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<mods:title id="E02075698D011CAA2D6180C014D57DFA">Reading between the lines: revealing cryptic species diversity and colour patterns in Hypselodoris nudibranchs (Mollusca: Heterobranchia: Chromodorididae)</mods:title>
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bullocki
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misidentification, not 
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<bibRefCitation id="AD233AE1FFC1FF9EFF08FA7FFEE06FB8" author="Collingwood C." box="[163,374,1490,1512]" journalOrPublisher="Transactions of the Linnean Society of London, Zoology" pageId="35" pageNumber="36" pagination="123 - 140" part="2" refId="ref41797" refString="Collingwood C. 1881. On some new species of nudibranchiate Mollusca from the eastern seas. Transactions of the Linnean Society of London, Zoology 2: 123 - 140." title="On some new species of nudibranchiate Mollusca from the eastern seas" type="journal article" year="1881">Collingwood, 1881</bibRefCitation>
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; 
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; 
<bibRefCitation id="AD233AE1FFC1FF9EFD99FA7FFD4A6FB8" author="Coleman N." box="[562,732,1490,1512]" journalOrPublisher="Springwood: Australia Neville Coleman's Underwater Geographic" pageId="35" pageNumber="36" refId="ref41724" refString="Coleman N. 2001. 1001 Nudibranchs: catalogue of Indo- Pacific sea slugs. Springwood: Australia Neville Coleman's Underwater Geographic." title="1001 Nudibranchs: catalogue of Indo- Pacific sea slugs" type="book" year="2001">Coleman, 2001</bibRefCitation>
: 79, lower right photograph; 
<bibRefCitation id="AD233AE1FFC1FF9EFE61FA5CFD066C56" author="Gaensslen RE" box="[458,656,1521,1543]" journalOrPublisher="Sydney: Australian Museum" pageId="35" pageNumber="36" publicationUrl="http://www.seaslugforum.net/find/20471" refId="ref42146" refString="Gaensslen RE 2007. Hypselodoris bullocki? from Papua New Guinea. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 20471" title="Hypselodoris bullocki? from Papua New Guinea" type="url" volumeTitle="Sea slug forum" year="2007">Gaensslen, 2007</bibRefCitation>
; 
<bibRefCitation id="AD233AE1FFC1FF9EFD34FA5CFF4C6C75" author="Rudman WB" journalOrPublisher="Sydney: Australian Museum" pageId="35" pageNumber="36" publicationUrl="http://www.seaslugforum.net/find/hypsbull" refId="ref45077" refString="Rudman WB. 2007. Hypselodoris bullocki - feeding record. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / hypsbull" title="Hypselodoris bullocki - feeding record" type="url" volumeTitle="Sea slug forum" year="2007">Rudman, 2007</bibRefCitation>
; 
<bibRefCitation id="AD233AE1FFC1FF9EFF4DF9BDFE0A6C75" author="Hanchard B." box="[230,412,1552,1574]" journalOrPublisher="Sydney: Australian Museum" pageId="35" pageNumber="36" publicationUrl="http://www.seaslugforum.net/find/22275" refId="ref42817" refString="Hanchard B. 2009. Blue Hypselodoris bullocki egg-laying. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 22275" title="Blue Hypselodoris bullocki egg-laying" type="url" volumeTitle="Sea slug forum" year="2009">Hanchard, 2009</bibRefCitation>
.
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sp. 7 
<bibRefCitation id="AD233AE1FFC1FF9EFE28F983FDE26C14" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." box="[387,628,1582,1604]" journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="35" pageNumber="36" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner 
<emphasis id="FBC69B02FFC1FF9EFE5AF982FDBA6C13" box="[497,556,1582,1604]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 2008
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: 268, second photograph from top.
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<emphasis id="FBC69B02FFC1FF9EFF08F9C1FEAF6CD1" box="[163,313,1644,1665]" italics="true" pageId="35" pageNumber="36">Hypselodoris</emphasis>
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sp. 16 
<bibRefCitation id="AD233AE1FFC1FF9EFE20F9C1FDE36CD2" author="Gosliner T &amp; Valdes A &amp; Behrens, DW" box="[395,629,1644,1666]" journalOrPublisher="Jacksonville: New World Press" pageId="35" pageNumber="36" refId="ref42644" refString="Gosliner T, Valdes A, Behrens, DW. 2015. Nudibranch and sea slug identification: Indo-Pacific. Jacksonville: New World Press." title="Nudibranch and sea slug identification: Indo-Pacific" type="book" year="2015">
Gosliner 
<emphasis id="FBC69B02FFC1FF9EFE5EF9C1FDB96CD1" box="[501,559,1644,1665]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 2015
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: 262, bottom left photograph.
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</paragraph>
<paragraph id="C90D4710FFC1FF9EFC90FF6AFBBC6F1B" blockId="35.[827,1443,198,1355]" pageId="35" pageNumber="36">
<materialsCitation id="79DA4D4DFFC1FF9EFC90FF6AFC306B68" ID-GBIF-Occurrence="2235579099" collectionCode="CASIZ" collectorName="R. Steene" elevation="9" location="Milne Bay" municipality="Milne Bay Province" pageId="35" pageNumber="36" specimenCode="CASIZ 185100" specimenCount="1" typeStatus="paratype">
<typeStatus id="1609F9B2FFC1FF9EFC90FF6AFC2D6A8C" box="[827,955,199,220]" pageId="35" pageNumber="36" type="paratype">
<emphasis id="FBC69B02FFC1FF9EFC90FF6AFC2D6A8C" box="[827,955,199,220]" italics="true" pageId="35" pageNumber="36">Paratypes</emphasis>
</typeStatus>
: 
<specimenCode id="9914EF6BFFC1FF9EFC7BFF6AFB076A8C" box="[976,1169,198,220]" collectionCode="CASIZ" pageId="35" pageNumber="36">CASIZ 185100</specimenCode>
, five specimens, one subsampled for molecular study, various locations, 
<location id="CC6D11CBFFC1FF9EFC90FEA9FC576B49" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFC90FEA9FC576B49" box="[827,961,260,282]" municipality="Milne Bay Province" name="Milne Bay" pageId="35" pageNumber="36">Milne Bay</location>
, 
<collectingMunicipality id="2969DD6AFFC1FF9EFC79FEA9FB4E6B4A" box="[978,1240,260,282]" pageId="35" pageNumber="36">Milne Bay Province</collectingMunicipality>
, 
<elevation id="429FA023FFC1FF9EFB4CFEA9FADE6B49" box="[1255,1352,260,282]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="35" pageNumber="36" unit="m" value="8.5" valueMax="12.0" valueMin="5.0">
<quantity id="0E4AEAF5FFC1FF9EFB4CFEA9FADE6B49" box="[1255,1352,260,282]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="35" pageNumber="36" unit="m" value="8.5" valueMax="12.0" valueMin="5.0">5–12 m</quantity>
</elevation>
depth, 
<collectorName id="644722C6FFC1FF9EFC90FE8EFC306B68" box="[827,934,291,312]" pageId="35" pageNumber="36">R. Steene</collectorName>
</materialsCitation>
. 
