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<mods:title id="E56BA85E607FDED290C8461D0154927D">Reading between the lines: revealing cryptic species diversity and colour patterns in Hypselodoris nudibranchs (Mollusca: Heterobranchia: Chromodorididae)</mods:title>
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<emphasis id="FBC69B02FFCCFF93FC67FE48FB7B6BAB" bold="true" box="[972,1261,485,508]" italics="true" pageId="46" pageNumber="47">HYPSELODORIS SKYLERI</emphasis>
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<emphasis id="FBC69B02FFCCFF93FC82FD68FBAF688A" box="[809,1081,709,730]" italics="true" pageId="46" pageNumber="47">Hypselodoris maculosa?</emphasis>
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.
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sp. 5
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Gosliner
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: 265, third photograph.
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sp. 10
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Gosliner
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: 259, middle right photograph.
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</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FDC8FB03FE356F70" ID-GBIF-Occurrence="2235579040" collectingDate="2014-05-04" collectionCode="CASIZ" collectorName="Tingloy &amp; T. Gosliner" country="Philippines" county="Luzon" latitude="13.62978" location="Bonito" longLatPrecision="1" longitude="120.94738" municipality="Culebra" pageId="47" pageNumber="48" specimenCode="CASIZ 200649" specimenCount="1" stateProvince="Batangas" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FDC8FB03FC916E94" box="[611,775,1198,1220]" collectionCode="CASIZ" pageId="47" pageNumber="48">
CASIZ
<date id="BD0C61D0FFCDFF92FD1FFB03FC916E94" box="[692,775,1198,1220]" pageId="47" pageNumber="48" value="49-06-20">200649</date>
</specimenCode>
, one specimen,
<location id="CC6D11CBFFCDFF92FEF9FB61FE366EB2" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FEF9FB61FE366EB2" box="[338,416,1228,1250]" country="Philippines" county="Luzon" latitude="13.62978" longLatPrecision="1" longitude="120.94738" municipality="Culebra" name="Bonito" pageId="47" pageNumber="48" stateProvince="Batangas">Bonito</location>
(
<collectingMunicipality id="2969DD6AFFCDFF92FE1AFB61FD876EB2" box="[433,529,1228,1250]" pageId="47" pageNumber="48">Culebra</collectingMunicipality>
) Island,
<geoCoordinate id="AC8621D7FFCDFF92FDD6FB61FC916EB2" box="[637,775,1228,1250]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="1" value="13.62978">13.62978°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFCDFF92FF08FB46FEA26F50" box="[163,308,1259,1280]" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="1" value="120.94738">120.94738°E</geoCoordinate>
,
<collectorName id="644722C6FFCDFF92FEEBFB46FE036F51" box="[320,405,1259,1281]" pageId="47" pageNumber="48">Tingloy</collectorName>
,
<collectingRegion id="0B7689F2FFCDFF92FE0BFB41FD9B6F51" box="[416,525,1260,1281]" country="Philippines" name="Batangas" pageId="47" pageNumber="48">Batangas</collectingRegion>
,
<collectingCounty id="206C3F9CFFCDFF92FDB1FB41FDF56F51" box="[538,611,1260,1281]" pageId="47" pageNumber="48">Luzon</collectingCounty>
,
<collectingCountry id="B1A50780FFCDFF92FDC5FB46FD676F51" box="[622,753,1259,1281]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FD56FB46FE8E6F4F" pageId="47" pageNumber="48" value="2014-05-04">
<date id="BD0C61D0FFCDFF92FD56FB46FE8E6F4F" pageId="47" pageNumber="48" value="2014-05-04">4 May 2014</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FE8FFAA7FE356F70" box="[292,419,1290,1312]" pageId="47" pageNumber="48">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FE04FAA7FC916F0D" ID-GBIF-Occurrence="2235579097" collectingDate="2014-05-29" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" latitude="13.77931" location="Binacas" longLatPrecision="1" longitude="120.10959" municipality="Lubang Island" pageId="47" pageNumber="48" specimenCode="CASIZ 200552" specimenCount="1" stateProvince="Mindoro Occidental" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FE04FAA7FDCD6F70" box="[431,603,1290,1312]" collectionCode="CASIZ" pageId="47" pageNumber="48">
CASIZ
<date id="BD0C61D0FFCDFF92FDAEFAA7FDCD6F70" box="[517,603,1290,1312]" pageId="47" pageNumber="48" value="52-05-20">200552</date>
</specimenCode>
, one specimen, off
<location id="CC6D11CBFFCDFF92FF6CFA85FEB46F6E" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FF6CFA85FEB46F6E" box="[199,290,1320,1342]" country="Philippines" latitude="13.77931" longLatPrecision="1" longitude="120.10959" municipality="Lubang Island" name="Binacas" pageId="47" pageNumber="48" stateProvince="Mindoro Occidental">Binacas</location>
,
