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<document id="E9DC2C6C295A460741DB5F10C725FB4F" ID-CLB-Dataset="177057" ID-DOI="10.5852/ejt.2022.845.1955" ID-GBIF-Dataset="93834279-6660-4c8b-90af-74cde9c0568f" ID-ISSN="2118-9773" ID-Zenodo-Dep="7258792" ID-ZooBank="CC9F793D-5AD6-4D7C-A25C-7B2C58CAB4C0" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666888781492" checkinUser="felipe" docAuthor="Enghoff, Henrik" docDate="2022" docId="03AA87BFFFA6C972FDFDFED68BB54B9F" docLanguage="en" docName="ejt-845_66-100.pdf" docOrigin="European Journal of Taxonomy 845" docSource="http://dx.doi.org/10.5852/ejt.2022.845.1955" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cryptocorypha exovo Enghoff 2022, sp. nov." docType="treatment" docUuid="35365AB4-87E7-4510-A5F9-E506BCFECE06" docUuidSource="ZooBank" docVersion="7" lastPageNumber="81" masterDocId="FF93FFC7FFA9C97DFF9AFFDE882C4D0E" masterDocTitle="A mountain of millipedes X: Species of Pyrgodesmidae and Cryptodesmidae in the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)" masterLastPageNumber="100" masterPageNumber="66" pageNumber="81" updateTime="1699408525271" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="55000351358B0CFD899FE04D6DA1D379">A mountain of millipedes X: Species of Pyrgodesmidae and Cryptodesmidae in the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="762F2E0722463B42B1AE0F130D29FCA5">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="5CC04D4914F0BF90A7C825159F6602C3">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen, Denmark.</mods:affiliation>
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<mods:title id="AD6519660023E9371496176747F673FC">European Journal of Taxonomy</mods:title>
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<mods:date id="052510E9DCA68E74C5A8539CFCC60D35">2022</mods:date>
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<treatment id="03AA87BFFFA6C972FDFDFED68BB54B9F" ID-DOI="http://doi.org/10.5281/zenodo.7258802" ID-GBIF-Taxon="203721834" ID-Zenodo-Dep="7258802" LSID="urn:lsid:zoobank.org:act:35365AB4-87E7-4510-A5F9-E506BCFECE06" httpUri="http://treatment.plazi.org/id/03AA87BFFFA6C972FDFDFED68BB54B9F" lastPageNumber="81" pageId="15" pageNumber="81">
<subSubSection id="C3196522FFA6C972FDFDFED68BE14C2D" box="[615,973,264,291]" pageId="15" pageNumber="81" type="nomenclature">
<paragraph id="8BBC36A9FFA6C972FDFDFED68BE14C2D" blockId="15.[368,1219,264,361]" box="[615,973,264,291]" pageId="15" pageNumber="81">
<heading id="D0F481C5FFA6C972FDFDFED68BE14C2D" box="[615,973,264,291]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="81" reason="2">
<taxonomicName id="4C034D2AFFA6C972FDFDFED68B474C2D" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[615,875,264,291]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="81" phylum="Arthropoda" rank="species" species="exovo" status="sp. nov.">
<emphasis id="B977EABBFFA6C972FDFDFED68B474C2D" bold="true" box="[615,875,264,291]" italics="true" pageId="15" pageNumber="81">Cryptocorypha exovo</emphasis>
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<taxonomicNameLabel id="A24457C0FFA6C972FCE8FED78BE14C2D" box="[882,973,265,291]" pageId="15" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BBC36A9FFA6C972FD43FE918B764C67" blockId="15.[368,1219,264,361]" box="[729,858,335,361]" pageId="15" pageNumber="81">
<figureCitation id="13382A2CFFA6C972FD43FE918B764C67" box="[729,858,335,361]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="16.[189,232,1857,1883]" captionStartId-1="17.[189,232,1893,1919]" captionTargetBox-0="[189,1398,303,1818]" captionTargetBox-1="[215,1372,265,1855]" captionTargetId-0="figure-12@16.[189,1398,303,1818]" captionTargetId-1="figure-14@17.[215,1372,265,1855]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="Fig. 10. Cryptocorypha exovo sp. nov., holotype (NHMD 621840), ♂. A. Head, frontal view. B. Collum and rings 23, dorsal view. C. Rings 23, dorsal view. D. Rings 16 (part), 17, 18 (apodous) and telson, oblique dorsal view, showing setae (long arrows) and a probably abraded seta (long stippled arrow) on posterior row of tubercles on apodous ring, and setae (short arrows) on paranota of apodous ring. E. Labrum and second coxae. Scale bars: AC = 0.1 mm; DE = 0.02 mm." captionText-1="Fig. 11. Cryptocorypha exovo sp. nov., holotype (NHMD 621840), ♂, right gonopod. A. Lateral view. B. Posterior view. C. Mesal view. D. Anterior view. E. Close-up of circled area in D, showing cannula. Abbreviations: bt = basal part of telopodite; ca = cannula; cx = coxa; de = denticles; ex = exomere; slm = solenomere; x = dirt, not part of the gonopod. Scale bars: AD = 0.02 mm; E = 0.01 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7258820" figureDoi-1="http://doi.org/10.5281/zenodo.7258824" httpUri-0="https://zenodo.org/record/7258820/files/figure.png" httpUri-1="https://zenodo.org/record/7258824/files/figure.png" pageId="15" pageNumber="81">Figs 1011</figureCitation>
