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<document id="1A0DB1D4D030363B9A01A9F9E7D52B31" ID-DOI="10.5281/zenodo.169638" ID-GBIF-Dataset="3ff15714-6d5c-4265-aff2-c7a19041a761" ID-ISSN="1175­5326" ID-Zenodo-Dep="169638" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461038850950" checkinUser="plazi" docAuthor="Springer, Victor G. &amp; Allen, Gerald R." docDate="2004" docId="E8605F443636FFFFFEBDEB8EDAD16D3D" docLanguage="en" docName="zt00791.pdf" docOrigin="Zootaxa 791" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Ecsenius caeruliventris Springer &amp; Allen, 2004, new species" docType="treatment" docUuid="6515A914-D516-4C35-959F-8310DAE39262" docUuidSource="ZooBank" docVersion="10" lastPageNumber="5" masterDocId="1459273C3634FFFBFFB5ED17DB5B6A73" masterDocTitle="Ecsenius caeruliventris and E. shirleyae, two new species of blenniid fishes from Indonesia, and new distribution records for other species of Ecsenius" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1698575124870" updateUser="plazi">
<mods:mods id="27686A68B838234E4822E58504B00A90" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="8DFB48DCDDAEB28BE3280311C6A63578">Ecsenius caeruliventris and E. shirleyae, two new species of blenniid fishes from Indonesia, and new distribution records for other species of Ecsenius</mods:title>
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<mods:name id="DB3431421C01856CAF57FB606E29B6D2" type="personal">
<mods:role id="3E49E7D1CD5EBA7CC971D5E4C1A617FD">
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<mods:namePart id="9E683A41EFA2EA897066623716F042D4">Springer, Victor G.</mods:namePart>
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<mods:namePart id="67FFCC72D427D39541456013ED913D7B">Allen, Gerald R.</mods:namePart>
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<mods:title id="DC52A6A1C7982C1F3E3DE40FC93AA97D">Zootaxa</mods:title>
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<mods:part id="026948374E78334B05B4EE9F99268BF7">
<mods:date id="E03A0EB09B571B5398B9322E55AFCE11">2004</mods:date>
<mods:detail id="CBCAEDACCAD3A7963F411988FC121FDC" type="volume">
<mods:number id="B1CE3B9EC9DBF8AC4B94F894CCC6D527">791</mods:number>
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<mods:classification id="00E570BDB893CA894F1FBE0B031198DA">journal article</mods:classification>
<mods:identifier id="A617909CD0CFAB4AFFE2F73F522077F0" type="DOI">10.5281/zenodo.169638</mods:identifier>
<mods:identifier id="F57245B9B71F559D5C1E16E301BBE900" type="GBIF-Dataset">3ff15714-6d5c-4265-aff2-c7a19041a761</mods:identifier>
<mods:identifier id="710C35B6A6F1E3842CBC6648B58715CF" type="ISSN">1175­5326</mods:identifier>
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<treatment id="E8605F443636FFFFFEBDEB8EDAD16D3D" ID-DOI="http://doi.org/10.5281/zenodo.6269360" ID-GBIF-Taxon="119609372" ID-Zenodo-Dep="6269360" LSID="urn:lsid:zoobank.org:act:6515A914-D516-4C35-959F-8310DAE39262" httpUri="http://treatment.plazi.org/id/E8605F443636FFFFFEBDEB8EDAD16D3D" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<subSubSection id="28D3BDD93636FFF9FEBDEB8EDA8A6CA5" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="6076EE523636FFF9FEBDEB8ED99A6CC0" blockId="2.[264,705,1689,1790]" box="[264,705,1689,1715]" pageId="2" pageNumber="3">
<heading id="3B3E593E3636FFF9FEBDEB8ED99A6CC0" bold="true" box="[264,705,1689,1715]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="52BD32403636FFF9FEBDEB8ED99A6CC0" bold="true" box="[264,705,1689,1715]" pageId="2" pageNumber="3">
<taxonomicName id="A7C995D13636FFF9FEBDEB8ED9796CC0" ID-CoL="38NYG" box="[264,546,1689,1715]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="caeruliventris" status="sp. nov.">
<emphasis id="52BD32403636FFF9FEBDEB8ED9796CC0" box="[264,546,1689,1715]" italics="true" pageId="2" pageNumber="3">Ecsenius caeruliventris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="498E8F3B3636FFF9FD84EB8ED99A6CC0" box="[561,705,1689,1715]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6076EE523636FFF9FEBDEBABDA8A6CA5" blockId="2.[264,705,1689,1790]" box="[264,465,1724,1750]" pageId="2" pageNumber="3">Bluebelly blenny</paragraph>
</subSubSection>
<subSubSection id="28D3BDD93636FFF9FEBDEBF3D9356C8D" box="[264,622,1764,1790]" pageId="2" pageNumber="3" type="description">
<paragraph id="6076EE523636FFF9FEBDEBF3D9356C8D" blockId="2.[264,705,1689,1790]" box="[264,622,1764,1790]" pageId="2" pageNumber="3">
(
<figureCitation id="F8F2F2D73636FFF9FEA5EBF3DA276C8D" box="[272,380,1764,1790]" captionStart="FIGURE 1" captionStartId="5.[264,368,1694,1718]" captionTargetBox="[312,1295,334,1610]" captionTargetId="figure@5.[264,1323,309,1627]" captionTargetPageId="5" captionText="FIGURE 1. Photographs of preserved specimens of three similar Ecsenius species. a – c, Ecsenius caeruliventris: a, b, MZB 12094, holotype, male, 24.3 mm SL (b, ventral surface of head); c, USNM 379435, female, 23.0 mm SL, ventral surface of head (note pale spots). d, Ecsenius shirleyae, USNM 211996, holotype, male, 28.4 mm SL; e, Ecsenius bandanus, USNM 221236, male, 31.7 mm SL, Biak, Irian Jaya, Indonesia." httpUri="https://zenodo.org/record/169639/files/figure.png" pageId="2" pageNumber="3">Figures 1</figureCitation>
a–c, 2 a, 3; Table 1)
</paragraph>
</subSubSection>
<subSubSection id="28D3BDD93636FFF9FEBDEA3DDF426D13" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="6076EE523636FFF9FEBDEA3DDF426D13" blockId="2.[264,1243,1833,1889]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="40D9C97C3636FFF9FEBDEA3DDF426D13" pageId="2" pageNumber="3">
<taxonomicName id="A7C995D13636FFF9FEBDEA3DDA8F6D32" ID-CoL="38NYB" box="[264,468,1834,1857]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="bandanus">
<emphasis id="52BD32403636FFF9FEBDEA3DDA8F6D32" box="[264,468,1834,1857]" italics="true" pageId="2" pageNumber="3">Ecsenius bandanus</emphasis>
</taxonomicName>
(not Springer), in part,
<treatmentCitation id="E168C8433636FFF9FD64EA3EDF486D33" author="Kuiter" box="[721,1043,1833,1856]" page="605" pageId="2" pageNumber="3" year="2001">
<bibRefCitation id="045893A33636FFF9FD64EA3ED8BD6D33" author="Kuiter" box="[721,998,1833,1856]" pageId="2" pageNumber="12" refString="Kuiter, R. H. &amp; Tonozuka, T. (2001) Pictorial Guide to Indonesian Reef Fishes. Zoonetics, Seaford, Victoria, Australia, 893 pp. Second edition, 2004, PT Dive &amp; Dive's, Bali, Indonesia, same pagination." type="book" year="2001">Kuiter &amp; Tonozuka, 2001</bibRefCitation>
:605
</treatmentCitation>
(Sangihe Islands).