<geoCoordinate id="AC8621D7FFCDFF92FE84FA84FE256F6E" box="[303,435,1321,1342]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="1" value="13.77931">13.77931°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFCDFF92FE6BFA85FDC76F6E" box="[448,593,1320,1342]" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="1" value="120.10959">120.10959°E</geoCoordinate>
,
<collectingMunicipality id="2969DD6AFFCDFF92FDF6FA85FC906F6E" box="[605,774,1320,1342]" pageId="47" pageNumber="48">Lubang Island</collectingMunicipality>
,
<collectingRegion id="0B7689F2FFCDFF92FF08FAEAFEEE6F0D" box="[163,376,1351,1373]" country="Philippines" name="Mindoro Occidental" pageId="47" pageNumber="48">Mindoro Occidental</collectingRegion>
,
<collectingCountry id="B1A50780FFCDFF92FE2BFAEAFE6A6F0D" box="[384,508,1351,1373]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FDAFFAEAFD1B6F0C" box="[516,653,1351,1373]" pageId="47" pageNumber="48" value="2014-05-29">
<date id="BD0C61D0FFCDFF92FDAFFAEAFD1B6F0C" box="[516,653,1351,1373]" pageId="47" pageNumber="48" value="2014-05-29">29 May 2014</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FD3EFAE5FC916F0D" box="[661,775,1351,1373]" pageId="47" pageNumber="48">T.Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FF08FACBFEB06FE9" ID-GBIF-Occurrence="2235579076" collectingDate="2016-04-10" collectionCode="CASIZ" collectorName="T. Gosliner" country="Philippines" latitude="9.0502" location="Tubod" longLatPrecision="7" longitude="123.5144" municipality="Island" pageId="47" pageNumber="48" specimenCode="CASIZ 217258" specimenCount="1" stateProvince="Siquijor" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FF08FACBFED66F2C" box="[163,320,1382,1404]" collectionCode="CASIZ" pageId="47" pageNumber="48">CASIZ 217258</specimenCode>
, one specimen, south side
<location id="CC6D11CBFFCDFF92FDF8FACBFD0F6F2C" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FDF8FACBFD0F6F2C" box="[595,665,1382,1404]" country="Philippines" latitude="9.0502" longLatPrecision="7" longitude="123.5144" municipality="Island" name="Tubod" pageId="47" pageNumber="48" stateProvince="Siquijor">Tubod</location>
,
<geoCoordinate id="AC8621D7FFCDFF92FD09FACBFC916F2C" box="[674,775,1382,1404]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="5" value="9.0502">9.0502°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFCDFF92FF08FA29FEB26FCA" box="[163,292,1412,1434]" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="5" value="123.5144">123.5144°E</geoCoordinate>
,
<collectingRegion id="0B7689F2FFCDFF92FE84FA29FE116FCA" box="[303,391,1412,1434]" country="Philippines" name="Siquijor" pageId="47" pageNumber="48">Siquijor</collectingRegion>
<collectingMunicipality id="2969DD6AFFCDFF92FE24FA29FE416FCA" box="[399,471,1412,1434]" pageId="47" pageNumber="48">Island</collectingMunicipality>
,
<collectingCountry id="B1A50780FFCDFF92FE49FA29FDF46FCA" box="[482,610,1412,1434]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FDC6FA28FC916FCA" box="[621,775,1412,1434]" pageId="47" pageNumber="48" value="2016-04-10">
<date id="BD0C61D0FFCDFF92FDC6FA28FC916FCA" box="[621,775,1412,1434]" pageId="47" pageNumber="48" value="2016-04-10">10 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FF08FA09FEB06FE9" box="[163,294,1443,1465]" pageId="47" pageNumber="48">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FE98FA0EFDB96FA6" ID-GBIF-Occurrence="2235579036" collectingDate="2016-04-05" collectionCode="CASIZ" collectorName="Siaton &amp; T. Gosliner" country="Philippines" latitude="9.1374" location="Tambobo Bay" longLatPrecision="7" longitude="123.1149" pageId="47" pageNumber="48" specimenCode="CASIZ 217424" specimenCount="1" stateProvince="Negros Oriental" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FE98FA0EFE756FE8" box="[307,483,1443,1464]" collectionCode="CASIZ" pageId="47" pageNumber="48">CASIZ 217424</specimenCode>
, one specimen,
<location id="CC6D11CBFFCDFF92FD0BFA0EFF466F87" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FD0BFA0EFF466F87" country="Philippines" latitude="9.1374" longLatPrecision="7" longitude="123.1149" name="Tambobo Bay" pageId="47" pageNumber="48" stateProvince="Negros Oriental">Tambobo Bay</location>
,
<geoCoordinate id="AC8621D7FFCDFF92FF77FA6FFEDC6F87" box="[220,330,1474,1495]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="5" value="9.1374">9.1374°N</geoCoordinate>
,
<geoCoordinate id="AC8621D7FFCDFF92FEFDFA6FFE486F87" box="[342,478,1474,1495]" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="5" value="123.1149">123.1149°E</geoCoordinate>
,
<collectorName id="644722C6FFCDFF92FE40FA6FFDAF6F88" box="[491,569,1474,1496]" pageId="47" pageNumber="48">Siaton</collectorName>
,
<collectingRegion id="0B7689F2FFCDFF92FDEDFA6FFC916F88" box="[582,775,1474,1496]" country="Philippines" name="Negros Oriental" pageId="47" pageNumber="48">Negros Oriental</collectingRegion>
,
<collectingCountry id="B1A50780FFCDFF92FF08FA4DFE896FA6" box="[163,287,1504,1526]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FE81FA4DFE276FA6" box="[298,433,1504,1526]" pageId="47" pageNumber="48" value="2016-04-05">
