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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1185.112099" ID-Pensoft-Pub="1313-2970-1185-1" ID-Pensoft-UUID="99994BD90C195466A249943D2E939FB4" ID-ZooBank="5B12796B210249948D3353FAD1A1B763" ModsDocID="1313-2970-1185-1" checkinTime="1701152605290" checkinUser="pensoft" docAuthor="Lee, Chi-Feng" docDate="2023" docId="370802DE1894541F8F5F500876CFFF2F" docLanguage="en" docName="ZooKeys 1185: 1-19" docOrigin="ZooKeys 1185" docPubDate="2023-11-27" docSource="http://dx.doi.org/10.3897/zookeys.1185.112099" docTitle="Sphaeroderma hsui Lee 2023, sp. nov." docType="treatment" docUuid="510555F9-6ABE-4D34-9789-0D27E498F030" docUuidSource="ZooBank" docVersion="1" id="99994BD90C195466A249943D2E939FB4" lastPageNumber="1" masterDocId="99994BD90C195466A249943D2E939FB4" masterDocTitle="Four new species of Sphaeroderma Stephens (Coleoptera, Chrysomelidae, Galerucinae, Alticini) from Taiwan, with discussion on genus boundaries based on S. flavonotatum Chujo and S. jungchani sp. nov." masterLastPageNumber="19" masterPageNumber="1" pageNumber="1" updateTime="1701152605290" updateUser="pensoft">
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<mods:title id="8962CB01144AA6FE58F4B455BE78C8F4">Four new species of Sphaeroderma Stephens (Coleoptera, Chrysomelidae, Galerucinae, Alticini) from Taiwan, with discussion on genus boundaries based on S. flavonotatum Chujo and S. jungchani sp. nov.</mods:title>
</mods:titleInfo>
<mods:name id="6AA72F2CA4C81BF73972B85D5286CFA4" type="personal">
<mods:role id="02C250DA35BEC3AD40D75CB719769998">
<mods:roleTerm id="10E3D7FF19246E821CAB4BDE452123BC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F69E9999B7D93E593995DAF54483BB8F">Lee, Chi-Feng</mods:namePart>
<mods:nameIdentifier id="DFED801AA0B432EBDFD56D73043B858E" type="ORCID">https://orcid.org/0000-0003-1996-0557</mods:nameIdentifier>
<mods:affiliation id="293A0F0EACCD83BD24CB3EA0970F0D95">Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan</mods:affiliation>
<mods:nameIdentifier id="0F7603FF934281C34A7CE997C2EAE301" type="email">chifeng@tari.gov.tw</mods:nameIdentifier>
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<mods:typeOfResource id="825D0DE68CEE241A7693B3A758E8FF82">text</mods:typeOfResource>
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<mods:title id="C1D144AA119CDA01644B1777AE470DE4">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="0DB351C9D61DCFAB21D46883117124B3">
<mods:date id="ED8B576C163AC97D5CA7783929E867D8">2023</mods:date>
<mods:detail id="A47EED20999C4B2475DCDA563C370FE3" type="pubDate">
<mods:number id="12FBE5E59E61AD2B955E0C367C4953C4">2023-11-27</mods:number>
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<mods:detail id="8FA84060CCFB60CD598F35584AE6C671" type="volume">
<mods:number id="CB842F6DE1D225511578AFD42F258237">1185</mods:number>
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<mods:url id="6D1E903900F70EFB1F9E2C70EA8B05B6">http://dx.doi.org/10.3897/zookeys.1185.112099</mods:url>
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<mods:classification id="2F8E4997EF21DD0914CB8B7FF82B9F01">journal article</mods:classification>
<mods:identifier id="00F7A4A752CFB133335B8ACEB90B0828" type="DOI">http://dx.doi.org/10.3897/zookeys.1185.112099</mods:identifier>
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<treatment id="370802DE1894541F8F5F500876CFFF2F" LSID="urn:lsid:zoobank.org:act:510555F9-6ABE-4D34-9789-0D27E498F030" httpUri="http://treatment.plazi.org/id/370802DE1894541F8F5F500876CFFF2F" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="B65FFA49F30903B6B9BAEC3865435D80" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="DA3605584123178ADCD3FF8474863D1C" pageId="0" pageNumber="1">
<taxonomicName id="EA2C4E8319BDF23BC9FA37BF328AF162" LSID="https://zoobank.org/510555F9-6ABE-4D34-9789-0D27E498F030" authority="Lee, 2023" authorityName="Lee" authorityYear="2023" class="Dothideomycetes" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma hsui" order="Coleoptera" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="hsui" status="sp. nov.">Sphaeroderma hsui</taxonomicName>