<materialsCitation id="79DA4D4DFFC1FF9EFC04FE8EFB916B07" ID-GBIF-Occurrence="2235579137" collectionCode="CASIZ" country="Iceland" location="Anemone Reef" municipality="Ruo" pageId="35" pageNumber="36" specimenCount="1" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFC04FE8EFC6C6B68" box="[943,1018,291,312]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 69754, one specimen, 
<location id="CC6D11CBFFC1FF9EFB56FE8EFA096B68" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFB56FE8EFA096B68" box="[1277,1439,290,312]" country="Iceland" municipality="Ruo" name="Anemone Reef" pageId="35" pageNumber="36">Anemone Reef</location>
, near 
<collectingMunicipality id="2969DD6AFFC1FF9EFCD0FEEFFC3A6B07" box="[891,940,322,343]" pageId="35" pageNumber="36">Ruo</collectingMunicipality>
, 
<collectingCountry id="B1A50780FFC1FF9EFC12FEECFB916B07" box="[953,1031,321,343]" name="Iceland" pageId="35" pageNumber="36">Island</collectingCountry>
</materialsCitation>
, 
<materialsCitation id="79DA4D4DFFC1FF9EFBBEFEECFBF66B25" ID-GBIF-Occurrence="2235579061" collectingDate="1989-07-20" country="Papua New Guinea" location="Lagoon" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectingRegion id="0B7689F2FFC1FF9EFBBEFEECFBE06B07" box="[1045,1142,321,343]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFB2BFEEFFB4C6B07" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFB2BFEEFFB4C6B07" box="[1152,1242,322,343]" country="Papua New Guinea" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
, 
<collectingRegion id="0B7689F2FFC1FF9EFB4CFEECFADC6B07" box="[1255,1354,321,343]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<collectingCountry id="B1A50780FFC1FF9EFAFCFEEFFC516B26" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
, 
<collectingDate id="AD489838FFC1FF9EFC79FECDFBF66B25" box="[978,1120,352,374]" pageId="35" pageNumber="36" value="1989-07-20">
<date id="BD0C61D0FFC1FF9EFC79FECDFBF66B25" box="[978,1120,352,374]" pageId="35" pageNumber="36" value="1989-07-20">20 July 1989</date>
</collectingDate>
</materialsCitation>
, M. Jebb. 
<materialsCitation id="79DA4D4DFFC1FF9EFB65FECDFBF46BE2" ID-GBIF-Occurrence="2235579029" collectingDate="1989-08" collectionCode="CASIZ" country="Papua New Guinea" location="Lagoon" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFB65FECDFA8F6B25" box="[1230,1305,352,373]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 69787, one specimen, dissected, 
<collectingRegion id="0B7689F2FFC1FF9EFB82FED3FB116BC4" box="[1065,1159,382,404]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFB25FED2FB726BC4" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFB25FED2FB726BC4" box="[1166,1252,383,404]" country="Papua New Guinea" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
, 
<collectingRegion id="0B7689F2FFC1FF9EFB44FED3FAD86BC4" box="[1263,1358,382,404]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<collectingCountry id="B1A50780FFC1FF9EFAF1FED2FC5E6BE3" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
, 
<collectingDate id="AD489838FFC1FF9EFC79FE30FBF46BE2" box="[978,1122,413,434]" pageId="35" pageNumber="36" value="1989-08">
<date id="BD0C61D0FFC1FF9EFC79FE30FBF46BE2" box="[978,1122,413,434]" pageId="35" pageNumber="36" value="1989-08">August 1989</date>
</collectingDate>
</materialsCitation>
, M. Gosliner. 
<materialsCitation id="79DA4D4DFFC1FF9EFB51FE30FBBB6BA0" ID-GBIF-Occurrence="2235579142" collectingDate="1989-08" collectionCode="CASIZ" country="Papua New Guinea" location="Lagoon" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFB51FE30FAD36BE2" box="[1274,1349,413,434]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 69785, three specimens, 
<collectingRegion id="0B7689F2FFC1FF9EFC50FE11FBC16B82" box="[1019,1111,444,466]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFBF6FE11FB266B81" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFBF6FE11FB266B81" box="[1117,1200,444,465]" country="Papua New Guinea" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
, 
<collectingRegion id="0B7689F2FFC1FF9EFB10FE11FA8E6B82" box="[1211,1304,444,466]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<collectingCountry id="B1A50780FFC1FF9EFA89FE11FC066BA0" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
, 
<collectingDate id="AD489838FFC1FF9EFC31FE76FBBB6BA0" box="[922,1069,475,496]" pageId="35" pageNumber="36" value="1989-08">
<date id="BD0C61D0FFC1FF9EFC31FE76FBBB6BA0" box="[922,1069,475,496]" pageId="35" pageNumber="36" value="1989-08">August 1989</date>
</collectingDate>
</materialsCitation>
, M. Gosliner. 
<materialsCitation id="79DA4D4DFFC1FF9EFB63FE76FA0B687D" ID-GBIF-Occurrence="2235579072" collectingDate="1988-02-03" collectionCode="CASIZ" country="Papua New Guinea" county="Hussein Village" elevation="20" location="Hole" municipality="Wall" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFB63FE76FA826BA0" box="[1224,1300,475,496]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 65356, one specimen, 
<location id="CC6D11CBFFC1FF9EFC1FFE54FC7C685F" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFC1FFE54FC7C685F" box="[948,1002,505,527]" country="Papua New Guinea" county="Hussein Village" municipality="Wall" name="Hole" pageId="35" pageNumber="36" stateProvince="Madang">Hole</location>
in the 
<collectingMunicipality id="2969DD6AFFC1FF9EFB95FE54FBE3685F" box="[1086,1141,505,527]" pageId="35" pageNumber="36">Wall</collectingMunicipality>
, north of 
<collectingCounty id="206C3F9CFFC1FF9EFB4EFE54FA09685F" box="[1253,1439,505,527]" pageId="35" pageNumber="36">Hussein Village</collectingCounty>
, north of 
<collectingRegion id="0B7689F2FFC1FF9EFC06FDB5FB82687E" box="[941,1044,536,558]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<elevation id="429FA023FFC1FF9EFB88FDB5FBF5687D" box="[1059,1123,536,557]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="m" value="20.0">
<quantity id="0E4AEAF5FFC1FF9EFB88FDB5FBF5687D" box="[1059,1123,536,557]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="m" value="20.0">20 m</quantity>
</elevation>
depth, 
<collectingDate id="AD489838FFC1FF9EFB63FDB5FA0B687D" box="[1224,1437,536,558]" pageId="35" pageNumber="36" value="1988-02-03">
<date id="BD0C61D0FFC1FF9EFB63FDB5FA0B687D" box="[1224,1437,536,558]" pageId="35" pageNumber="36" value="1988-02-03">3 February 1988</date>
</collectingDate>
</materialsCitation>
, T. Gosliner. 
<materialsCitation id="79DA4D4DFFC1FF9EFC15FD9AFA09683A" ID-GBIF-Occurrence="2235579119" collectingDate="1988-01-24" collectionCode="CASIZ" country="Papua New Guinea" elevation="24" location="Kranket Island" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFC15FD9AFB9F681C" box="[958,1033,567,588]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 71237, one specimen, patch reef off N side 
<location id="CC6D11CBFFC1FF9EFC27FDF8FBA9683B" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFC27FDF8FBA9683B" box="[908,1087,597,619]" country="Papua New Guinea" name="Kranket Island" pageId="35" pageNumber="36" stateProvince="Madang">Kranket Island</location>
, 
<elevation id="429FA023FFC1FF9EFBE0FDF8FB12683B" box="[1099,1156,597,619]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="35" pageNumber="36" unit="m" value="24.0">
<quantity id="0E4AEAF5FFC1FF9EFBE0FDF8FB12683B" box="[1099,1156,597,619]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="35" pageNumber="36" unit="m" value="24.0">24 m</quantity>
</elevation>
depth, 
<collectingDate id="AD489838FFC1FF9EFB70FDF8FA09683A" box="[1243,1439,597,619]" pageId="35" pageNumber="36" value="1988-01-24">
<date id="BD0C61D0FFC1FF9EFB70FDF8FA09683A" box="[1243,1439,597,619]" pageId="35" pageNumber="36" value="1988-01-24">24 January 1988</date>
</collectingDate>
</materialsCitation>
, T. Gosliner. 