<emphasis id="52BD32403636FFF9FEBDEA5DDA446D12" box="[264,287,1866,1889]" italics="true" pageId="2" pageNumber="3">E.</emphasis>
sp.
<treatmentCitation id="E168C8433636FFF9FEFEEA5ED9086D13" author="Allen" box="[331,595,1865,1888]" page="54" pageId="2" pageNumber="3" year="2003">
<bibRefCitation id="045893A33636FFF9FEFEEA5ED9686D13" author="Allen" box="[331,563,1865,1888]" pageId="2" pageNumber="12" refString="Allen, G. R. &amp; Adrim, M. (2003) Coral-reef fishes of Indonesia. Zoological Studies, 42 (1), 1 - 72." type="journal article" year="2003">Allen &amp; Adrim, 2003</bibRefCitation>
:54
</treatmentCitation>
&amp; 19, fig. 71 (Togean &amp; Sangihe islands).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="28D3BDD93637FFF8FEBDEC0BDFEF6BDD" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="6076EE523637FFF8FEBDEC0BDFEF6BDD" blockId="3.[264,1323,284,670]" pageId="3" pageNumber="4">
<typeStatus id="BF7250F03637FFF8FEBDEC0BDA286B46" box="[264,371,284,309]" pageId="3" pageNumber="4" type="holotype">
<emphasis id="52BD32403637FFF8FEBDEC0BDA286B46" box="[264,371,284,309]" italics="true" pageId="3" pageNumber="4">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="06D876973637FFF8FE3FEC0BDA916B45" box="[394,458,284,310]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="4">MZB</collectionCode>
12094, male,
<quantity id="A73143B73637FFF8FD33EC0BD9AD6B45" box="[646,758,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="3" pageNumber="4" unit="mm" value="24.3">24.3 mm</quantity>
SL, S entrance to Batudak Passage, Togean Islands, Sulawesi,
<collectingCountry id="18DEAEC23637FFF8FE6EEC53D9166B2D" box="[475,589,324,350]" name="Indonesia" pageId="3" pageNumber="4">Indonesia</collectingCountry>
(00.27’S,
<geoCoordinate id="05FD88953637FFF8FD08EC53D8096B2D" box="[701,850,322,350]" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="9" value="121.93633">121°56.18’E</geoCoordinate>
,
<quantity id="A73143B73637FFF8FCEEEC53D8F86B2D" box="[859,931,324,350]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="4" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">3–5 m</quantity>
,quinaldine, G. Allen,
<date id="1477C8923637FFF8FB2FEC53DE7D6B2D" box="[1178,1318,324,350]" pageId="3" pageNumber="4" value="1998-10-31">31Oct 1998</date>
.
<typeStatus id="BF7250F03637FFF8FEBDEC7BDA256BF6" box="[264,382,364,389]" pageId="3" pageNumber="4" type="paratype">
<emphasis id="52BD32403637FFF8FEBDEC7BDA256BF6" box="[264,382,364,389]" italics="true" pageId="3" pageNumber="4">Paratypes</emphasis>
</typeStatus>
(five specimens, all with same collection data as
<typeStatus id="BF7250F03637FFF8FC70EC7BDF696BF5" box="[965,1074,364,390]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
):
<collectionCode id="06D876973637FFF8FBF0EC7BDFD76BF5" LSID="urn:lsid:biocol.org:col:34981" box="[1093,1164,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="3" pageNumber="4">WAM</collectionCode>
P 31492­001 (
<specimenCount id="76CF25DB3637FFF8FEA4EC83DADD6BDD" box="[273,390,404,430]" pageId="3" pageNumber="4" type="female">2 females</specimenCount>
, 22.8–23.0;
<specimenCount id="76CF25DB3637FFF8FDA2EC83D92E6BDD" box="[535,629,404,430]" pageId="3" pageNumber="4" type="male">2 males</specimenCount>
, 15.6–20.1) and
<collectionCode id="06D876973637FFF8FC83EC83D8D06BDD" LSID="urn:lsid:biocol.org:col:34871" box="[822,907,404,430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="4">USNM</collectionCode>
379435 (
<specimenCount id="76CF25DB3637FFF8FC48EC83DF3F6BDD" box="[1021,1124,404,430]" pageId="3" pageNumber="4" type="female">1 female</specimenCount>
, 23.7).
