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<document id="453E68A6F365527817803D843A6C204E" ID-DOI="10.11646/zootaxa.5507.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13747694" ID-ZooBank="09AC0223-FC47-4C34-8719-0376CD92C837" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726055664737" checkinUser="plazi" docAuthor="Viraktamath, C. A., Webb, M. D. &amp; Yeshwanth, H. M." docDate="2024" docId="7E7D0B1F766C381DFF1EF9A9FB4FFCCC" docLanguage="en" docName="zootaxa.5507.1.4.pdf" docOrigin="Zootaxa 5507 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Libengaia (Nandiana) lamerallis Sunil &amp; Meshram 2023" docType="treatment" docVersion="2" lastPageNumber="115" masterDocId="8244736776673816FF89FFB7F83CFB36" masterDocTitle="Opsiini leafhopper genus Libengaia Linnavuori (Hemiptera: Cicadellidae: Deltocephalinae) in Africa and India with description of one new subgenus and six new species" masterLastPageNumber="122" masterPageNumber="104" pageNumber="115" updateTime="1726056483101" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AD4C310E8803DA5857D296C7F8B2CE32">Opsiini leafhopper genus Libengaia Linnavuori (Hemiptera: Cicadellidae: Deltocephalinae) in Africa and India with description of one new subgenus and six new species</mods:title>
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<mods:namePart id="DD1C0D722107E00D09DD883CFDB40E03">Viraktamath, C. A.</mods:namePart>
<mods:affiliation id="F386711209C7AA35256AFC5E83F0D463">Department of Entomology, University of Agricultural Sciences, GKVK, Bengaluru 560065, India</mods:affiliation>
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<mods:namePart id="8D550D20A145D247D4BE98153C54AFB3">Webb, M. D.</mods:namePart>
<mods:affiliation id="7E72D4535537F98C43BD9BB7616EA692">Department of Life Science, The Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom</mods:affiliation>
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<mods:namePart id="B453B9408ECEFB45EA19A2F95107C4C2">Yeshwanth, H. M.</mods:namePart>
<mods:affiliation id="262019BDE96FF1455DC52203B8ACD3D4">Department of Entomology, University of Agricultural Sciences, GKVK, Bengaluru 560065, India</mods:affiliation>
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<mods:title id="C767F52ABAB12EA99D0B6F9ED08A4F94">Zootaxa</mods:title>
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<mods:date id="C40FBE64B3CC9BF426CAE028A3486257">2024</mods:date>
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<mods:number id="DBF4EF5E0FB5D88AE012238B2C2A6EEF">5507</mods:number>
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<treatment id="7E7D0B1F766C381DFF1EF9A9FB4FFCCC" ID-DOI="http://doi.org/10.5281/zenodo.13747632" ID-Zenodo-Dep="13747632" LSID="urn:lsid:plazi:treatment:7E7D0B1F766C381DFF1EF9A9FB4FFCCC" httpUri="http://treatment.plazi.org/id/7E7D0B1F766C381DFF1EF9A9FB4FFCCC" lastPageNumber="115" pageId="11" pageNumber="115">
<subSubSection id="BECEE982766C381DFF1EF9A9FB32FD0F" box="[151,782,1566,1593]" pageId="11" pageNumber="115" type="nomenclature">
<paragraph id="F66BBA09766C381DFF1EF9A9FB32FD0F" blockId="11.[151,782,1566,1629]" box="[151,782,1566,1593]" pageId="11" pageNumber="115">
<heading id="AD230D65766C381DFF1EF9A9FB32FD0F" bold="true" box="[151,782,1566,1593]" fontSize="11" level="1" pageId="11" pageNumber="115" reason="1">
<taxonomicName id="31D4C18A766C381DFF1EF9A9FB32FD0F" ID-CoL="BV683" authority="Sunil &amp; Meshram" authorityName="Sunil &amp; Meshram" authorityYear="2023" box="[151,782,1566,1593]" class="Insecta" family="Cicadellidae" genus="Libengaia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="115" phylum="Arthropoda" rank="species" species="lamerallis" subGenus="Nandiana">
<emphasis id="C4A0661B766C381DFF1EF9A9FB32FD0F" bold="true" box="[151,782,1566,1593]" pageId="11" pageNumber="115">
<emphasis id="C4A0661B766C381DFF1EF9A9F92DFD0F" bold="true" box="[151,273,1566,1593]" italics="true" pageId="11" pageNumber="115">Libengaia</emphasis>
