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<document id="13FEE5EB483DCB6E4B4287BAFE59DD9C" ID-DOI="10.5281/zenodo.274091" ID-GBIF-Dataset="b7200336-d4b8-40e8-a706-1613e492922b" ID-ISSN="1175-5326" ID-Zenodo-Dep="274091" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459816383984" checkinUser="plazi" docAuthor="Naskrecki, Piotr" docDate="2008" docId="AA75ED36FFC7FF990C9DFDE4FD90FDED" docLanguage="en" docName="zt01698p064.pdf" docOrigin="Zootaxa 1698" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ricinoides atewa Naskrecki, 2008, n. sp." docType="treatment" docVersion="9" lastPageNumber="63" masterDocId="564C954EFFC6FF9E0C0AFFABFC43FFAF" masterDocTitle="A new ricinuleid of the genus Ricinoides Ewing (Arachnida, Ricinulei) from Ghana" masterLastPageNumber="64" masterPageNumber="57" pageNumber="58" updateTime="1698221437621" updateUser="plazi">
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<mods:title id="BB3FAAF9A44C5F8D2230DF6A826CBD66">A new ricinuleid of the genus Ricinoides Ewing (Arachnida, Ricinulei) from Ghana</mods:title>
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<mods:namePart id="97959497C9EAA027A95E516AB9DDB76F">Naskrecki, Piotr</mods:namePart>
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<mods:date id="10DCD72D2E9D84C6D6C83EFAD18BAEA9">2008</mods:date>
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<treatment id="AA75ED36FFC7FF990C9DFDE4FD90FDED" ID-DOI="http://doi.org/10.5281/zenodo.5619185" ID-GBIF-Taxon="119354777" ID-Zenodo-Dep="5619185" LSID="urn:lsid:zoobank.org:act:CE9D8B39-045B-479F-8CE3-E1BCF72DA516" httpUri="http://treatment.plazi.org/id/AA75ED36FFC7FF990C9DFDE4FD90FDED" lastPageId="7" lastPageNumber="63" pageId="1" pageNumber="58">
<subSubSection id="6AC60FABFFC7FF9F0C9DFDE4FDB7FD23" pageId="1" pageNumber="58" type="nomenclature">
<paragraph id="22635C20FFC7FF9F0C9DFDE4FDEFFDC6" blockId="1.[151,500,591,652]" box="[151,428,591,617]" pageId="1" pageNumber="58">
<heading id="792BEB4CFFC7FF9F0C9DFDE4FDEFFDC6" bold="true" box="[151,428,591,617]" fontSize="11" level="1" pageId="1" pageNumber="58" reason="1">
<emphasis id="10A88032FFC7FF9F0C9DFDE4FDEFFDC6" bold="true" box="[151,428,591,617]" pageId="1" pageNumber="58">
<taxonomicName id="E5DC27A3FFC7FF9F0C9DFDE4FD20FDC6" ID-CoL="4T24F" box="[151,355,591,617]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="atewa" status="sp. nov.">
<emphasis id="10A88032FFC7FF9F0C9DFDE4FD20FDC6" bold="true" box="[151,355,591,617]" italics="true" pageId="1" pageNumber="58">Ricinoides atewa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0B9B3D49FFC7FF9F0D60FDE4FDEFFDC6" box="[362,428,591,617]" pageId="1" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="22635C20FFC7FF9F0C9DFDD9FDB7FD23" blockId="1.[151,500,591,652]" box="[151,500,626,652]" pageId="1" pageNumber="58">
(
<figureCitation id="BAE740A5FFC7FF9F0C95FDD9FD7FFD23" box="[159,316,626,652]" captionStart-0="FIGURES 1 7" captionStart-1="FIGURES 8 13" captionStartId-0="2.[151,269,1867,1891]" captionStartId-1="4.[151,269,1842,1866]" captionTargetBox-0="[157,1436,313,1841]" captionTargetBox-1="[172,1422,295,1803]" captionTargetId-0="figure@2.[151,1436,304,1843]" captionTargetId-1="figure@4.[151,1436,277,1816]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 1 7. Ricinoides atewa sp. n. 1, male, dorsal view; 2, male, ventral view; 3, male cucullus, dorsal view; 4 6, female left chelicera, ventral, lateral, and dorsal views; 7, penis." captionText-1="FIGURES 8 13. Ricinoides atewa sp. n.: 8, female, dorsal view; 9, ventral view; 10, female posterior genital lip, showing genital sacks (spermathecae), internal view; 11, close-up of female posterior genital lip, showing genital sacks (spermathecae) and accessory spermathecae (a. s.); 12, tubercles and navicular setae on opisthosomal tergite; 13, female telson." httpUri-0="https://zenodo.org/record/274092/files/figure.png" httpUri-1="https://zenodo.org/record/274093/files/figure.png" pageId="1" pageNumber="58">Figures 113</figureCitation>
,
<figureCitation id="BAE740A5FFC7FF9F0D43FDD9FDD6FD23" box="[329,405,626,652]" captionStart="FIGURES 20 25. R" captionStartId="5.[151,269,1860,1884]" captionTargetBox="[182,1400,205,1727]" captionTargetId="figure@5.[151,1436,194,1733]" captionTargetPageId="5" captionText="FIGURES 20 25. R. atewa n. sp.: 20 23, tarsal process, frontal, lateral, posterior, and dorsal views; 24, male tarsus III, posterior view (bt basitarsus; bp basitarsal process; lc lamina cyathiformis; tt 1 tt 4 tarsomeres); 25, male tibia III." httpUri="https://zenodo.org/record/274094/files/figure.png" pageId="1" pageNumber="58">2025</figureCitation>
,
<figureCitation id="BAE740A5FFC7FF9F0DABFDD9FDA8FD23" box="[417,491,626,652]" captionStart="FIGURES 26 28" captionStartId="6.[151,269,1680,1704]" captionTargetBox="[151,1436,274,1655]" captionTargetId="figure@6.[151,1436,274,1656]" captionTargetPageId="6" captionText="FIGURES 26 28. Ricinoides atewa sp. n. and habitat. 26, an old, abandoned mound of soil termites Cubitermes sp., a typical habitat of R. atewa in the Upland Evergreen Forest of Ghana; 27, female R. atewa; 28, tritonymph of R. atewa, note the reddish coloration typical of larval and nymphal stages of Ricinulei." httpUri="https://zenodo.org/record/274095/files/figure.png" pageId="1" pageNumber="58">2728</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6AC60FABFFC7FF9D0C9DFD69F906F8AD" lastPageId="3" lastPageNumber="60" pageId="1" pageNumber="58" type="description">
<paragraph id="22635C20FFC7FF9F0C9DFD69FE1DFB13" blockId="1.[151,1436,706,1252]" pageId="1" pageNumber="58">
<emphasis id="10A88032FFC7FF9F0C9DFD69FDEAFD73" bold="true" box="[151,425,706,732]" pageId="1" pageNumber="58">Differential diagnosis.</emphasis>
This species can be easily recognized by the combination of its large size (body length up to
<quantity id="E524F1C5FFC7FF9F0CD0FD41FD6FFCAB" box="[218,300,746,772]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="58" unit="mm" value="11.0">11 mm</quantity>
in the male; see