<date id="BD0C61D0FFCDFF92FE81FA4DFE276FA6" box="[298,433,1504,1526]" pageId="47" pageNumber="48" value="2016-04-05">5 April 2016</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FE11FA4CFDB96FA6" box="[442,559,1504,1526]" pageId="47" pageNumber="48">T. Gosliner</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FD92FA4CFDB06C64" ID-GBIF-Occurrence="2235579095" collectingDate="1983-04-21" collectionCode="CASIZ" collectorName="S. Johnson" country="Marshall Islands" location="Munjor Pinnacle" pageId="47" pageNumber="48" specimenCode="CASIZ 120921" specimenCount="1" stateProvince="Enewetak Atoll" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FD92FA4CFD4F6FA6" box="[569,729,1505,1526]" collectionCode="CASIZ" pageId="47" pageNumber="48">
CASIZ
<date id="BD0C61D0FFCDFF92FD22FA4CFD4F6FA6" box="[649,729,1505,1526]" pageId="47" pageNumber="48" value="21-12-09">120921</date>
</specimenCode>
, one specimen,
<location id="CC6D11CBFFCDFF92FEB1FA52FE4F6C45" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FEB1FA52FE4F6C45" box="[282,473,1535,1557]" country="Marshall Islands" name="Munjor Pinnacle" pageId="47" pageNumber="48" stateProvince="Enewetak Atoll">Munjor Pinnacle</location>
,
<collectingRegion id="0B7689F2FFCDFF92FE4EFA52FD016C45" box="[485,663,1535,1557]" inferredBySuffix="true" pageId="47" pageNumber="48">Enewetak Atoll</collectingRegion>
,
<collectingCountry id="B1A50780FFCDFF92FD08FA52FF616C64" name="Marshall Islands" pageId="47" pageNumber="48">Marshall Islands</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FEA9F9B3FE086C63" box="[258,414,1566,1588]" pageId="47" pageNumber="48" value="1983-04-21">
<date id="BD0C61D0FFCDFF92FEA9F9B3FE086C63" box="[258,414,1566,1588]" pageId="47" pageNumber="48" value="1983-04-21">21 April 1983</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FE01F9B3FDB06C64" box="[426,550,1566,1588]" pageId="47" pageNumber="48">S. Johnson</collectorName>
</materialsCitation>
.
<materialsCitation id="79DA4D4DFFCDFF92FD9AF9B3FDE46C21" ID-GBIF-Occurrence="2235579050" collectingDate="2013-07-22" collectionCode="CASIZ" collectorName="S. Johnson" country="Marshall Islands" location="Pinnacle" pageId="47" pageNumber="48" specimenCode="CASIZ 194004" specimenCount="1" stateProvince="Kwajalein" typeStatus="paratype">
<specimenCode id="9914EF6BFFCDFF92FD9AF9B3FD416C63" box="[561,727,1566,1587]" collectionCode="CASIZ" pageId="47" pageNumber="48">CASIZ 194004</specimenCode>
, one specimen, under rock, K9
<location id="CC6D11CBFFCDFF92FE73F991FDA96C02" LSID="urn:lsid:plazi:treatment:411BF606FFCCFF8DFC67FE48FAFB6E67:CC6D11CBFFCDFF92FE73F991FDA96C02" box="[472,575,1596,1618]" country="Marshall Islands" name="Pinnacle" pageId="47" pageNumber="48" stateProvince="Kwajalein">Pinnacle</location>
,
<collectingRegion id="0B7689F2FFCDFF92FDE6F991FC906C02" box="[589,774,1596,1618]" country="Marshall Islands" name="Kwajalein" pageId="47" pageNumber="48">Kwajalein Atoll</collectingRegion>
,
<collectingCountry id="B1A50780FFCDFF92FF08F9F6FECA6C21" box="[163,348,1627,1649]" name="Marshall Islands" pageId="47" pageNumber="48">Marshall Islands</collectingCountry>
,
<collectingDate id="AD489838FFCDFF92FECDF9F6FE676C20" box="[358,497,1627,1649]" pageId="47" pageNumber="48" value="2013-07-22">
<date id="BD0C61D0FFCDFF92FECDF9F6FE676C20" box="[358,497,1627,1649]" pageId="47" pageNumber="48" value="2013-07-22">22 July 2013</date>
</collectingDate>
,
<collectorName id="644722C6FFCDFF92FE50F9F6FDE46C21" box="[507,626,1627,1649]" pageId="47" pageNumber="48">S. Johnson</collectorName>
</materialsCitation>
.
</paragraph>
<caption id="9DCD1798FFCCFF93FF3AF8B7FDBA6D3B" httpUri="https://zenodo.org/record/2632840/files/figure.png" pageId="46" pageNumber="47" startId="46.[145,223,1818,1840]" targetBox="[145,1425,1158,1778]" targetPageId="46">
<paragraph id="C90D4710FFCCFF93FF3AF8B7FDBA6D3B" blockId="46.[145,1425,1818,1899]" pageId="46" pageNumber="47">
<emphasis id="FBC69B02FFCCFF93FF3AF8B7FE936D7F" bold="true" box="[145,261,1818,1840]" pageId="46" pageNumber="47">Figure 27.</emphasis>
<taxonomicName id="0EB23C93FFCCFF93FEBBF8B7FD5C6D60" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[272,714,1818,1840]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="skyleri" status="sp. nov.">
<emphasis id="FBC69B02FFCCFF93FEBBF8B7FE6F6D60" bold="true" box="[272,505,1818,1840]" italics="true" pageId="46" pageNumber="47">Hypselodoris skyleri</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCCFF93FD64F8B6FCB26D60" box="[719,804,1819,1840]" pageId="46" pageNumber="47" rank="species">
<emphasis id="FBC69B02FFCCFF93FD64F8B6FCB26D60" bold="true" box="[719,804,1819,1840]" pageId="46" pageNumber="47">sp. nov.</emphasis>
</taxonomicNameLabel>
, living animals. A, paratype, CASIZ 217258, Tubod, Siquijor, Philippines. B, paratype, CASIZ 217424, Siaton, Negros Oriental, Philippines. C, holotype, NMP 0 41284. D, paratype, CASIZ 177305, Tingloy, Batangas, Philippines.
