treatments-xml/data/E8/60/5F/E8605F443633FFF3FEBDEC0BD8B56F6D.xml

<document id="E5F969D83D233D055D3222BFBF195054" ID-DOI="10.5281/zenodo.169638" ID-GBIF-Dataset="3ff15714-6d5c-4265-aff2-c7a19041a761" ID-ISSN="1175­5326" ID-Zenodo-Dep="169638" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461038850950" checkinUser="plazi" docAuthor="Springer, Victor G. &amp; Allen, Gerald R." docDate="2004" docId="E8605F443633FFF3FEBDEC0BD8B56F6D" docLanguage="en" docName="zt00791.pdf" docOrigin="Zootaxa 791" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Ecsenius shirleyae Springer &amp; Allen, 2004, new species" docType="treatment" docUuid="DE3BA438-05EF-49DC-A1C8-3BC6A77D7F00" docUuidSource="ZooBank" docVersion="10" lastPageNumber="9" masterDocId="1459273C3634FFFBFFB5ED17DB5B6A73" masterDocTitle="Ecsenius caeruliventris and E. shirleyae, two new species of blenniid fishes from Indonesia, and new distribution records for other species of Ecsenius" masterLastPageNumber="12" masterPageNumber="1" pageNumber="8" updateTime="1698575124870" updateUser="plazi">
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<mods:title id="D8C2762615FFA3B510C98803FD4E05F5">Ecsenius caeruliventris and E. shirleyae, two new species of blenniid fishes from Indonesia, and new distribution records for other species of Ecsenius</mods:title>
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<mods:namePart id="502C58D1E8E23A46F433C17AF7A0A217">Springer, Victor G.</mods:namePart>
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<mods:date id="ADEBE7507904134C0412D35EC3664958">2004</mods:date>
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<taxonomicName id="A7C995D13633FFFCFEBDEC0BDAB26B45" ID-CoL="38P23" box="[264,489,284,310]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="shirleyae" status="sp. nov.">
<emphasis id="52BD32403633FFFCFEBDEC0BDAB26B45" bold="true" box="[264,489,284,310]" italics="true" pageId="7" pageNumber="8">Ecsenius shirleyae</emphasis>
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, 
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<paragraph id="6076EE523633FFFCFEBDEC28DA9C6B2A" blockId="7.[264,648,283,385]" box="[264,455,319,345]" pageId="7" pageNumber="8">Shirley’s blenny</paragraph>
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<figureCitation id="F8F2F2D73633FFFCFEBDEC70DA286BF2" box="[264,371,359,385]" captionStart="FIGURE 1" captionStartId="5.[264,368,1694,1718]" captionTargetBox="[312,1295,334,1610]" captionTargetId="figure@5.[264,1323,309,1627]" captionTargetPageId="5" captionText="FIGURE 1. Photographs of preserved specimens of three similar Ecsenius species. a – c, Ecsenius caeruliventris: a, b, MZB 12094, holotype, male, 24.3 mm SL (b, ventral surface of head); c, USNM 379435, female, 23.0 mm SL, ventral surface of head (note pale spots). d, Ecsenius shirleyae, USNM 211996, holotype, male, 28.4 mm SL; e, Ecsenius bandanus, USNM 221236, male, 31.7 mm SL, Biak, Irian Jaya, Indonesia." httpUri="https://zenodo.org/record/169639/files/figure.png" pageId="7" pageNumber="8">Figures 1</figureCitation>
d, 2b, 3
</paragraph>
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<taxonomicName id="A7C995D13633FFFCFEBDECA0DA826BBD" ID-CoL="38NYB" box="[264,473,439,462]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bandanus">
<emphasis id="52BD32403633FFFCFEBDECA0DA826BBD" box="[264,473,439,462]" italics="true" pageId="7" pageNumber="8">Ecsenius bandanus</emphasis>
</taxonomicName>
(not Springer), in part: 
<treatmentCitation id="E168C8433633FFFCFD5FECA1D8E66BBE" author="Springer" box="[746,957,438,461]" page="106" pageId="7" pageNumber="8" year="1988">