<tableCitation id="6F5E699BFFC7FF9F0DF8FD41FE04FCAB" box="[498,583,746,772]" captionStart="TABLE 1" captionStartId="1.[151,239,1306,1330]" captionTargetBox="[159,1380,1362,1505]" captionTargetPageId="1" captionText="TABLE 1. Body measurements of R. atewa n. sp." httpUri="http://table.plazi.org/id/76A30CA8FFC7FF9F0C9DFAB1FEEEFA9D" pageId="1" pageNumber="58" tableUuid="76A30CA8FFC7FF9F0C9DFAB1FEEEFA9D">Table 1</tableCitation>
), the enlarged femur II and the presence of a short spur on tibia II in the male, a strongly elongated, thin and sickle-shaped apical lobe
<emphasis id="10A88032FFC7FF9F0F7EFCB9FFC2FC84" box="[884,897,786,811]" italics="true" pageId="1" pageNumber="58">c</emphasis>
of the tarsal process, and the presence of 67 genital sacks in the female. From
<taxonomicName id="E5DC27A3FFC7FF9F0E2CFC91FF09FCFB" authority="Thorell, 1892" authorityName="Thorell" authorityYear="1892" box="[550,842,826,852]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="afzelii">
<emphasis id="10A88032FFC7FF9F0E2CFC91FED3FCFC" box="[550,656,826,851]" italics="true" pageId="1" pageNumber="58">R. afzelii</emphasis>
(Thorell, 1892)
</taxonomicName>
it can be distinguished by the overall shape of the tarsal process, in particular the extreme elongation of the apical lobe
<emphasis id="10A88032FFC7FF9F0FDAFCC9FF9EFCD4" box="[976,989,866,891]" italics="true" pageId="1" pageNumber="58">c</emphasis>
(lobe
<emphasis id="10A88032FFC7FF9F0821FCC9F87BFCD4" box="[1067,1080,866,891]" italics="true" pageId="1" pageNumber="58">c</emphasis>
much shorter and wider in
<taxonomicName id="E5DC27A3FFC7FF9F0989FCC9FC9CFC0C" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="afzelii">
<emphasis id="10A88032FFC7FF9F0989FCC9FC9CFC0C" italics="true" pageId="1" pageNumber="58">R. afzelii</emphasis>
</taxonomicName>
; figs. 1417), the proportions of the lamina cyathiformis
<emphasis id="10A88032FFC7FF9F0F89FC21FFE8FC0C" box="[899,939,906,931]" italics="true" pageId="1" pageNumber="58">(lc)</emphasis>
on tarsus III of the male (in
<emphasis id="10A88032FFC7FF9F08F7FC21F9C2FC0C" box="[1277,1409,906,931]" italics="true" pageId="1" pageNumber="58">
<taxonomicName id="E5DC27A3FFC7FF9F08F7FC21F927FC0C" box="[1277,1380,906,931]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="atewa">R. atewa</taxonomicName>
lc
</emphasis>
is about as long as the basal part of tarsomere 2
<emphasis id="10A88032FFC7FF9F0EA7FC19FEA3FC64" box="[685,736,946,971]" italics="true" pageId="1" pageNumber="58">(tt2)</emphasis>
and about twice as long as wide; in
<emphasis id="10A88032FFC7FF9F0882FC19F94EFC64" box="[1160,1293,946,971]" italics="true" pageId="1" pageNumber="58">
<taxonomicName id="E5DC27A3FFC7FF9F0882FC19F8B2FC64" box="[1160,1265,946,971]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="afzelii">R. afzelii</taxonomicName>
lc
</emphasis>
is about 0.8 times as long as the basal part of
<emphasis id="10A88032FFC7FF9F0E1CFC71FE75FC5C" box="[534,566,986,1011]" italics="true" pageId="1" pageNumber="58">tt2</emphasis>
and
<emphasis id="10A88032FFC7FF9F0E66FC71FEC2FC5C" box="[620,641,986,1011]" italics="true" pageId="1" pageNumber="58">lc</emphasis>
is only about 1.6 times as long as wide), and the development of the tibia on leg II (the spur in
<taxonomicName id="E5DC27A3FFC7FF9F0DDEFBA9FE7CFBB4" box="[468,575,1026,1051]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="afzelii">
<emphasis id="10A88032FFC7FF9F0DDEFBA9FE7CFBB4" box="[468,575,1026,1051]" italics="true" pageId="1" pageNumber="58">R. afzelii</emphasis>
</taxonomicName>
is much larger, slightly longer than the width of the tibia, whereas in
<taxonomicName id="E5DC27A3FFC7FF9F0989FBA9FC9EFBEC" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="atewa">
<emphasis id="10A88032FFC7FF9F0989FBA9FC9EFBEC" italics="true" pageId="1" pageNumber="58">R. atewa</emphasis>
</taxonomicName>
the spur is only as long as half the width of the tibia). From
<taxonomicName id="E5DC27A3FFC7FF9F0FAEFB81F82BFBEB" authority="Hansen" authorityName="Hansen" box="[932,1128,1066,1092]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="feae">
<emphasis id="10A88032FFC7FF9F0FAEFB81FFB5FBEC" box="[932,1014,1066,1091]" italics="true" pageId="1" pageNumber="58">R. feae</emphasis>
(Hansen)
</taxonomicName>
it differs in its larger size, shorter and stouter palpal tibia, and the shape of the tarsal process, especially the size and shape of the lateral lobe
<emphasis id="10A88032FFC7FF9F0CDAFBD0FCB9FB3B" box="[208,250,1147,1172]" italics="true" pageId="1" pageNumber="58">(lb)</emphasis>
. It can be distinguished from all other African
<taxonomicName id="E5DC27A3FFC7FF9F0F15FBD1FFC9FB3B" box="[799,906,1146,1172]" class="Arachnida" kingdom="Animalia" order="Ricinulei" pageId="1" pageNumber="58" phylum="Arthropoda" rank="order">Ricinulei</taxonomicName>
by its large size and the strong sexual dimorphism in the development of the leg II.
</paragraph>
<paragraph id="22635C20FFC7FF9F0CCCFB61F91DFB4B" blockId="1.[151,1436,706,1252]" box="[198,1374,1226,1252]" pageId="1" pageNumber="58">
<emphasis id="10A88032FFC7FF9F0CCCFB61FD1DFB4B" bold="true" box="[198,350,1226,1252]" pageId="1" pageNumber="58">Description.</emphasis>
<emphasis id="10A88032FFC7FF9F0D6CFB60FDE1FB4B" box="[358,418,1227,1252]" italics="true" pageId="1" pageNumber="58">Male</emphasis>
. Body large for the genus, robust and stout (
<tableCitation id="6F5E699BFFC7FF9F0FA1FB61F84CFB4B" box="[939,1039,1226,1252]" captionStart="TABLE 1" captionStartId="1.[151,239,1306,1330]" captionTargetBox="[159,1380,1362,1505]" captionTargetPageId="1" captionText="TABLE 1. Body measurements of R. atewa n. sp." httpUri="http://table.plazi.org/id/76A30CA8FFC7FF9F0C9DFAB1FEEEFA9D" pageId="1" pageNumber="58" tableUuid="76A30CA8FFC7FF9F0C9DFAB1FEEEFA9D">Tables 1</tableCitation>
and
<tableCitation id="6F5E699BFFC7FF9F0842FB61F81AFB4B" box="[1096,1113,1226,1252]" captionStart="TABLE 2" captionStartId="3.[151,239,866,890]" captionTargetBox="[159,1339,922,1332]" captionTargetPageId="3" captionText="TABLE 2. Leg measurements of R. atewa n. sp. (number above male; number below female)." httpUri="http://table.plazi.org/id/76A30CA8FFC5FF9D0C9DFCC9F8EEFCD6" pageId="1" pageNumber="58" tableUuid="76A30CA8FFC5FF9D0C9DFCC9F8EEFCD6">2</tableCitation>
); legs relatively thick.