</paragraph>
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<subSubSection id="C3196522FFA6C972FF27FE4E8B594CF2" pageId="15" pageNumber="81" type="diagnosis">
<paragraph id="8BBC36A9FFA6C972FF27FE4E891B4CA4" blockId="15.[189,311,400,426]" box="[189,311,400,426]" pageId="15" pageNumber="81">
<heading id="D0F481C5FFA6C972FF27FE4E891B4CA4" bold="true" box="[189,311,400,426]" fontSize="11" level="3" pageId="15" pageNumber="81" reason="3">
<emphasis id="B977EABBFFA6C972FF27FE4E891B4CA4" bold="true" box="[189,311,400,426]" pageId="15" pageNumber="81">Diagnosis</emphasis>
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Differs from all congeneric species by the unique shape of the snakelike gonopod telopodite. Agrees with
<taxonomicName id="4C034D2AFFA6C972FF62FE3D899A4CF2" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[248,438,483,509]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="81" phylum="Arthropoda" rank="species" species="geminiramus" status="sp. nov.">
<emphasis id="B977EABBFFA6C972FF62FE3D899A4CF2" box="[248,438,483,509]" italics="true" pageId="15" pageNumber="81">C. geminiramus</emphasis>
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<taxonomicNameLabel id="A24457C0FFA6C972FE27FE3D8A344CF3" box="[445,536,483,509]" pageId="15" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
in non-gonopodal characters.
</paragraph>
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<subSubSection id="C3196522FFA6C972FF27FDFA8BA84F9E" pageId="15" pageNumber="81" type="etymology">
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<heading id="D0F481C5FFA6C972FF27FDFA89694F30" bold="true" box="[189,325,548,574]" fontSize="11" level="3" pageId="15" pageNumber="81" reason="3">
<emphasis id="B977EABBFFA6C972FF27FDFA89694F30" bold="true" box="[189,325,548,574]" pageId="15" pageNumber="81">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC36A9FFA6C972FF27FD8C8BA84F9E" blockId="15.[189,1399,594,656]" pageId="15" pageNumber="81">The species epithet means out of the egg and refers to the shape of the gonopod which somewhat resembles a snake (telopodite) hatching from the egg (coxa).</paragraph>
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<subSubSection id="C3196522FFA6C972FF27FD6989314E67" pageId="15" pageNumber="81" type="materials_examined">
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<emphasis id="B977EABBFFA6C972FF27FD6989814FDF" bold="true" box="[189,429,695,721]" pageId="15" pageNumber="81">Material examined</emphasis>
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<materialsCitation id="3B6B3CF4FFA6C972FF70FD3889314E67" ID-GBIF-Occurrence="3948948346" collectingDate="2013-12-16" collectingMethod="pitfall trap" collectionCode="T" collectorName="T. Pape &amp; N. Scharff" country="Tanzania" county="Udzungwa Mountains Natl Park" elevation="1207" latitude="-7.8373055" location="Mito Mitatu" longLatPrecision="1" longitude="36.846333" municipality="Udzungwa Mountains" pageId="15" pageNumber="81" specimenCode="NHMD 621840" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="holotype">
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<typeStatus id="54B8880BFFA6C972FF70FD3889714E0E" box="[234,349,742,768]" pageId="15" pageNumber="81" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3147639FFA6C972FF27FCD7897D4E2D" box="[189,337,777,803]" name="Tanzania" pageId="15" pageNumber="81">TANZANIA</collectingCountry>
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;
<collectingRegion id="49C7F84BFFA6C972FE06FCD78A5A4E2D" box="[412,630,777,803]" country="Tanzania" name="Morogoro" pageId="15" pageNumber="81">Morogoro Region</collectingRegion>
,
<collectingCounty id="62DD4E25FFA6C972FD1CFCD78C3D4E2D" box="[646,1041,777,803]" pageId="15" pageNumber="81">Udzungwa Mountains Natl Park</collectingCounty>
,
<location id="8EDC6072FFA6C972FBBAFCD78C9B4E2D" LSID="urn:lsid:plazi:treatment:03AA87BFFFA6C972FDFDFED68BB54B9F:8EDC6072FFA6C972FBBAFCD78C9B4E2D" box="[1056,1207,777,803]" country="Tanzania" county="Udzungwa Mountains Natl Park" latitude="-7.8373055" longLatPrecision="1" longitude="36.846333" municipality="Udzungwa Mountains" name="Mito Mitatu" pageId="15" pageNumber="81" stateProvince="Morogoro Region">Mito Mitatu</location>
;
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,
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;
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<quantity id="4CFB9B4CFFA6C972FEEBFCF289E64E49" box="[369,458,812,839]" metricMagnitude="3" metricUnit="m" metricValue="1.207" pageId="15" pageNumber="81" unit="m" value="1207.0">1207 m</quantity>
a.s.l.