<bibRefCitation id="045893A33633FFFCFD5FECA1D8D46BBE" author="Springer" box="[746,911,438,461]" pageId="7" pageNumber="12" refString="Springer, V. G. (1988) The Indo-Pacific blenniid fish genus Ecsenius. Smithsonian Contribtions to Zoology, 465, 1 - 134, 14 pls." type="journal article" year="1988">Springer, 1988</bibRefCitation>
:106
</treatmentCitation>
, fig. 57, pl.13, fig. 5; 
<treatmentCitation id="E168C8433633FFFCFB72ECA1D94D6B9E" author="Kuiter" page="606" pageId="7" pageNumber="8" year="2001">
<bibRefCitation id="045893A33633FFFCFB72ECA1DAB26B9E" author="Kuiter" pageId="7" pageNumber="12" refString="Kuiter, R. H. &amp; Tonozuka, T. (2001) Pictorial Guide to Indonesian Reef Fishes. Zoonetics, Seaford, Victoria, Australia, 893 pp. Second edition, 2004, PT Dive &amp; Dive's, Bali, Indonesia, same pagination." type="book" year="2001">Kuiter &amp; Tonozuka, 2001</bibRefCitation>
:606
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(Maumere, 
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); 
<treatmentCitation id="E168C8433633FFFCFD4AECC1DF516B9E" author="Allen" box="[767,1034,470,493]" page="54" pageId="7" pageNumber="8" year="2003">
<bibRefCitation id="045893A33633FFFCFD4AECC1D8B06B9E" author="Allen" box="[767,1003,470,493]" pageId="7" pageNumber="12" refString="Allen, G. R. &amp; Adrim, M. (2003) Coral-reef fishes of Indonesia. Zoological Studies, 42 (1), 1 - 72." type="journal article" year="2003">Allen &amp; Adrin, 2003</bibRefCitation>
:54
</treatmentCitation>
&amp; 18, fig. 65 (Maumere, 
<collectingCountry id="18DEAEC23633FFFCFE8DECE1DA24687E" box="[312,383,502,525]" name="Indonesia" pageId="7" pageNumber="8">Flores</collectingCountry>
); 
<treatmentCitation id="E168C8433633FFFCFE27ECE1D913687E" author="Myers" box="[402,584,502,525]" page="224" pageId="7" pageNumber="8" year="1999">
<bibRefCitation id="045893A33633FFFCFE27ECE1D941687E" author="Myers" box="[402,538,502,525]" pageId="7" pageNumber="12" refString="Myers, R. F. (1999) Micronesian Reef Fishes, 3 rd edition. Coral Graphics, Guam, 330 pp., 132 pls." type="book" year="1999">Myers, 1999</bibRefCitation>
:224
</treatmentCitation>
&amp; pl. 146K.
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</paragraph>
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<paragraph id="6076EE523633FFFCFEBDEF28DF4268DA" blockId="7.[264,1324,575,1881]" pageId="7" pageNumber="8">
<typeStatus id="BF7250F03633FFFCFEBDEF28DA28682B" box="[264,371,575,600]" pageId="7" pageNumber="8" type="holotype">
<emphasis id="52BD32403633FFFCFEBDEF28DA28682B" box="[264,371,575,600]" italics="true" pageId="7" pageNumber="8">Holotype</emphasis>
</typeStatus>
: 
<collectionCode id="06D876973633FFFCFE32EF28DA87682A" LSID="urn:lsid:biocol.org:col:34871" box="[391,476,575,601]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8">USNM</collectionCode>
211996, male, 
<quantity id="A73143B73633FFFCFD2EEF28D85C682A" box="[667,775,575,601]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="7" pageNumber="8" unit="mm" value="28.4">28.4 mm</quantity>
SL, off NW side of 
<collectingCountry id="18DEAEC23633FFFCFBB8EF28DF0B682A" box="[1037,1104,575,601]" name="Iceland" pageId="7" pageNumber="8">Pulau</collectingCountry>
Tikus, 
<collectingCountry id="18DEAEC23633FFFCFB1BEF28DFAA682A" box="[1198,1265,575,601]" name="Iceland" pageId="7" pageNumber="8">Pulau</collectingCountry>
Pari group, 
<collectingCountry id="18DEAEC23633FFFCFEEEEF70DAC668F2" box="[347,413,615,641]" name="Iceland" pageId="7" pageNumber="8">Pulau</collectingCountry>
Seribu, 
<collectingCountry id="18DEAEC23633FFFCFDB4EF70D92868F2" box="[513,627,615,641]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
(ca. 