</paragraph>
</subSubSection>
<subSubSection id="28D3BDD93637FFF8FE8DECABDA8E68ED" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="6076EE523637FFF8FE8DECABDA8E68ED" blockId="3.[264,1323,284,670]" pageId="3" pageNumber="4">
<emphasis id="52BD32403637FFF8FE8DECABDAED6BA6" box="[312,438,444,469]" italics="true" pageId="3" pageNumber="4">Diagnosis.</emphasis>
A species of the
<taxonomicName id="A7C995D13637FFF8FD31ECABD9A96BA5" box="[644,754,444,470]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="9" phylum="Chordata" rank="species" species="prooculis">Prooculis</taxonomicName>
group of
<taxonomicName id="A7C995D13637FFF8FCDCECABD88A6BA6" box="[873,977,444,469]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
<emphasis id="52BD32403637FFF8FCDCECABD88A6BA6" box="[873,977,444,469]" italics="true" pageId="3" pageNumber="4">Ecsenius</emphasis>
</taxonomicName>
with the following combination of characters: Body without alternating dark and pale stripes or distinct dark spots; dark stripe extending posteriorly from orbital margin across head and variably for short distance onto body anteriorly. In life, dark post­orbital stripe bordered ventrally by bright white stripe; other bright markings on head, white with at most a very faint tinge of yellow; belly bluish.
</paragraph>
</subSubSection>
<paragraph id="6076EE523637FFF8FEBDEFF1DE7A6EEF" pageId="3" pageNumber="4">
<table id="12C91CF236370004FEBDEFF1DE706EEE" box="[264,1323,742,1181]" gridcols="3" gridrows="12" pageId="3" pageNumber="4">
<tr id="DEF9EC1036370004FEBDEFF1DE70688D" box="[264,1323,742,766]" gridrow="0" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDEFF1D9E7688D" box="[264,700,742,766]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="52BD32403637FFF8FEBDEFF1DA25688D" bold="true" box="[264,382,742,766]" pageId="3" pageNumber="4">TABLE 1.</emphasis>
Frequency distributions for
</th>
<th id="9D28856C36370004FD73EFF1DF41688D" box="[710,1050,742,766]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">
selected characters in
<taxonomicName id="A7C995D13637FFF8FC0EEFF0DF41688D" box="[955,1050,743,766]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
<emphasis id="52BD32403637FFF8FC0EEFF0DF41688D" box="[955,1050,743,766]" italics="true" pageId="3" pageNumber="4">Ecsenius</emphasis>
</taxonomicName>
</th>
<th id="9D28856C36370004FB91EFF1DE70688D" box="[1060,1323,742,766]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">
species of the
<taxonomicName id="A7C995D13637FFF8FB73EFF1DE70688E" box="[1222,1323,742,765]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="9" phylum="Chordata" rank="species" species="prooculis">Prooculis</taxonomicName>
</th>
</tr>
<tr id="DEF9EC1036370004FEBDEE1ADE706957" box="[264,1323,781,804]" gridrow="1" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1">
<th id="9D28856C36370004FEBDEE1AD9E76957" box="[264,700,781,804]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">group.</th>
</tr>
<tr id="DEF9EC1036370004FEBDEE54DE70692B" box="[264,1323,835,856]" gridrow="2" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDEE54D9E7692B" box="[264,700,835,856]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">Species Segmented dorsal­fin rays</th>
<td id="9D28856C36370004FD73EE54DF41692B" box="[710,1050,835,856]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">Segmented anal­fin rays</td>
<td id="9D28856C36370004FB91EE54DE70692B" box="[1060,1323,835,856]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">Caudal vertebrae</td>
</tr>
<tr id="DEF9EC1036370004FEBDEE70DE70690F" box="[264,1323,871,892]" gridrow="3" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDEE70D9E7690F" box="[264,700,871,892]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">13 14 15</th>
<td id="9D28856C36370004FD73EE70DF41690F" box="[710,1050,871,892]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">15 16 17</td>
<td id="9D28856C36370004FB91EE70DE70690F" box="[1060,1323,871,892]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">21 22 23</td>
</tr>
<tr id="DEF9EC1036370004FEBDEE99DE7069D0" box="[264,1323,910,931]" gridrow="4" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDEE99D9E769D0" box="[264,700,910,931]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9EE99DACA69D0" box="[268,401,910,931]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="caeruliventris">
<emphasis id="52BD32403637FFF8FEB9EE99DACA69D0" box="[268,401,910,931]" italics="true" pageId="3" pageNumber="4">caeruliventris</emphasis>
</taxonomicName>
2 4* 13.7
</th>
<td id="9D28856C36370004FD73EE99DF4169D0" box="[710,1050,910,931]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">2 4* 15.7</td>
<td id="9D28856C36370004FB91EE99DE7069D0" box="[1060,1323,910,931]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">1 5* 21.8</td>
</tr>
<tr id="DEF9EC1036370004FEBDEEA5DE7069B4" box="[264,1323,946,967]" gridrow="5" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDEEA5D9E769B4" box="[264,700,946,967]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9EEA5DA3F69B4" box="[268,356,946,967]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="shirleyae">
<emphasis id="52BD32403637FFF8FEB9EEA5DA3F69B4" box="[268,356,946,967]" italics="true" pageId="3" pageNumber="4">shirleyae</emphasis>