<materialsCitation id="79DA4D4DFFC1FF9EFC6DFDD9FAD06897" ID-GBIF-Occurrence="2235579124" collectingDate="1986-09-01" collectionCode="CASIZ" country="Solomon Islands" county="Florida Group" elevation="19" location="Takahate Bay" municipality="Big Nggela Island" pageId="35" pageNumber="36" specimenCount="1" typeStatus="paratype">
<collectionCode id="AFA3DFD5FFC1FF9EFC6DFDD9FB8568D9" box="[966,1043,628,649]" pageId="35" pageNumber="36">CASIZ</collectionCode>
0 71474, two specimens, 
<location id="CC6D11CBFFC1FF9EFA9FFDD9FCFD68F8" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFA9FFDD9FCFD68F8" country="Solomon Islands" county="Florida Group" municipality="Big Nggela Island" name="Takahate Bay" pageId="35" pageNumber="36">Takahate Bay</location>
, 
<collectingMunicipality id="2969DD6AFFC1FF9EFCD1FD3FFBF268F8" box="[890,1124,658,680]" pageId="35" pageNumber="36">Big Nggela Island</collectingMunicipality>
, 
<collectingCounty id="206C3F9CFFC1FF9EFBDFFD3FFABC68F8" box="[1140,1322,658,680]" pageId="35" pageNumber="36">Florida Group</collectingCounty>
, 
<collectingCountry id="B1A50780FFC1FF9EFA91FD3FFC036897" name="Solomon Islands" pageId="35" pageNumber="36">Solomon Islands</collectingCountry>
, 
<elevation id="429FA023FFC1FF9EFC09FD1CFB9D6897" box="[930,1035,689,711]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="35" pageNumber="36" unit="m" value="19.0" valueMax="23.0" valueMin="15.0">
<quantity id="0E4AEAF5FFC1FF9EFC09FD1CFB9D6897" box="[930,1035,689,711]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="35" pageNumber="36" unit="m" value="19.0" valueMax="23.0" valueMin="15.0">15–23 m</quantity>
</elevation>
depth, 
<collectingDate id="AD489838FFC1FF9EFBC3FD1CFAD06897" box="[1128,1350,689,711]" pageId="35" pageNumber="36" value="1986-09-01">
<date id="BD0C61D0FFC1FF9EFBC3FD1CFAD06897" box="[1128,1350,689,711]" pageId="35" pageNumber="36" value="1986-09-01">1 September 1986</date>
</collectingDate>
</materialsCitation>
, R. Van Syoc. 
<materialsCitation id="79DA4D4DFFC1FF9EFCD7FD7DFA926973" ID-GBIF-Occurrence="2235579067" collectingDate="1922-11" collectionCode="CASIZ" collectorName="N. Tadwai Island" country="Papua New Guinea" elevation="11" latitude="-4.985" location="Lagoon" longLatPrecision="55" longitude="145.7915" pageId="35" pageNumber="36" specimenCode="CASIZ 191392" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFCD7FD7DFBB368B5" box="[892,1061,720,741]" collectionCode="CASIZ" pageId="35" pageNumber="36">CASIZ 191392</specimenCode>
, one specimen, 
<collectorName id="644722C6FFC1FF9EFB72FD7DFA0868B6" box="[1241,1438,720,742]" pageId="35" pageNumber="36">N. Tadwai Island</collectorName>
, 
<geoCoordinate id="AC8621D7FFC1FF9EFC90FD43FC056954" box="[827,915,750,772]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="55" value="-4.985">4.985°S</geoCoordinate>
, 
<geoCoordinate id="AC8621D7FFC1FF9EFC35FD43FBB76954" box="[926,1057,750,772]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.7915">145.7915°E</geoCoordinate>
, 
<collectingRegion id="0B7689F2FFC1FF9EFB86FD43FB1C6954" box="[1069,1162,750,772]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFB3AFD42FB706954" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFB3AFD42FB706954" box="[1169,1254,751,772]" country="Papua New Guinea" latitude="-4.985" longLatPrecision="55" longitude="145.7915" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
, 
<collectingRegion id="0B7689F2FFC1FF9EFB5AFD43FAD96954" box="[1265,1359,750,772]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<collectingCountry id="B1A50780FFC1FF9EFAF0FD42FC5B6973" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
, 
<elevation id="429FA023FFC1FF9EFC72FCA0FB856973" box="[985,1043,781,803]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="35" pageNumber="36" unit="m" value="11.0">
<quantity id="0E4AEAF5FFC1FF9EFC72FCA0FB856973" box="[985,1043,781,803]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="35" pageNumber="36" unit="m" value="11.0">11 m</quantity>
</elevation>
depth, 
<collectingDate id="AD489838FFC1FF9EFBC0FCA0FA926973" box="[1131,1284,781,803]" pageId="35" pageNumber="36" value="1922-11">
<date id="BD0C61D0FFC1FF9EFBC0FCA0FA926973" box="[1131,1284,781,803]" pageId="35" pageNumber="36" value="1922-11">22 November</date>
</collectingDate>
</materialsCitation>
, 2012. 
<materialsCitation id="79DA4D4DFFC1FF9EFAFDFCA0FAD96930" ID-GBIF-Occurrence="2235579035" collectingDate="2012-11-12" collectionCode="CASIZ" country="Papua New Guinea" location="Sek Island" municipality="Lagoon" pageId="35" pageNumber="36" specimenCode="CASIZ 191160" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFAFDFCA0FC056912" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ 
<date id="BD0C61D0FFC1FF9EFC90FC81FC056912" box="[827,915,812,834]" pageId="35" pageNumber="36" value="60-11-19">191160</date>
</specimenCode>
, one specimen, 
<location id="CC6D11CBFFC1FF9EFBE5FC81FB466912" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFBE5FC81FB466912" box="[1102,1232,812,834]" country="Papua New Guinea" municipality="Lagoon" name="Sek Island" pageId="35" pageNumber="36" stateProvince="Madang">Sek Island</location>
, 
<collectingRegion id="0B7689F2FFC1FF9EFB76FC81FAAB6912" box="[1245,1341,812,834]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<collectingMunicipality id="2969DD6AFFC1FF9EFAEDFC81FA086911" box="[1350,1438,812,833]" pageId="35" pageNumber="36">Lagoon</collectingMunicipality>
, 
<collectingRegion id="0B7689F2FFC1FF9EFC90FCE7FC0E6930" box="[827,920,842,864]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<collectingCountry id="B1A50780FFC1FF9EFC0BFCE6FBE16930" box="[928,1143,842,864]" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
, 
<collectingDate id="AD489838FFC1FF9EFB2BFCE6FAD96930" box="[1152,1359,842,864]" pageId="35" pageNumber="36" value="2012-11-12">
<date id="BD0C61D0FFC1FF9EFB2BFCE6FAD96930" box="[1152,1359,842,864]" pageId="35" pageNumber="36" value="2012-11-12">12 November 2012</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="79DA4D4DFFC1FF9EFAF3FCE6FB2E69EC" ID-GBIF-Occurrence="2235579100" collectingDate="2012-11-15" collectionCode="CASIZ" country="Papua New Guinea" elevation="8" latitude="-5.0985" location="Southern Sek Island" longLatPrecision="7" longitude="145.821" municipality="Lagoon" pageId="35" pageNumber="36" specimenCode="CASIZ 191227" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFAF3FCE6FC19692E" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ 
<date id="BD0C61D0FFC1FF9EFC90FCC4FC19692E" box="[827,911,873,894]" pageId="35" pageNumber="36" value="27-12-19">191227</date>
</specimenCode>
, one specimen, 
<location id="CC6D11CBFFC1FF9EFBEBFCC4FABB692F" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFBEBFCC4FABB692F" box="[1088,1325,873,895]" country="Papua New Guinea" latitude="-5.0985" longLatPrecision="7" longitude="145.821" municipality="Lagoon" name="Southern Sek Island" pageId="35" pageNumber="36" stateProvince="Madang">Southern Sek Island</location>
, 
<geoCoordinate id="AC8621D7FFC1FF9EFA92FCC4FA08692F" box="[1337,1438,873,895]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="5" value="-5.0985">5.0985°S</geoCoordinate>
, 
<geoCoordinate id="AC8621D7FFC1FF9EFC90FC25FC2A69CE" box="[827,956,904,926]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.821">145.8210°E</geoCoordinate>
, 
<elevation id="429FA023FFC1FF9EFC6EFC25FC7B69CD" box="[965,1005,904,925]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="36" unit="m" value="8.0">
<quantity id="0E4AEAF5FFC1FF9EFC6EFC25FC7B69CD" box="[965,1005,904,925]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="36" unit="m" value="8.0">8 m</quantity>
