treatments-xml/data/9B/6A/FA/9B6AFA91B4245D419EB13B4232A47FFA.xml

<document id="5273750087F6BA878A60A4F67B934A96" ID-CLB-Dataset="299410" ID-DOI="10.3897/asp.82.e113041" ID-GBIF-Dataset="1c9fb08b-57c8-491c-bb13-aa763dcbb1c8" ID-ZooBank="B2966C0A-75B7-4B73-A45C-0A2EFB159715" ID-publisher-id="113041" URI-arpha="E3A8F026-F964-5504-980C-341627EF4BE6" URI-zoobank="http://zoobank.org/B2966C0A-75B7-4B73-A45C-0A2EFB159715" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1719937550960" checkinUser="pensoft" docAuthor="Wang, Jin, Yao, Chong-Hui, Jiang, Chao &amp; Li, Wei-Chun" docDate="2024" docId="9B6AFA91B4245D419EB13B4232A47FFA" docLanguage="en" docName="Arthropod Systematics &amp; Phylogeny 82: 527-549" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 82" docSource="https://arthropod-systematics.arphahub.com/article/113041/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Lucasioides formosus Li &amp; Wang 2024, sp. nov." docType="treatment" docUuid="989DCD87-7808-4D8F-B7AD-49FA36C7A95C" docUuidSource="ZooBank" docVersion="5" dtd-version="3.0" lastPageNumber="549" masterDocId="E3A8F026F9645504980C341627EF4BE6" masterDocTitle="Integrative taxonomy reveals exceptional species diversity of Lucasioides from China (Isopoda: Oniscidea: Agnaridae)" masterLastPageNumber="549" masterPageNumber="527" pageNumber="527" updateTime="1719955579240" updateUser="ExternalLinkService">
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<mods:title id="553E2C35743112257C795BB30A437445">Integrative taxonomy reveals exceptional species diversity of Lucasioides from China (Isopoda: Oniscidea: Agnaridae)</mods:title>
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<mods:namePart id="E91A90DB805CC78FF4709D874E857B92">Wang, Jin</mods:namePart>
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<mods:namePart id="BB358ADC82823BDE06027911D6678B22">Yao, Chong-Hui</mods:namePart>
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<mods:namePart id="87A27A34A469275C423E55EE4986B745">Jiang, Chao</mods:namePart>
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<mods:affiliation id="A22EEB2FE7198109C46A920091C5B32A">State Key Laboratory for Quality Ensurance and Sustainable Use of Dao-di Herbs, National Resource Center for Chinese Materia Medica, China Academy of Chinese Medical Sciences, Beijing 100700, China</mods:affiliation>
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<mods:namePart id="6A5FB11252ED5BF778894991E6F726D8">Li, Wei-Chun</mods:namePart>
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<subSubSection id="07747159E8BE85674C6F66001D158248" type="nomenclature">
<paragraph id="2A4F833B99C02969FC128DF71A3046F0">
<heading id="2F1F320884C54FC70EC8C445504FEFCB" reason="label">3.3. 5.</heading>
<taxonomicName id="9A99D6153976FEE3EB6BE2C06ACCA273" authority="Li &amp; Wang" authorityName="Li &amp; Wang" authorityYear="2024" class="Malacostraca" family="Agnaridae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="260F0E86E474DA532FE226499F154031" italics="true">Lucasioides formosus</emphasis>
Li &amp; Wang
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<taxonomicNameLabel id="08F0FC342DD760EE3539875631D0DBCA" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="F6232B656FFF5BD1976805C6815EC2E5" type="description">
<paragraph id="EC08740B0388612F1BC4410A928C4EB5">
<figureCitation id="2A3FE7C6224BBA3BF7DA4F12DDA26221" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of male Lucasioides species in dorsal view. A L. dissectus sp. nov.; B L. isseli (Arcangeli, 1927); C L. dianensis sp. nov.; D L. digitatus sp. nov.; E L. schmidti sp. nov.; F L. nudus Li, 2017; G L. formosus sp. nov.; H L. gracilentus sp. nov.; I L. subcurvatus sp. nov. (Scale: 1 mm)." figureDoi="10.3897/asp.82.e113041.figure6" httpUri="https://binary.pensoft.net/fig/1083536">Figures 6 G</figureCitation>
, 
<figureCitation id="D23874D2D2EAD348AE54F549024E53CB" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11</figureCitation>
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<heading id="5739D47EC9568AE3B37B12DD04372B55" reason="title">Type material.</heading>
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.