</taxonomicName>
25 53* 13.7
</th>
<td id="9D28856C36370004FD73EEA5DF4169B4" box="[710,1050,946,967]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">16 54* 15.8</td>
<td id="9D28856C36370004FB91EEA5DE7069B4" box="[1060,1323,946,967]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">15 49* 3 21.8</td>
</tr>
<tr id="DEF9EC1036370004FEBDEEC1DE706998" box="[264,1323,982,1003]" gridrow="6" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDEEC1D9E76998" box="[264,700,982,1003]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9EEC1DA326998" box="[268,361,982,1003]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="bandanus">
<emphasis id="52BD32403637FFF8FEB9EEC1DA326998" box="[268,361,982,1003]" italics="true" pageId="3" pageNumber="4">bandanus</emphasis>
</taxonomicName>
3 45* 2 14
</th>
<td id="9D28856C36370004FD73EEC1DF416998" box="[710,1050,982,1003]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">1 47* 4 16.1</td>
<td id="9D28856C36370004FB91EEC1DE706998" box="[1060,1323,982,1003]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">6* 43 5 22</td>
</tr>
<tr id="DEF9EC1036370004FEBDEEEEDE706E7D" box="[264,1323,1017,1038]" gridrow="7" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDEEEED9E76E7D" box="[264,700,1017,1038]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9EEEEDADA6E7D" box="[268,385,1017,1038]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="52BD32403637FFF8FEB9EEEEDADA6E7D" box="[268,385,1017,1038]" italics="true" pageId="3" pageNumber="4">bimaculatus</emphasis>
</taxonomicName>
5 10* 13.7
</th>
<td id="9D28856C36370004FD73EEEEDF416E7D" box="[710,1050,1017,1038]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">3 11* 1 15.9</td>
<td id="9D28856C36370004FB91EEEEDE706E7D" box="[1060,1323,1017,1038]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">2 12* 21.9</td>
</tr>
<tr id="DEF9EC1036370004FEBDE90ADE706E41" box="[264,1323,1053,1074]" gridrow="8" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDE90AD9E76E41" box="[264,700,1053,1074]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">
<emphasis id="52BD32403637FFF8FEB9E90ADA026E41" box="[268,345,1053,1074]" italics="true" pageId="3" pageNumber="4">collettei</emphasis>
6* 2 14.2
</th>
<td id="9D28856C36370004FD73E90ADF416E41" box="[710,1050,1053,1074]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">8* 16</td>
<td id="9D28856C36370004FB91E90ADE706E41" box="[1060,1323,1053,1074]" gridcol="2" gridrow="8" pageId="3" pageNumber="4">2 6* 21.8</td>
</tr>
<tr id="DEF9EC1036370004FEBDE957DE706E26" box="[264,1323,1088,1109]" gridrow="9" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDE957D9E76E26" box="[264,700,1088,1109]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9E957DA3D6E26" box="[268,358,1088,1109]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="prooculis">
<emphasis id="52BD32403637FFF8FEB9E957DA3D6E26" box="[268,358,1088,1109]" italics="true" pageId="3" pageNumber="4">prooculis</emphasis>
</taxonomicName>
1 66* 7 14,1
</th>
<td id="9D28856C36370004FD73E957DF416E26" box="[710,1050,1088,1109]" gridcol="1" gridrow="9" pageId="3" pageNumber="4">5* 60 3 16</td>
<td id="9D28856C36370004FB91E957DE706E26" box="[1060,1323,1088,1109]" gridcol="2" gridrow="9" pageId="3" pageNumber="4">10* 58 6 21.9</td>
</tr>
<tr id="DEF9EC1036370004FEBDE973DE706E0A" box="[264,1323,1124,1145]" gridrow="10" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDE973D9E76E0A" box="[264,700,1124,1145]" gridcol="0" gridrow="10" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9E973DA026E0A" box="[268,345,1124,1145]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="randalli">
<emphasis id="52BD32403637FFF8FEB9E973DA026E0A" box="[268,345,1124,1145]" italics="true" pageId="3" pageNumber="4">randalli</emphasis>
</taxonomicName>
1*
</th>
<td id="9D28856C36370004FD73E973DF416E0A" box="[710,1050,1124,1145]" gridcol="1" gridrow="10" pageId="3" pageNumber="4">1*</td>
<td id="9D28856C36370004FB91E973DE706E0A" box="[1060,1323,1124,1145]" gridcol="2" gridrow="10" pageId="3" pageNumber="4">1*</td>
</tr>
<tr id="DEF9EC1036370004FEBDE990DE706EEE" box="[264,1323,1159,1181]" gridrow="11" pageId="3" pageNumber="4">
<th id="9D28856C36370004FEBDE990D9E76EEE" box="[264,700,1159,1181]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">
<emphasis id="52BD32403637FFF8FEB9E99FDA3F6EEE" box="[268,356,1160,1181]" italics="true" pageId="3" pageNumber="4">taeniatus</emphasis>
3* 14
</th>
<td id="9D28856C36370004FD73E990DF416EEE" box="[710,1050,1159,1181]" gridcol="1" gridrow="11" pageId="3" pageNumber="4">3* 16</td>
<td id="9D28856C36370004FB91E990DE706EEE" box="[1060,1323,1159,1181]" gridcol="2" gridrow="11" pageId="3" pageNumber="4">2* 1 22.3</td>
</tr>
</table>
</paragraph>
<paragraph id="6076EE523637FFF8FDCCE9C4DAF26C82" pageId="3" pageNumber="4">
<table id="12C91CF236370004FEBDE9C4DE7A6C82" box="[264,1313,1235,1777]" gridcols="6" gridrows="15" pageId="3" pageNumber="4">
<tr id="DEF9EC1036370004FEBDE9C4DE7A6E9B" box="[264,1313,1235,1256]" gridrow="0" pageId="3" pageNumber="4" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<th id="9D28856C36370004FDC2E9C4DF016E9B" box="[631,1114,1235,1256]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">Total dentary teeth (excluding posterior canines)</th>