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;
<date id="FFBD1069FFA6C972FD8DFCF28A9A4E48" box="[535,694,812,839]" pageId="15" pageNumber="81" value="2013-12-16">
<collectingDate id="EFF9E981FFA6C972FD8DFCF28A9A4E48" box="[535,694,812,839]" pageId="15" pageNumber="81" value="2013-12-16">16 Dec. 2013</collectingDate>
</date>
;
<collectorName id="26F6537FFFA6C972FD58FCF38B354E49" box="[706,793,813,839]" pageId="15" pageNumber="81">T. Pape</collectorName>
and
<collectorName id="26F6537FFFA6C972FCCBFCF38BE74E48" box="[849,971,812,839]" pageId="15" pageNumber="81">N. Scharff</collectorName>
leg.;
<collectingMethod id="52424EBEFFA6C972FB96FCF28CA64E49" box="[1036,1162,812,839]" pageId="15" pageNumber="81">pitfall trap</collectingMethod>
, Plot MM1;
<specimenCode id="DBA59ED2FFA6C972FA87FCF389354E67" collectionCode="NHMD" pageId="15" pageNumber="81">NHMD 621840</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3196522FFA6C972FF27FC4E8A34485C" pageId="15" pageNumber="81" type="description">
<paragraph id="8BBC36A9FFA6C972FF27FC4E89634EA4" blockId="15.[189,335,912,938]" box="[189,335,912,938]" pageId="15" pageNumber="81">
<heading id="D0F481C5FFA6C972FF27FC4E89634EA4" bold="true" box="[189,335,912,938]" fontSize="11" level="3" pageId="15" pageNumber="81" reason="3">
<emphasis id="B977EABBFFA6C972FF27FC4E89634EA4" bold="true" box="[189,335,912,938]" pageId="15" pageNumber="81">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC36A9FFA6C972FF27FC618A104EF2" blockId="15.[189,1399,959,1021]" pageId="15" pageNumber="81">
SIZE. Male: Length
<quantity id="4CFB9B4CFFA6C972FE3FFC618A2F4ED4" box="[421,515,959,986]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="15" pageNumber="81" unit="mm" value="5.3">5.3 mm</quantity>
; maximal body width (across metazona)
<quantity id="4CFB9B4CFFA6C972FC70FC618C7A4ED4" box="[1002,1110,959,986]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="15" pageNumber="81" unit="mm" value="0.78">0.78 mm</quantity>
; 17 podous + 1 apodous ring + telson. Female: unknown.