</paragraph>
</caption>
<caption id="9DCD1798FFCDFF92FF08FBFBFCF36ED9" httpUri="https://zenodo.org/record/2632844/files/figure.png" pageId="47" pageNumber="48" startId="47.[163,241,1110,1132]" targetBox="[227,1379,195,1069]" targetPageId="47">
<paragraph id="C90D4710FFCDFF92FF08FBFBFCF36ED9" blockId="47.[163,1444,1110,1161]" pageId="47" pageNumber="48">
<emphasis id="FBC69B02FFCDFF92FF08FBFBFE8E6E3B" bold="true" box="[163,280,1110,1132]" pageId="47" pageNumber="48">Figure 28.</emphasis>
<taxonomicName id="0EB23C93FFCDFF92FE88FBFBFD756E3C" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[291,739,1110,1132]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="skyleri" status="sp. nov.">
<emphasis id="FBC69B02FFCDFF92FE88FBFBFD986E3C" bold="true" box="[291,526,1110,1132]" italics="true" pageId="47" pageNumber="48">Hypselodoris skyleri</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFCDFF92FD42FBFBFCA96E3B" box="[745,831,1110,1131]" pageId="47" pageNumber="48" rank="species">
<emphasis id="FBC69B02FFCDFF92FD42FBFBFCA96E3B" bold="true" box="[745,831,1110,1131]" pageId="47" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature, paratype, CASIZ 177305. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth.
</paragraph>
</caption>
<paragraph id="C90D4710FFCDFF92FF08F934FEAB6CE0" blockId="47.[163,779,1688,1773]" box="[163,317,1688,1712]" pageId="47" pageNumber="48">
<heading id="9245F07CFFCDFF92FF08F934FEAB6CE0" box="[163,317,1688,1712]" fontSize="10" level="4" pageId="47" pageNumber="48" reason="1">
<emphasis id="FBC69B02FFCDFF92FF08F934FEAB6CE0" box="[163,317,1688,1712]" italics="true" pageId="47" pageNumber="48">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCDFF92FF08F914FEBD6CBD" blockId="47.[163,779,1688,1773]" pageId="47" pageNumber="48">
Sea Pen, Tingloy, Maricaban Island,
<collectingRegion id="0B7689F2FFCDFF92FD27F914FC906C9E" box="[652,774,1721,1742]" country="Philippines" name="Batangas" pageId="47" pageNumber="48">Batangas</collectingRegion>
,
<collectingCountry id="B1A50780FFCDFF92FF08F97AFEBE6CBD" box="[163,296,1751,1773]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCDFF92FF08F8B9FDB66D39" pageId="47" pageNumber="48" type="distribution">
<paragraph id="C90D4710FFCDFF92FF08F8B9FE766D7C" blockId="47.[163,779,1812,1897]" box="[163,480,1812,1836]" pageId="47" pageNumber="48">
<heading id="9245F07CFFCDFF92FF08F8B9FE766D7C" box="[163,480,1812,1836]" fontSize="10" level="4" pageId="47" pageNumber="48" reason="1">
<emphasis id="FBC69B02FFCDFF92FF08F8B9FE766D7C" box="[163,480,1812,1836]" italics="true" pageId="47" pageNumber="48">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCDFF92FF08F898FDB66D39" blockId="47.[163,779,1812,1897]" pageId="47" pageNumber="48">
Known from the
<collectingCountry id="B1A50780FFCDFF92FED4F899FD846D1A" box="[383,530,1844,1866]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingCountry id="B1A50780FFCDFF92FD89F899FD096D1A" box="[546,671,1844,1866]" name="Indonesia" pageId="47" pageNumber="48">Indonesia</collectingCountry>
and the
<collectingCountry id="B1A50780FFCDFF92FF08F8FEFEF36D39" box="[163,357,1875,1897]" name="Marshall Islands" pageId="47" pageNumber="48">Marshall Islands</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCDFF92FC90FB00FA176F2D" pageId="47" pageNumber="48" type="etymology">
<paragraph id="C90D4710FFCDFF92FC90FB00FC2A6E95" blockId="47.[827,1443,1197,1405]" box="[827,956,1197,1221]" pageId="47" pageNumber="48">
<heading id="9245F07CFFCDFF92FC90FB00FC2A6E95" box="[827,956,1197,1221]" fontSize="10" level="4" pageId="47" pageNumber="48" reason="1">
<emphasis id="FBC69B02FFCDFF92FC90FB00FC2A6E95" box="[827,956,1197,1221]" italics="true" pageId="47" pageNumber="48">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCDFF92FC90FB63FA176F2D" blockId="47.[827,1443,1197,1405]" pageId="47" pageNumber="48">This species is named for author Rebecca Johnsons son, Skyler (Sky) Rodgers. The beautiful bright white spots on the mantle of this species are reminiscent of bright stars in the sky on a clear night. Sky, thanks for your patience, kindness and thoughtfulness. May you always stay curious about the stars and our place in the universe.</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFCDFF8DFC90FA08FF416C8E" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="description">
<paragraph id="C90D4710FFCDFF92FC90FA08FC516FEC" blockId="47.[827,1444,1445,1897]" box="[827,967,1445,1468]" pageId="47" pageNumber="48">
<heading id="9245F07CFFCDFF92FC90FA08FC516FEC" box="[827,967,1445,1468]" fontSize="10" level="4" pageId="47" pageNumber="48" reason="1">
<emphasis id="FBC69B02FFCDFF92FC90FA08FC516FEC" box="[827,967,1445,1468]" italics="true" pageId="47" pageNumber="48">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFCDFF8DFC90FA69FEF46AAB" blockId="47.[827,1444,1445,1897]" lastBlockId="48.[145,760,198,251]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="FBC69B02FFCDFF92FC90FA69FBAF6F89" box="[827,1081,1476,1497]" italics="true" pageId="47" pageNumber="48">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFCDFF92FAB9FA69FAF56F8A" box="[1298,1379,1476,1498]" captionStart="Figure 27" captionStartId="46.[145,223,1818,1840]" captionTargetBox="[145,1425,1158,1778]" captionTargetId="figure@46.[145,1425,1158,1778]" captionTargetPageId="46" captionText="Figure 27. Hypselodoris skyleri Gosliner &amp; Johnson sp. nov., living animals. A, paratype, CASIZ 217258, Tubod, Siquijor, Philippines. B, paratype, CASIZ 217424, Siaton, Negros Oriental, Philippines. C, holotype, NMP 041284. D, paratype, CASIZ 177305, Tingloy, Batangas, Philippines." httpUri="https://zenodo.org/record/2632840/files/figure.png" pageId="47" pageNumber="48">Fig. 27</figureCitation>