</tr>
<tr id="DEF9EC1036370004FEBDE9E0DE7A6F7F" box="[264,1313,1271,1292]" gridrow="1" pageId="3" pageNumber="4" rowspan-5="1">
<th id="9D28856C36370004FEBDE9E0DA9D6F7F" box="[264,454,1271,1292]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">39</th>
<td id="9D28856C36370004FE44E9E0D9526F7F" box="[497,521,1271,1292]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">40</td>
<td id="9D28856C36370004FD81E9E0D9176F7F" box="[564,588,1271,1292]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">41</td>
<td id="9D28856C36370004FDC2E9E0DF016F7F" box="[631,1114,1271,1292]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">42 43 44 45 46 47 48 49</td>
<td id="9D28856C36370004FB31E9E0DFC76F7F" box="[1156,1180,1271,1292]" gridcol="4" gridrow="1" pageId="3" pageNumber="4">50</td>
</tr>
<tr id="DEF9EC1036370004FEBDE80ADE7A6F41" box="[264,1313,1309,1330]" gridrow="2" pageId="3" pageNumber="4" rowspan-4="1">
<th id="9D28856C36370004FEBDE80ADA9D6F41" box="[264,454,1309,1330]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9E80ADACA6F41" box="[268,401,1309,1330]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="caeruliventris">
<emphasis id="52BD32403637FFF8FEB9E80ADACA6F41" box="[268,401,1309,1330]" italics="true" pageId="3" pageNumber="4">caeruliventris</emphasis>
</taxonomicName>
1
</th>
<td id="9D28856C36370004FE44E80AD9526F41" box="[497,521,1309,1330]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">1</td>
<td id="9D28856C36370004FD81E80AD9176F41" box="[564,588,1309,1330]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">­</td>
<td id="9D28856C36370004FDC2E80ADF016F41" box="[631,1114,1309,1330]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">3 ­ ­ 1*</td>
<td id="9D28856C36370004FB42E80ADE7A6F41" box="[1271,1313,1309,1330]" gridcol="5" gridrow="2" pageId="3" pageNumber="4">41.7</td>
</tr>
<tr id="DEF9EC1036370004FEBDE855DE7A6F24" box="[264,1313,1346,1367]" gridrow="3" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="9D28856C36370004FEBDE855DA9D6F24" box="[264,454,1346,1367]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9E855DA3F6F24" box="[268,356,1346,1367]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="shirleyae">
<emphasis id="52BD32403637FFF8FEB9E855DA3F6F24" box="[268,356,1346,1367]" italics="true" pageId="3" pageNumber="4">shirleyae</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="DEF9EC1036370004FEBDE871DE7A6F08" box="[264,1313,1382,1403]" gridrow="4" pageId="3" pageNumber="4" rowspan-4="1">
<th id="9D28856C36370004FEBDE871DA9D6F08" box="[264,454,1382,1403]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">Males 1</th>
<td id="9D28856C36370004FE44E871D9526F08" box="[497,521,1382,1403]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">2</td>
<td id="9D28856C36370004FD81E871D9176F08" box="[564,588,1382,1403]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">3*</td>
<td id="9D28856C36370004FDC2E871DF016F08" box="[631,1114,1382,1403]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">9 7 6 1</td>
<td id="9D28856C36370004FB42E871DE7A6F08" box="[1271,1313,1382,1403]" gridcol="5" gridrow="4" pageId="3" pageNumber="4">42.4</td>
</tr>
<tr id="DEF9EC1036370004FEBDE89CDE7A6FD3" box="[264,1313,1419,1440]" gridrow="5" pageId="3" pageNumber="4" rowspan-4="1">
<th id="9D28856C36370004FEBDE89CDA9D6FD3" box="[264,454,1419,1440]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">Females</th>
<td id="9D28856C36370004FE44E89CD9526FD3" box="[497,521,1419,1440]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">1</td>
<td id="9D28856C36370004FD81E89CD9176FD3" box="[564,588,1419,1440]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">1</td>
<td id="9D28856C36370004FDC2E89CDF016FD3" box="[631,1114,1419,1440]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">1 4 6 7 5 1</td>
<td id="9D28856C36370004FB42E89CDE7A6FD3" box="[1271,1313,1419,1440]" gridcol="5" gridrow="5" pageId="3" pageNumber="4">44.3</td>
</tr>
<tr id="DEF9EC1036370004FEBDE8B8DE7A6FB7" box="[264,1313,1455,1476]" gridrow="6" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="9D28856C36370004FEBDE8B8DA9D6FB7" box="[264,454,1455,1476]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9E8B8DA326FB7" box="[268,361,1455,1476]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="bandanus">
<emphasis id="52BD32403637FFF8FEB9E8B8DA326FB7" box="[268,361,1455,1476]" italics="true" pageId="3" pageNumber="4">bandanus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="DEF9EC1036370004FEBDE8C4DE7A6F9B" box="[264,1313,1491,1512]" gridrow="7" pageId="3" pageNumber="4" rowspan-4="1">
<th id="9D28856C36370004FEBDE8C4DA9D6F9B" box="[264,454,1491,1512]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">Males</th>
<td id="9D28856C36370004FE44E8C4D9526F9B" box="[497,521,1491,1512]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">1</td>
<td id="9D28856C36370004FD81E8C4D9176F9B" box="[564,588,1491,1512]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">3</td>
<td id="9D28856C36370004FDC2E8C4DF016F9B" box="[631,1114,1491,1512]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">9 4* 4 1</td>