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• 
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, 
<geoCoordinate id="C5DBD318162AB95AAC1650C3B06C75B9" degrees="99.8193" direction="east" orientation="longitude" precision="5" value="99.8193">99.8193 ° E</geoCoordinate>
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<collectorName id="3F4613FED4F180F21AF07A893972BD59">J. Wang</collectorName>
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<collectorName id="D5FD0313A0090EF695C83DC3A64E31EB">X. K. Hong</collectorName>
leg. (DNA no. 
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, Prep. slide no. 
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. — 
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, 
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; same data as the holotype (DNA nos. 
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<collectorName id="D5EAA081BD046107F5EBEC6CD18A6F11">J. Wang</collectorName>
, 
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leg. (DNA nos. 
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- 
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, Prep. slide no. 
<specimenCode id="B53A5E3E3B17BAED2241BC15A49A80B9">L 22082</specimenCode>
)
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. • 
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<location id="026991CA9A7E6B42E2AAB14AD2B98333" LSID="urn:lsid:plazi:treatment:9B6AFA91B4245D419EB13B4232A47FFA:026991CA9A7E6B42E2AAB14AD2B98333" country="Hong Kong" county="Jianchuan County" latitude="26.5328" longLatPrecision="6" longitude="99.8877" municipality="Dali City" name="Qianshi Mountain" stateProvince="Yunnan Province">Qianshi Mountain</location>
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<collectorName id="21C2CEE219000C6E2380DF183B3C4C37">J. Wang</collectorName>
, 
<collectorName id="2285D7A272D0F801F998914AF4BEF63C">X. K. Hong</collectorName>
leg. (DNA nos. 
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- 
<specimenCode id="A432B790BDBC492420FCBACA81FBA5FB">QSS 2206</specimenCode>
, Prep. slide no. 
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</materialsCitation>
. • 
<materialsCitation id="0C2AD0B736A16288308CECAF6C9E3A70" ID-GBIF-Occurrence="4902831319" collectingDate="2022-07-07" collectionCode="LJGC" collectorName="J. Wang &amp; X. K. Hong" country="Hong Kong" elevation="2465" latitude="26.9258" location="Gucheng District" longLatPrecision="6" longitude="100.2079" municipality="Lijiang City" specimenCode="LJGC 2201, LJGC 2204" specimenCount="4" specimenCount-female="4" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount id="04E05384CFE66BE825AE4FCE3630C9D4" count="4" type="female">4 ♀♀</specimenCount>
; 
<collectingRegion id="0B5B825408B53366D39B0254AE093971" country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingMunicipality id="41DF96CB631E6206B4F5E527B8383ED4">Lijiang City</collectingMunicipality>
, 
<location id="3B17F375ED3DC2418419A80BD8372C89" LSID="urn:lsid:plazi:treatment:9B6AFA91B4245D419EB13B4232A47FFA:3B17F375ED3DC2418419A80BD8372C89" country="Hong Kong" latitude="26.9258" longLatPrecision="6" longitude="100.2079" municipality="Lijiang City" name="Gucheng District" stateProvince="Yunnan Province">Gucheng District</location>
; 
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<geoCoordinate id="3F6ED07976AD829637EF95A158F91B6F" degrees="26.9258" direction="north" orientation="latitude" precision="5" value="26.9258">26.9258 ° N</geoCoordinate>
, 
<geoCoordinate id="BBA5014A1310ECEE773D8F0F84D66B7C" degrees="100.2079" direction="east" orientation="longitude" precision="5" value="100.2079">100.2079 ° E</geoCoordinate>
</named-content>
, el. 