AD) small, reaching
<quantity id="0E4AEAF5FFCDFF92FC5DFA4EFBD16FA9" box="[1014,1095,1507,1529]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="47" pageNumber="48" unit="mm" value="15.0">15 mm</quantity>
in length. Body long, slender, translucent pink to peach. Fine parallel or intersecting brown longitudinal lines on notum sides of body and foot. Small, scattered opaque white spot situated on small tubercles on notum sides of body and foot. Scattered bluepurple spots found around margin of notum and foot; pigment more dense in centre of spot. Seven to eight unipinnate gill branches having a translucent white base and inner margin and bright redorange pigment on outer surface of rachis and some gill filaments. Inner surface with opaque white. Bulb of perfoliate rhinophores opaque white with three red transverse bands and bearing about seven to ten densely arranged lamellae. Base of rhinophores translucent white.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFF3AFE89FECC6BE4" blockId="48.[145,761,292,436]" pageId="48" pageNumber="49">
<emphasis id="FBC69B02FFD2FF8DFF3AFE89FEAA6B69" box="[145,316,292,313]" italics="true" pageId="48" pageNumber="49">Mantle glands</emphasis>
: Subcutaneous mantle glands simple and rounded in shape (
<figureCitation id="51895B95FFD2FF8DFE12FEEFFD806B07" box="[441,534,322,344]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="48" pageNumber="49">Fig. 2O</figureCitation>
). Glands situated around entire margin of body in six specimens
<quantity id="0E4AEAF5FFD2FF8DFD35FECCFD6F6B26" box="[670,761,353,374]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="48" pageNumber="49" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">48 mm</quantity>
preserved length. Anterior and posterior glands larger than lateral ones.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFF3AFE70FEE36E4A" blockId="48.[145,761,477,1051]" pageId="48" pageNumber="49">
<emphasis id="FBC69B02FFD2FF8DFF3AFE70FECA6BA3" box="[145,348,477,499]" italics="true" pageId="48" pageNumber="49">Buccal armature</emphasis>
: Muscular portion of buccal mass about equal in length to oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFD2FF8DFD20FD94FD7D681E" box="[651,747,569,591]" captionStart="Figure 28" captionStartId="47.[163,241,1110,1132]" captionTargetBox="[227,1379,195,1069]" captionTargetId="figure@47.[227,1379,195,1069]" captionTargetPageId="47" captionText="Figure 28. Hypselodoris skyleri Gosliner &amp; Johnson sp. nov., buccal armature, paratype, CASIZ 177305. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632844/files/figure.png" pageId="48" pageNumber="49">Fig. 28A</figureCitation>
). Rodlets sharply angled, acutely pointed with one to three apices and posterior lateral extensions. Radular formula of
<typeStatus id="1609F9B2FFD2FF8DFEB2FD38FE1068FA" box="[281,390,661,682]" pageId="48" pageNumber="49" type="paratype">paratype</typeStatus>
(CASIZ 177305) 53 × 38.0.38. Rachidian row of teeth absent (
<figureCitation id="51895B95FFD2FF8DFDAEFD19FDFE6899" box="[517,616,692,714]" captionStart="Figure 28" captionStartId="47.[163,241,1110,1132]" captionTargetBox="[227,1379,195,1069]" captionTargetId="figure@47.[227,1379,195,1069]" captionTargetPageId="47" captionText="Figure 28. Hypselodoris skyleri Gosliner &amp; Johnson sp. nov., buccal armature, paratype, CASIZ 177305. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632844/files/figure.png" pageId="48" pageNumber="49">Fig. 28B</figureCitation>
). Innermost lateral teeth having two or three large, triangular denticles on inner side of bifid primary cusp, with another two or three outer denticles. Next several laterals lacking inner triangular denticle but possessing two or three denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFD2FF8DFDCFFCC1FD5F69D1" box="[612,713,876,898]" captionStart="Figure 28" captionStartId="47.[163,241,1110,1132]" captionTargetBox="[227,1379,195,1069]" captionTargetId="figure@47.[227,1379,195,1069]" captionTargetPageId="47" captionText="Figure 28. Hypselodoris skyleri Gosliner &amp; Johnson sp. nov., buccal armature, paratype, CASIZ 177305. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632844/files/figure.png" pageId="48" pageNumber="49">Fig. 28C</figureCitation>
) all lacking inner denticles but having three or four sharply pointed, triangular outer denticles. Outermost teeth having a narrower base and somewhat shorter tooth shape, with five or six rounded to pointed outer denticles (
<figureCitation id="51895B95FFD2FF8DFEAEFBA8FEF36E4A" box="[261,357,1029,1051]" captionStart="Figure 28" captionStartId="47.[163,241,1110,1132]" captionTargetBox="[227,1379,195,1069]" captionTargetId="figure@47.[227,1379,195,1069]" captionTargetPageId="47" captionText="Figure 28. Hypselodoris skyleri Gosliner &amp; Johnson sp. nov., buccal armature, paratype, CASIZ 177305. A, jaw rodlets. B, inner lateral teeth. C, middle lateral teeth. D, outer lateral teeth." httpUri="https://zenodo.org/record/2632844/files/figure.png" pageId="48" pageNumber="49">Fig. 28D</figureCitation>
).