<td id="9D28856C36370004FB42E8C4DE7A6F9B" box="[1271,1313,1491,1512]" gridcol="5" gridrow="7" pageId="3" pageNumber="4">42.4</td>
</tr>
<tr id="DEF9EC1036370004FEBDE8EFDE7A6C7E" box="[264,1313,1528,1549]" gridrow="8" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<th id="9D28856C36370004FEBDE8EFDA9D6C7E" box="[264,454,1528,1549]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">Females</th>
<td id="9D28856C36370004FDC2E8EFDF016C7E" box="[631,1114,1528,1549]" gridcol="3" gridrow="8" pageId="3" pageNumber="4">2 4 5 2 1</td>
<td id="9D28856C36370004FB42E8EFDE7A6C7E" box="[1271,1313,1528,1549]" gridcol="5" gridrow="8" pageId="3" pageNumber="4">44.7</td>
</tr>
<tr id="DEF9EC1036370004FEBDEB0BDE7A6C42" box="[264,1313,1564,1585]" gridrow="9" pageId="3" pageNumber="4" rowspan-1="1" rowspan-4="1">
<th id="9D28856C36370004FEBDEB0BDA9D6C42" box="[264,454,1564,1585]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9EB0BDADA6C42" box="[268,385,1564,1585]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="52BD32403637FFF8FEB9EB0BDADA6C42" box="[268,385,1564,1585]" italics="true" pageId="3" pageNumber="4">bimaculatus</emphasis>
</taxonomicName>
</th>
<td id="9D28856C36370004FD81EB0BD9176C42" box="[564,588,1564,1585]" gridcol="2" gridrow="9" pageId="3" pageNumber="4">2</td>
<td id="9D28856C36370004FDC2EB0BDF016C42" box="[631,1114,1564,1585]" gridcol="3" gridrow="9" pageId="3" pageNumber="4">5* 2 2 2</td>
<td id="9D28856C36370004FB42EB0BDE7A6C42" box="[1271,1313,1564,1585]" gridcol="5" gridrow="9" pageId="3" pageNumber="4">42.8</td>
</tr>
<tr id="DEF9EC1036370004FEBDEB57DE7A6C25" box="[264,1313,1600,1622]" gridrow="10" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1">
<th id="9D28856C36370004FEBDEB57DA9D6C25" box="[264,454,1600,1622]" gridcol="0" gridrow="10" pageId="3" pageNumber="4">
<emphasis id="52BD32403637FFF8FEB9EB56DA026C25" box="[268,345,1601,1622]" italics="true" pageId="3" pageNumber="4">collettei</emphasis>
</th>
<td id="9D28856C36370004FDC2EB57DF016C25" box="[631,1114,1600,1622]" gridcol="3" gridrow="10" pageId="3" pageNumber="4">2 2 ­ 2* 1</td>
<td id="9D28856C36370004FB31EB57DFC76C25" box="[1156,1180,1600,1622]" gridcol="4" gridrow="10" pageId="3" pageNumber="4">1</td>
<td id="9D28856C36370004FB42EB57DE7A6C25" box="[1271,1313,1600,1622]" gridcol="5" gridrow="10" pageId="3" pageNumber="4">47.1</td>
</tr>
<tr id="DEF9EC1036370004FEBDEB72DE7A6C09" box="[264,1313,1637,1658]" gridrow="11" pageId="3" pageNumber="4" rowspan-4="1">
<th id="9D28856C36370004FEBDEB72DA9D6C09" box="[264,454,1637,1658]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9EB72DA3D6C09" box="[268,358,1637,1658]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="prooculis">
<emphasis id="52BD32403637FFF8FEB9EB72DA3D6C09" box="[268,358,1637,1658]" italics="true" pageId="3" pageNumber="4">prooculis</emphasis>
</taxonomicName>
1
</th>
<td id="9D28856C36370004FE44EB72D9526C09" box="[497,521,1637,1658]" gridcol="1" gridrow="11" pageId="3" pageNumber="4">2</td>
<td id="9D28856C36370004FD81EB72D9176C09" box="[564,588,1637,1658]" gridcol="2" gridrow="11" pageId="3" pageNumber="4">7*</td>
<td id="9D28856C36370004FDC2EB72DF016C09" box="[631,1114,1637,1658]" gridcol="3" gridrow="11" pageId="3" pageNumber="4">12 11 13 5 5 8 3</td>
<td id="9D28856C36370004FB42EB72DE7A6C09" box="[1271,1313,1637,1658]" gridcol="5" gridrow="11" pageId="3" pageNumber="4">43.7</td>
</tr>
<tr id="DEF9EC1036370004FEBDEB9EDE7A6CEC" box="[264,1313,1673,1695]" gridrow="12" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<th id="9D28856C36370004FEBDEB9EDA9D6CEC" box="[264,454,1673,1695]" gridcol="0" gridrow="12" pageId="3" pageNumber="4">
<taxonomicName id="A7C995D13637FFF8FEB9EB9DDA026CEC" box="[268,345,1674,1695]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="randalli">
<emphasis id="52BD32403637FFF8FEB9EB9DDA026CEC" box="[268,345,1674,1695]" italics="true" pageId="3" pageNumber="4">randalli</emphasis>
</taxonomicName>
</th>
<td id="9D28856C36370004FDC2EB9EDF016CEC" box="[631,1114,1673,1695]" gridcol="3" gridrow="12" pageId="3" pageNumber="4">1*</td>
</tr>
<tr id="DEF9EC1036370004FEBDEBB9DE7A6CB0" box="[264,1313,1710,1731]" gridrow="13" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<th id="9D28856C36370004FEBDEBB9DA9D6CB0" box="[264,454,1710,1731]" gridcol="0" gridrow="13" pageId="3" pageNumber="4">
<emphasis id="52BD32403637FFF8FEB9EBB9DA3F6CB0" box="[268,356,1710,1731]" italics="true" pageId="3" pageNumber="4">taeniatus</emphasis>
</th>
<td id="9D28856C36370004FDC2EBB9DF016CB0" box="[631,1114,1710,1731]" gridcol="3" gridrow="13" pageId="3" pageNumber="4">1 ­ 1 1*</td>
<td id="9D28856C36370004FB42EBB9DE7A6CB0" box="[1271,1313,1710,1731]" gridcol="5" gridrow="13" pageId="3" pageNumber="4">45.7</td>
</tr>
<tr id="DEF9EC1036370004FEBDEBC9DE7A6C82" box="[264,1313,1758,1777]" gridrow="14" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="9D28856C36370004FEBDEBC9DA9D6C82" box="[264,454,1758,1777]" gridcol="0" gridrow="14" pageId="3" pageNumber="4">*includes holotype</th>
</tr>
</table>
</paragraph>