</elevation>
depth, 
<collectingRegion id="0B7689F2FFC1FF9EFB90FC25FB0169CE" box="[1083,1175,904,926]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<collectingMunicipality id="2969DD6AFFC1FF9EFB30FC25FB7869CD" box="[1179,1262,904,925]" pageId="35" pageNumber="36">Lagoon</collectingMunicipality>
, 
<collectingRegion id="0B7689F2FFC1FF9EFB5DFC25FAC569CE" box="[1270,1363,904,926]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<collectingCountry id="B1A50780FFC1FF9EFAF0FC25FC5869EC" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
, 
<collectingDate id="AD489838FFC1FF9EFC71FC0BFB2E69EC" box="[986,1208,934,956]" pageId="35" pageNumber="36" value="2012-11-15">
<date id="BD0C61D0FFC1FF9EFC71FC0BFB2E69EC" box="[986,1208,934,956]" pageId="35" pageNumber="36" value="2012-11-15">15 November 2012</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="79DA4D4DFFC1FF9EFB68FC0AFBFF6E48" ID-GBIF-Occurrence="2235579059" collectingDate="2012-12-12" collectionCode="CASIZ" collectorName="Cement Mixer Reef" country="Papua New Guinea" elevation="13" latitude="-5.15176" location="Lagoon" longLatPrecision="1" longitude="145.81831" pageId="35" pageNumber="36" specimenCode="CASIZ 190823" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFB68FC0AFAF869EC" box="[1219,1390,935,956]" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ 
<date id="BD0C61D0FFC1FF9EFAB3FC0AFAF869EC" box="[1304,1390,935,956]" pageId="35" pageNumber="36" value="23-08-19">190823</date>
</specimenCode>
, one specimen, 
<collectorName id="644722C6FFC1FF9EFC04FC68FB1F698B" box="[943,1161,965,987]" pageId="35" pageNumber="36">Cement Mixer Reef</collectorName>
, 
<geoCoordinate id="AC8621D7FFC1FF9EFB38FC68FA93698B" box="[1171,1285,965,987]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="1" value="-5.15176">5.15176°S</geoCoordinate>
, 
<geoCoordinate id="AC8621D7FFC1FF9EFAA4FC68FA09698B" box="[1295,1439,965,987]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="1" value="145.81831">145.81832°E</geoCoordinate>
, 
<elevation id="429FA023FFC1FF9EFC90FC49FC0669A9" box="[827,912,996,1017]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="2.3" metricValueMin="0.2" pageId="35" pageNumber="36" unit="m" value="12.5" valueMax="23.0" valueMin="2.0">
<quantity id="0E4AEAF5FFC1FF9EFC90FC49FC0669A9" box="[827,912,996,1017]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="2.3" metricValueMin="0.2" pageId="35" pageNumber="36" unit="m" value="12.5" valueMax="23.0" valueMin="2.0">2–23 m</quantity>
</elevation>
depth, 
<collectingRegion id="0B7689F2FFC1FF9EFC42FC49FBDE69AA" box="[1001,1096,996,1018]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFBE4FC49FB3069A9" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFBE4FC49FB3069A9" box="[1103,1190,996,1017]" country="Papua New Guinea" latitude="-5.15176" longLatPrecision="1" longitude="145.81831" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
, 
<collectingRegion id="0B7689F2FFC1FF9EFB19FC49FA8469AA" box="[1202,1298,996,1018]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<collectingCountry id="B1A50780FFC1FF9EFAB6FC49FC186E48" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
<collectingDate id="AD489838FFC1FF9EFC3EFBAEFBFF6E48" box="[917,1129,1026,1048]" pageId="35" pageNumber="36" value="2012-12-12">
<date id="BD0C61D0FFC1FF9EFC3EFBAEFBFF6E48" box="[917,1129,1026,1048]" pageId="35" pageNumber="36" value="2012-12-12">12 December 2012</date>
</collectingDate>
</materialsCitation>
, T. Gosliner. 
<materialsCitation id="79DA4D4DFFC1FF9EFB53FBAEFA096E05" ID-GBIF-Occurrence="2235579039" collectingDate="2012-11-19" collectionCode="CASIZ" country="Papua New Guinea" latitude="-5.0367" location="south Rempi" longLatPrecision="7" longitude="145.8066" pageId="35" pageNumber="36" specimenCode="CASIZ 191326" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFB53FBAEFA096E48" box="[1272,1439,1026,1048]" collectionCode="CASIZ" pageId="35" pageNumber="36">CASIZ 191326</specimenCode>
, one specimen, 
<location id="CC6D11CBFFC1FF9EFC5BFB8CFB1A6E67" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFC5BFB8CFB1A6E67" box="[1008,1164,1057,1079]" country="Papua New Guinea" latitude="-5.0367" longLatPrecision="7" longitude="145.8066" name="south Rempi" pageId="35" pageNumber="36" stateProvince="Madang">south Rempi</location>
<geoCoordinate id="AC8621D7FFC1FF9EFB3DFB8CFA956E67" box="[1174,1283,1057,1079]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="5" value="-5.0367">5.0367°S</geoCoordinate>
, 
<geoCoordinate id="AC8621D7FFC1FF9EFABAFB8CFA086E67" box="[1297,1438,1057,1079]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.8066">145.8066°E</geoCoordinate>
, 
<collectingRegion id="0B7689F2FFC1FF9EFC90FBEDFC376E06" box="[827,929,1088,1110]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<collectingCountry id="B1A50780FFC1FF9EFC04FBEDFB326E06" box="[943,1188,1088,1110]" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
, 
<collectingDate id="AD489838FFC1FF9EFB19FBEDFA096E05" box="[1202,1439,1088,1110]" pageId="35" pageNumber="36" value="2012-11-19">
<date id="BD0C61D0FFC1FF9EFB19FBEDFA096E05" box="[1202,1439,1088,1110]" pageId="35" pageNumber="36" value="2012-11-19">19 November 2012</date>
</collectingDate>
</materialsCitation>
, Francois Michonneau. 
<materialsCitation id="79DA4D4DFFC1FF9EFBEDFBF2FB606EE1" ID-GBIF-Occurrence="2235579088" collectingDate="2012-11-12" collectionCode="CASIZ" country="Papua New Guinea" elevation="22" latitude="-5.2967" location="Bilbil Island" longLatPrecision="7" longitude="145.7816" pageId="35" pageNumber="36" specimenCode="CASIZ 191139" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFBEDFBF2FB646E24" box="[1094,1266,1119,1140]" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ 
<date id="BD0C61D0FFC1FF9EFB37FBF2FB646E24" box="[1180,1266,1119,1140]" pageId="35" pageNumber="36" value="39-11-19">191139</date>
</specimenCode>
, one specimen, 
<location id="CC6D11CBFFC1FF9EFC90FBD0FC556EC3" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFC90FBD0FC556EC3" box="[827,963,1149,1171]" country="Papua New Guinea" latitude="-5.2967" longLatPrecision="7" longitude="145.7816" name="Bilbil Island" pageId="35" pageNumber="36" stateProvince="Madang">Bilbil Island</location>
<geoCoordinate id="AC8621D7FFC1FF9EFC63FBD0FBBD6EC3" box="[968,1067,1149,1171]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="5" value="-5.2967">5.2967°S</geoCoordinate>
, 
<geoCoordinate id="AC8621D7FFC1FF9EFB9FFBD0FB206EC3" box="[1076,1206,1149,1171]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.7816">145.7816°E</geoCoordinate>
, 
<collectingRegion id="0B7689F2FFC1FF9EFB15FBD0FA8D6EC3" box="[1214,1307,1149,1171]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<collectingCountry id="B1A50780FFC1FF9EFA8FFBD3FC186EE2" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
, 
<elevation id="429FA023FFC1FF9EFC32FB31FC466EE1" box="[921,976,1180,1201]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="35" pageNumber="36" unit="m" value="22.0">
<quantity id="0E4AEAF5FFC1FF9EFC32FB31FC466EE1" box="[921,976,1180,1201]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="35" pageNumber="36" unit="m" value="22.0">22 m</quantity>
</elevation>
depth, 
<collectingDate id="AD489838FFC1FF9EFB88FB31FB606EE1" box="[1059,1270,1180,1202]" pageId="35" pageNumber="36" value="2012-11-12">
<date id="BD0C61D0FFC1FF9EFB88FB31FB606EE1" box="[1059,1270,1180,1202]" pageId="35" pageNumber="36" value="2012-11-12">12 November 2012</date>
</collectingDate>
</materialsCitation>
, Heok Hui Tan. 