<geoCoordinate id="05FD88953633FFFCFD00EF70D81368F2" box="[693,840,613,641]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="15" value="-5.8569446">05°51’25’’ S</geoCoordinate>
, ca. 
<geoCoordinate id="05FD88953633FFFCFCCAEF70DF7968F2" box="[895,1058,613,641]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="15" value="106.57083">106°34’15’’ E</geoCoordinate>
), to ca. 
<quantity id="A73143B73633FFFCFB35EF70DFEA68F2" box="[1152,1201,615,641]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="m" value="5.0">5 m</quantity>
, rotenone, V.G. Springer, M.F. Gomon, &amp; Sukarno, 
<date id="1477C8923633FFFCFD53EF98D82468DA" box="[742,895,655,681]" pageId="7" pageNumber="8" value="1974-04-05">5 April 1974</date>
; VGS 74­34.
</paragraph>
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<typeStatus id="BF7250F03633FFFCFE8DEFA0DAF468A3" box="[312,431,695,720]" pageId="7" pageNumber="8" type="paratype">
<emphasis id="52BD32403633FFFCFE8DEFA0DAF468A3" box="[312,431,695,720]" italics="true" pageId="7" pageNumber="8">Paratypes</emphasis>
</typeStatus>
: 
<collectionCode id="06D876973633FFFCFE08EFA0D94968A2" LSID="urn:lsid:biocol.org:col:34871" box="[445,530,695,721]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8">USNM</collectionCode>
379409 (15:15.6 33.5) and 
<collectionCode id="06D876973633FFFCFCD3EFA0D8F668A2" LSID="urn:lsid:biocol.org:col:34981" box="[870,941,695,721]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="8">WAM</collectionCode>
P32602.001 (2: 22.2–29.1), collected with 
<typeStatus id="BF7250F03633FFFCFE27EFC8DAA7688A" box="[402,508,735,761]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
. AMS I.18490001 (10:17.3–28.6), Tallabassi Bay, just off NW shore of Big Damalawa Islet, Kabaena Island, Celebes, 
<collectingCountry id="18DEAEC23633FFFCFCE2EE10D8926952" box="[855,969,775,801]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
(ca. 
<geoCoordinate id="05FD88953633FFFCFBB3EE10DFCC6952" box="[1030,1175,773,801]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="15" value="-5.288889">05°17’20’’ S</geoCoordinate>
, ca. 
<geoCoordinate id="05FD88953633FFFCFB7AEE10DA46693A" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="122.066666">122°04’ E</geoCoordinate>
), 
<quantity id="A73143B73633FFFCFE86EE38DA24693A" box="[307,383,815,841]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="7" pageNumber="8" unit="m" value="6.0" valueMax="8.0" valueMin="4.0">4–8 m</quantity>
, rotenone, V.G. Springer, M.F. Gomon, &amp; Indonesians, 
<date id="1477C8923633FFFCFBBAEE38DFB8693A" box="[1039,1251,815,841]" pageId="7" pageNumber="8" value="1974-02-24">24 February 1974</date>
, VGS 74­1. 
<collectionCode id="06D876973633FFFCFEF9EE40DAFA6902" LSID="urn:lsid:biocol.org:col:34871" box="[332,417,855,881]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8">USNM</collectionCode>
211916 (14:22.5–29.2, includes one specimen now lost), Tallabassi Bay, just off NE tip of Big Damalawa Islet, Kabaena Island, Celebes, 
<collectingCountry id="18DEAEC23633FFFCFC54EE68DF0869EA" box="[993,1107,895,921]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
(ca. 