</paragraph>
<caption id="76A30CA8FFC7FF9F0C9DFAB1FEEEFA9D" ID-Table-UUID="76A30CA8FFC7FF9F0C9DFAB1FEEEFA9D" box="[151,685,1306,1330]" httpUri="http://table.plazi.org/id/76A30CA8FFC7FF9F0C9DFAB1FEEEFA9D" pageId="1" pageNumber="58" targetBox="[159,1380,1362,1505]" targetIsTable="true" targetPageId="1">
<paragraph id="22635C20FFC7FF9F0C9DFAB1FEEEFA9D" blockId="1.[151,685,1306,1330]" box="[151,685,1306,1330]" pageId="1" pageNumber="58">
<emphasis id="10A88032FFC7FF9F0C9DFAB1FD4BFA9D" bold="true" box="[151,264,1306,1330]" pageId="1" pageNumber="58">TABLE 1.</emphasis>
Body measurements of
<taxonomicName id="E5DC27A3FFC7FF9F0E07FAB1FE28FA9E" box="[525,619,1306,1329]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="atewa" status="sp. nov.">
<emphasis id="10A88032FFC7FF9F0E07FAB1FE28FA9E" box="[525,619,1306,1329]" italics="true" pageId="1" pageNumber="58">R. atewa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0B9B3D49FFC7FF9F0E7BFAB1FEEEFA9D" box="[625,685,1306,1330]" pageId="1" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="22635C20FFC7FF9F0D16FAF9F8BAFA4E" pageId="1" pageNumber="58">
<table id="50DCAE80FFC700610C95FAF9F927FA4E" box="[159,1380,1362,1505]" gridcols="7" gridrows="3" pageId="1" pageNumber="58">
<tr id="9CEC5E62FFC700610C95FAF9F927FA23" box="[159,1380,1362,1420]" gridrow="0" pageId="1" pageNumber="58" rowspan-0="1">
<th id="DF3D371EFFC700610D11FAF9FDC5FA23" box="[283,390,1362,1420]" gridcol="1" gridrow="0" pageId="1" pageNumber="58">Body</th>
<th id="DF3D371EFFC700610DB9FAF9FE51FA23" box="[435,530,1362,1420]" gridcol="2" gridrow="0" pageId="1" pageNumber="58">Cucullus width</th>
<th id="DF3D371EFFC700610E42FAF9FEA6FA23" box="[584,741,1362,1420]" gridcol="3" gridrow="0" pageId="1" pageNumber="58">Cephalothorax width</th>
<th id="DF3D371EFFC700610F23FAF9FF85FA23" box="[809,966,1362,1420]" gridcol="4" gridrow="0" pageId="1" pageNumber="58">Cephalothorax length</th>
<th id="DF3D371EFFC700610801FAF9F8DBFA23" box="[1035,1176,1362,1420]" gridcol="5" gridrow="0" pageId="1" pageNumber="58">Opisthosoma width</th>
<th id="DF3D371EFFC7006108DDFAF9F927FA23" box="[1239,1380,1362,1420]" gridcol="6" gridrow="0" pageId="1" pageNumber="58">Opisthosoma length</th>
</tr>
<tr id="9CEC5E62FFC700610C95FA0BF927FA18" box="[159,1380,1440,1463]" gridrow="1" pageId="1" pageNumber="58">
<th id="DF3D371EFFC700610C95FA0BFCAEFA18" box="[159,237,1440,1463]" gridcol="0" gridrow="1" pageId="1" pageNumber="58">Male</th>
<td id="DF3D371EFFC700610D11FA0BFDC5FA18" box="[283,390,1440,1463]" gridcol="1" gridrow="1" pageId="1" pageNumber="58">10.011.0</td>
<td id="DF3D371EFFC700610DB9FA0BFE51FA18" box="[435,530,1440,1463]" gridcol="2" gridrow="1" pageId="1" pageNumber="58">3.1</td>
<td id="DF3D371EFFC700610E42FA0BFEA6FA18" box="[584,741,1440,1463]" gridcol="3" gridrow="1" pageId="1" pageNumber="58">3.8</td>
<td id="DF3D371EFFC700610F23FA0BFF85FA18" box="[809,966,1440,1463]" gridcol="4" gridrow="1" pageId="1" pageNumber="58">3.4</td>
<td id="DF3D371EFFC700610801FA0BF8DBFA18" box="[1035,1176,1440,1463]" gridcol="5" gridrow="1" pageId="1" pageNumber="58">4.04.2</td>
<td id="DF3D371EFFC7006108DDFA0BF927FA18" box="[1239,1380,1440,1463]" gridcol="6" gridrow="1" pageId="1" pageNumber="58">6.26.4</td>
</tr>
<tr id="9CEC5E62FFC700610C95FA61F927FA4E" box="[159,1380,1482,1505]" gridrow="2" pageId="1" pageNumber="58">
<th id="DF3D371EFFC700610C95FA61FCAEFA4E" box="[159,237,1482,1505]" gridcol="0" gridrow="2" pageId="1" pageNumber="58">Female</th>
<td id="DF3D371EFFC700610D11FA61FDC5FA4E" box="[283,390,1482,1505]" gridcol="1" gridrow="2" pageId="1" pageNumber="58">9.7</td>
<td id="DF3D371EFFC700610DB9FA61FE51FA4E" box="[435,530,1482,1505]" gridcol="2" gridrow="2" pageId="1" pageNumber="58">3.1</td>
<td id="DF3D371EFFC700610E42FA61FEA6FA4E" box="[584,741,1482,1505]" gridcol="3" gridrow="2" pageId="1" pageNumber="58">3.6</td>
<td id="DF3D371EFFC700610F23FA61FF85FA4E" box="[809,966,1482,1505]" gridcol="4" gridrow="2" pageId="1" pageNumber="58">3.2</td>
<td id="DF3D371EFFC700610801FA61F8DBFA4E" box="[1035,1176,1482,1505]" gridcol="5" gridrow="2" pageId="1" pageNumber="58">4.5</td>
<td id="DF3D371EFFC7006108DDFA61F927FA4E" box="[1239,1380,1482,1505]" gridcol="6" gridrow="2" pageId="1" pageNumber="58">6.1</td>
</tr>
</table>
</paragraph>
<paragraph id="22635C20FFC7FF9F0CCFF998F9D8F8C8" blockId="1.[151,1436,1586,2015]" pageId="1" pageNumber="58">
<emphasis id="10A88032FFC7FF9F0CCFF998FD6EF9E3" box="[197,301,1587,1612]" italics="true" pageId="1" pageNumber="58">Prosoma</emphasis>
. Cucullus about 1.5 times wider than long (fig. 3); anteromedian knob (
<bibRefCitation id="464D21D1FFC7FF9F08A9F999F975F9E3" author="Legg" box="[1187,1334,1586,1612]" pageId="1" pageNumber="64" refString="Legg, G. (1978 b) The status of Ricinoides plebejus and a note on R. afzelii (Ricinulei, Arachnida). Bulletin of the British Arachnological Society, 4, 124 - 125." type="journal article" year="1978" yearSuffix="b">Legg 1978b</bibRefCitation>
) absent. Cephalothorax slightly wider than long, ratio width/length 1.1, dorsally with distinct median furrow and two oblique lateral depressions about half way back from front; no ocelli present; dorsal surface densely covered with rounded tubercles (density 3439 tubercles/
<quantity id="E524F1C5FFC7FF9F0ED4F906FF5EF968" box="[734,797,1709,1735]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="58" unit="mm" value="1.0">1 mm</quantity>
2) (fig. 12); posterior margin of cephalothorax broadly rounded, with distinct, long furrow, parallel to posterior edge (fig. 1). Sternal region with coxa I not meeting tritosternum; coxa II longest, meeting coxa III along their posterior 3/4; coxa IV about as long as coxa III, but distinctly wider distally and directed posteriad (fig. 2). Ventral surface of coxae covered mostly with short navicular setae and few, scattered tubercles (greatest concentration of tubercles on posterior edge of coxa III).
</paragraph>
<paragraph id="22635C20FFC7FF9C0CCFF8DDFF88FEAB" blockId="1.[151,1436,1586,2015]" lastBlockId="2.[151,1436,152,260]" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58">
<emphasis id="10A88032FFC7FF9F0CCFF8DDFD23F820" box="[197,352,1910,1935]" italics="true" pageId="1" pageNumber="58">Opisthosoma</emphasis>
. About 1.4 times as long as wide, its margins nearly parallel (fig. 1). Tergites 1113 large, rectangular; ratio of widths of median portions of tergites 1113: 1/0.83/0.64; tergite 10 very wide and short, 1.8 times wider than tergite 13. Surface of tergites covered with tubercles similar in size and density to those on cephalothorax. Sternites 1113 also covered with tubercles, their density slightly higher than on tergites (about 40 tubercles/
<quantity id="E524F1C5FFC4FF9C0D8CFF69FD86FF73" box="[390,453,194,220]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="59" unit="mm" value="1.0">1 mm</quantity>
2); sternites 1113 with pair of shallow depressions near anterior margin of sternite. Basal segment of telson (sternite 14) with small dorsal notch (fig. 13).
</paragraph>
<caption id="76A30CA8FFC4FF9C0C9DF8E0FF02F82A" httpUri="https://zenodo.org/record/274092/files/figure.png" pageId="2" pageNumber="59" targetBox="[157,1436,313,1841]" targetPageId="2">
<paragraph id="22635C20FFC4FF9C0C9DF8E0FF02F82A" blockId="2.[151,1435,1867,1925]" pageId="2" pageNumber="59">
<emphasis id="10A88032FFC4FF9C0C9DF8E0FD00F8CC" bold="true" box="[151,323,1867,1891]" pageId="2" pageNumber="59">FIGURES 17.</emphasis>
<taxonomicName id="E5DC27A3FFC4FF9C0D40F8E0FE41F8CD" box="[330,514,1867,1890]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="atewa" status="sp. nov.">
<emphasis id="10A88032FFC4FF9C0D40F8E0FE41F8CD" box="[330,514,1867,1890]" italics="true" pageId="2" pageNumber="59">Ricinoides atewa</emphasis>
</taxonomicName>
<emphasis id="10A88032FFC4FF9C0E03F8E0FE05F8CC" bold="true" box="[521,582,1867,1891]" pageId="2" pageNumber="59">
<taxonomicNameLabel id="0B9B3D49FFC4FF9C0E03F8E0FE05F8CC" box="[521,582,1867,1891]" pageId="2" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
1, male, dorsal view; 2, male, ventral view; 3, male cucullus, dorsal view; 46, female left chelicera, ventral, lateral, and dorsal views; 7, penis.