<materialsCitation id="79DA4D4DFFC1FF9EFC90FB16FCE56F5D" ID-GBIF-Occurrence="2235579123" collectingDate="2012-11-09" collectionCode="CASIZ" collectorName="N. Siar Island" country="Papua New Guinea" latitude="-5.1967" location="Madang" longLatPrecision="7" longitude="145.8067" pageId="35" pageNumber="36" specimenCode="CASIZ 191066" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFC90FB16FC766E80" box="[827,992,1210,1232]" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ 
<date id="BD0C61D0FFC1FF9EFC26FB17FC766E80" box="[909,992,1210,1232]" pageId="35" pageNumber="36" value="66-10-19">191066</date>
</specimenCode>
, one specimen, 
<collectorName id="644722C6FFC1FF9EFB3BFB16FAB96E80" box="[1168,1327,1210,1232]" pageId="35" pageNumber="36">N. Siar Island</collectorName>
, 
<geoCoordinate id="AC8621D7FFC1FF9EFA91FB17FA086E80" box="[1338,1438,1210,1232]" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="5" value="-5.1967">5.1967°S</geoCoordinate>
, 
<geoCoordinate id="AC8621D7FFC1FF9EFC90FB74FC296EBF" box="[827,959,1241,1263]" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="145.8067">145.8067°E</geoCoordinate>
, 
<collectingRegion id="0B7689F2FFC1FF9EFC61FB74FBBE6EBF" box="[970,1064,1241,1263]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
, 
<collectingCountry id="B1A50780FFC1FF9EFB9FFB77FA876EBF" box="[1076,1297,1241,1263]" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
<collectingDate id="AD489838FFC1FF9EFAB2FB74FCE56F5D" pageId="35" pageNumber="36" value="2012-11-09">
<date id="BD0C61D0FFC1FF9EFAB2FB74FCE56F5D" pageId="35" pageNumber="36" value="2012-11-09">9 November 2012</date>
</collectingDate>
</materialsCitation>
, Heok Hui Tan. 
<materialsCitation id="79DA4D4DFFC1FF9EFB89FB55FBB36F1B" ID-GBIF-Occurrence="2235579079" collectionCode="CASIZ" country="Papua New Guinea" county="Marco Oliverio" location="Lagoon" municipality="November-December" pageId="35" pageNumber="36" specimenCode="CASIZ 190842" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<specimenCode id="9914EF6BFFC1FF9EFB89FB55FB536F5D" box="[1058,1221,1272,1293]" collectionCode="CASIZ" pageId="35" pageNumber="36">
CASIZ 
<date id="BD0C61D0FFC1FF9EFBD9FB55FB536F5D" box="[1138,1221,1272,1293]" pageId="35" pageNumber="36" value="42-08-19">190842</date>
</specimenCode>
, one specimen, from orange sponge, 
<collectingRegion id="0B7689F2FFC1FF9EFC41FABBFBD16F7C" box="[1002,1095,1302,1324]" country="Papua New Guinea" name="Madang" pageId="35" pageNumber="36">Madang</collectingRegion>
<location id="CC6D11CBFFC1FF9EFBE4FABAFB326F7C" LSID="urn:lsid:plazi:treatment:411BF606FFC1FF9BFE86FB79FDAE6B68:CC6D11CBFFC1FF9EFBE4FABAFB326F7C" box="[1103,1188,1303,1324]" country="Papua New Guinea" county="Marco Oliverio" municipality="November-December" name="Lagoon" pageId="35" pageNumber="36" stateProvince="Madang">Lagoon</location>
, 
<collectingMunicipality id="2969DD6AFFC1FF9EFB04FABBFA356F7C" box="[1199,1443,1302,1324]" pageId="35" pageNumber="36">November–December</collectingMunicipality>
2012, 
<collectingCounty id="206C3F9CFFC1FF9EFCD6FA9BFBB36F1B" box="[893,1061,1333,1355]" pageId="35" pageNumber="36">Marco Oliverio</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC1FF9EFC90FADFFB276F97" pageId="35" pageNumber="36" type="distribution">
<paragraph id="C90D4710FFC1FF9EFC90FADFFBEE6FDA" blockId="35.[827,1443,1394,1479]" box="[827,1144,1394,1418]" pageId="35" pageNumber="36">
<heading id="9245F07CFFC1FF9EFC90FADFFBEE6FDA" box="[827,1144,1394,1418]" fontSize="10" level="4" pageId="35" pageNumber="36" reason="1">
<emphasis id="FBC69B02FFC1FF9EFC90FADFFBEE6FDA" box="[827,1144,1394,1418]" italics="true" pageId="35" pageNumber="36">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC1FF9EFC90FA3EFB276F97" blockId="35.[827,1443,1394,1479]" pageId="35" pageNumber="36">
Known only from the 
<collectingCountry id="B1A50780FFC1FF9EFBE7FA3EFAAD6FF8" box="[1100,1339,1426,1448]" name="Papua New Guinea" pageId="35" pageNumber="36">Papua New Guinea</collectingCountry>
and the 
<collectingCountry id="B1A50780FFC1FF9EFC90FA1CFC616F97" box="[827,1015,1457,1479]" name="Solomon Islands" pageId="35" pageNumber="36">Solomon Islands</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC1FF9EFC90FA43FA326C13" pageId="35" pageNumber="36" type="etymology">
<paragraph id="C90D4710FFC1FF9EFC90FA43FC2A6C56" blockId="35.[827,1444,1518,1603]" box="[827,956,1518,1542]" pageId="35" pageNumber="36">
<heading id="9245F07CFFC1FF9EFC90FA43FC2A6C56" box="[827,956,1518,1542]" fontSize="10" level="4" pageId="35" pageNumber="36" reason="1">
<emphasis id="FBC69B02FFC1FF9EFC90FA43FC2A6C56" box="[827,956,1518,1542]" italics="true" pageId="35" pageNumber="36">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC1FF9EFC90F9A3FA326C13" blockId="35.[827,1444,1518,1603]" pageId="35" pageNumber="36">
<taxonomicName id="0EB23C93FFC1FF9EFC90F9A3FBCE6C73" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,1112,1550,1571]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC1FF9EFC90F9A3FBCE6C73" box="[827,1112,1550,1571]" italics="true" pageId="35" pageNumber="36">Hypselodoris melanesica</emphasis>
</taxonomicName>
is named for Melanesia, the region to which this species is geographically restricted.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC1FF98FC90F9C6FD7D6B07" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36" type="description">
<paragraph id="C90D4710FFC1FF9EFC90F9C6FC516CD2" blockId="35.[827,1443,1643,1880]" box="[827,967,1643,1666]" pageId="35" pageNumber="36">
<heading id="9245F07CFFC1FF9EFC90F9C6FC516CD2" box="[827,967,1643,1666]" fontSize="10" level="4" pageId="35" pageNumber="36" reason="1">
<emphasis id="FBC69B02FFC1FF9EFC90F9C6FC516CD2" box="[827,967,1643,1666]" italics="true" pageId="35" pageNumber="36">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC1FF99FC90F927FD3B6F4E" blockId="35.[827,1443,1643,1880]" lastBlockId="36.[145,762,1226,1310]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="FBC69B02FFC1FF9EFC90F927FBAE6CCF" box="[827,1080,1674,1695]" italics="true" pageId="35" pageNumber="36">External morphology</emphasis>
: Living animals (
