm</quantity>
(mean
<quantity id="91719B34F52F8749A2F0F67E902F5E52" box="[714,795,591,618]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="mm" value="60.0">60 µm</quantity>
).
</paragraph>
<paragraph id="563636D1F52F8749A0ADF68E92DB5EDA" blockId="15.[151,495,672,738]" box="[151,495,672,738]" pageId="15" pageNumber="16">
Distribution Mossel Bay, WCP,
<specimenCount id="408FFD58F52F8749A1A4F6E692DB5EDA" box="[414,495,712,738]" pageId="15" pageNumber="16" type="generic">
<quantity id="91719B34F52F8749A1A4F6E692DF5EDA" box="[414,491,712,738]" metricMagnitude="2" metricUnit="m" metricValue="1.06" pageId="15" pageNumber="16" unit="m" value="106.0">106 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="563636D1F52F8749A0ADF73693CB5F0A" blockId="15.[151,1436,792,1058]" box="[151,255,792,818]" pageId="15" pageNumber="16">Remarks</paragraph>
<paragraph id="563636D1F52F8749A084F76E93CA581A" blockId="15.[151,1436,792,1058]" pageId="15" pageNumber="16">
Because of the form of the calcareous ring and the lack of elongated plates in the body wall, the new species is here referred to
<taxonomicName id="91894D52F52F8749A198F746910E5FB9" authorityName="Thandar" authorityYear="1989" box="[418,570,872,897]" class="Holothuroidea" family="Sclerodactylidae" genus="Sclerothyone" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="genus">
<emphasis id="64FDEAC3F52F8749A198F746910E5FB9" box="[418,570,872,897]" italics="true" pageId="15" pageNumber="16">Sclerothyone</emphasis>
</taxonomicName>
, whose diagnosis is modified to accommodate it. The absence of deposits in the podia and introvert, may be age-related, as all
<specimenCount id="408FFD58F52F8749A33BF7BE90895F92" box="[769,957,912,938]" pageId="15" pageNumber="16" type="generic">three specimens</specimenCount>
are juvenile. However, in the absence of regular four-holed table discs, the presence of fused pillars, tri-forked crowns with bifurcating teeth, and
<specimenCount id="408FFD58F52F8749A55BF79693E15FC2" pageId="15" pageNumber="16" type="generic" typeStatus="types">three types</specimenCount>
of tentacle deposits, it differs substantially from the
<typeStatus id="89328873F52F8749A366F7CE90BA5FC2" box="[860,910,992,1018]" pageId="15" pageNumber="16">type</typeStatus>
species,
<taxonomicName id="91894D52F52F8749A43AF7CE93CE581A" authority="(Ludwig &amp; Heding)" baseAuthorityName="Ludwig &amp; Heding" baseAuthorityYear="1935" class="Holothuroidea" family="Sclerodactylidae" genus="Sclerothyone" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="velligera">
<emphasis id="64FDEAC3F52F8749A43AF7CE963F5FC1" box="[1024,1291,992,1017]" italics="true" pageId="15" pageNumber="16">Sclerothyone velligera</emphasis>
(Ludwig &amp; Heding)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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