<taxonomicNameLabel id="74BDC490A266C5D37913906FB82C30BA" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="09E7FF6DDCE30A099210655CB862D16B" pageId="0" pageNumber="1" type="description">
<paragraph id="38A4561CB54C14BAB29B19695333D411" pageId="0" pageNumber="1">
<figureCitation id="3A0B4E7C0007C3F3DCCFAC9D73415170" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Sphaeroderma species in frontal view A S. hsui sp. nov. B S. changi sp. nov. C S. sheipaensis sp. nov. D S. flavonotatum Chujo E S. jungchani sp. nov." figureDoi="10.3897/zookeys.1185.112099.figure2" httpUri="https://binary.pensoft.net/fig/942952" pageId="0" pageNumber="1">Figs 2A</figureCitation>
<figureCitation id="DE9F6EF19332E909283DC37A627A71B9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Sphaeroderma hsui sp. nov. A typical form, male, dorsal view B ditto, ventral view C ditto, lateral view D color variation, male, dorsal view E ditto, ventral view F ditto, lateral view G color variation, female, dorsal view H ditto, ventral view I ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure3" httpUri="https://binary.pensoft.net/fig/942953" pageId="0" pageNumber="1">, 3</figureCitation>
<figureCitation id="4B74CAE5A700724DC6160C6768B4526C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FFFB67EB881A4578DC2F54FF8644C655" pageId="0" pageNumber="1" type="types">
<paragraph id="F6C2A53EE7E9238E34C3BD8EE69F6B0C" pageId="0" pageNumber="1">Types.</paragraph>
<paragraph id="7C15CA8280A82705EA4B1E22F88A5013" pageId="0" pageNumber="1">
<emphasis id="F5D2F010BD67BDD9A085B148BA563D6F" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
♂ (TARI). Taiwan: Miaoli, Hsiaopangchih (小胖池), 28.VIII.2021-16.XI.2021, leg. Y.-F. Hsu.
<emphasis id="11537890C21966B21C586E4955D23BFE" bold="true" pageId="0" pageNumber="1">
<emphasis id="D9CA9C9EF1A174C9AAB1D10F461DF6C0" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
.
</emphasis>
29♂, 22♀ (TARI: 26♂, 19♀; BMNH: 3♂, 3♀), same data as holotype; 4♂, 2♀ (TARI), same but with
<normalizedToken id="785D42B3BAAF7B216A74DE708FE62029" originalValue="“28.VIII.2021”">&quot;28.VIII.2021&quot;</normalizedToken>
; 2♂, 3♀ (TARI), same but with
<normalizedToken id="926453A46D19AAEE4F892DA38EBDA618" originalValue="“21.V.-28.VIII.2021”">&quot;21.V.-28.VIII.2021&quot;</normalizedToken>
; 1♀ (TARI): Hsishihshan (西勢山), 16.XI.2021-21.IV.2022, leg. Y.-F. Hsu.
</paragraph>
<caption id="73DCA2E72E24FF0C7C92C9371363B06E" doi="10.3897/zookeys.1185.112099.figure2" httpUri="https://binary.pensoft.net/fig/942952" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph id="234E1F6EBC7A3FCA830D984F7660CA20" pageId="0" pageNumber="1">
<emphasis id="16884FF125B535B519EBB6360175290E" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
Heads of
<taxonomicName id="BFBAAE525072217F565EDF8C0CC081C7" authorityName="Stephens" authorityYear="1831" class="Dothideomycetes" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma" order="Coleoptera" pageId="0" pageNumber="1" phylum="Ascomycota" rank="genus">
<emphasis id="CF9BF4C4ED66B82C58488C1D0A0D46A4" italics="true" pageId="0" pageNumber="1">Sphaeroderma</emphasis>
</taxonomicName>
species in frontal view
<emphasis id="AD94565E148CBE2AB3BC3AA946699D2F" bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName id="4C9E435724CA330AC6227451328B1B50" lsidName="S. hsui" pageId="0" pageNumber="1" rank="species" species="hsui">
<emphasis id="F4CD6D50834CDE36482E5DB520810742" italics="true" pageId="0" pageNumber="1">S. hsui</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="E524F38A26629E847280AA02AF3B3E3D" bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName id="9636FC872C21F62261767E8DD0EC85A8" lsidName="S. changi" pageId="0" pageNumber="1" rank="species" species="changi">
<emphasis id="EF94CEB34B27F8D19AF8FEAA8D898112" italics="true" pageId="0" pageNumber="1">S. changi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="B59B13865F46322F996E6DC9ECC1DF90" bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName id="55B559744234EDA0E70526098AB3B710" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="AFFBF6B96656E0D967E5C4BFD1C68AA6" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="F081E7BB3B65C0E52B82A60182F77D2A" bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName id="C97BBE8F66284ABA953EC256849A2F72" lsidName="S. flavonotatum" pageId="0" pageNumber="1" rank="species" species="flavonotatum">