<figureCitation id="51895B95FFC1FF9EFABEF927FAEA6CF0" box="[1301,1404,1674,1696]" captionStart="Figure 18" captionStartId="33.[163,244,1577,1599]" captionTargetBox="[323,1283,196,1537]" captionTargetId="figure@33.[323,1283,196,1537]" captionTargetPageId="33" captionText="Figure 18. Living animals. A, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 177617, Tingloy, Batangas, Philippines. B, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208188, Puerto Galera, Mindoro Oriental, Philippines. C, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 217345, Tingloy, Batangas, Philippines. D, H. lacuna Gosliner &amp; Johnson sp. nov., paratype, CASIZ 208652, Puerto Galera, Mindoro Oriental, Philippines. E, Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., paratype, CASIZ 190823, Madang, Papua New Guinea. F, Hypselodoris paradisa Gosliner &amp; Johnson sp. nov., holotype, CASIZ 191464, Madang, Papua New Guinea, photograph by Vanessa Knutson. G, Hypselodoris perii Gosliner &amp; Johnson sp. nov., NMP 041281, Mabini, Batangas, Philippines. H, Hypselodoris roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 204801, Puerto Galera, Mindoro Oriental, Philippines. I, H. roo Gosliner &amp; Johnson sp. nov., CASIZ 208193, paratype, Puerto Galera, Mindoro Oriental, Philippines. J, H. roo Gosliner &amp; Johnson sp. nov., paratype, CASIZ 186098, Mabini, Tingloy, Philippines." httpUri="https://zenodo.org/record/2632810/files/figure.png" pageId="35" pageNumber="36">Fig. 18E</figureCitation>
) of moderately size, reaching 
<quantity id="0E4AEAF5FFC1FF9EFB2FF904FB4F6CEF" box="[1156,1241,1705,1727]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="mm" value="30.0">30 mm</quantity>
in length. Body translucent purple, with thin white band encircling the margin of notum and foot. Five unipinnate gill branches on notum. One large specimen (CASIZ 
<date id="BD0C61D0FFC1FF9EFAA2F8A8FAC96D4B" box="[1289,1375,1797,1819]" pageId="35" pageNumber="36" value="37-07-12">071237</date>
) with seven gill branches. Gill branches light orange, with darker orange at their common base. Base of gill pocket deep violet. Bulb of rhinophores bright orange, with redder apex. Bulb with ~25 densely packed lamellae. Base of rhinophore sheath deep violet to purple.
</paragraph>
<caption id="9DCD1798FFC6FF99FF38FBF8FE4A6EF6" httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="36" pageNumber="37" startId="36.[147,227,1109,1131]" targetBox="[211,1363,195,1068]" targetPageId="36">
<paragraph id="C90D4710FFC6FF99FF38FBF8FE4A6EF6" blockId="36.[147,1428,1109,1190]" pageId="36" pageNumber="37">
<emphasis id="FBC69B02FFC6FF99FF38FBF8FE9D6E3A" bold="true" box="[147,267,1109,1131]" pageId="36" pageNumber="37">Figure 20.</emphasis>
<taxonomicName id="0EB23C93FFC6FF99FEBDFBF8FC8F6E3B" authority="Gosliner &amp; Johnson" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[278,793,1109,1131]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="melanesica" status="sp. nov.">
<emphasis id="FBC69B02FFC6FF99FEBDFBF8FDAB6E3B" bold="true" box="[278,573,1109,1131]" italics="true" pageId="36" pageNumber="37">Hypselodoris melanesica</emphasis>
Gosliner &amp; Johnson
</taxonomicName>
<taxonomicNameLabel id="E0F52679FFC6FF99FC8BFBFBFCEF6E3B" box="[800,889,1110,1131]" pageId="36" pageNumber="37" rank="species">
<emphasis id="FBC69B02FFC6FF99FC8BFBFBFCEF6E3B" bold="true" box="[800,889,1110,1131]" pageId="36" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
, buccal armature, paratype CASIZ 0 69787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587.
</paragraph>
</caption>
<paragraph id="C90D4710FFC6FF99FF3AFAEBFF726F2B" blockId="36.[145,761,1350,1403]" pageId="36" pageNumber="37">
<emphasis id="FBC69B02FFC6FF99FF3AFAEBFEA16F0B" box="[145,311,1350,1371]" italics="true" pageId="36" pageNumber="37">Mantle glands</emphasis>
: Subcutaneous mantle glands entirely absent.
</paragraph>
<paragraph id="C90D4710FFC6FF99FF3AFA33FB7C6F6C" blockId="36.[145,761,1438,1890]" lastBlockId="36.[809,1425,1226,1340]" pageId="36" pageNumber="37">
<emphasis id="FBC69B02FFC6FF99FF3AFA33FECD6FE4" box="[145,347,1438,1460]" italics="true" pageId="36" pageNumber="37">Buccal armature</emphasis>
: Muscular portion of buccal mass about twice the length of oral tube. Chitinous labial cuticle found at anterior end of muscular portion of the buccal mass bearing numerous jaw rodlets (
<figureCitation id="51895B95FFC6FF99FD20FA57FD7C6C5F" box="[651,746,1530,1552]" captionStart="Figure 20" captionStartId="36.[147,227,1109,1131]" captionTargetBox="[211,1363,195,1068]" captionTargetId="figure@36.[211,1363,195,1068]" captionTargetPageId="36" captionText="Figure 20. Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., buccal armature, paratype CASIZ 069787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="36" pageNumber="37">Fig. 20A</figureCitation>
). Rodlets narrow with short base and evenly curved, with single, acutely pointed apex. Radular formula of one 
<typeStatus id="1609F9B2FFC6FF99FF3AF9FAFF606C3C" box="[145,246,1623,1644]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
(CASIZ 069787) 57 × 59.0.59. Rachidian row of teeth absent (
<figureCitation id="51895B95FFC6FF99FE80F9D8FE1F6CDA" box="[299,393,1653,1675]" captionStart="Figure 20" captionStartId="36.[147,227,1109,1131]" captionTargetBox="[211,1363,195,1068]" captionTargetId="figure@36.[211,1363,195,1068]" captionTargetPageId="36" captionText="Figure 20. Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., buccal armature, paratype CASIZ 069787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="36" pageNumber="37">Fig. 20B</figureCitation>
). Innermost lateral teeth having one pointed denticle on inner side of bifid primary cusp, with another one or two outer denticles. Denticles not extending far beyond middle of elongate primary cusp. Next several laterals lacking inner triangular denticle but possessing three or four denticles on outer side of primary bifid cusps. Midlateral teeth (
<figureCitation id="51895B95FFC6FF99FDC1F880FD5D6D12" box="[618,715,1837,1859]" captionStart="Figure 20" captionStartId="36.[147,227,1109,1131]" captionTargetBox="[211,1363,195,1068]" captionTargetId="figure@36.[211,1363,195,1068]" captionTargetPageId="36" captionText="Figure 20. Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., buccal armature, paratype CASIZ 069787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="36" pageNumber="37">Fig. 20C</figureCitation>
) all lacking inner denticles but having four or five rounded to triangular outer denticles and extended primary cusp. Outermost teeth having a narrower base and shorter tooth shape, with three rounded outer denticles (
<figureCitation id="51895B95FFC6FF99FC9AFA8BFC066F6B" box="[817,912,1318,1340]" captionStart="Figure 20" captionStartId="36.[147,227,1109,1131]" captionTargetBox="[211,1363,195,1068]" captionTargetId="figure@36.[211,1363,195,1068]" captionTargetPageId="36" captionText="Figure 20. Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., buccal armature, paratype CASIZ 069787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="36" pageNumber="37">Fig. 20D</figureCitation>
), often larger than bifid cusps.