</paragraph>
</caption>
<paragraph id="22635C20FFC4FF9D0CCFF818FF68FC9D" blockId="2.[151,1436,1971,2037]" lastBlockId="3.[151,1436,152,818]" lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59">
<emphasis id="10A88032FFC4FF9C0CCFF818FD17F863" box="[197,340,1971,1996]" italics="true" pageId="2" pageNumber="59">Appendages</emphasis>
. Basal part of chelicera robust, nearly as thick as long (fig. 5); cheliceral fixed digit with 5 large, rounded denticles; proximal denticles about 1/3 size of distal denticles; movable cheliceral digit strongly sickle-shaped and with 89 shallow denticles (denticles appear to be worn off in older individuals), about 2.5 times as long as fixed digit; base of movable digit with 1518 setae (setal formula 1+ 14 to 1 + 17; figs. 46). Palpal femur short and stout, distinctly bent, at base nearly twice as thick as at apex; ratio length/ maximum width 1.7; ventral surface of femur with distinct groove devoid of setae or tubercles. Palpal tibia long and straight, with parallel edges; ratio length/width 6.1 (length measured from base to tip of movable digit). Leg I short and stout; all podomeres densely covered with tubercles but relatively few setae (except for densely setose tarsus); femur without well developed dorsal groove; basitarsus grooved dorsally, tarsus consisting of single teleomere. Leg II greatly enlarged, robust; femur strongly thickened, nearly 3 times as thick as femur of legs III and IV, ratio length/maximum width 1.81 (figs. 12); not grooved dorsally. Tibia of leg II with large, internal tooth about half as long as tibias width (fig. 25). Basitarsus II and 3 basal teleomeres with distinct dorsal groove devoid of setae or tubercles. Leg III slender, with reproductive modifications of tarsus typical of all male
<taxonomicName id="E5DC27A3FFC5FF9D0D70FDFBFDAAFDC5" box="[378,489,592,618]" class="Arachnida" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="60" phylum="Arthropoda" rank="order">Ricinulei</taxonomicName>
; femur grooved dorsally; lamina cyathiformis
<emphasis id="10A88032FFC5FF9D081FFDFBF87EFDC6" box="[1045,1085,592,617]" italics="true" pageId="3" pageNumber="60">(lc)</emphasis>
on 2nd tarsomere
<emphasis id="10A88032FFC5FF9D0915FDFBF912FDC6" box="[1311,1361,592,617]" italics="true" pageId="3" pageNumber="60">(tt2)</emphasis>
about twice as long as wide, and about as long as basal portion of
<emphasis id="10A88032FFC5FF9D0F67FDD3FFCEFD3E" box="[877,909,632,657]" italics="true" pageId="3" pageNumber="60">tt2</emphasis>
(fig. 24); 4th tarsomere
<emphasis id="10A88032FFC5FF9D08BEFDD3F8A4FD3E" box="[1204,1255,632,657]" italics="true" pageId="3" pageNumber="60">(tt4)</emphasis>
about twice as long as 3rd tarsomere
<emphasis id="10A88032FFC5FF9D0D91FD0BFD8EFD16" box="[411,461,672,697]" italics="true" pageId="3" pageNumber="60">(tt3)</emphasis>
. Tarsal process with large, wide, lateral lobe
<emphasis id="10A88032FFC5FF9D0FEAFD0BF849FD16" box="[992,1034,672,697]" italics="true" pageId="3" pageNumber="60">(lb)</emphasis>
; apical lobe
<emphasis id="10A88032FFC5FF9D08AAFD0BF8EEFD16" box="[1184,1197,672,697]" italics="true" pageId="3" pageNumber="60">c</emphasis>
strongly developed, long and sickle-shaped when seen laterally (figs. 2023); lobe
<emphasis id="10A88032FFC5FF9D0F7AFD63FF3EFD4E" box="[880,893,712,737]" italics="true" pageId="3" pageNumber="60">c</emphasis>
clearly extends beyond proximal edge of lamina cyathiformis even when tarsal process is folded (fig. 24). Leg IV slender, unmodified; femur, tibia, basitarsus and 4 basal teleomeres with distinct groove dorsally.
</paragraph>
<caption id="76A30CA8FFC5FF9D0C9DFCC9F8EEFCD6" ID-Table-UUID="76A30CA8FFC5FF9D0C9DFCC9F8EEFCD6" box="[151,1197,866,890]" httpUri="http://table.plazi.org/id/76A30CA8FFC5FF9D0C9DFCC9F8EEFCD6" pageId="3" pageNumber="60" targetBox="[159,1339,922,1332]" targetIsTable="true" targetPageId="3">
<paragraph id="22635C20FFC5FF9D0C9DFCC9F8EEFCD6" blockId="3.[151,1197,866,890]" box="[151,1197,866,890]" pageId="3" pageNumber="60">
<emphasis id="10A88032FFC5FF9D0C9DFCC9FD4BFCD5" bold="true" box="[151,264,866,890]" pageId="3" pageNumber="60">TABLE 2.</emphasis>
Leg measurements of
<taxonomicName id="E5DC27A3FFC5FF9D0DF6FCC9FE19FCD6" box="[508,602,866,889]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="atewa" status="sp. nov.">
<emphasis id="10A88032FFC5FF9D0DF6FCC9FE19FCD6" box="[508,602,866,889]" italics="true" pageId="3" pageNumber="60">R. atewa</emphasis>
</taxonomicName>
<emphasis id="10A88032FFC5FF9D0E6BFCC9FEDFFCD5" bold="true" box="[609,668,866,890]" pageId="3" pageNumber="60">
<taxonomicNameLabel id="0B9B3D49FFC5FF9D0E6BFCC9FEDFFCD5" box="[609,668,866,890]" pageId="3" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(number above male; number below female).