<geoCoordinate id="05FD88953633FFFCFB24EE68DE7E69EA" box="[1169,1317,893,921]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="15" value="-5.288889">05°17’20’’ S</geoCoordinate>
, ca. 
<geoCoordinate id="05FD88953633FFFCFE84EEB0DAF269B2" box="[305,425,933,961]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="122.066666">122°04’ E</geoCoordinate>
), 
<quantity id="A73143B73633FFFCFE0AEEB0D94769B2" box="[447,540,935,961]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="7" pageNumber="8" unit="m" value="8.5" valueMax="15.0" valueMin="2.0">2–15 m</quantity>
, rotenone, V. G. Springer, M. F. Gomon, &amp; Indonesians, 
<date id="1477C8923633FFFCFB0DEEB0DA2D699A" pageId="7" pageNumber="8" value="1974-02-25">25 February 1974</date>
, VGS 74­2. 
<collectionCode id="06D876973633FFFCFDAAEED8D92F699A" LSID="urn:lsid:biocol.org:col:34871" box="[543,628,975,1001]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8">USNM</collectionCode>
211993 (18:13.3–29.2), off W side of 
<collectingCountry id="18DEAEC23633FFFCFBE1EED8DFCC699A" box="[1108,1175,975,1001]" name="Iceland" pageId="7" pageNumber="8">Pulau</collectingCountry>
Ajer, 
<collectingCountry id="18DEAEC23633FFFCFB5DEED8DE70699A" box="[1256,1323,975,1001]" name="Iceland" pageId="7" pageNumber="8">Pulau</collectingCountry>
Seribu, 
<collectingCountry id="18DEAEC23633FFFCFED1EEE0DA8D6E62" box="[356,470,1015,1041]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
(ca. 
<geoCoordinate id="05FD88953633FFFCFDA7EEE0D9206E62" box="[530,635,1013,1041]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="925" value="-5.7666664">05°46’ S</geoCoordinate>
, ca. 
<geoCoordinate id="05FD88953633FFFCFD06EEE0D8776E62" box="[691,812,1013,1041]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="106.583336">106°35’ E</geoCoordinate>
), to ca. 
<quantity id="A73143B73633FFFCFC36EEE0D8986E62" box="[899,963,1015,1041]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="8" unit="m" value="19.0">19 m</quantity>
, rotenone, V.G. Springer, M.F. Gomon, &amp; Sukarno, 
<date id="1477C8923633FFFCFDB2E908D9FC6E4A" box="[519,679,1055,1081]" pageId="7" pageNumber="8" value="1974-04-04">4 April 1974</date>
; VGS 74­32. 
<collectionCode id="06D876973633FFFCFCE7E908D8FC6E4A" LSID="urn:lsid:biocol.org:col:34871" box="[850,935,1055,1081]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8">USNM</collectionCode>
211979 (11:23.7–31.2), off SW coast Karimundjawa Island, 
<collectingCountry id="18DEAEC23633FFFCFDEEE950D98B6E12" box="[603,720,1095,1121]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
, ca. 
<geoCoordinate id="05FD88953633FFFCFCB3E950D8CD6E12" box="[774,918,1093,1121]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="15" value="-5.875">05°52’30” S</geoCoordinate>
, 
<geoCoordinate id="05FD88953633FFFCFC11E950DF1F6E12" box="[932,1092,1093,1121]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="15" value="110.42777">110°25’40” E</geoCoordinate>
, ca. 
<quantity id="A73143B73633FFFCFBC9E950DF8C6E12" box="[1148,1239,1095,1121]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="12.0" metricValueMin="1.0" pageId="7" pageNumber="8" unit="m" value="6.5" valueMax="12.0" valueMin="1.0">1–12 m</quantity>
, rotenone, V.G. Springer &amp; Indonesians, 
<date id="1477C8923633FFFCFD2AE978D80F6EFA" box="[671,852,1135,1161]" pageId="7" pageNumber="8" value="1974-03-29">29 March 1974</date>
, VGS 74–28.