<subSubSection id="28D3BDD93637FFFFFE8DEA3EDF336C7D" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="6076EE523637FFFFFE8DEA3ED8E0694D" blockId="3.[264,1323,1833,1899]" lastBlockId="4.[264,1323,284,1870]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="52BD32403637FFF8FE8DEA3EDA996D31" box="[312,450,1833,1858]" italics="true" pageId="3" pageNumber="4">Description</emphasis>
(only differentiating characters or characters not otherwise mentioned in the introduction, identification key, and Table 1 are discussed). Preserved specimens (Figure 1a–c) are most noteworthy for their general lack of distinguishing characters. There is a diffusely dusky spot on the orbital margin at about the three o’clock position. On each side of the ventral surface of the head of the male
<typeStatus id="BF7250F03630FFFFFCDEEC7BD8896BF5" box="[875,978,364,390]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
(
<figureCitation id="F8F2F2D73630FFFFFC56EC7BDF1C6BF5" box="[995,1095,364,390]" captionStart="FIGURE 1" captionStartId="5.[264,368,1694,1718]" captionTargetBox="[312,1295,334,1610]" captionTargetId="figure@5.[264,1323,309,1627]" captionTargetPageId="5" captionText="FIGURE 1. Photographs of preserved specimens of three similar Ecsenius species. a – c, Ecsenius caeruliventris: a, b, MZB 12094, holotype, male, 24.3 mm SL (b, ventral surface of head); c, USNM 379435, female, 23.0 mm SL, ventral surface of head (note pale spots). d, Ecsenius shirleyae, USNM 211996, holotype, male, 28.4 mm SL; e, Ecsenius bandanus, USNM 221236, male, 31.7 mm SL, Biak, Irian Jaya, Indonesia." httpUri="https://zenodo.org/record/169639/files/figure.png" pageId="4" pageNumber="5">Figure 1</figureCitation>
b), there is a small dark spot, surrounded by a pale halo, in the area external to articulations of the anterior branchiostegals with the ceratohyal. A second, inconspicuous, faintly dusky spot is present near the anteriormost mandibular sensory pore. The body is darkly dusky anteriorly below the spinous dorsal fin, but becomes gradually paler posteriorly. The fins are unremarkable except for an immaculate basal stripe­like area in the segmented­ray portion of the dorsalfin rays (fine melanophores are on the fin rays dorsal to the stripe, and in the posteriormore interradial membranes) and the anal fin has a general covering of fine melanophores. The other two, smaller, males lack ventral head markings and the immaculate basal area in the segmented­ray portion of the dorsal fin. Females also lack the fin stripe, and the dark spots on the ventral head surface are replaced by one or two pairs of larger, pale spots (
<figureCitation id="F8F2F2D73630FFFFFB69EFEBDA48694D" captionStart="FIGURE 1" captionStartId="5.[264,368,1694,1718]" captionTargetBox="[312,1295,334,1610]" captionTargetId="figure@5.[264,1323,309,1627]" captionTargetPageId="5" captionText="FIGURE 1. Photographs of preserved specimens of three similar Ecsenius species. a – c, Ecsenius caeruliventris: a, b, MZB 12094, holotype, male, 24.3 mm SL (b, ventral surface of head); c, USNM 379435, female, 23.0 mm SL, ventral surface of head (note pale spots). d, Ecsenius shirleyae, USNM 211996, holotype, male, 28.4 mm SL; e, Ecsenius bandanus, USNM 221236, male, 31.7 mm SL, Biak, Irian Jaya, Indonesia." httpUri="https://zenodo.org/record/169639/files/figure.png" pageId="4" pageNumber="5">Figure 1</figureCitation>
c) in the positions occupied by the dark spots of the male.
</paragraph>
<paragraph id="6076EE523630FFFFFE8DEE5BDE5F6ED5" blockId="4.[264,1323,284,1870]" pageId="4" pageNumber="5">
In life (
<figureCitation id="F8F2F2D73630FFFFFE39EE5BDAAA6915" box="[396,497,844,870]" captionStart="FIGURE 2" captionStartId="6.[264,368,1765,1789]" captionTargetBox="[318,1268,315,1758]" captionTargetId="figure@6.[318,1268,315,1762]" captionTargetPageId="6" captionText="FIGURE 2. Photographs in nature of four similar Ecsenius species of the Prooculis group. a, Ecsenius caeruliventris, Togean Islands, Sulawesi (G. R. Allen); b, Ecsenius shirleyae, off Ujung Padang, Sulawesi (J. E. Randall); c, Ecsenius bimaculatus, Batangas, Luzon, Philippine Islands (J. E. Randall); d, Ecsenius bandanus, Rani Island, off Biak, Irian Jaya (G. E. Barrall)." httpUri="https://zenodo.org/record/169640/files/figure.png" pageId="4" pageNumber="5">Figure 2</figureCitation>
a), the most obvious color markings are the deep blue of the abdominal area, a vertical pair of white stripes extending through the orbit dorsal and ventral to the pupil, a pair of small white spots in the interorbital region anteriorly, and a small, mid­dorsal white spot between the dorsoposterior margins of the orbits. The dusky post­orbital spot is bordered dorsally by a fine white spot and ventrally by a fine, pale, whitish stripe that continues faintly across the head and separates the pale yellowish cheeks dorsally from the brownish color of the head dorsally. The blue of the abdomen pales posteriorly and the body becomes faintly yellow similar to the cheeks. Specimens may or may not bear evidence of broad dusky bands on the body (see
<bibRefCitation id="045893A33630FFFFFCC9E99BDF266ED5" author="Allen" box="[892,1149,1164,1190]" pageId="4" pageNumber="12" refString="Allen, G. R. &amp; Adrim, M. (2003) Coral-reef fishes of Indonesia. Zoological Studies, 42 (1), 1 - 72." type="journal article" year="2003">Allen &amp; Adrim, 2003</bibRefCitation>
: figure 71).