<elevation id="5F4D29CC730BEFE078D805AF6659F6E9" metricMagnitude="3" metricUnit="m" metricValue="2.465" unit="m" value="2465.0">
<quantity id="C204AFA1305995A49830DAAFA5E10CA2" metricMagnitude="3" metricUnit="m" metricValue="2.465" unit="m" value="2465.0">2465 m</quantity>
a. s. l.
</elevation>
; 
<date id="957B26E17C038CC27693E9CD3469321E" value="2022-07-07">
<collectingDate id="A518FB9EE9C6C611936B400E321497DF" value="2022-07-07">7. vii. 2022</collectingDate>
</date>
; 
<collectorName id="3C16F1F78AFB5A779332C74B7EC23BCD">J. Wang</collectorName>
, 
<collectorName id="DA8CBC70BE852346047929BDD8CA3051">X. K. Hong</collectorName>
leg. (DNA nos. 
<specimenCode id="A861F96D76722DF05FC857C2F2BCA1B7" collectionCode="LJGC">LJGC 2201</specimenCode>
- 
<specimenCode id="78F2E2F8B58ECC9BB9F72880E61547A8" collectionCode="LJGC">LJGC 2204</specimenCode>
)
</materialsCitation>
. • 
<materialsCitation id="E80585BDF459C18473E80DFA2ED9A9B8" ID-GBIF-Occurrence="4902831301" collectingDate="2022-07-06" collectionCode="MQC" collectorName="J. Wang &amp; X. K. Hong" country="Hong Kong" elevation="2486" latitude="26.9058" location="Meiquan Village" longLatPrecision="6" longitude="100.1499" municipality="Lijiang City" specimenCode="MQC 2201, MQC 22032205" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount id="195E654FD68033CFF0B4518E8A83585E" count="3" type="female">3 ♀♀</specimenCount>
; 
<collectingRegion id="8D9D53096A975A40E9718B3B20E718E7" country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingMunicipality id="07C1720382A0932FB68A1279F7B4CE4B">Lijiang City</collectingMunicipality>
, 
<location id="9C86DEA00D0785E7B9FA72096B3FA2E5" LSID="urn:lsid:plazi:treatment:9B6AFA91B4245D419EB13B4232A47FFA:9C86DEA00D0785E7B9FA72096B3FA2E5" country="Hong Kong" latitude="26.9058" longLatPrecision="6" longitude="100.1499" municipality="Lijiang City" name="Meiquan Village" stateProvince="Yunnan Province">Meiquan Village</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EEIBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[100.149900,26.905800]}">
<geoCoordinate id="E54F8D38D3317B12BC4D7BEC2BC3441B" degrees="26.9058" direction="north" orientation="latitude" precision="5" value="26.9058">26.9058 ° N</geoCoordinate>
, 
<geoCoordinate id="73AE4429FB9D292F086160ED8DC47B16" degrees="100.1499" direction="east" orientation="longitude" precision="5" value="100.1499">100.1499 ° E</geoCoordinate>
</named-content>
, el. 
<elevation id="800BA1C2B469DF49378DADD47B9D5383" metricMagnitude="3" metricUnit="m" metricValue="2.4859999999999998" unit="m" value="2486.0">
<quantity id="6DD54D1B1D6B6112653F3A3AE3CD37E4" metricMagnitude="3" metricUnit="m" metricValue="2.4859999999999998" unit="m" value="2486.0">2486 m</quantity>
a. s. l.
</elevation>
; 
<date id="0E67C65A86F9B635207A7B2A3E678719" value="2022-07-06">
<collectingDate id="5569E4F504BA93A98AB0FF887826AF9E" value="2022-07-06">6. vii. 2022</collectingDate>
</date>
; 
<collectorName id="57255EA746FF9E850D3F246A223162A1">J. Wang</collectorName>
, 
<collectorName id="4E6F2ACC004CED239C92BEE2E9B89D4B">X. K. Hong</collectorName>
leg. (DNA nos. 