</paragraph>
</subSubSection>
<subSubSection id="28D3BDD93633FFFCFE8DE980D80C6EAA" pageId="7" pageNumber="8" type="description">
<paragraph id="6076EE523633FFFCFE8DE980D80C6EAA" blockId="7.[264,1324,575,1881]" pageId="7" pageNumber="8">
<emphasis id="52BD32403633FFFCFE8DE980D9716EC3" box="[312,554,1175,1200]" italics="true" pageId="7" pageNumber="8">Additional specimen</emphasis>
: AMS I.34501010 (1:15.6), off Wailiti, 
<quantity id="A73143B73633FFFCFBA6E980DF046EC2" box="[1043,1119,1175,1201]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="8" unit="km" value="11.0">11 km</quantity>
NW Sao Wisata Resort, 
<collectingCountry id="18DEAEC23633FFFCFED6E9A8D96B6EAA" box="[355,560,1215,1241]" name="Indonesia" pageId="7" pageNumber="8">Flores, Indonesia</collectingCountry>
, 
<geoCoordinate id="05FD88953633FFFCFD8EE9A8D9E26EAA" box="[571,697,1213,1241]" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="-8.571667">08°34.3’ S</geoCoordinate>
, 
<geoCoordinate id="05FD88953633FFFCFD73E9A8D8086EAA" box="[710,851,1213,1241]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="122.195">122°11.7’ E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="28D3BDD93633FFFCFE8DE9F0D9E76FD2" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="6076EE523633FFFCFE8DE9F0D9E76FD2" blockId="7.[264,1324,575,1881]" pageId="7" pageNumber="8">
<emphasis id="52BD32403633FFFCFE8DE9F0DAED6F73" box="[312,438,1255,1280]" italics="true" pageId="7" pageNumber="8">Diagnosis.</emphasis>
A species of the 
<taxonomicName id="A7C995D13633FFFCFD31E9F0D9A96F72" box="[644,754,1255,1281]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="9" phylum="Chordata" rank="species" species="prooculis">Prooculis</taxonomicName>
group of 
<taxonomicName id="A7C995D13633FFFCFCDCE9F0D88A6F73" box="[873,977,1255,1280]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="52BD32403633FFFCFCDCE9F0D88A6F73" box="[873,977,1255,1280]" italics="true" pageId="7" pageNumber="8">Ecsenius</emphasis>
</taxonomicName>
with the following combination of characters: Body without alternating dark and pale stripes or distinct dark spots; head with dark postorbital stripe extending across head onto body anteriorly. In life, dark postorbital stripe bordered ventrally by bright white stripe; other bright markings on head white or pale yellowish; belly bluish.
</paragraph>
</subSubSection>
<subSubSection id="28D3BDD93633FFF3FE8DE8B8DAE36EBD" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="6076EE523633FFF3FE8DE8B8DA126B2D" blockId="7.[264,1324,575,1881]" lastBlockId="8.[264,1323,284,1310]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="52BD32403633FFFCFE8DE8B8DA996FBB" box="[312,450,1455,1480]" italics="true" pageId="7" pageNumber="8">Description</emphasis>
(only differentiating characters or characters not otherwise mentioned in the introduction, identification key, and Table 1 are discussed). Preserved specimens (Figure 1d) are most notable for the black stripe extending across the head posteriorly from the postorbital margin at about the three o’clock position. The stripe may extend for a variable, usually short, distance, onto the body anteriorly. On each side of the ventral surface of the head of the male 
<typeStatus id="BF7250F03633FFFCFD9EEB60D9C96CE2" box="[555,658,1655,1681]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
there are two pairs of dark spots similar to those of 
<taxonomicName id="A7C995D13633FFFCFAA4EB60DAF06CCB" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="caeruliventris">
<emphasis id="52BD32403633FFFCFAA4EB60DAF06CCB" italics="true" pageId="7" pageNumber="8">E. caeruliventris</emphasis>
</taxonomicName>
, although the halo is not evident (presence or absence of the spots varies in other males). The body is not so darkly dusky anteriorly as in 