</paragraph>
<paragraph id="C90D4710FFC6FF98FC82FAC8FD7D6B07" blockId="36.[809,1426,1381,1894]" lastBlockId="37.[163,779,198,343]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<emphasis id="FBC69B02FFC6FF99FC82FAC8FBB16F2B" box="[809,1063,1381,1403]" italics="true" pageId="36" pageNumber="37">Reproductive system</emphasis>
: Reproductive organs of the 
<typeStatus id="1609F9B2FFC6FF99FC82FA29FC006FC9" box="[809,918,1412,1433]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
(CASIZ 069787) fully mature (
<figureCitation id="51895B95FFC6FF99FAB1FA29FA146FC9" box="[1306,1410,1412,1434]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="36" pageNumber="37">Fig. 13E</figureCitation>
). Ampulla thick, tubular and slightly curved, narrowing somewhat before bifurcating into oviduct and vas deferens. Short oviduct entering female gland mass near albumen gland. Prostatic proximal portion of vas deferens convoluted, curved and thick, and narrowing slightly as it transitions into muscular ejaculatory portion. Prostatic portion enveloping bursa copulatrix. Ejaculatory portion convoluted, narrow, entering short, wider penial bulb. Penial bulb adjacent to straight, moderately wide vaginal duct at common gonopore. Distal end of vas deferens devoid of penial hooks. Female gland mass consisting of large mucous gland and small membrane and albumen glands. Large, lobate vestibular gland situated near exit of mucous gland. Relatively long vagina leading to small, straight receptaculum seminis and larger spherical, thin-walled receptaculum seminis. Receptaculum seminis appressed against vagina in distal half of vaginal length. Moderately short uterine duct emerging from vagina close to base of bursa and entering female gland mass near the albumen gland.
</paragraph>
</subSubSection>
<subSubSection id="81A8149BFFC7FF9BFF08FED3FDAE6B68" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="discussion">
<paragraph id="C90D4710FFC7FF98FF08FED3FE866BC6" blockId="37.[163,779,382,896]" box="[163,272,382,406]" pageId="37" pageNumber="38">
<heading id="9245F07CFFC7FF98FF08FED3FE866BC6" box="[163,272,382,406]" fontSize="10" level="4" pageId="37" pageNumber="38" reason="1">
<emphasis id="FBC69B02FFC7FF98FF08FED3FE866BC6" box="[163,272,382,406]" italics="true" pageId="37" pageNumber="38">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C90D4710FFC7FF98FF08FE32FB1A683B" blockId="37.[163,779,382,896]" lastBlockId="37.[827,1443,198,895]" pageId="37" pageNumber="38">
In our phylogenetic analyses, 
<taxonomicName id="0EB23C93FFC7FF98FE51FE32FD0C6BE3" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[506,666,414,436]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FE51FE32FD0C6BE3" box="[506,666,414,436]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
is always sister to 
<taxonomicName id="0EB23C93FFC7FF98FEBBFE13FE086B82" authorityName="Collingwood" authorityYear="1881" box="[272,414,445,467]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FEBBFE13FE086B82" box="[272,414,445,467]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
but forms a distinct clade. In the ABGD analysis, 
<taxonomicName id="0EB23C93FFC7FF98FE39FE71FDA06BA1" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[402,566,476,497]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FE39FE71FDA06BA1" box="[402,566,476,497]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
is included in the same group as 
<taxonomicName id="0EB23C93FFC7FF98FEFBFE56FE43685F" authorityName="Collingwood" authorityYear="1881" box="[336,469,506,528]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FEFBFE56FE43685F" box="[336,469,506,528]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
, thus suggesting that they should be considered as conspecific. Their uncorrected p-distances for the 
<emphasis id="FBC69B02FFC7FF98FEDAFD95FE36681D" box="[369,416,568,589]" italics="true" pageId="37" pageNumber="38">COI</emphasis>
gene range between 2.0 and 2.6% different, right near the boundary for consideration of these two as distinct species. Although 
<taxonomicName id="0EB23C93FFC7FF98FDC2FDDBFC9C68DA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[617,778,629,651]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FDC2FDDBFC9C68DA" box="[617,778,629,651]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="0EB23C93FFC7FF98FE89FD39FE3B68F9" authorityName="Collingwood" authorityYear="1881" box="[290,429,660,681]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FE89FD39FE3B68F9" box="[290,429,660,681]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
in its shape and body colour, there are consistent morphological differences. In 
<taxonomicName id="0EB23C93FFC7FF98FF08FD7FFEB968B6" authorityName="Collingwood" authorityYear="1881" box="[163,303,721,743]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FF08FD7FFEB968B6" box="[163,303,721,743]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
, the body colour is generally light pink to a deep purple. There may or may not be darker pigment at the base of the gill and the rhinophores. When present, the purple pigment is a wide band that is very diffuse, without distinctly defined edges. In contrast, 
<taxonomicName id="0EB23C93FFC7FF98FEA7FCC6FE3C692F" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[268,426,874,896]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FEA7FCC6FE3C692F" box="[268,426,874,896]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
is always light purple in colour and always has well-defined narrow bands of darker pigment at the base of the rhinophores and gill. The bands at the base of the rhinophores of 
<taxonomicName id="0EB23C93FFC7FF98FAA9FEA9FA346B49" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1282,1442,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FAA9FEA9FA346B49" box="[1282,1442,260,281]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
always have a break in the band on the posterior side of the rhinophores. Internally, 
<taxonomicName id="0EB23C93FFC7FF98FB09FEEFFAD36B06" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1186,1349,321,343]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FB09FEEFFAD36B06" box="[1186,1349,321,343]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
(CASIZ 069787) has a radular formula of 65 × 68.0.68 vs. 77 × 97.0. 