<specimenCode id="389CD3AEDA7EA297BA606FE2564725EB" collectionCode="MQC">MQC 2201</specimenCode>
- 
<specimenCode id="4B1DF788C17AB95C2115B3526A1030DE">MQC 22032205</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIIBG" type="diagnosis">
<paragraph id="CEA46EA0CCEAA7D2949F6BF3C134194C">
<heading id="10C06E864ED766EF7D86839480F6D534" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="FF1097326BE3819D159DEFF880CDC3D1">Pereonite 1 slightly sinuous on posterior margin of epimeron, postero-lateral corner nearly right-angled; noduli laterales on pereonites 1-4 shifted from lateral margins than those of pereonites 5-7; pereopod 7 remarkably expanded on dorsal margin of carpus; pleopod 1 exopod slightly concave at posterior tip.</paragraph>
</subSubSection>
<subSubSection id="SECID0ENIBG" type="description">
<paragraph id="1C41CAAD2CE2C51743003FCFF9D63ADE">
<heading id="33871BE3AA1D57C66B1DC4BEB765468B" reason="title">Description.</heading>
</paragraph>
<paragraph id="D4BFF66FCD0B88D533A5056690850B27">
Body length of males 6.3-8.0 mm, of females 5.5-11.0 mm. Color in alcohol blackish brown, dorsum granulated, bearing irregular yellowish white muscle spots (living individuals with yellowish blue muscle spots). Pereonite 1 slightly sinuous on posterior margin of epimeron, postero-lateral corner nearly right-angled (Fig. 
<figureCitation id="357FF659D8A3C303256030C0AF8909D3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of male Lucasioides species in dorsal view. A L. dissectus sp. nov.; B L. isseli (Arcangeli, 1927); C L. dianensis sp. nov.; D L. digitatus sp. nov.; E L. schmidti sp. nov.; F L. nudus Li, 2017; G L. formosus sp. nov.; H L. gracilentus sp. nov.; I L. subcurvatus sp. nov. (Scale: 1 mm)." figureDoi="10.3897/asp.82.e113041.figure6" httpUri="https://binary.pensoft.net/fig/1083536">6 G</figureCitation>
). Noduli laterales on pereonites 1-4 shifted from lateral margins than those of pereonites 5-7 (Fig. 
<figureCitation id="C0E74D0776D2E36BE75CD8B2B7925577" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11 A</figureCitation>
). Telson triangular, width about twice as long as length, lateral margin slightly concave, apex blunted; uropod exopodite about three times as long as basipodite (Figs 
<figureCitation id="EFB026CE69AE0D2D4E226653674338E1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of male Lucasioides species in dorsal view. A L. dissectus sp. nov.; B L. isseli (Arcangeli, 1927); C L. dianensis sp. nov.; D L. digitatus sp. nov.; E L. schmidti sp. nov.; F L. nudus Li, 2017; G L. formosus sp. nov.; H L. gracilentus sp. nov.; I L. subcurvatus sp. nov. (Scale: 1 mm)." figureDoi="10.3897/asp.82.e113041.figure6" httpUri="https://binary.pensoft.net/fig/1083536">6 G</figureCitation>
, 
<figureCitation id="38186B29EC3626AB8D5DDAD1121FCE97" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11 A</figureCitation>
). — 
<emphasis id="EA893446CDA410A5FD6A35185C1EC9A0" italics="true">Cephalon</emphasis>
with arched median lobe, median lobe longer than lateral lobes (Figs 
<figureCitation id="B8124E376087F6AF1FABC919A45AF657" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus of male Lucasioides species in dorsal view. A L. dissectus sp. nov.; B L. isseli (Arcangeli, 1927); C L. dianensis sp. nov.; D L. digitatus sp. nov.; E L. schmidti sp. nov.; F L. nudus Li, 2017; G L. formosus sp. nov.; H L. gracilentus sp. nov.; I L. subcurvatus sp. nov. (Scale: 1 mm)." figureDoi="10.3897/asp.82.e113041.figure6" httpUri="https://binary.pensoft.net/fig/1083536">6 G</figureCitation>
, 
<figureCitation id="11FD1BD2A95B04CFF730C74DF71C7190" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11 A</figureCitation>
). Antennula with several aesthetascs on distal tip of third article (Fig. 