<emphasis id="52BD32403633FFFCFC42EBD0DF4A6C93" box="[1015,1041,1735,1760]" italics="true" pageId="7" pageNumber="8">E.</emphasis>
<taxonomicName id="A7C995D13633FFFCFBACEBD0DFE76C92" box="[1049,1212,1735,1761]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="caeruliventris">caeruliventris</taxonomicName>
. The fins are similar to those of 
<taxonomicName id="A7C995D13633FFFCFDBAEBF8D9896D7B" box="[527,722,1775,1800]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="caeruliventris">
<emphasis id="52BD32403633FFFCFDBAEBF8D9896D7B" box="[527,722,1775,1800]" italics="true" pageId="7" pageNumber="8">E. caeruliventris</emphasis>
</taxonomicName>
, except that an immaculate stripe­like basal area in the segmented­ray portion of the dorsal fin is not evident. Females are generally more pale than males, and the dark spots on the ventral head surface are absent or replaced by one or two pairs of larger, pale spots (
<figureCitation id="F8F2F2D7363CFFF3FDDBEC0BD9956B45" box="[622,718,284,310]" captionStart="FIGURE 1" captionStartId="5.[264,368,1694,1718]" captionTargetBox="[312,1295,334,1610]" captionTargetId="figure@5.[264,1323,309,1627]" captionTargetPageId="5" captionText="FIGURE 1. Photographs of preserved specimens of three similar Ecsenius species. a – c, Ecsenius caeruliventris: a, b, MZB 12094, holotype, male, 24.3 mm SL (b, ventral surface of head); c, USNM 379435, female, 23.0 mm SL, ventral surface of head (note pale spots). d, Ecsenius shirleyae, USNM 211996, holotype, male, 28.4 mm SL; e, Ecsenius bandanus, USNM 221236, male, 31.7 mm SL, Biak, Irian Jaya, Indonesia." httpUri="https://zenodo.org/record/169639/files/figure.png" pageId="8" pageNumber="9">Figure 1</figureCitation>
c) in the positions occupied by the dark spots of the male.
</paragraph>
<paragraph id="6076EE52363CFFF3FE8DEC7BD85368ED" blockId="8.[264,1323,284,1310]" pageId="8" pageNumber="9">
In life (
<figureCitation id="F8F2F2D7363CFFF3FE24EC7BDAA36BF5" box="[401,504,364,390]" captionStart="FIGURE 2" captionStartId="6.[264,368,1765,1789]" captionTargetBox="[318,1268,315,1758]" captionTargetId="figure@6.[318,1268,315,1762]" captionTargetPageId="6" captionText="FIGURE 2. Photographs in nature of four similar Ecsenius species of the Prooculis group. a, Ecsenius caeruliventris, Togean Islands, Sulawesi (G. R. Allen); b, Ecsenius shirleyae, off Ujung Padang, Sulawesi (J. E. Randall); c, Ecsenius bimaculatus, Batangas, Luzon, Philippine Islands (J. E. Randall); d, Ecsenius bandanus, Rani Island, off Biak, Irian Jaya (G. E. Barrall)." httpUri="https://zenodo.org/record/169640/files/figure.png" pageId="8" pageNumber="9">Figure 2</figureCitation>
b), the most obvious color markings are the bluish abdominal area, a horizontal pair of white stripes extending through the orbit dorsal and ventral to the pupil, a pair of small white spots in the interorbital region anteriorly, a fine, white mid­predorsal stripe (all of these white stripes may be very faintly tinged with yellow), the black stripe extending posteriorly from the orbit margined ventrally by a bright white stripe, which continues well out onto the body. The head and body are brownish dorsal to the black stripe, and may show indications of a few broad, slightly darker bands anteriorly; posteriorly the body is similar to 
<taxonomicName id="A7C995D1363CFFF3FD88EF93D85A68EE" box="[573,769,644,669]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="caeruliventris">
<emphasis id="52BD3240363CFFF3FD88EF93D85A68EE" box="[573,769,644,669]" italics="true" pageId="8" pageNumber="9">E. caeruliventris</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="6076EE52363CFFF3FE8DEFBBDAE36EBD" blockId="8.[264,1323,284,1310]" pageId="8" pageNumber="9">
<emphasis id="52BD3240363CFFF3FE8DEFBBDA8768B6" box="[312,476,684,709]" italics="true" pageId="8" pageNumber="9">Comparisons.</emphasis>