</paragraph>
<paragraph id="C90D4710FFD2FF8DFF3AFBE9FF416C8E" blockId="48.[145,761,1092,1758]" pageId="48" pageNumber="49">
<emphasis id="FBC69B02FFD2FF8DFF3AFBE9FEE26E09" box="[145,372,1092,1113]" italics="true" pageId="48" pageNumber="49">Reproductive system</emphasis>
: Reproductive organs fully mature in one specimen examined (CASIZ 177305;
<figureCitation id="51895B95FFD2FF8DFD23FBCFFD7D6E27" box="[648,747,1122,1144]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="48" pageNumber="49">Fig. 13K</figureCitation>
). Ampulla thick, convoluted, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens convoluted, curved and thick and narrowing slightly as it transitions into long, convoluted, muscular ejaculatory portion. Ejaculatory portion widening into much enlarged penial bulb. Penial papilla distinctly curved with broad base, devoid of penial hooks. Penial bulb adjacent to straight, slightly narrow, widening into penial sac. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina widening proximally, leading to minute receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Receptaculum situated near base of bursa. Moderately short, narrow uterine duct emerging from vagina close to base of bursa and female gland mass, near albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFD2FF8DFF3AF8A9FAFB6E67" pageId="48" pageNumber="49" type="discussion">
<paragraph id="C90D4710FFD2FF8DFF3AF8A9FF686D4C" blockId="48.[145,761,1796,1882]" box="[145,254,1796,1820]" pageId="48" pageNumber="49">
<heading id="9245F07CFFD2FF8DFF3AF8A9FF686D4C" box="[145,254,1796,1820]" fontSize="10" level="4" pageId="48" pageNumber="49" reason="1">
<emphasis id="FBC69B02FFD2FF8DFF3AF8A9FF686D4C" box="[145,254,1796,1820]" italics="true" pageId="48" pageNumber="49">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFD2FF8DFF3AF888FA1C68D9" blockId="48.[145,761,1796,1882]" lastBlockId="48.[809,1426,198,1079]" pageId="48" pageNumber="49">
<taxonomicName id="0EB23C93FFD2FF8DFF3AF888FE1D6D6A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[145,395,1829,1850]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFF3AF888FE1D6D6A" box="[145,395,1829,1850]" italics="true" pageId="48" pageNumber="49">Hypselodoris skyleri</emphasis>
</taxonomicName>
is a member of a large clade that includes
<taxonomicName id="0EB23C93FFD2FF8DFE9EF8E9FE406D09" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[309,470,1860,1881]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="katherinae" status="sp. nov.">
<emphasis id="FBC69B02FFD2FF8DFE9EF8E9FE406D09" box="[309,470,1860,1881]" italics="true" pageId="48" pageNumber="49">H. katherinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD2FF8DFE74F8E9FDAE6D09" box="[479,568,1860,1881]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="0EB23C93FFD2FF8DFDEFF8E9FD466D09" authority="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner, 2018" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[580,720,1860,1881]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="paradisa" status="sp. nov.">
<emphasis id="FBC69B02FFD2FF8DFDEFF8E9FD466D09" box="[580,720,1860,1881]" italics="true" pageId="48" pageNumber="49">H. paradisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFD2FF8DFD72F8E9FCCC6A8C" box="[729,858,199,1881]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="FBC69B02FFD2FF8DFCCDFF6AFC6F6A8C" box="[870,1017,199,220]" italics="true" pageId="48" pageNumber="49">H. maritima</emphasis>
,
<taxonomicName id="0EB23C93FFD2FF8DFBA3FF6AFB026A8B" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1032,1172,198,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="rudmani">
<emphasis id="FBC69B02FFD2FF8DFBA3FF6AFB026A8B" box="[1032,1172,198,220]" italics="true" pageId="48" pageNumber="49">H. rudmani</emphasis>
</taxonomicName>
and
<emphasis id="FBC69B02FFD2FF8DFB7BFF6AFACF6A8B" box="[1232,1369,198,220]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="0EB23C93FFD2FF8DFB7BFF6AFAC36A8B" authorityName="Gosliner &amp; Johnson" authorityYear="1999" box="[1232,1365,198,220]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="bertschi">H. bertschi</taxonomicName>
.