<emphasis id="0C0ACE92FCE86164DC335C4B7C55A8ED" italics="true" pageId="0" pageNumber="1">S. flavonotatum</emphasis>
</taxonomicName>
<normalizedToken id="6F60D5BC3B743561F07B560132275F8F" originalValue="Chûjô">Chujo</normalizedToken>
<emphasis id="CA1F8AD3996F33F27F7F3C9227014BDD" bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName id="3DBF55A7F8F9C910D599BC70B3DC9F0F" lsidName="S. jungchani" pageId="0" pageNumber="1" rank="species" species="jungchani">
<emphasis id="60C74D61674CA20B7BA69A2913FE41B6" italics="true" pageId="0" pageNumber="1">S. jungchani</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4233A8B0030514ADED5BF9BC047A3749" pageId="0" pageNumber="1" type="description">
<paragraph id="F3AC184AE423C5AF6AEF115505D3C590" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="0AE31489869D01B8A84D41A14941C6EF" pageId="0" pageNumber="1">
Length 2.5-2.8 mm, width 1.6-1.9 mm. Body color (Fig.
<figureCitation id="A209FCBFD4B3EF28F36EB17D052855FB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Sphaeroderma hsui sp. nov. A typical form, male, dorsal view B ditto, ventral view C ditto, lateral view D color variation, male, dorsal view E ditto, ventral view F ditto, lateral view G color variation, female, dorsal view H ditto, ventral view I ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure3" httpUri="https://binary.pensoft.net/fig/942953" pageId="0" pageNumber="1">3A-C</figureCitation>
) yellowish-brown. Antennae filiform in males (Fig.
<figureCitation id="67AED0CFDFAA98B9C7B3C6DFAF74B042" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4A</figureCitation>
), antennomeres VIII-X wider, length ratios of antennomeres I-XI 1.0:0.6:0.4:0.5:0.7:0.6:0.7:0.8:0.7:0.7:1.0, length to width ratios of antennomeres I-XI 2.9:2.1:1.8:2.0:2.5:2.4:2.5:2.6:2.3:2.3:3.0; similar shape in females (Fig.
<figureCitation id="50302C1575FEE5389DD55BEC1A901E75" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4B</figureCitation>
), but antennomeres VIII-X narrower, length ratios of antennomeres I-XI 1.0:0.6:0.4:0.5:0.8:0.6:0.8:0.7:0.7:0.7:1.0, length to width ratios of antennomeres I-XI 3.1:2.3:1.9:2.2:3.2:2.6:2.9:2.5:2.4:2.2:3.3. Antennal calli longitudinal, with supracallinal sulci poorly delimited. Pronotum 1.7-1.9
<normalizedToken id="17FD4E637CC5DC6FC30D82F6A2864377" originalValue="×">x</normalizedToken>
wider than long, disc with fine, scattered punctures, same size as punctures on elytra; lateral margins almost straight; anterolateral callosity protruding forward; posterolateral callosity poorly developed. Elytra 1.3 times longer than wide, sides widely rounded; disc with punctures entirely confused; humeral calli well developed. Abdominal ventrite V without internal median ridge in both sexes. Male genitalia: aedeagus (Fig.
<figureCitation id="F2CB3513571A002F001B24F13D7E8E1B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4C, D</figureCitation>
) slender in dorsal view, 4.0
<normalizedToken id="BBF678339689EB5EDB37B5AC7DFED3B1" originalValue="×">x</normalizedToken>
longer than wide; parallel-sided, but slightly narrowed at apical 1/4, apex widely rounded; moderately curved at middle in lateral view, apex slightly recurved, ventral margin with densely, tiny teeth at middle; ostium membranous and with Y-shaped sclerotized area. Female genitalia: ventrite VIII (Fig.
<figureCitation id="88E3DD562DA87643690B8A0FF8DD2BD4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4E</figureCitation>
) with apical part triangular, apical margin smooth and lacking setae, disc with medial part membranous, spiculum short; gonocoxae (Fig.
<figureCitation id="FF5CED7DB4F3028740161B84D9EF6B68" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4G</figureCitation>
) separated, transversely triangular, with dense, long setae along apical margins, with one slender and transverse basal sclerite; receptacle of spermatheca (Fig.