Within the 
<taxonomicName id="A7C995D1363CFFF3FDD3EFBBD98E68B5" box="[614,725,684,710]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="prooculis">Prooculis</taxonomicName>
group, 
<taxonomicName id="A7C995D1363CFFF3FC85EFBBDF5768B6" box="[816,1036,684,709]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="shirleyae">
<emphasis id="52BD3240363CFFF3FC85EFBBDF5768B6" box="[816,1036,684,709]" italics="true" pageId="8" pageNumber="9">Ecsenius shirleyae</emphasis>
</taxonomicName>
appears to be most similar to 
<taxonomicName id="A7C995D1363CFFF3FEEBEFC3D9DE689D" authority="Springer" authorityName="Springer" box="[350,645,724,750]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="52BD3240363CFFF3FEEBEFC3D949689E" box="[350,530,724,749]" italics="true" pageId="8" pageNumber="9">E. bimaculatus</emphasis>
Springer
</taxonomicName>
, from which it differs mainly in lacking the two black abdominal spots and in that some individuals have a bluish abdomen. It is similar to 
<taxonomicName id="A7C995D1363CFFF3FAA4EFEBDAF0694E" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="caeruliventris">
<emphasis id="52BD3240363CFFF3FAA4EFEBDE786966" box="[1297,1315,764,789]" italics="true" pageId="8" pageNumber="9">E</emphasis>
. 
<emphasis id="52BD3240363CFFF3FEBDEE33DAF0694E" box="[264,427,804,829]" italics="true" pageId="8" pageNumber="9">caeruliventris</emphasis>
</taxonomicName>
and 
<taxonomicName id="A7C995D1363CFFF3FE53EE33D921694E" box="[486,634,804,829]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="bandanus">
<emphasis id="52BD3240363CFFF3FE53EE33D921694E" box="[486,634,804,829]" italics="true" pageId="8" pageNumber="9">E. bandanus</emphasis>
</taxonomicName>
in having a bluish abdomen (at least in some specimens), in having broad, faintly dusky bands on the body, and in lacking distinct dark spots or stripes on the entire body. It differs from both species in having a dark postorbital stripe that frequently extends well out onto the body and in having the stripe margined below by a white stripe that also extends well out onto the body. It further differs from 
<taxonomicName id="A7C995D1363CFFF3FB39EED3DA746E76" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="caeruliventris">
<emphasis id="52BD3240363CFFF3FB39EED3DA746E76" italics="true" pageId="8" pageNumber="9">E caeruliventris</emphasis>
</taxonomicName>
in that the blue of the abdomen is not as strongly manifested, and from 
<taxonomicName id="A7C995D1363CFFF3FBCFEEFBDE566E76" box="[1146,1293,1004,1029]" class="Actinopterygii" family="Blenniidae" genus="Ecsenius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="bandanus">
<emphasis id="52BD3240363CFFF3FBCFEEFBDE566E76" box="[1146,1293,1004,1029]" italics="true" pageId="8" pageNumber="9">E. bandanus</emphasis>
</taxonomicName>
in that the pale markings on the head and eye are bright white (or very faintly tinged with yellow) rather than bright yellow, and that the head portion of the dark post­orbital stripe is generally less deep (depth measurement is difficult because margins of the stripe are not sharply demarcated, but the difference is usually apparent when specimens of equal size are compared).
</paragraph>
</subSubSection>
<subSubSection id="28D3BDD9363CFFF3FE8DE9CBD8B56F6D" pageId="8" pageNumber="9" type="etymology">
<paragraph id="6076EE52363CFFF3FE8DE9CBD8B56F6D" blockId="8.[264,1323,284,1310]" pageId="8" pageNumber="9">
<emphasis id="52BD3240363CFFF3FE8DE9CBDAEE6E86" box="[312,437,1244,1269]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
: This species is named for my (VGS) wife, Shirley, in recognition of her unstinting support for my research for the past almost 40 years.
</paragraph>
</subSubSection>
</treatment>
</document>