</paragraph>
</caption>
<paragraph id="22635C20FFC5FF9D0D48FC31F967FA9B" pageId="3" pageNumber="60">
<table id="50DCAE80FFC500610C95FC31F978FA9B" box="[159,1339,922,1332]" gridcols="7" gridrows="6" pageId="3" pageNumber="60">
<tr id="9CEC5E62FFC500610C95FC31F978FC1E" box="[159,1339,922,945]" gridrow="0" pageId="3" pageNumber="60" rowspan-0="1">
<th id="DF3D371EFFC500610D48FC31FDF2FC1E" box="[322,433,922,945]" gridcol="1" gridrow="0" pageId="3" pageNumber="60">Basifemur</th>
<th id="DF3D371EFFC500610E12FC31FE1DFC1E" box="[536,606,922,945]" gridcol="2" gridrow="0" pageId="3" pageNumber="60">Femur</th>
<th id="DF3D371EFFC500610EC2FC31FF53FC1E" box="[712,784,922,945]" gridcol="3" gridrow="0" pageId="3" pageNumber="60">Patella</th>
<th id="DF3D371EFFC500610F70FC31FFF2FC1E" box="[890,945,922,945]" gridcol="4" gridrow="0" pageId="3" pageNumber="60">Tibia</th>
<th id="DF3D371EFFC500610828FC31F8D3FC1E" box="[1058,1168,922,945]" gridcol="5" gridrow="0" pageId="3" pageNumber="60">Basitarsus</th>
<th id="DF3D371EFFC5006108FFFC31F978FC1E" box="[1269,1339,922,945]" gridcol="6" gridrow="0" pageId="3" pageNumber="60">Tarsus</th>
</tr>
<tr id="9CEC5E62FFC500610C95FC6EF978FC50" box="[159,1339,965,1023]" gridrow="1" pageId="3" pageNumber="60">
<th id="DF3D371EFFC500610C95FC6EFCA8FC50" box="[159,235,965,1023]" gridcol="0" gridrow="1" pageId="3" pageNumber="60">Palp</th>
<td id="DF3D371EFFC500610D48FC6EFDF2FC50" box="[322,433,965,1023]" gridcol="1" gridrow="1" pageId="3" pageNumber="60"></td>
<td id="DF3D371EFFC500610E12FC6EFE1DFC50" box="[536,606,965,1023]" gridcol="2" gridrow="1" pageId="3" pageNumber="60">1.70 1.50</td>
<td id="DF3D371EFFC500610EC2FC6EFF53FC50" box="[712,784,965,1023]" gridcol="3" gridrow="1" pageId="3" pageNumber="60"></td>
<td id="DF3D371EFFC500610F70FC6EFFF2FC50" box="[890,945,965,1023]" gridcol="4" gridrow="1" pageId="3" pageNumber="60">2.48 2.50</td>
<td id="DF3D371EFFC500610828FC6EF8D3FC50" box="[1058,1168,965,1023]" gridcol="5" gridrow="1" pageId="3" pageNumber="60"></td>
<td id="DF3D371EFFC5006108FFFC6EF978FC50" box="[1269,1339,965,1023]" gridcol="6" gridrow="1" pageId="3" pageNumber="60"></td>
</tr>
<tr id="9CEC5E62FFC500610C95FBB9F978FBE3" box="[159,1339,1042,1100]" gridrow="2" pageId="3" pageNumber="60">
<th id="DF3D371EFFC500610C95FBB9FCA8FBE3" box="[159,235,1042,1100]" gridcol="0" gridrow="2" pageId="3" pageNumber="60">Leg I</th>
<td id="DF3D371EFFC500610D48FBB9FDF2FBE3" box="[322,433,1042,1100]" gridcol="1" gridrow="2" pageId="3" pageNumber="60"></td>
<td id="DF3D371EFFC500610E12FBB9FE1DFBE3" box="[536,606,1042,1100]" gridcol="2" gridrow="2" pageId="3" pageNumber="60">2.39 2.15</td>
<td id="DF3D371EFFC500610EC2FBB9FF53FBE3" box="[712,784,1042,1100]" gridcol="3" gridrow="2" pageId="3" pageNumber="60">1.54 1.51</td>
<td id="DF3D371EFFC500610F70FBB9FFF2FBE3" box="[890,945,1042,1100]" gridcol="4" gridrow="2" pageId="3" pageNumber="60">1.93 1.89</td>
<td id="DF3D371EFFC500610828FBB9F8D3FBE3" box="[1058,1168,1042,1100]" gridcol="5" gridrow="2" pageId="3" pageNumber="60">2.13 1.96</td>
<td id="DF3D371EFFC5006108FFFBB9F978FBE3" box="[1269,1339,1042,1100]" gridcol="6" gridrow="2" pageId="3" pageNumber="60">0.83 0.81</td>
</tr>
<tr id="9CEC5E62FFC500610C95FBCBF978FB36" box="[159,1339,1120,1177]" gridrow="3" pageId="3" pageNumber="60">
<th id="DF3D371EFFC500610C95FBCBFCA8FB36" box="[159,235,1120,1177]" gridcol="0" gridrow="3" pageId="3" pageNumber="60">Leg II</th>
<td id="DF3D371EFFC500610D48FBCBFDF2FB36" box="[322,433,1120,1177]" gridcol="1" gridrow="3" pageId="3" pageNumber="60"></td>
<td id="DF3D371EFFC500610E12FBCBFE1DFB36" box="[536,606,1120,1177]" gridcol="2" gridrow="3" pageId="3" pageNumber="60">3.97 3.40</td>
<td id="DF3D371EFFC500610EC2FBCBFF53FB36" box="[712,784,1120,1177]" gridcol="3" gridrow="3" pageId="3" pageNumber="60">2.17 1.80</td>
<td id="DF3D371EFFC500610F70FBCBFFF2FB36" box="[890,945,1120,1177]" gridcol="4" gridrow="3" pageId="3" pageNumber="60">3.06 2.90</td>
<td id="DF3D371EFFC500610828FBCBF8D3FB36" box="[1058,1168,1120,1177]" gridcol="5" gridrow="3" pageId="3" pageNumber="60">3.13 2.60</td>
<td id="DF3D371EFFC5006108FFFBCBF978FB36" box="[1269,1339,1120,1177]" gridcol="6" gridrow="3" pageId="3" pageNumber="60">2.67 2.05</td>
</tr>
<tr id="9CEC5E62FFC500610C95FB06F978FB48" box="[159,1339,1197,1255]" gridrow="4" pageId="3" pageNumber="60">
<th id="DF3D371EFFC500610C95FB06FCA8FB48" box="[159,235,1197,1255]" gridcol="0" gridrow="4" pageId="3" pageNumber="60">Leg III</th>
<td id="DF3D371EFFC500610D48FB06FDF2FB48" box="[322,433,1197,1255]" gridcol="1" gridrow="4" pageId="3" pageNumber="60">1.24 1.06</td>
<td id="DF3D371EFFC500610E12FB06FE1DFB48" box="[536,606,1197,1255]" gridcol="2" gridrow="4" pageId="3" pageNumber="60">2.93 2.45</td>
<td id="DF3D371EFFC500610EC2FB06FF53FB48" box="[712,784,1197,1255]" gridcol="3" gridrow="4" pageId="3" pageNumber="60">1.62 1.50</td>
<td id="DF3D371EFFC500610F70FB06FFF2FB48" box="[890,945,1197,1255]" gridcol="4" gridrow="4" pageId="3" pageNumber="60">2.01 1.89</td>
<td id="DF3D371EFFC500610828FB06F8D3FB48" box="[1058,1168,1197,1255]" gridcol="5" gridrow="4" pageId="3" pageNumber="60">1.40 1.78</td>
<td id="DF3D371EFFC5006108FFFB06F978FB48" box="[1269,1339,1197,1255]" gridcol="6" gridrow="4" pageId="3" pageNumber="60">2.51 1.51</td>
</tr>
<tr id="9CEC5E62FFC500610C95FB51F978FA9B" box="[159,1339,1274,1332]" gridrow="5" pageId="3" pageNumber="60">
<th id="DF3D371EFFC500610C95FB51FCA8FA9B" box="[159,235,1274,1332]" gridcol="0" gridrow="5" pageId="3" pageNumber="60">Leg IV</th>
<td id="DF3D371EFFC500610D48FB51FDF2FA9B" box="[322,433,1274,1332]" gridcol="1" gridrow="5" pageId="3" pageNumber="60">0.95 1.00</td>
<td id="DF3D371EFFC500610E12FB51FE1DFA9B" box="[536,606,1274,1332]" gridcol="2" gridrow="5" pageId="3" pageNumber="60">2.90 2.73</td>
<td id="DF3D371EFFC500610EC2FB51FF53FA9B" box="[712,784,1274,1332]" gridcol="3" gridrow="5" pageId="3" pageNumber="60">1.62 1.28</td>
<td id="DF3D371EFFC500610F70FB51FFF2FA9B" box="[890,945,1274,1332]" gridcol="4" gridrow="5" pageId="3" pageNumber="60">2.06 1.96</td>
<td id="DF3D371EFFC500610828FB51F8D3FA9B" box="[1058,1168,1274,1332]" gridcol="5" gridrow="5" pageId="3" pageNumber="60">1.85 1.80</td>
<td id="DF3D371EFFC5006108FFFB51F978FA9B" box="[1269,1339,1274,1332]" gridcol="6" gridrow="5" pageId="3" pageNumber="60">1.86 1.74</td>
</tr>
</table>
</paragraph>
<paragraph id="22635C20FFC5FF9D0CCFFA2BFF34F925" blockId="3.[151,1437,1408,2034]" pageId="3" pageNumber="60">
<emphasis id="10A88032FFC5FF9D0CCFFA2BFD5FFA36" box="[197,284,1408,1433]" italics="true" pageId="3" pageNumber="60">Female</emphasis>
. Similar to male in size, but opisthosoma shorter and wider, ratio length/maximum width 1.3 (fig. 8). Legs not enlarged; leg I with distinct dorsal groove only on basitarsus; legs IIIV with distinct dorsal grooves on all podomeres except patella and terminal teleomeres. Anterior genital lip (sternite VIII) of genital opening
<quantity id="E524F1C5FFC5FF9D0CF6FA53FD27F9BD" box="[252,356,1528,1554]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="3" pageNumber="60" unit="mm" value="0.76">0.76 mm</quantity>
wide; posterior genital lip (sternite IX)
<quantity id="E524F1C5FFC5FF9D0F3CFA53FFDDF9BD" box="[822,926,1528,1554]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="60" unit="mm" value="1.03">1.03 mm</quantity>
wide (fig. 10); posterior genital sacks (possibly functioning as spermathecae [
<bibRefCitation id="464D21D1FFC5FF9D0E32F98BFEFEF995" author="Legg" box="[568,701,1568,1594]" pageId="3" pageNumber="64" refString="Legg, G. (1976) The external morphology of a new species of ricinuleid (Arachnida) from Sierra Leone. Zoological Journal of the Linnean Society, 59, 1 - 58." type="journal article" year="1976">Legg 1976</bibRefCitation>
]) tubular, arranged in 2 groups on both sides of midline; each group with 67 sacks; outermost sacks longest, 130150 µm long (fig. 11); accessory spermathecae
<emphasis id="10A88032FFC5FF9D0949F9E3F932F9CE" box="[1347,1393,1608,1633]" italics="true" pageId="3" pageNumber="60">(as)</emphasis>
(as defined by
<bibRefCitation id="464D21D1FFC5FF9D0D11F9DBFD84F925" author="Harvey" box="[283,455,1648,1674]" pageId="3" pageNumber="64" refString="Harvey, M. S. (1984) The female of Ricinoides westermannii (Guerin-Meneville), with notes on those of R. afzelii (Thorell) and R. karschii (Hansen &amp; Sorensen) (Ricinulei). Bulletin of the British Arachnological Society, 6, 205 - 210." type="journal article" year="1984">Harvey [1984]</bibRefCitation>
) well developed, 340350 µm long.