</paragraph>
<paragraph id="8BBC36A9FFA6C972FF27FBFA8CC34930" blockId="15.[189,1263,1060,1086]" box="[189,1263,1060,1086]" pageId="15" pageNumber="81">
OTHER NON- GONOPODAL CHARACTERS (
<figureCitation id="13382A2CFFA6C972FDF3FBFA8AED4930" box="[617,705,1060,1086]" captionStart="Fig" captionStartId="16.[189,232,1857,1883]" captionTargetBox="[189,1398,303,1818]" captionTargetId="figure-12@16.[189,1398,303,1818]" captionTargetPageId="16" captionText="Fig. 10. Cryptocorypha exovo sp. nov., holotype (NHMD 621840), ♂. A. Head, frontal view. B. Collum and rings 23, dorsal view. C. Rings 23, dorsal view. D. Rings 16 (part), 17, 18 (apodous) and telson, oblique dorsal view, showing setae (long arrows) and a probably abraded seta (long stippled arrow) on posterior row of tubercles on apodous ring, and setae (short arrows) on paranota of apodous ring. E. Labrum and second coxae. Scale bars: AC = 0.1 mm; DE = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.7258820" httpUri="https://zenodo.org/record/7258820/files/figure.png" pageId="15" pageNumber="81">Fig. 10</figureCitation>
). Virtually identical to
<taxonomicName id="4C034D2AFFA6C972FC4AFBFA8CA14930" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[976,1165,1060,1086]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="81" phylum="Arthropoda" rank="species" species="geminiramus" status="sp. nov.">
<emphasis id="B977EABBFFA6C972FC4AFBFA8CA14930" box="[976,1165,1060,1086]" italics="true" pageId="15" pageNumber="81">C. geminiramus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA6C972FB0EFBFA8CC34930" box="[1172,1263,1060,1086]" pageId="15" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BBC36A9FFA6C972FF27FBBB8A34485C" blockId="15.[188,1400,1125,1363]" pageId="15" pageNumber="81">
GONOPODS (
<figureCitation id="13382A2CFFA6C972FED9FBBB89B44971" box="[323,408,1125,1151]" captionStart="Fig" captionStartId="17.[189,232,1893,1919]" captionTargetBox="[215,1372,265,1855]" captionTargetId="figure-14@17.[215,1372,265,1855]" captionTargetPageId="17" captionText="Fig. 11. Cryptocorypha exovo sp. nov., holotype (NHMD 621840), ♂, right gonopod. A. Lateral view. B. Posterior view. C. Mesal view. D. Anterior view. E. Close-up of circled area in D, showing cannula. Abbreviations: bt = basal part of telopodite; ca = cannula; cx = coxa; de = denticles; ex = exomere; slm = solenomere; x = dirt, not part of the gonopod. Scale bars: AD = 0.02 mm; E = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.7258824" httpUri="https://zenodo.org/record/7258824/files/figure.png" pageId="15" pageNumber="81">Fig. 11</figureCitation>
). Coxa (
<emphasis id="B977EABBFFA6C972FE60FBB88A384971" box="[506,532,1126,1151]" italics="true" pageId="15" pageNumber="81">cx</emphasis>
) moderately globose, its surface scaly and with scattered level 3 spherules, mesally forming a shallow gonocoel. Cannula (
<figureCitation id="13382A2CFFA6C972FD7EFB568B4249AD" box="[740,878,1160,1187]" captionStart="Fig" captionStartId="17.[189,232,1893,1919]" captionTargetBox="[215,1372,265,1855]" captionTargetId="figure-14@17.[215,1372,265,1855]" captionTargetPageId="17" captionText="Fig. 11. Cryptocorypha exovo sp. nov., holotype (NHMD 621840), ♂, right gonopod. A. Lateral view. B. Posterior view. C. Mesal view. D. Anterior view. E. Close-up of circled area in D, showing cannula. Abbreviations: bt = basal part of telopodite; ca = cannula; cx = coxa; de = denticles; ex = exomere; slm = solenomere; x = dirt, not part of the gonopod. Scale bars: AD = 0.02 mm; E = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.7258824" httpUri="https://zenodo.org/record/7258824/files/figure.png" pageId="15" pageNumber="81">Fig. 11C, E</figureCitation>
:
<emphasis id="B977EABBFFA6C972FCE3FB578BB949AC" box="[889,917,1161,1186]" italics="true" pageId="15" pageNumber="81">ca</emphasis>
, not seen in the other two species) with a few tiny denticles (
<emphasis id="B977EABBFFA6C972FE07FB72899549C8" box="[413,441,1196,1222]" italics="true" pageId="15" pageNumber="81">de</emphasis>
).Telopodite projecting strongly from gonocoel, very evident in lateral and ventral views. Basal (prefemoral) part of telopodite (
<emphasis id="B977EABBFFA6C972FD74FB118B2949E7" box="[750,773,1231,1257]" italics="true" pageId="15" pageNumber="81">bt</emphasis>
) massive, with several distolateral setae, giving rise to a very long, sinous shaft which slightly beyond its midlength divides into a slightly curved branch, probably corresponding to an exomere (
<emphasis id="B977EABBFFA6C972FD03FAC88A9F4821" box="[665,691,1302,1327]" italics="true" pageId="15" pageNumber="81">ex</emphasis>
), and a strongly S-shaped branch, probably the solenomere (
<emphasis id="B977EABBFFA6C972FF5DFAE688DC485C" box="[199,240,1336,1362]" italics="true" pageId="15" pageNumber="81">slm</emphasis>
). A velum is not evident.