</emphasis>
It is sister to
<taxonomicName id="0EB23C93FFD2FF8DFC38FF4BFBA16AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[915,1079,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFD2FF8DFC38FF4BFBA16AAA" box="[915,1079,229,251]" italics="true" pageId="48" pageNumber="49">H. katherinae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFBDFFF4BFA946AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1140,1282,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFD2FF8DFBDFFF4BFA946AAA" box="[1140,1282,229,251]" italics="true" pageId="48" pageNumber="49">H. paradisa</emphasis>
</taxonomicName>
. The ABGD analysis indicates that
<taxonomicName id="0EB23C93FFD2FF8DFBEFFEA9FB2E6B49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1092,1208,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFBEFFEA9FB2E6B49" box="[1092,1208,260,281]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
is clearly distinct from
<taxonomicName id="0EB23C93FFD2FF8DFCC5FE8EFB886B67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[878,1054,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFD2FF8DFCC5FE8EFB886B67" box="[878,1054,290,312]" italics="true" pageId="48" pageNumber="49">H. katherinae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFBC8FE8EFB6A6B67" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1123,1276,290,312]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFD2FF8DFBC8FE8EFB6A6B67" box="[1123,1276,290,312]" italics="true" pageId="48" pageNumber="49">H. paradisa</emphasis>
</taxonomicName>
. A detailed comparison of these species is found in the remarks section of
<taxonomicName id="0EB23C93FFD2FF8DFC0BFECDFBD76B25" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[928,1089,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="katherinae">
<emphasis id="FBC69B02FFD2FF8DFC0BFECDFBD76B25" box="[928,1089,352,373]" italics="true" pageId="48" pageNumber="49">H. katherinae</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFBD7FECDFA9E6B25" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1148,1288,352,373]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="paradisa">
<emphasis id="FBC69B02FFD2FF8DFBD7FECDFA9E6B25" box="[1148,1288,352,373]" italics="true" pageId="48" pageNumber="49">H. paradisa</emphasis>
</taxonomicName>
. The colour pattern of
<taxonomicName id="0EB23C93FFD2FF8DFC08FED2FB856BC3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[931,1043,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFC08FED2FB856BC3" box="[931,1043,382,404]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="0EB23C93FFD2FF8DFAAAFED2FA076BC3" authorityName="? Rudman" authorityYear="2003" box="[1281,1425,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFAAAFED2FA076BC3" box="[1281,1425,382,404]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFCF5FE33FC7E6BE2" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[862,1000,413,435]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFCF5FE33FC7E6BE2" box="[862,1000,413,435]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
but differs in having brown rather than opaque white lines. In
<taxonomicName id="0EB23C93FFD2FF8DFB2EFE11FB6D6B81" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1157,1275,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFB2EFE11FB6D6B81" box="[1157,1275,444,465]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
, the opaque white spots are scattered over the surface of the body and elevated on small tubercles. In
<taxonomicName id="0EB23C93FFD2FF8DFB67FE57FACB685E" authorityName="? Rudman" authorityYear="2003" box="[1228,1373,505,527]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFB67FE57FACB685E" box="[1228,1373,505,527]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFC82FDB5FC3C687D" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[809,938,536,557]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFC82FDB5FC3C687D" box="[809,938,536,557]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
, the spots are found only on the margins of the body and are flush with the body surface. In both
<taxonomicName id="0EB23C93FFD2FF8DFC82FDFBFC03683A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,917,597,619]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFC82FDFBFC03683A" box="[809,917,597,619]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFC65FDFBFBC6683A" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[974,1104,597,619]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFC65FDFBFBC6683A" box="[974,1104,597,619]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
there are three rhinophoral red rings, whereas there are only two in
<taxonomicName id="0EB23C93FFD2FF8DFB5DFDD9FA1568D9" authorityName="? Rudman" authorityYear="2003" box="[1270,1411,628,649]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFB5DFDD9FA1568D9" box="[1270,1411,628,649]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFD2FF8DFCEAFD3FFAFB6E67" blockId="48.[809,1426,198,1079]" pageId="48" pageNumber="49">
Both
<taxonomicName id="0EB23C93FFD2FF8DFCD6FD3EFC6B68F7" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[893,1021,658,680]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFCD6FD3EFC6B68F7" box="[893,1021,658,680]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFD2FF8DFBA0FD3FFBCD68F7" box="[1035,1115,658,680]" captionStart="Figure 2" captionStartId="8.[144,228,1783,1805]" captionTargetBox="[146,1425,197,1740]" captionTargetId="figure@8.[144,1427,195,1742]" captionTargetPageId="8" captionText="Figure 2. Mantle glands. A, Hypselodoris alburtuqali Gosliner &amp; Johnson sp. nov., CASIZ 192295, holotype. B, Hypselodoris cerisae Gosliner &amp; Johnson sp. nov., holotype, CASIZ 178350. C, Hypselodoris confetti Gosliner &amp; Johnson sp. nov., CASIZ 191070. D, Hypselodoris decorata (Risbec, 1928), CASIZ 184316. E, Hypselodoris ghardaqana (Gohar &amp; Abu-Ela, 1957), CASIZ 192282. F, Hypselodoris iba Gosliner &amp; Johnson sp. nov., holotype, NMP 041279." httpUri="https://zenodo.org/record/2632775/files/figure.png" pageId="48" pageNumber="49">Fig. 2D</figureCitation>
) and