<figureCitation id="EB28416BD0F7CC127CF6F3B433D89132" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4F</figureCitation>
) moderately swollen; pump short and strongly curved, transverse wrinkles present on entire pump and extending into half of receptacle; sclerotized proximal spermathecal duct long, with ramus oblong.
</paragraph>
<caption id="B43C598AA0B2725470AF596BD3283996" doi="10.3897/zookeys.1185.112099.figure3" httpUri="https://binary.pensoft.net/fig/942953" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph id="7B5448F1EC1D9D8317B9CE7AE6AFFE97" pageId="0" pageNumber="1">
<emphasis id="58B49DF98FA1901548BA44EB48DAE5B2" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
Habitus of
<taxonomicName id="FB17FF75565C4797CC19F02C72615414" authorityName="Lee" authorityYear="2023" class="Dothideomycetes" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma hsui" order="Coleoptera" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="hsui">
<emphasis id="E75278DA48FD4406B7BEEA6FB474AAB6" italics="true" pageId="0" pageNumber="1">Sphaeroderma hsui</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="DDA274696F5F8AE6A55AA09BC0B8F429" bold="true" pageId="0" pageNumber="1">A</emphasis>
typical form, male, dorsal view
<emphasis id="2D04536ABF7595E3C376A7F0D0D97497" bold="true" pageId="0" pageNumber="1">B</emphasis>
ditto, ventral view
<emphasis id="90AA319238870FAFBF9C57D280E2FA65" bold="true" pageId="0" pageNumber="1">C</emphasis>
ditto, lateral view
<emphasis id="566C042A125951DEA3094DCE7BF45E93" bold="true" pageId="0" pageNumber="1">D</emphasis>
color variation, male, dorsal view
<emphasis id="7DA379F25BBF9A2A50AF7ADEEC12D2B6" bold="true" pageId="0" pageNumber="1">E</emphasis>
ditto, ventral view
<emphasis id="C1FDE5A6261623593E87B15FC9DC467D" bold="true" pageId="0" pageNumber="1">F</emphasis>
ditto, lateral view
<emphasis id="5C2993E9C40E5212A6EF53C99FCD0FF2" bold="true" pageId="0" pageNumber="1">G</emphasis>
color variation, female, dorsal view
<emphasis id="BC374C70740DD6B499B2CF28F62167A3" bold="true" pageId="0" pageNumber="1">H</emphasis>
ditto, ventral view
<emphasis id="62CD266BB1F7E7920956387B738F7043" bold="true" pageId="0" pageNumber="1">I</emphasis>
ditto, lateral view.
</paragraph>
</caption>
<caption id="EBAE2588E2F8ECE0224EAD3F515713EE" doi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="C772089B772D01440EBE58723E7B2F79" pageId="0" pageNumber="1">
<emphasis id="D3B884C620CE2643F7106E66444C62CC" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
Diagnostic characters of
<taxonomicName id="62F7FCE98409432B3181AD3717DC3C9B" authorityName="Lee" authorityYear="2023" class="Dothideomycetes" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma hsui" order="Coleoptera" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="hsui">
<emphasis id="9EDD323DE5071ECCF9926AC385662D3A" italics="true" pageId="0" pageNumber="1">Sphaeroderma hsui</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="7934C69736FA541341518D8715851B4A" bold="true" pageId="0" pageNumber="1">A</emphasis>
antenna, male
<emphasis id="62E1DEF3558F23B59062787E6305734C" bold="true" pageId="0" pageNumber="1">B</emphasis>
antenna, female
<emphasis id="494F50AEAFB8173673DA058E2BE591C7" bold="true" pageId="0" pageNumber="1">C</emphasis>
aedeagus, dorsal view
<emphasis id="670E4587B70B3024A8A7DD2B3673E5D5" bold="true" pageId="0" pageNumber="1">D</emphasis>
aedeagus, lateral view
<emphasis id="E32FE3BAF5D42FD2161E408CBF170124" bold="true" pageId="0" pageNumber="1">E</emphasis>
abdominal ventrite VIII, female
<emphasis id="23E0C56B5C5A4287665B719019A6CB57" bold="true" pageId="0" pageNumber="1">F</emphasis>
spermatheca
<emphasis id="2A7936912E0F5EBD045F288C7CE57A3A" bold="true" pageId="0" pageNumber="1">G</emphasis>
gonocoxae.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="10E73D3EDB0C1D7BFB6246F06141040A" pageId="0" pageNumber="1" type="color variation">
<paragraph id="21805853757B9344682021F833136746" pageId="0" pageNumber="1">Color variation.</paragraph>
<paragraph id="4B9590E9FDEA7B29CA0634DD17E251E6" pageId="0" pageNumber="1">
Some adults have black bodies (Fig.