<quantity id="0E4AEAF5FFC7FF98FC1BFED2FC7B6BC4" box="[944,1005,382,404]" metricMagnitude="0" metricUnit="m" metricValue="2.4638" pageId="37" pageNumber="38" unit="in" value="97.0">97 in</quantity>
one specimen of 
<taxonomicName id="0EB23C93FFC7FF98FB11FED2FAD26BC3" authorityName="Collingwood" authorityYear="1881" box="[1210,1348,382,404]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FB11FED2FAD26BC3" box="[1210,1348,382,404]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
(CASIZ 083685). Also, the primary cusps of the radular teeth of 
<taxonomicName id="0EB23C93FFC7FF98FC90FE11FC4F6B81" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[827,985,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FC90FE11FC4F6B81" box="[827,985,444,465]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
appear shorter than those of 
<taxonomicName id="0EB23C93FFC7FF98FA8AFE11FA346B81" authorityName="Collingwood" authorityYear="1881" box="[1313,1442,444,465]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FA8AFE11FA346B81" box="[1313,1442,444,465]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFC7FF98FCE8FE77FC026BA0" box="[835,916,474,496]" captionStart="Figure 20" captionStartId="36.[147,227,1109,1131]" captionTargetBox="[211,1363,195,1068]" captionTargetId="figure@36.[211,1363,195,1068]" captionTargetPageId="36" captionText="Figure 20. Hypselodoris melanesica Gosliner &amp; Johnson sp. nov., buccal armature, paratype CASIZ 069787. A, jaw rodlets. B, inner lateral teeth, paratype, CASIZ 177617. C, middle lateral teeth, paratype, CASIZ 177617. D, outer lateral teeth, paratype, CASIZ 208587." httpUri="https://zenodo.org/record/2632816/files/figure.png" pageId="37" pageNumber="38">Figs 20</figureCitation>
, 
<figureCitation id="51895B95FFC7FF98FC0BFE76FC2D6BA0" box="[928,955,475,496]" captionStart="Figure 21" captionStartId="37.[163,242,1850,1872]" captionTargetBox="[227,1380,938,1809]" captionTargetId="figure@37.[227,1380,938,1809]" captionTargetPageId="37" captionText="Figure 21. Hypselodoris bullockii (Collingwood, 1881), buccal mass. A, jaw rodlets, CASIZ 083685. B, inner lateral teeth, CASIZ 085905. C, middle lateral teeth, CASIZ 085905. D, outer lateral teeth, CASIZ 085905." httpUri="https://zenodo.org/record/2632820/files/figure.png" pageId="37" pageNumber="38">21</figureCitation>
). The reproductive system of 
<taxonomicName id="0EB23C93FFC7FF98FAAEFE76FA356BBF" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1285,1443,474,496]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FAAEFE76FA356BBF" box="[1285,1443,474,496]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
has a shorter penial bulb and a longer ejaculatory duct that is more convoluted than that found in 
<taxonomicName id="0EB23C93FFC7FF98FA8BFDB5FA34687D" authorityName="Collingwood" authorityYear="1881" box="[1312,1442,536,557]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FA8BFDB5FA34687D" box="[1312,1442,536,557]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
(
<figureCitation id="51895B95FFC7FF98FCE8FD9BFC30681C" box="[835,934,566,588]" captionStart="Figure 13" captionStartId="24.[143,224,1826,1848]" captionTargetBox="[207,1359,196,1783]" captionTargetId="figure@24.[206,1361,195,1785]" captionTargetPageId="24" captionText="Figure 13. Reproductive systems. A, Hypselodoris ghardaqana (Gohar &amp; Aboul-Ela, 1957), CASIZ 192282, scale bar: 3.5 mm. B, Hypselodoris iba Gosliner &amp; Johnson sp. nov., CASIZ 177777, scale bar: 3.0 mm. C, Hypselodoris katherinae Gosliner &amp; Johnson sp. nov., CASIZ 181257, scale bar: 1.0 mm. D, Hypselodoris lacuna Gosliner &amp; Johnson sp. nov.," httpUri="https://zenodo.org/record/2632797/files/figure.png" pageId="37" pageNumber="38">Fig. 13F</figureCitation>
). Also, 
<taxonomicName id="0EB23C93FFC7FF98FC54FD9AFB30681B" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[1023,1190,566,588]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FC54FD9AFB30681B" box="[1023,1190,566,588]" italics="true" pageId="37" pageNumber="38">H. melanesica</emphasis>
</taxonomicName>
has a longer vaginal duct than that of 
<taxonomicName id="0EB23C93FFC7FF98FBA9FDFBFB13683A" authorityName="Collingwood" authorityYear="1881" box="[1026,1157,597,619]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FBA9FDFBFB13683A" box="[1026,1157,597,619]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C90D4710FFC7FF9BFCF8FDD9FDAE6B68" blockId="37.[827,1443,198,895]" lastBlockId="38.[145,761,198,312]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<taxonomicName id="0EB23C93FFC7FF98FCF8FDD9FBF868D9" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[851,1134,628,649]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FCF8FDD9FBF868D9" box="[851,1134,628,649]" italics="true" pageId="37" pageNumber="38">Hypselodoris melanesica</emphasis>
</taxonomicName>
is known only from 
<collectingCountry id="B1A50780FFC7FF98FAF1FDD9FC4C68F8" name="Papua New Guinea" pageId="37" pageNumber="38">Papua New Guinea</collectingCountry>
and the 
<collectingCountry id="B1A50780FFC7FF98FBFFFD3FFABD68F8" box="[1108,1323,658,680]" name="Solomon Islands" pageId="37" pageNumber="38">Solomon Islands</collectingCountry>
, whereas 
<taxonomicName id="0EB23C93FFC7FF98FC90FD1FFC5B6896" authorityName="Collingwood" authorityYear="1881" box="[827,973,689,711]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FC90FD1FFC5B6896" box="[827,973,689,711]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
is known from the western Pacific of 
<collectingCountry id="B1A50780FFC7FF98FC90FD7DFC3F68B6" box="[827,937,720,742]" name="Australia" pageId="37" pageNumber="38">Australia</collectingCountry>
, 
<collectingCountry id="B1A50780FFC7FF98FC19FD7DFBF668B6" box="[946,1120,720,742]" name="New Caledonia" pageId="37" pageNumber="38">New Caledonia</collectingCountry>
, 
<collectingCountry id="B1A50780FFC7FF98FBC3FD7DFB4468B6" box="[1128,1234,720,742]" name="Malaysia" pageId="37" pageNumber="38">Malaysia</collectingCountry>
, 
<collectingCountry id="B1A50780FFC7FF98FB77FD7DFAD868B6" box="[1244,1358,720,742]" name="Indonesia" pageId="37" pageNumber="38">Indonesia</collectingCountry>
, 
<collectingCountry id="B1A50780FFC7FF98FAFCFD7DFA0868B5" box="[1367,1438,720,741]" name="Japan" pageId="37" pageNumber="38">Japan</collectingCountry>
, 
<collectingCountry id="B1A50780FFC7FF98FC90FD43FC0C6954" box="[827,922,750,772]" name="Taiwan" pageId="37" pageNumber="38">Taiwan</collectingCountry>
, 
<collectingCountry id="B1A50780FFC7FF98FC02FD43FBAA6954" box="[937,1084,750,772]" name="Philippines" pageId="37" pageNumber="38">Philippines</collectingCountry>
, 
<collectingCountry id="B1A50780FFC7FF98FBE1FD43FAB26954" box="[1098,1316,750,772]" name="Marshall Islands" pageId="37" pageNumber="38">Marshall Islands</collectingCountry>
(
<bibRefCitation id="AD233AE1FFC7FF98FA93FD43FC2D6972" author="Gosliner TM &amp; Behrens, DW &amp; Valdes, A." journalOrPublisher="Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences" pageId="37" pageNumber="38" refId="ref42554" refString="Gosliner TM, Behrens, DW, Valdes, A. 2008. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Gig Harbor / San Francisco: Sea Challengers / California Academy of Sciences." title="Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna" type="book" year="2008">
Gosliner 
<emphasis id="FBC69B02FFC7FF98FC90FCA3FCE36972" box="[827,885,781,803]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
, 2008
</bibRefCitation>
) and 
<collectingCountry id="B1A50780FFC7FF98FBABFCA0FBD26973" box="[1024,1092,781,803]" name="Palau" pageId="37" pageNumber="38">Palau</collectingCountry>
(present study). 
<taxonomicName id="0EB23C93FFC7FF98FAA7FCA0FC2F6911" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC7FF98FAA7FCA0FC2F6911" italics="true" pageId="37" pageNumber="38">Hypselodoris melanesica</emphasis>
</taxonomicName>
and 
<taxonomicName id="0EB23C93FFC7FF98FC46FC81FBF96911" authorityName="Collingwood" authorityYear="1881" box="[1005,1135,812,833]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC7FF98FC46FC81FBF96911" box="[1005,1135,812,833]" italics="true" pageId="37" pageNumber="38">H. bullockii</emphasis>
</taxonomicName>
are geographically isolated and have minor but consistent differences in their coloration and internal morphology spanning several organ systems. Based on these consistent differences, we consider 
<taxonomicName id="0EB23C93FFC4FF9BFE88FF4BFE516AAA" authorityName="Epstein &amp; Hallas &amp; Johnson &amp; Lopez &amp; Gosliner" authorityYear="2018" box="[291,455,229,251]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="melanesica">
<emphasis id="FBC69B02FFC4FF9BFE88FF4BFE516AAA" box="[291,455,229,251]" italics="true" pageId="38" pageNumber="39">H. melanesica</emphasis>
</taxonomicName>
as a distinct species from 
<taxonomicName id="0EB23C93FFC4FF9BFF3AFEA9FE8F6B49" authorityName="Collingwood" authorityYear="1881" box="[145,281,260,281]" class="Gastropoda" family="Chromodorididae" genus="Hypselodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="bullockii">
<emphasis id="FBC69B02FFC4FF9BFF3AFEA9FE8F6B49" box="[145,281,260,281]" italics="true" pageId="38" pageNumber="39">H. bullockii</emphasis>
</taxonomicName>
despite the fact that the ABGD analysis clusters these species as conspecifics.
</paragraph>
</subSubSection>
</treatment>
</document>