<taxonomicName id="0EB23C93FFD2FF8DFB33FD3EFAB568F7" authorityName="? Rudman" authorityYear="2003" box="[1176,1315,658,680]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFB33FD3EFAB568F7" box="[1176,1315,658,680]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD233AE1FFD2FF8DFA84FD3FFBDA6896" author="Gosliner T &amp; Johnson, R." journalOrPublisher="Zoological Journal of the Linnean Society" pageId="48" pageNumber="49" pagination="1 - 114" part="125" refId="ref42597" refString="Gosliner T, Johnson, R. 1999. Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species. Zoological Journal of the Linnean Society 125: 1 - 114." title="Phylogeny of Hypselodoris (Nudibranchia: Chromodorididae) with a review of the monophyletic clade of Indo-Pacific species, including descriptions of twelve new species" type="journal article" year="1999">Gosliner &amp; Johnson, 1999: fig. 29D</bibRefCitation>
) have mantle glands only at the anterior and posterior ends of the body, whereas
<taxonomicName id="0EB23C93FFD2FF8DFC82FD42FC356953" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[809,931,750,772]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFC82FD42FC356953" box="[809,931,750,772]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
also has many lateral glands. The jaw rodlets of
<taxonomicName id="0EB23C93FFD2FF8DFC37FCA3FB9A6972" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[924,1036,781,803]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFC37FCA3FB9A6972" box="[924,1036,781,803]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
have one to three cusps, whereas
<taxonomicName id="0EB23C93FFD2FF8DFC82FC81FC3D6911" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[809,939,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFC82FC81FC3D6911" box="[809,939,812,833]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFD2FF8DFC11FC81FB936911" box="[954,1029,812,834]" captionStart="Figure 10" captionStartId="21.[163,244,1584,1606]" captionTargetBox="[227,1379,195,1544]" captionTargetId="figure@21.[227,1379,195,1544]" captionTargetPageId="21" captionText="Figure 10. Hypselodoris decorata (Risbec, 1928). A, entire jaw, CASIZ 190661. B, jaw rodlets, CASIZ 191214. C, entire radula, CASIZ 190661. D, inner lateral teeth, CASIZ 190661. E, middle lateral teeth, CASIZ 190661. F, outer lateral teeth, CASIZ 190661." httpUri="https://zenodo.org/record/2632791/files/figure.png" pageId="48" pageNumber="49">Fig. 10</figureCitation>
) and
<taxonomicName id="0EB23C93FFD2FF8DFBEDFC81FB456911" authorityName="? Rudman" authorityYear="2003" box="[1094,1235,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFBEDFC81FB456911" box="[1094,1235,812,833]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFD2FF8DFB4AFC81FABB6911" box="[1249,1325,812,834]" captionStart="Figure 11" captionStartId="22.[145,225,1849,1871]" captionTargetBox="[209,1361,487,1808]" captionTargetId="figure@22.[209,1361,487,1808]" captionTargetPageId="22" captionText="Figure 11. Hypselodoris maculosa (Pease, 1871), CASIZ 139545. A, entire jaw. B, jaw rodlets. C, entire radula. D, inner lateral teeth. E, middle lateral teeth. F, outer lateral teeth." httpUri="https://zenodo.org/record/2632793/files/figure.png" pageId="48" pageNumber="49">Fig. 11</figureCitation>
) possess a single cusp. The radulae of
<taxonomicName id="0EB23C93FFD2FF8DFB2AFCE6FB64690F" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1153,1266,842,864]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFB2AFCE6FB64690F" box="[1153,1266,842,864]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
,
<taxonomicName id="0EB23C93FFD2FF8DFAABFCE6FA04690F" authorityName="? Rudman" authorityYear="2003" box="[1280,1426,842,864]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFAABFCE6FA04690F" box="[1280,1426,842,864]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFCF7FCC7FC74692E" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[860,994,873,895]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFCF7FCC7FC74692E" box="[860,994,873,895]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
all have a similar formula and teeth of a similar shape. One notable difference is that the cusps and adjacent denticles on the inner and middle lateral teeth are more elongate in
<taxonomicName id="0EB23C93FFD2FF8DFB6AFC6BFAA4698A" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1217,1330,965,987]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFB6AFC6BFAA4698A" box="[1217,1330,965,987]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
than in
<taxonomicName id="0EB23C93FFD2FF8DFC82FC49FC2F69A9" authorityName="? Rudman" authorityYear="2003" box="[809,953,996,1017]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFC82FC49FC2F69A9" box="[809,953,996,1017]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFC5FFC49FBEF69A9" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[1012,1145,996,1017]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFC5FFC49FBEF69A9" box="[1012,1145,996,1017]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
. In
<taxonomicName id="0EB23C93FFD2FF8DFB03FC49FA8E69A9" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1192,1304,996,1017]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="skyleri">
<emphasis id="FBC69B02FFD2FF8DFB03FC49FA8E69A9" box="[1192,1304,996,1017]" italics="true" pageId="48" pageNumber="49">H. skyleri</emphasis>
</taxonomicName>
, the penis is proportionately wider than in
<taxonomicName id="0EB23C93FFD2FF8DFB69FBAEFACC6E47" authorityName="? Rudman" authorityYear="2003" box="[1218,1370,1026,1048]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="maculosa">
<emphasis id="FBC69B02FFD2FF8DFB69FBAEFACC6E47" box="[1218,1370,1026,1048]" italics="true" pageId="48" pageNumber="49">H. maculosa</emphasis>
</taxonomicName>
and
<taxonomicName id="0EB23C93FFD2FF8DFC82FB8FFC3D6E66" baseAuthorityName="Risbec" baseAuthorityYear="1928" box="[809,939,1057,1079]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="decorata">
<emphasis id="FBC69B02FFD2FF8DFC82FB8FFC3D6E66" box="[809,939,1057,1079]" italics="true" pageId="48" pageNumber="49">H. decorata</emphasis>
</taxonomicName>
and possesses a distinct penial papilla.
</paragraph>
</subSubSection>
</treatment>
</document>