<figureCitation id="E87AF31574A2D7EC8793C6352D8C67D1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Sphaeroderma hsui sp. nov. A typical form, male, dorsal view B ditto, ventral view C ditto, lateral view D color variation, male, dorsal view E ditto, ventral view F ditto, lateral view G color variation, female, dorsal view H ditto, ventral view I ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure3" httpUri="https://binary.pensoft.net/fig/942953" pageId="0" pageNumber="1">3D-F</figureCitation>
) with legs and antennae that are yellowish-brown, with metafemora darkener; some adults (Fig.
<figureCitation id="DDB3C2C279740957664B1075F944A701" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Sphaeroderma hsui sp. nov. A typical form, male, dorsal view B ditto, ventral view C ditto, lateral view D color variation, male, dorsal view E ditto, ventral view F ditto, lateral view G color variation, female, dorsal view H ditto, ventral view I ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure3" httpUri="https://binary.pensoft.net/fig/942953" pageId="0" pageNumber="1">3G-I</figureCitation>
) similar in this color form, but the elytra are yellowish-brown except darker base that extends to lateral margins, then abbreviated at apical 1/3.
</paragraph>
</subSubSection>
<subSubSection id="2A86C9C746A0361A8C15C9CAB3BF2DB5" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="D833EC7EFED0D4E6DCDC7BB1BF18F584" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="157FCA6C9930119B2CCBF70CDA30EC4D" pageId="0" pageNumber="1">
Although adults of
<taxonomicName id="39AD4D882AABBD186A4666491D412AC1" authorityName="Lee" authorityYear="2023" class="Dothideomycetes" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma hsui" order="Coleoptera" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="hsui">
<emphasis id="FF5390E23238C5ABCC8B55A79495324A" italics="true" pageId="0" pageNumber="1">Sphaeroderma hsui</emphasis>
</taxonomicName>
sp. nov. display diverse color patterns (Fig.
<figureCitation id="8EFFC1A259FB9EE1705E94B7FF1C91B7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Sphaeroderma hsui sp. nov. A typical form, male, dorsal view B ditto, ventral view C ditto, lateral view D color variation, male, dorsal view E ditto, ventral view F ditto, lateral view G color variation, female, dorsal view H ditto, ventral view I ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure3" httpUri="https://binary.pensoft.net/fig/942953" pageId="0" pageNumber="1">3</figureCitation>
), they are characteristic and diagnostic. In addition, this new species differs from the two other Taiwanese species (
<taxonomicName id="844F4EDE941632056A811E8DB3E61C17" lsidName="S. changi" pageId="0" pageNumber="1" rank="species" species="changi">
<emphasis id="17C4136544A1C9BDB2897975FB2452CB" italics="true" pageId="0" pageNumber="1">S. changi</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="8D60533A5E56EBC50D9100B58D8B4D22" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="9855A0DD071382DE9533377C2AE6E900" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov.) based the following combination characters: entire yellowish-brown antennae (Fig.
<figureCitation id="CC061D725F06164E811806288B7E3922" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Sphaeroderma hsui sp. nov. A typical form, male, dorsal view B ditto, ventral view C ditto, lateral view D color variation, male, dorsal view E ditto, ventral view F ditto, lateral view G color variation, female, dorsal view H ditto, ventral view I ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure3" httpUri="https://binary.pensoft.net/fig/942953" pageId="0" pageNumber="1">3</figureCitation>
) [yellowish-brown antennomeres I-IV and black antennomeres V-XI in other species (Fig.
<figureCitation id="40B23AEFD2FBC22391BDA90528DA2E6E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus of Sphaeroderma changi sp. nov. and S. sheipaensis sp. nov. A S. changi sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D S. sheipaensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure5" httpUri="https://binary.pensoft.net/fig/942955" pageId="0" pageNumber="1">5</figureCitation>
)]; longitudinal antennal calli with poorly delimited supracallinal sulci (Fig.