<figureCitation id="F8D427C203AAAE08087C49BBCDA28C3F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11 B</figureCitation>
). Antennal flagellum with second segment twice as long as first one (Fig. 
<figureCitation id="4EADF7AE2A6F682507B69263A58A2533" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11 C</figureCitation>
). — 
<emphasis id="41EBAB45225E4A0885346C8F08FD0E31" italics="true">Pereopod</emphasis>
1 with brush of long setae on carpus and merus (Fig. 
<figureCitation id="8C0EFD472ACCED9A6E6A5B4DB0382FEB" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11 D</figureCitation>
). Pereopod 7 ischium without deep depression on rostral surface, carpus remarkably expanded near middle on dorsal margin (Fig. 
<figureCitation id="84C8F71002C6137E4853EE866650D1C4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11 E</figureCitation>
). — 
<emphasis id="4ABE42152167359ECF35E977D580F45C" italics="true">Pleopods</emphasis>
1-5 exopods with 
<taxonomicName id="C4783B8611533EC39B637DEBDB9C8688" authorityName="Verhoeff" authorityYear="1917" class="Malacostraca" family="Agnaridae" genus="Protracheoniscus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="genus">
<emphasis id="49C03118E432C3E67C99400A70426979" italics="true">Protracheoniscus</emphasis>
</taxonomicName>
- type pseudotrachea (Fig. 
<figureCitation id="1E14AC5B9A11DD19F4E99E5D59CDB5AF" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11 F-J</figureCitation>
). — MALE: pleopods 1 exopod slightly concave at posterior tip, forming two inconspicuous lobes, endopod with broad basal part, narrowed towards apex, posterior tip beak-shaped and bent outwards (Fig. 
<figureCitation id="1AA72E561E263B8443587C3BB5AC4866" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11 F</figureCitation>
); pleopod 2 exopoddistinctive concave on outer margin, posterior two fifths with several setae, endopod styliform, slightly longer than exopod (Fig. 
<figureCitation id="21128E747B4A24D9EC6EA6DFB7D23D98" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lucasioides formosus sp. nov. A Holotype in dorsal view; B antennula; C antenna; D pereopod 1; E pereopod 7; F pleopod 1; G pleopod 2; H pleopod 3 exopod; I pleopod 4 exopod; J pleopod 5 exopod." figureDoi="10.3897/asp.82.e113041.figure11" httpUri="https://binary.pensoft.net/fig/1083541">11 G</figureCitation>
).
</paragraph>
<caption id="3D249E853AF4706AFB7755D8A2FCA71E" ID-DOI="10.3897/asp.82.e113041.figure11" ID-arpha="DB64C7A7-8AED-55AC-9578-16ECEFB2BA68" httpUri="https://binary.pensoft.net/fig/1083541" startId="F11">
<paragraph id="E487FEE6E74327AEC9582C0BDF04004B">
<label id="601191E0CC7F4B9C85CD497774FAC828">Figure 11.</label>
</paragraph>
<paragraph id="6E281EE6A72E7F237A50CD9B5E7C3CC7">
<taxonomicName id="CA45E8E8EA611F3F3445872865A4333E" authorityName="Wang &amp; Yao &amp; Jiang &amp; Li" authorityYear="2024" class="Malacostraca" family="Agnaridae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="CEC8FCE69FEB0293F2C25A2C3F31924F" italics="true">Lucasioides formosus</emphasis>
</taxonomicName>
<emphasis id="C33BC2BBF39C2DFBCAB8A87BA34FC213" bold="true">
<taxonomicNameLabel id="EB5FAC017FAFCF0053FBB997CA976AD5" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
Holotype in dorsal view; 
<emphasis id="80486DFC64648AD9C2AFC066EB78AE8B" bold="true">B</emphasis>
antennula; 
<emphasis id="90BB7329A2A36EB368E8B34F345813FD" bold="true">C</emphasis>
antenna; 
<emphasis id="825A711E3BE6E3B02AAF86F4842D92F0" bold="true">D</emphasis>