</paragraph>
</subSubSection>
<subSubSection id="C3196522FFA6C972FF27FAA48BB54B9F" pageId="15" pageNumber="81" type="discussion">
<paragraph id="8BBC36A9FFA6C972FF27FAA4891C489A" blockId="15.[189,304,1402,1428]" box="[189,304,1402,1428]" pageId="15" pageNumber="81">
<heading id="D0F481C5FFA6C972FF27FAA4891C489A" bold="true" box="[189,304,1402,1428]" fontSize="11" level="3" pageId="15" pageNumber="81" reason="3">
<emphasis id="B977EABBFFA6C972FF27FAA4891C489A" bold="true" box="[189,304,1402,1428]" pageId="15" pageNumber="81">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC36A9FFA6C972FF27FA77893448E8" blockId="15.[189,1398,1448,1510]" pageId="15" pageNumber="81">
A male of
<taxonomicName id="4C034D2AFFA6C972FEAFFA7789DE48CC" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[309,498,1449,1475]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="81" phylum="Arthropoda" rank="species" species="geminiramus" status="sp. nov.">
<emphasis id="B977EABBFFA6C972FEAFFA7789DE48CC" box="[309,498,1449,1475]" italics="true" pageId="15" pageNumber="81">C. geminiramus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA6C972FE63FA778A7F48CD" box="[505,595,1449,1475]" pageId="15" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
was found in the same pitfall trap as the unique
<typeStatus id="54B8880BFFA6C972FB19FA768CC648CC" box="[1155,1258,1448,1474]" pageId="15" pageNumber="81" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C034D2AFFA6C972FA95FA778D5A48CC" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[1295,1398,1449,1475]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="81" phylum="Arthropoda" rank="species" species="exovo" status="sp. nov.">
<emphasis id="B977EABBFFA6C972FA95FA778D5A48CC" box="[1295,1398,1449,1475]" italics="true" pageId="15" pageNumber="81">C. exovo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA6C972FF27FA12893448E8" box="[189,280,1484,1510]" pageId="15" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BBC36A9FFA6C972FF27F9D38BB54B9F" blockId="15.[189,1398,1549,1681]" pageId="15" pageNumber="81">
If the Afrotropical alternative in the first couplet of the key provided by
<bibRefCitation id="EF924B58FFA6C972FBBFF9D38CDB4B29" author="Golovatch S. I." box="[1061,1271,1549,1575]" pageId="15" pageNumber="81" pagination="179 - 190" refId="ref14226" refString="Golovatch S. I. 2019. The millipede genus Cryptocorypha Attems, 1907 revisited, with descriptions of two new Oriental species. Arthropoda Selecta 28: 179 - 190." type="journal article" year="2019">Golovatch (2019)</bibRefCitation>
is chosen,
<taxonomicName id="4C034D2AFFA6C972FF27F9EF89044B44" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[189,296,1585,1611]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="81" phylum="Arthropoda" rank="species" species="exovo" status="sp. nov.">
<emphasis id="B977EABBFFA6C972FF27F9EF89044B44" box="[189,296,1585,1611]" italics="true" pageId="15" pageNumber="81">C. exovo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA6C972FEA9F9EF89BD4B45" box="[307,401,1585,1611]" pageId="15" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
runs to
<taxonomicName id="4C034D2AFFA6C972FE60F9EF8B734B44" authority="(Brolemann, 1920)" baseAuthorityName="Brolemann" baseAuthorityYear="1920" box="[506,863,1584,1611]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="81" phylum="Arthropoda" rank="species" species="diffusa">
<emphasis id="B977EABBFFA6C972FE60F9EF8A5E4B44" box="[506,626,1584,1611]" italics="true" pageId="15" pageNumber="81">C. diffusa</emphasis>
(Brolemann, 1920)
</taxonomicName>
from which it differs by several characters, especially by strongly different gonopods. If the Asian alternative in the first couplet is chosen,
<taxonomicName id="4C034D2AFFA6C972FA8AF98A8D5A4B63" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[1296,1398,1620,1646]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="81" phylum="Arthropoda" rank="species" species="exovo" status="sp. nov.">
<emphasis id="B977EABBFFA6C972FA8AF98A8D5A4B63" box="[1296,1398,1620,1646]" italics="true" pageId="15" pageNumber="81">C. exovo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA6C972FF27F9A989354B9F" box="[189,281,1655,1681]" pageId="15" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
runs to couplet 17(18) where it fits neither alternative.
</paragraph>
</subSubSection>
</treatment>
</document>