<figureCitation id="1C7ACDFA66BFE6015FAA973BA4610B9F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Sphaeroderma species in frontal view A S. hsui sp. nov. B S. changi sp. nov. C S. sheipaensis sp. nov. D S. flavonotatum Chujo E S. jungchani sp. nov." figureDoi="10.3897/zookeys.1185.112099.figure2" httpUri="https://binary.pensoft.net/fig/942952" pageId="0" pageNumber="1">2A</figureCitation>
) [transverse antennal calli with well-developed supracallinal sulci in
<taxonomicName id="67EA2FF99BDBB37D66829D5539E85A46" lsidName="S. changi" pageId="0" pageNumber="1" rank="species" species="changi">
<emphasis id="D5895B1252B4ADF79EFA3A37380E0ED8" italics="true" pageId="0" pageNumber="1">S. changi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="64BC32B960973C8DB051F5F5AE8D6F4C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Sphaeroderma species in frontal view A S. hsui sp. nov. B S. changi sp. nov. C S. sheipaensis sp. nov. D S. flavonotatum Chujo E S. jungchani sp. nov." figureDoi="10.3897/zookeys.1185.112099.figure2" httpUri="https://binary.pensoft.net/fig/942952" pageId="0" pageNumber="1">2B</figureCitation>
) and
<taxonomicName id="5D3CF2FFF5B10A419937CC24DD171A8C" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="E1E3ECEE943C7F1BFF2E4860DAB27A96" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="C1F655C48803FFB99B3FC89D707E3416" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Sphaeroderma species in frontal view A S. hsui sp. nov. B S. changi sp. nov. C S. sheipaensis sp. nov. D S. flavonotatum Chujo E S. jungchani sp. nov." figureDoi="10.3897/zookeys.1185.112099.figure2" httpUri="https://binary.pensoft.net/fig/942952" pageId="0" pageNumber="1">2C</figureCitation>
)]; slender aedeagus, 4.0
<normalizedToken id="30DD9938575A020D7F7F80AE70F12AE4" originalValue="×">x</normalizedToken>
longer than wide, with cluster of setae at middle of inner margin in lateral view (Fig.
<figureCitation id="BA87496D2EA159D132B065B8418EECD8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4D</figureCitation>
) [wide aedeagus, 3.1
<normalizedToken id="313E569374E1C4762D82E36908B09F58" originalValue="×">x</normalizedToken>
longer than wide, and inner margin lacking setae in lateral view in
<taxonomicName id="57729FC23DB6FCF741AF62A439CAF7AA" lsidName="S. changi" pageId="0" pageNumber="1" rank="species" species="changi">
<emphasis id="E30C6CFF6D311210B4FE467319BF5059" italics="true" pageId="0" pageNumber="1">S. changi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="AF922FF79A624ADFDCD37973F764139F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6D</figureCitation>
)]; triangular abdominal ventrite VIII in females with apical margin lacking setae, subapically and moderately narrowed sides (Fig.
<figureCitation id="3CAF281EB48C336E6FB6DC15C1E150A3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4E</figureCitation>
) [apical margin with seven pairs of setae at medial part in
<taxonomicName id="9D345D1EED61DEDF53AE4B6BFD7CF071" lsidName="S. changi" pageId="0" pageNumber="1" rank="species" species="changi">
<emphasis id="8A5EC4A8E5A9FE6D7F2151613E6D3171" italics="true" pageId="0" pageNumber="1">S. changi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="9FA4D6552470E2C0D21DEE55932E3D5A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6E</figureCitation>
), subapically and slightly narrowed sides in
<taxonomicName id="CA7D36E42FD52ABA2E59BCE26DE46AE9" lsidName="S. changi" pageId="0" pageNumber="1" rank="species" species="changi">
<emphasis id="9EAB6152DFC4AFCC8410B0E5BF2A92EB" italics="true" pageId="0" pageNumber="1">S. changi</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="D9BBC2FB8A5FA30796B2E85DA37BB749" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="10D53DE865347A26100BE269857AE914" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov. (Figs
<figureCitation id="357673F8CBB675481B08D7BD063966B2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6E</figureCitation>
,
<figureCitation id="8269FE370784E1604C2B213665D1D65E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Sphaeroderma sheipaensis sp. nov. and S. jungchani Lee, sp. nov. A antenna, male, S. jungchani sp. nov. B antenna, female, Sphaeroderma sheipaensis sp. nov. C aedeagus, dorsal view, S. jungchani sp. nov. D ditto, lateral view E abdominal ventrite VIII, female, S. sheipaensis sp. nov. F spermatheca, S. sheipaensis sp. nov. G gonocoxae, S. sheipaensis sp. nov." figureDoi="10.3897/zookeys.1185.112099.figure7" httpUri="https://binary.pensoft.net/fig/942957" pageId="0" pageNumber="1">7E</figureCitation>
)]; transversely triangular gonocoxae (Fig.
<figureCitation id="EF66872AED8A4D485838991F05B7CA6D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4G</figureCitation>
) [longitudinally triangular gonocoxae in
<taxonomicName id="28B6417C337F95C760401490BFB55004" lsidName="S. changi" pageId="0" pageNumber="1" rank="species" species="changi">
<emphasis id="CD0EDA5714E6C6B6E812156B96B28FBE" italics="true" pageId="0" pageNumber="1">S. changi</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="AEB3FF9FD0BB8B25E63C92A17CFB79CC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6G</figureCitation>
)].