pereopod 1; 
<emphasis id="B6CAAF89FB3FC357B5DE35591E6898BC" bold="true">E</emphasis>
pereopod 7; 
<emphasis id="BE3716474013233C54886EA66920FA2E" bold="true">F</emphasis>
pleopod 1; 
<emphasis id="D11C07B0A104BBC6E0A7521014BAAFAB" bold="true">G</emphasis>
pleopod 2; 
<emphasis id="9F5A2B89093772B85BEEDACA86C4EA66" bold="true">H</emphasis>
pleopod 3 exopod; 
<emphasis id="2A687E60A1890B6987D094AB6F1AFFA1" bold="true">I</emphasis>
pleopod 4 exopod; 
<emphasis id="24D9770E3E08746D6BEAFC7519403DE0" bold="true">J</emphasis>
pleopod 5 exopod.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EAMBG" type="etymology">
<paragraph id="CF2D9311EF973D057EEA959143E16D45">
<heading id="94F1FA72BBB0BC66D484D1ECF76A0A36" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="ABE6BEF7BBECB2CBCBD4411B1DEA6650">
Latin 
<taxonomicName id="78F383D7B1CB362AAB30A156D9A7666E" authorityName="Wang &amp; Yao &amp; Jiang &amp; Li" authorityYear="2024" class="Malacostraca" family="Agnaridae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="formosus">
“ 
<emphasis id="917BD266B8863DA0D534AB5D384B9540" italics="true">formosus</emphasis>
</taxonomicName>
” = beautiful. The new species name is an allusion to beautiful yellowish blue muscle spots on the dorsum of the living individuals.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQMBG" type="remarks">
<paragraph id="246E803C7F5C9DB769EF77E68E7270EC">
<heading id="AE198AAE8ADADBCB24176B00F427CA50" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="708F26959A159E067E9C75D238F22A46">
The new species is not strictly consistent with the generic characters of 
<taxonomicName id="49636394226A7B924BB078971B617EB5" authorityName="Kwon" authorityYear="1993" class="Malacostraca" family="Agnaridae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="genus">
<emphasis id="E04753DCC8A8B9F4F29A671798CD6AF0" italics="true">Lucasioides</emphasis>
</taxonomicName>
because of the noduli laterales on pereonites 1-4 shifted from lateral margins than those of pereonites 5-7 instead of the noduli laterales on pereonites 2-4 much farther from lateral margins than those of pereonites 1, 5-7. It is difficult to give a ready answer for this dilemma based on the morphological evidence alone. In the phylogenetic analyses, the relationship between this species and the other 
<taxonomicName id="D05A47CCE95C155BA9CC9501534063D9">
<emphasis id="F5A65D1778C99505F9F2E0F13B8A3E50" italics="true">Lucasioides</emphasis>
species
</taxonomicName>
is well-supported (Fig. 
<figureCitation id="332AAE5E4CF0CD05E52130300BB86AE8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bayesian Consensus tree based on four-gene (COI + 18 S + 28 S + NaK) sequences. Asterisks and circles represent high node support with Bayesian posterior probabilities &gt; 0.95 and maximum bootstrap values &gt; 95. Coloured bars represent hypothesised species groupings." figureDoi="10.3897/asp.82.e113041.figure5" httpUri="https://binary.pensoft.net/fig/1083535">5</figureCitation>
). Hence, we assigned this species as a member of 
<taxonomicName id="7589CC1A2CB07829C8ED8CFF79E3C2C1" authorityName="Kwon" authorityYear="1993" class="Malacostraca" family="Agnaridae" genus="Lucasioides" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="genus">
<emphasis id="FB9846079BB6D6E9EC6A822832F1E141" italics="true">Lucasioides</emphasis>
</taxonomicName>
by integrating molecular and morphological evidence in this context.
</paragraph>
</subSubSection>
</treatment>
</document>