</paragraph>
<paragraph id="22635C20FFC5FF9D0CCFF933F906F8AD" blockId="3.[151,1437,1408,2034]" pageId="3" pageNumber="60">
<emphasis id="10A88032FFC5FF9D0CCFF933FD05F91E" box="[197,326,1688,1713]" italics="true" pageId="3" pageNumber="60">Coloration</emphasis>
. General coloration of adult animals gray-brown, body often covered with soil and mud, obscuring underlying body coloration; freshly molted adults reddish; as in most
<taxonomicName id="E5DC27A3FFC5FF9D087BF96BF89CF975" box="[1137,1247,1728,1754]" class="Arachnida" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="60" phylum="Arthropoda" rank="order">Ricinulei</taxonomicName>
, the larvae and nymphs are orange-red to red, with opisthosomal tergites darker than surrounding membrane (fig. 28).
</paragraph>
</subSubSection>
<subSubSection id="6AC60FABFFC5FF9A0CCFF8A4F88DFF75" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60" type="materials_examined">
<paragraph id="22635C20FFC5FF9A0CCFF8A4F88DFF75" blockId="3.[151,1437,1408,2034]" lastBlockId="4.[151,1436,152,218]" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
<emphasis id="10A88032FFC5FF9D0CCFF8A4FDFDF886" bold="true" box="[197,446,1807,1833]" pageId="3" pageNumber="60">Material examined.</emphasis>
<collectingCountry id="5ACB1CB0FFC5FF9D0DC0F8BBFE77F885" box="[458,564,1808,1834]" name="Ghana" pageId="3" pageNumber="60">GHANA</collectingCountry>
: Eastern Region,
<taxonomicName id="E5DC27A3FFC5FF9D0F1EF8BBFF23F885" box="[788,864,1808,1834]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="atewa">Atewa</taxonomicName>
Range Forest Reserve, Asiakwa (
<geoCoordinate id="47E83AE7FFC5FF9D090EF8BBF9D4F885" box="[1284,1431,1808,1834]" direction="north" orientation="latitude" pageId="3" pageNumber="60" precision="1" value="6.2621946">6°15'43.9''N</geoCoordinate>
,
<geoCoordinate id="47E83AE7FFC5FF9D0C9DF893FD70F8FD" box="[151,307,1848,1874]" direction="west" orientation="longitude" pageId="3" pageNumber="60" precision="1" value="-0.5545834">0°33'16.5”W</geoCoordinate>
), elev.
<quantity id="E524F1C5FFC5FF9D0D80F893FD99F8FD" box="[394,474,1848,1874]" metricMagnitude="2" metricUnit="m" metricValue="8.27" pageId="3" pageNumber="60" unit="m" value="827.0">827 m</quantity>
,
<date id="56627AE0FFC5FF9D0DEDF893FE2BF8FD" box="[487,616,1848,1874]" pageId="3" pageNumber="60" value="2007-12-08">8.XII.2007</date>
(leg. P. Naskrecki, V. Awotwe-Pratt and N. Jengre)
<typeStatus id="FD67E282FFC5FF9D08F9F893F919F8FD" box="[1267,1370,1848,1874]" pageId="3" pageNumber="60" type="holotype">holotype</typeStatus>
male (
<collectionCode id="44CDC4E5FFC5FF9D0C95F8CBFCBCF8D5" LSID="urn:lsid:biocol.org:col:34925" box="[159,255,1888,1914]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="60">AMNH</collectionCode>
),
<specimenCount id="34DA97A9FFC5FF9D0D18F8CBFD22F8D5" box="[274,353,1888,1914]" pageId="3" pageNumber="60" type="male">1 male</specimenCount>
(
<typeStatus id="FD67E282FFC5FF9D0D7AF8CBFD98F8D5" box="[368,475,1888,1914]" pageId="3" pageNumber="60" type="paratype">paratype</typeStatus>
),
<specimenCount id="34DA97A9FFC5FF9D0DE4F8CBFE17F8D5" box="[494,596,1888,1914]" pageId="3" pageNumber="60" type="female">2 female</specimenCount>
(
<typeStatus id="FD67E282FFC5FF9D0E68F8CBFE9BF8D5" box="[610,728,1888,1914]" pageId="3" pageNumber="60" type="paratype">paratypes</typeStatus>
), 1 deutonymph, 2 tritonymphs (
<collectionCode id="44CDC4E5FFC5FF9D085FF8CBF8DAF8D5" LSID="urn:lsid:biocol.org:col:33791" box="[1109,1177,1888,1914]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="60">MCZ</collectionCode>
,
<collectionCode id="44CDC4E5FFC5FF9D08AEF8CBF940F8D5" LSID="urn:lsid:biocol.org:col:34925" box="[1188,1283,1888,1914]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="60">AMNH</collectionCode>
); same locality,
<date id="56627AE0FFC5FF9D0CCEF823FD37F80D" box="[196,372,1928,1954]" pageId="3" pageNumber="60" value="2006-06-11" valueMax="2006-06-16" valueMin="2006-06-11">1116.VI.2006</date>
(leg. P. Naskrecki)
<specimenCount id="34DA97A9FFC5FF9D0E7DF823FE9DF80D" box="[631,734,1928,1954]" pageId="3" pageNumber="60" type="female">1 female</specimenCount>
(
<typeStatus id="FD67E282FFC5FF9D0EE5F823FF19F80D" box="[751,858,1928,1954]" pageId="3" pageNumber="60" type="paratype">paratype</typeStatus>
), 1 deutonymph, 1 tritonymph (
<collectionCode id="44CDC4E5FFC5FF9D08D9F823F95AF80D" LSID="urn:lsid:biocol.org:col:33791" box="[1235,1305,1928,1954]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="60">MCZ</collectionCode>
); Asiakwa, along main reserve road (6°15'.7''N,
<geoCoordinate id="47E83AE7FFC5FF9D0E4BF81BFE98F865" box="[577,731,1968,1994]" direction="west" orientation="longitude" pageId="3" pageNumber="60" precision="1" value="-0.5649167">0°33'53.7''W</geoCoordinate>
), elev.