</paragraph>
<paragraph id="6076EE523630FFFFFE8DE9A3DF336C7D" blockId="4.[264,1323,284,1870]" pageId="4" pageNumber="5">
<emphasis id="52BD32403630FFFFFE8DE9A3DA876EBE" box="[312,476,1204,1229]" italics="true" pageId="4" pageNumber="5">Comparisons.</emphasis>
Within the
<taxonomicName id="A7C995D13630FFFFFDDDE9A3D98C6EBD" box="[616,727,1204,1230]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="9" phylum="Chordata" rank="species" species="prooculis">Prooculis</taxonomicName>
group,
<taxonomicName id="A7C995D13630FFFFFC87E9A3DF1F6EBE" box="[818,1092,1204,1229]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="caeruliventris">
<emphasis id="52BD32403630FFFFFC87E9A3DF1F6EBE" box="[818,1092,1204,1229]" italics="true" pageId="4" pageNumber="5">Ecsenius caeruliventris</emphasis>
</taxonomicName>
appears to be most similar to
<taxonomicName id="A7C995D13630FFFFFECAE9CBD9506E86" box="[383,523,1244,1270]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="shirleyae">
<emphasis id="52BD32403630FFFFFECAE9CBDACA6E86" box="[383,401,1244,1269]" italics="true" pageId="4" pageNumber="5">E</emphasis>
.
<emphasis id="52BD32403630FFFFFE2AE9CBD9506E86" box="[415,523,1244,1269]" italics="true" pageId="4" pageNumber="5">shirleyae</emphasis>
</taxonomicName>
and
<taxonomicName id="A7C995D13630FFFFFDF1E9CBD98D6E86" box="[580,726,1244,1269]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="bandanus">
<emphasis id="52BD32403630FFFFFDF1E9CBD98D6E86" box="[580,726,1244,1269]" italics="true" pageId="4" pageNumber="5">E. bandanus</emphasis>
</taxonomicName>
in having a blue abdomen, in that some specimens have broad, very faintly dusky bands on the body, and in lacking distinct dark spots or stripes on the body. It differs from both species in lacking a distinct dark stripe that extends posteriorly from the posterior margin of the orbit across the head. It further differs from
<taxonomicName id="A7C995D13630FFFFFAA4E843DA286FE6" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="shirleyae">
<emphasis id="52BD32403630FFFFFAA4E843DA286FE6" italics="true" pageId="4" pageNumber="5">E. shirleyae</emphasis>
</taxonomicName>
in that the pale stripe margining the dusky postorbital spot is not intensely white and does not clearly extend across the head and enter the body, and in the more intense manifestation of the blue color of the abdomen. It further differs from
<taxonomicName id="A7C995D13630FFFFFBF0E8DBDF826F96" box="[1093,1241,1484,1509]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="bandanus">
<emphasis id="52BD32403630FFFFFBF0E8DBDF826F96" box="[1093,1241,1484,1509]" italics="true" pageId="4" pageNumber="5">E. bandanus</emphasis>
</taxonomicName>
in that the pale markings on the head and iris are white rather than bright yellow.
</paragraph>
</subSubSection>
<subSubSection id="28D3BDD93630FFFFFE8DEB0BD9476CA5" pageId="4" pageNumber="5" type="discussion">
<paragraph id="6076EE523630FFFFFE8DEB0BD9476CA5" blockId="4.[264,1323,284,1870]" pageId="4" pageNumber="5">
<emphasis id="52BD32403630FFFFFE8DEB0BDAFD6C46" box="[312,422,1564,1589]" italics="true" pageId="4" pageNumber="5">Remarks.</emphasis>
This is one of the smallest species of
<taxonomicName id="A7C995D13630FFFFFC33EB0BD8B56C46" box="[902,1006,1564,1589]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
<emphasis id="52BD32403630FFFFFC33EB0BD8B56C46" box="[902,1006,1564,1589]" italics="true" pageId="4" pageNumber="5">Ecsenius</emphasis>
</taxonomicName>
; the
<quantity id="A73143B73630FFFFFB84EB0BDFC56C45" box="[1073,1182,1564,1590]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" pageId="4" pageNumber="5" unit="mm" value="23.7">23.7 mm</quantity>
female has well­developed, apparently ripe, ovaries. The
<quantity id="A73143B73630FFFFFC87EB53D8C76C2D" box="[818,924,1604,1630]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="4" pageNumber="5" unit="mm" value="15.6">15.6 mm</quantity>
male has well­developed dentary teeth, including obvious posterior canines. In most
<taxonomicName id="A7C995D13630FFFFFCEAEB7BD89C6CF6" box="[863,967,1644,1669]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
<emphasis id="52BD32403630FFFFFCEAEB7BD89C6CF6" box="[863,967,1644,1669]" italics="true" pageId="4" pageNumber="5">Ecsenius</emphasis>
</taxonomicName>
species this size, the posterior canines are not present and the number of other dentary teeth does not appear to have been completely established.
</paragraph>
</subSubSection>
<subSubSection id="28D3BDD93630FFFFFE8DEBF3DAD16D3D" pageId="4" pageNumber="5" type="etymology">
<paragraph id="6076EE523630FFFFFE8DEBF3DAD16D3D" blockId="4.[264,1323,284,1870]" pageId="4" pageNumber="5">
<emphasis id="52BD32403630FFFFFE8DEBF3DAEE6C8E" box="[312,437,1764,1789]" italics="true" pageId="4" pageNumber="5">Etymology</emphasis>
: The specific name is derived from the Latin
<emphasis id="52BD32403630FFFFFC54EBF3DF086C8E" box="[993,1107,1764,1789]" italics="true" pageId="4" pageNumber="5">caeruleus</emphasis>
(sky­blue) +
<emphasis id="52BD32403630FFFFFB4DEBF3DA746D56" italics="true" pageId="4" pageNumber="5">ventris</emphasis>
, (belly), and refers to the blue abdominal area of the species; used here as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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