</paragraph>
<caption id="A6B108D479E752E0B780DDF2302A1125" doi="10.3897/zookeys.1185.112099.figure5" httpUri="https://binary.pensoft.net/fig/942955" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph id="16B5CFDF8F1F5B5EE7F2EE26918443CC" pageId="0" pageNumber="1">
<emphasis id="86D1370C1DA66325B0BE54D0EA74FBDC" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
Habitus of
<taxonomicName id="4626139300AA37C6E19E920D8250EE48" authorityName="Lee" authorityYear="2023" class="Dothideomycetes" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma changi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="changi">
<emphasis id="381DC7B4D97FE2C817333560FB9E4C6A" italics="true" pageId="0" pageNumber="1">Sphaeroderma changi</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="77ECDC9E13E10E9C233F4DCB32A1653C" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="8A78DF2E28ED5A9B55CFF79D7B649F4C" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="6995FFCD90171ECCA81AB8B26F90B491" bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName id="6D92C062F274673E28227FE673583935" lsidName="S. changi" pageId="0" pageNumber="1" rank="species" species="changi">
<emphasis id="1BB2983055604E46CD8797967CA6ACEE" italics="true" pageId="0" pageNumber="1">S. changi</emphasis>
</taxonomicName>
sp. nov., female, dorsal view
<emphasis id="D9854332C6C221D095165800C0FEE62C" bold="true" pageId="0" pageNumber="1">B</emphasis>
ditto, ventral view
<emphasis id="49AFF16A6FEB8B1C3C1CA27232A281A5" bold="true" pageId="0" pageNumber="1">C</emphasis>
ditto, lateral view
<emphasis id="633B8BE3FDE9584E0A0C06422F3B0E2E" bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName id="F06C0DDCAFD03BD130B6F4AE6087A54A" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="D1D9744AE1158FE314A1A183A4D6DA6A" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov., female, dorsal view
<emphasis id="F4205F6ADAF32AE5CFA2E10053C5E62B" bold="true" pageId="0" pageNumber="1">E</emphasis>
ditto, ventral view
<emphasis id="EA0730424CAABA4C6C949C07E320F229" bold="true" pageId="0" pageNumber="1">F</emphasis>
ditto, lateral view.
</paragraph>
</caption>
<caption id="456E2176A823B0C04C064AF4B2586DFF" doi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
<paragraph id="E0CB31D95B760B29162107C2530036D0" pageId="0" pageNumber="1">
<emphasis id="0F22CBD6BF788AC680CF0EBADED3472C" bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
Diagnostic characters of
<taxonomicName id="64DE4E216FDF44F5813917C23DAD5F43" authorityName="Lee" authorityYear="2023" class="Dothideomycetes" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma changi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="changi">
<emphasis id="23246FEAD8EDA391627B51E74B7C7136" italics="true" pageId="0" pageNumber="1">Sphaeroderma changi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="A95CD65B77E47C22570B3D4F17E4356A" bold="true" pageId="0" pageNumber="1">A</emphasis>
antenna, male
<emphasis id="6B8E4F002C0DE133FDA41D10B8C6EAB2" bold="true" pageId="0" pageNumber="1">B</emphasis>
antenna, female
<emphasis id="453FC37FC6CF7166F23704FD56E8F73B" bold="true" pageId="0" pageNumber="1">C</emphasis>
aedeagus, dorsal view
<emphasis id="88C8FC59961B680ACA193A42B336C34B" bold="true" pageId="0" pageNumber="1">D</emphasis>
aedeagus, lateral view
<emphasis id="EDAD18B379AC7591BBF9B71C7006BB52" bold="true" pageId="0" pageNumber="1">E</emphasis>
abdominal ventrite VIII, female
<emphasis id="62FF2C161BF5CC2AE9787F99A2AE8344" bold="true" pageId="0" pageNumber="1">F</emphasis>
spermatheca
<emphasis id="4CABF314DD39594BC622C04534B0CC48" bold="true" pageId="0" pageNumber="1">G</emphasis>
gonocoxae.