<quantity id="E524F1C5FFC5FF9D0F26F81BFF3AF865" box="[812,889,1968,1994]" metricMagnitude="2" metricUnit="m" metricValue="8.17" pageId="3" pageNumber="60" unit="m" value="817.0">817 m</quantity>
,
<date id="56627AE0FFC5FF9D0F8EF81BF86CF865" box="[900,1071,1968,1994]" pageId="3" pageNumber="60" value="2007-12-07" valueMax="2007-12-11" valueMin="2007-12-07">711.XII.2007</date>
(leg. P. Naskrecki, V. Awotwe- Pratt and N. Jengre)
<specimenCount id="34DA97A9FFC5FF9D0D91F873FDB4F85D" box="[411,503,2008,2034]" pageId="3" pageNumber="60" type="male">4 males</specimenCount>
,
<specimenCount id="34DA97A9FFC5FF9D0E08F873FE30F85D" box="[514,627,2008,2034]" pageId="3" pageNumber="60" type="female">2 females</specimenCount>
(
<typeStatus id="FD67E282FFC5FF9D0E8BF873FEB4F85D" box="[641,759,2008,2034]" pageId="3" pageNumber="60" type="paratype">paratypes</typeStatus>
), 3 deutonymphs, 6 tritonymphs (
<collectionCode id="44CDC4E5FFC5FF9D0877F873F882F85D" LSID="urn:lsid:biocol.org:col:33791" box="[1149,1217,2008,2034]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="60">MCZ</collectionCode>
,
<collectionCode id="44CDC4E5FFC5FF9D08C7F873F96FF85D" LSID="urn:lsid:biocol.org:col:34925" box="[1229,1324,2008,2034]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="60">AMNH</collectionCode>
); Ajenjua Bepo Forest Reserve (
<geoCoordinate id="47E83AE7FFC2FF9A0DA1FF33FE73FF1D" box="[427,560,152,178]" direction="north" orientation="latitude" pageId="4" pageNumber="61" precision="1" value="6.3673058">6°22'2.3''N</geoCoordinate>
,
<geoCoordinate id="47E83AE7FFC2FF9A0E4BFF33FEF6FF1D" box="[577,693,152,178]" direction="west" orientation="longitude" pageId="4" pageNumber="61" precision="15" value="-1.0327778">1°1'58''W</geoCoordinate>
), elev.
<quantity id="E524F1C5FFC2FF9A0F19FF33FFE3FF1D" box="[787,928,152,178]" metricMagnitude="2" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="4" pageNumber="61" unit="m" value="315.0" valueMax="330.0" valueMin="300.0">300330 m</quantity>
,
<date id="56627AE0FFC2FF9A0FBBFF33F835FF1D" box="[945,1142,152,178]" pageId="4" pageNumber="61" value="2006-08-26" valueMax="2006-08-30" valueMin="2006-08-26">2630.VIII.2006</date>
(leg. P. Naskrecki)
<specimenCount id="34DA97A9FFC2FF9A0987FF33FC9DFF75" pageId="4" pageNumber="61" type="male">3 males</specimenCount>
,
<specimenCount id="34DA97A9FFC2FF9A0CE0FF6BFD18FF75" box="[234,347,192,218]" pageId="4" pageNumber="61" type="female">4 females</specimenCount>
(
<typeStatus id="FD67E282FFC2FF9A0D60FF6BFDA2FF75" box="[362,481,192,218]" pageId="4" pageNumber="61" type="paratype">paratypes</typeStatus>
), 1 protonymph, 1 deutonymph, 2 tritonymphs (
<collectionCode id="44CDC4E5FFC2FF9A0818FF6BF815FF75" LSID="urn:lsid:biocol.org:col:33791" box="[1042,1110,192,218]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="4" pageNumber="61">MCZ</collectionCode>
,
<collectionCode id="44CDC4E5FFC2FF9A0868FF6BF882FF75" LSID="urn:lsid:biocol.org:col:34925" box="[1122,1217,192,218]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="61">AMNH</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="76A30CA8FFC2FF9A0C9DF899FC9DF81E" httpUri="https://zenodo.org/record/274093/files/figure.png" pageId="4" pageNumber="61" targetBox="[172,1422,295,1803]" targetPageId="4">
<paragraph id="22635C20FFC2FF9A0C9DF899FC9DF81E" blockId="4.[151,1436,1842,1969]" pageId="4" pageNumber="61">
<emphasis id="10A88032FFC2FF9A0C9DF899FD13F8E5" bold="true" box="[151,336,1842,1866]" pageId="4" pageNumber="61">FIGURES 813.</emphasis>
<taxonomicName id="E5DC27A3FFC2FF9A0D5CF899FE4DF8E6" box="[342,526,1842,1865]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="atewa" status="sp. nov.">
<emphasis id="10A88032FFC2FF9A0D5CF899FE4DF8E6" box="[342,526,1842,1865]" italics="true" pageId="4" pageNumber="61">Ricinoides atewa</emphasis>
</taxonomicName>
<emphasis id="10A88032FFC2FF9A0E1EF899FE13F8E5" bold="true" box="[532,592,1842,1866]" pageId="4" pageNumber="61">
<taxonomicNameLabel id="0B9B3D49FFC2FF9A0E1EF899FE13F8E5" box="[532,592,1842,1866]" pageId="4" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: 8, female, dorsal view; 9, ventral view; 10, female posterior genital lip, showing genital sacks (spermathecae), internal view; 11, close-up of female posterior genital lip, showing genital sacks (spermathecae) and accessory spermathecae (
<emphasis id="10A88032FFC2FF9A0E53F8D3FE3DF820" box="[601,638,1912,1935]" italics="true" pageId="4" pageNumber="61">a.s.</emphasis>
); 12, tubercles and navicular setae on opisthosomal tergite; 13, female telson.
</paragraph>
</caption>
<caption id="76A30CA8FFC3FF9B0C9DF977FE63F896" pageId="5" pageNumber="62">
<paragraph id="22635C20FFC3FF9B0C9DF977FE63F896" blockId="5.[151,1436,1756,1919]" pageId="5" pageNumber="62">
<emphasis id="10A88032FFC3FF9B0C9DF977FD1CF95B" bold="true" box="[151,351,1756,1780]" pageId="5" pageNumber="62">FIGURES 1419.</emphasis>
<taxonomicName id="E5DC27A3FFC3FF9B0D62F976FEC8F95B" authority="Thorell" authorityName="Thorell" box="[360,651,1757,1780]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="afzelii">
<emphasis id="10A88032FFC3FF9B0D62F976FD9AF95B" box="[360,473,1757,1780]" italics="true" pageId="5" pageNumber="62">Ricinoides</emphasis>
<emphasis id="10A88032FFC3FF9B0DEBF976FE61F95B" box="[481,546,1757,1780]" italics="true" pageId="5" pageNumber="62">afzelii</emphasis>
(Thorell)
</taxonomicName>
(specimen from Guinea, Diecke): 1417, tarsal process, frontal, lateral, posterior, and dorsal views (
<emphasis id="10A88032FFC3FF9B0DC9F8ABFD93F8B8" box="[451,464,1792,1815]" italics="true" pageId="5" pageNumber="62">a</emphasis>
,
<emphasis id="10A88032FFC3FF9B0DD6F8ABFDAAF8B8" box="[476,489,1792,1815]" italics="true" pageId="5" pageNumber="62">b</emphasis>
,
<emphasis id="10A88032FFC3FF9B0DFDF8ABFE40F8B8" box="[503,515,1792,1815]" italics="true" pageId="5" pageNumber="62">c</emphasis>
apical lobes of tarsal process;
<emphasis id="10A88032FFC3FF9B0F50F8ABFF30F8B8" box="[858,883,1792,1815]" italics="true" pageId="5" pageNumber="62">ac</emphasis>
accessory piece;
<emphasis id="10A88032FFC3FF9B084BF8ABF815F8B8" box="[1089,1110,1792,1815]" italics="true" pageId="5" pageNumber="62">lb</emphasis>
lateral lobe); 18, male tarsus III, posterior view; 19, male tibia III.
</paragraph>
</caption>
<caption id="76A30CA8FFC3FF9B0C9DF8EFF9DFF8D1" httpUri="https://zenodo.org/record/274094/files/figure.png" pageId="5" pageNumber="62" targetBox="[182,1400,205,1727]" targetPageId="5">
<paragraph id="22635C20FFC3FF9B0C9DF8EFF9DFF8D1" blockId="5.[151,1436,1756,1919]" pageId="5" pageNumber="62">
<emphasis id="10A88032FFC3FF9B0C9DF8EFFD1DF8F3" bold="true" box="[151,350,1860,1884]" pageId="5" pageNumber="62">FIGURES 2025.</emphasis>
<taxonomicName id="E5DC27A3FFC3FF9B0D6EF8EEFD80F8F3" box="[356,451,1861,1884]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="atewa" status="sp. nov.">
<emphasis id="10A88032FFC3FF9B0D6EF8EEFD80F8F3" box="[356,451,1861,1884]" italics="true" pageId="5" pageNumber="62">R. atewa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0B9B3D49FFC3FF9B0DC3F8EEFE45F8F3" box="[457,518,1861,1884]" pageId="5" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
: 2023, tarsal process, frontal, lateral, posterior, and dorsal views; 24, male tarsus III, posterior view (
<emphasis id="10A88032FFC3FF9B0D35F8C3FD10F8D0" box="[319,339,1896,1919]" italics="true" pageId="5" pageNumber="62">bt</emphasis>
basitarsus;
<emphasis id="10A88032FFC3FF9B0DECF8C3FE42F8D0" box="[486,513,1896,1919]" italics="true" pageId="5" pageNumber="62">bp</emphasis>
basitarsal process;
<emphasis id="10A88032FFC3FF9B0EEFF8C3FEBBF8D0" box="[741,760,1896,1919]" italics="true" pageId="5" pageNumber="62">lc</emphasis>
lamina cyathiformis;
<emphasis id="10A88032FFC3FF9B0FFDF8C3F87FF8D0" box="[1015,1084,1896,1919]" italics="true" pageId="5" pageNumber="62">tt1tt4</emphasis>
tarsomeres); 25, male tibia III.
</paragraph>
</caption>
<subSubSection id="6AC60FABFFC3FF990CCFF819FD90FDED" lastPageId="7" lastPageNumber="64" pageId="5" pageNumber="62" type="etymology">
<paragraph id="22635C20FFC3FF980CCFF819FE1EFF76" blockId="5.[151,1437,1970,2036]" lastBlockId="6.[151,1436,152,218]" lastPageId="6" lastPageNumber="63" pageId="5" pageNumber="62">
<emphasis id="10A88032FFC3FF9B0CCFF819FD11F863" bold="true" box="[197,338,1970,1996]" pageId="5" pageNumber="62">Etymology.</emphasis>
The specific epithet refers to the
<typeStatus id="FD67E282FFC3FF9B0EE6F819FF5DF863" box="[748,798,1970,1996]" pageId="5" pageNumber="62">type</typeStatus>
locality of the new species, the
<taxonomicName id="E5DC27A3FFC3FF9B08A2F819F8B6F863" box="[1192,1269,1970,1996]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="atewa">Atewa</taxonomicName>
Range Forest Reserve, one of the last remaining sites of the Upland Evergreen Forest in West Africa. The
<taxonomicName id="E5DC27A3FFC3FF9B08C5F871F95FF85B" box="[1231,1308,2010,2036]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="atewa">Atewa</taxonomicName>
forests are threatened by mining, logging, and bushmeat hunting, and their preservation is vital for the survival of many species, including this new
<emphasis id="10A88032FFC0FF980DD0FF6BFE1EFF76" box="[474,605,192,217]" italics="true" pageId="6" pageNumber="63">
<taxonomicName id="E5DC27A3FFC0FF980DD0FF6BFE1AFF76" box="[474,601,192,217]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="63" phylum="Arthropoda" rank="genus">Ricinoides</taxonomicName>
.
</emphasis>
</paragraph>
<caption id="76A30CA8FFC0FF980C9DF93BFF8AF943" httpUri="https://zenodo.org/record/274095/files/figure.png" pageId="6" pageNumber="63" targetBox="[151,1436,274,1655]" targetPageId="6">
<paragraph id="22635C20FFC0FF980C9DF93BFF8AF943" blockId="6.[151,1436,1680,1772]" pageId="6" pageNumber="63">
<emphasis id="10A88032FFC0FF980C9DF93BFD1CF907" bold="true" box="[151,351,1680,1704]" pageId="6" pageNumber="63">FIGURES 2628.</emphasis>
<taxonomicName id="E5DC27A3FFC0FF980D6CF93BFE5CF908" box="[358,543,1680,1703]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="atewa" status="sp. nov.">
<emphasis id="10A88032FFC0FF980D6CF93BFE5CF908" box="[358,543,1680,1703]" italics="true" pageId="6" pageNumber="63">Ricinoides atewa</emphasis>
</taxonomicName>
<emphasis id="10A88032FFC0FF980E2CF93BFE21F907" bold="true" box="[550,610,1680,1704]" pageId="6" pageNumber="63">
<taxonomicNameLabel id="0B9B3D49FFC0FF980E2CF93BFE21F907" box="[550,610,1680,1704]" pageId="6" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and habitat. 26, an old, abandoned mound of soil termites
<taxonomicName id="E5DC27A3FFC0FF9808EBF93BF91FF908" box="[1249,1372,1680,1703]" class="Insecta" family="Termitidae" genus="Cubitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="6" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="10A88032FFC0FF9808EBF93BF91FF908" box="[1249,1372,1680,1703]" italics="true" pageId="6" pageNumber="63">Cubitermes</emphasis>
</taxonomicName>
sp., a typical habitat of
<taxonomicName id="E5DC27A3FFC0FF980D5CF918FDF7F965" box="[342,436,1715,1738]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="atewa">
<emphasis id="10A88032FFC0FF980D5CF918FDF7F965" box="[342,436,1715,1738]" italics="true" pageId="6" pageNumber="63">R. atewa</emphasis>
</taxonomicName>
in the Upland Evergreen Forest of Ghana; 27, female
<taxonomicName id="E5DC27A3FFC0FF980809F918F821F965" box="[1027,1122,1715,1738]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="atewa">
<emphasis id="10A88032FFC0FF980809F918F821F965" box="[1027,1122,1715,1738]" italics="true" pageId="6" pageNumber="63">R. atewa</emphasis>
</taxonomicName>
; 28, tritonymph of
<taxonomicName id="E5DC27A3FFC0FF98093DF918F9D6F965" box="[1335,1429,1715,1738]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="atewa">
<emphasis id="10A88032FFC0FF98093DF918F9D6F965" box="[1335,1429,1715,1738]" italics="true" pageId="6" pageNumber="63">R. atewa</emphasis>
</taxonomicName>
, note the reddish coloration typical of larval and nymphal stages of
<taxonomicName id="E5DC27A3FFC0FF980F55F97EFF87F943" box="[863,964,1749,1772]" class="Arachnida" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="63" phylum="Arthropoda" rank="order">Ricinulei</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="22635C20FFC0FF990CCFF88BFD90FDED" blockId="6.[151,1436,1824,2010]" lastBlockId="7.[151,1437,152,578]" lastPageId="7" lastPageNumber="64" pageId="6" pageNumber="63">
<emphasis id="10A88032FFC0FF980CCFF88BFDD1F895" bold="true" box="[197,402,1824,1850]" pageId="6" pageNumber="63">Natural history.</emphasis>
Although searches were conducted in both the rainy and dry seasons, seasonality appeared to have no effect on the abundance of this species, perhaps because of the nearly constant high humidity of the soil in the closed canopy forest. Their abundance was relatively low, and a three-person team usually managed to collect only 23 individuals in an hour of searching. Their distribution in the forest appeared to be patchy, and restricted only to undisturbed, shady parts of the forest, with little or no undergrowth but well developed leaf litter.
<taxonomicName id="E5DC27A3FFC1FF990E51FF33FE80FF1E" box="[603,707,152,177]" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="atewa">
<emphasis id="10A88032FFC1FF990E51FF33FE80FF1E" box="[603,707,152,177]" italics="true" pageId="7" pageNumber="64">R. atewa</emphasis>
</taxonomicName>
shows a degree of association with termites, and individuals were often collected near termite colonies (mostly nasutitermites) and small colonies of ants. Nymphs were frequently encountered in old, abandoned mounds of
<taxonomicName id="E5DC27A3FFC1FF990F0DFF43FFCDFEAE" box="[775,910,232,257]" class="Insecta" family="Termitidae" genus="Cubitermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="7" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="10A88032FFC1FF990F0DFF43FFCDFEAE" box="[775,910,232,257]" italics="true" pageId="7" pageNumber="64">Cubitermes</emphasis>
</taxonomicName>
sp., both in the soil under the nest and inside empty chambers of the nest. While such nests had no active termite colonies, small colonies of ants were always present there, providing a potential source of food for the ricinuleids. In captivity individuals of
<taxonomicName id="E5DC27A3FFC1FF990989FE93FC9EFED6" class="Arachnida" family="Ricinoididae" genus="Ricinoides" kingdom="Animalia" order="Ricinulei" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="atewa">
<emphasis id="10A88032FFC1FF990989FE93FC9EFED6" italics="true" pageId="7" pageNumber="64">R. atewa</emphasis>
</taxonomicName>
were observed feeding both on nymphs and workers of termites, and larvae of ants. The density of individuals in abandoned termite mounds was usually high, reaching up to 8 individuals per nest (not all nymphal specimens were collected). However, only single individuals were found in individual chambers, whereas multiple individuals could be found sitting close together under logs and rocks (both nymphs and adults). While ricinuleids show nocturnal activity, on one occasion a single nymph was found during the day walking on the surface of leaf litter.
</paragraph>
</subSubSection>
</treatment>
</document>