</paragraph>
</caption>
<caption id="CC1130357D60B667D74199B2FFBD8D0B" doi="10.3897/zookeys.1185.112099.figure7" httpUri="https://binary.pensoft.net/fig/942957" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph id="31E993E2F5816489A980AAE3F9D5E945" pageId="0" pageNumber="1">
<emphasis id="3681A5F460A15BFAEDADDDE5E2B2F318" bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
Diagnostic characters of
<taxonomicName id="4A77253FB4F4885ED69FB74A0B3D5EBA" authorityName="Lee" authorityYear="2023" class="Dothideomycetes" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma sheipaensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="sheipaensis">
<emphasis id="B9653C97B9581682B7E4C24BE2B81241" italics="true" pageId="0" pageNumber="1">Sphaeroderma sheipaensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="EE0D9926F8AB60186713E4882A5AE8AF" lsidName="S. jungchani" pageId="0" pageNumber="1" rank="species" species="jungchani">
<emphasis id="8F165BC54B43A7785D6AAE645ECC47D4" italics="true" pageId="0" pageNumber="1">S. jungchani</emphasis>
</taxonomicName>
Lee, sp. nov.
<emphasis id="0B468C51765A88A45CDAB50B2ABBDEA6" bold="true" pageId="0" pageNumber="1">A</emphasis>
antenna, male,
<taxonomicName id="CC02108E6573A2705F5C7AE7502DE9FF" lsidName="S. jungchani" pageId="0" pageNumber="1" rank="species" species="jungchani">
<emphasis id="1F77A9B1FE5798CB2470F08764F7B929" italics="true" pageId="0" pageNumber="1">S. jungchani</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="F304C6A9000DD9BBC23569817B4B00EE" bold="true" pageId="0" pageNumber="1">B</emphasis>
antenna, female,
<taxonomicName id="304E33C12C0069808146384AFFADB74A" authorityName="Lee" authorityYear="2023" class="Dothideomycetes" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma sheipaensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="sheipaensis">
<emphasis id="A868204DD788F1229AD2C4CD89F3C3D8" italics="true" pageId="0" pageNumber="1">Sphaeroderma sheipaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="F7B923BD21B64C1355F3FDF34CDE09EF" bold="true" pageId="0" pageNumber="1">C</emphasis>
aedeagus, dorsal view,
<taxonomicName id="3C5236DE2D757EA132107C954544A023" lsidName="S. jungchani" pageId="0" pageNumber="1" rank="species" species="jungchani">
<emphasis id="C895A6B33A3318A4B22AF57B18235EAB" italics="true" pageId="0" pageNumber="1">S. jungchani</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="CB2A01B592E7A5358678FBAFC5B3A617" bold="true" pageId="0" pageNumber="1">D</emphasis>
ditto, lateral view
<emphasis id="BEC9E4547DBC3D6385E7BE3966655F72" bold="true" pageId="0" pageNumber="1">E</emphasis>
abdominal ventrite VIII, female,
<taxonomicName id="31980A652290D5FEFC2A9E6517CDFF51" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="52060228454615578F502D8045486A03" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="3A37CFD491BEC455715E1B59BCD3435E" bold="true" pageId="0" pageNumber="1">F</emphasis>
spermatheca,
<taxonomicName id="54B98C28F882801815F4DAC7184BFFEA" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="BB7F0EE55ED990D52E260A20C4B13AEA" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="2B383BC9EFC72868310EDE59F70DFC64" bold="true" pageId="0" pageNumber="1">G</emphasis>
gonocoxae,
<taxonomicName id="72FD39201B1011FFEB1CD0894B874CA2" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="28465C518520C38767AD1DE5297C221D" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="6FF60A9F9D7D4DDFD1D7595FB32FB377" pageId="0" pageNumber="1" type="etymology">
<paragraph id="FC58610369607C21D9A3D418E00F900A" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="3E2B0F9564181E83DD389A9E544C87BC" pageId="0" pageNumber="1">The name is dedicated to Dr Yu-Feng Hsu (徐堉峰), who is the director for the insect survey project at Shei-Pa National Park.</paragraph>
</subSubSection>
<subSubSection id="44B29BB6B0A401ED610304B27D2BE5AE" pageId="0" pageNumber="1" type="distribution">
<paragraph id="2638DE3A45ACFC24129106FB5B3C40E3" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="A9F26AEFF547626A75FE6FC6C39C65BD" pageId="0" pageNumber="1">
<taxonomicName id="5BDD1CDA913F7BFB2F728D365C58B500" authorityName="Lee" authorityYear="2023" class="Dothideomycetes" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma hsui" order="Coleoptera" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="hsui">
<emphasis id="C5DB5D7CF9724DD66F6F712DA20C8CB2" italics="true" pageId="0" pageNumber="1">Sphaeroderma hsui</emphasis>
</taxonomicName>
sp. nov. seems to be the dominant species at the Shei-Pa National Park.
</paragraph>
</subSubSection>
</treatment>
</document>