(
<emphasis id="C4A0661B766C381DFEAAF9A9F9A0FD0F" bold="true" box="[291,412,1566,1593]" italics="true" pageId="11" pageNumber="115">Nandiana</emphasis>
)
<emphasis id="C4A0661B766C381DFE25F9A9FA19FD0F" bold="true" box="[428,549,1566,1593]" italics="true" pageId="11" pageNumber="115">lamerallis</emphasis>
Sunil &amp; Meshram
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BECEE982766C381DFF1EF9F4F991FD6B" box="[151,429,1603,1629]" pageId="11" pageNumber="115" type="description">
<paragraph id="F66BBA09766C381DFF1EF9F4F991FD6B" blockId="11.[151,782,1566,1629]" box="[151,429,1603,1629]" pageId="11" pageNumber="115">
<figureCitation id="6EEFA68C766C381DFF1EF9F4F929FD6B" box="[151,277,1603,1629]" captionStart="FIGURE 6" captionStartId="12.[151,250,1885,1909]" captionTargetBox="[156,1417,186,1857]" captionTargetId="figure-18@12.[151,1435,181,1862]" captionTargetPageId="12" captionText="FIGURE 6 A–P. Species of Libengaia (Nandiana), habitus dorsal and lateral views.A– J, color variation in L. (N.) bharatiensis (Malhotra &amp; Shrma): specimens from Bandipur (AB), Khandala (CD), Mudigere (EF), Thirthahalli (GH) and Waghai (IJ); K–N, L. (N.) lamerallis Sunil &amp; Meshram, male (KL), female (MN); O–P, L. (N.) ramamurthyi sp. nov., holotype male." figureDoi="http://doi.org/10.5281/zenodo.13747706" httpUri="https://zenodo.org/record/13747706/files/figure.png" pageId="11" pageNumber="115">Figs 6K–N</figureCitation>
,
<figureCitation id="6EEFA68C766C381DFEA8F9F4F912FD6B" box="[289,302,1603,1629]" captionStart="FIGURE 7" captionStartId="13.[151,250,1866,1890]" captionTargetBox="[164,1414,195,1837]" captionTargetId="figure-28@13.[151,1436,181,1842]" captionTargetPageId="13" captionText="FIGURE 7 A–R. Species of Libengaia (Nandiana), faces and female genitalia.A–E, L. (N.) bharatiensis (Malhotra &amp; Sharma), color variation in face: specimens from Bandipur (A), Khandala (B), Mudigere (C), Thirthahalli (D) and Waghai (E); F–G, L. (N.) lamerallis Sunil &amp; Meshram, male and female; H, L. (N.) ramamurthyi sp. nov., holotype male face; I–M, L. (N.) bharatiensis: I, female sternite VII; JK, valvula I; LM, valvula II; N–R, L. (N.) lamerallis: N, female sternite VII; OP, valvula I; QR, valvula II." figureDoi="http://doi.org/10.5281/zenodo.13747708" httpUri="https://zenodo.org/record/13747708/files/figure.png" pageId="11" pageNumber="115">7</figureCitation>
FG, 9A–H.
</paragraph>
</subSubSection>
<subSubSection id="BECEE982766C381DFF1EF937FB01FDAE" box="[151,829,1664,1689]" pageId="11" pageNumber="115" type="reference_group">
<paragraph id="F66BBA09766C381DFF1EF937FB01FDAE" blockId="11.[151,829,1664,1689]" box="[151,829,1664,1689]" pageId="11" pageNumber="115">
<treatmentCitationGroup id="D6C49D27766C381DFF1EF937FB01FDAE" box="[151,829,1664,1689]" pageId="11" pageNumber="115">
<treatmentCitation id="77759C18766C381DFF1EF937FAA9FDAE" author="Sunil, S. &amp; Meshram, N. M." box="[151,661,1664,1689]" page="186" pageId="11" pageNumber="115" year="2024">
<taxonomicName id="31D4C18A766C381DFF1EF937FAA9FDAE" ID-CoL="BV683" authority="Sunil &amp; Meshram 2024: 186" authorityName="Sunil &amp; Meshram" authorityPageNumber="186" authorityYear="2023" box="[151,661,1664,1689]" class="Insecta" family="Cicadellidae" genus="Libengaia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="115" phylum="Arthropoda" rank="species" species="lamerallis">
<emphasis id="C4A0661B766C381DFF1EF937F957FDAE" box="[151,363,1664,1688]" italics="true" pageId="11" pageNumber="115">Libengaia lamerallis</emphasis>
<bibRefCitation id="9245C7F8766C381DFEF8F937FAA9FDAE" author="Sunil, S. &amp; Meshram, N. M." box="[369,661,1664,1689]" pageId="11" pageNumber="115" pagination="183 - 189" refId="ref8863" refString="Sunil, S. &amp; Meshram, N. M. (2024) First record of the leafhopper genus Libengaia from India, with description of new species (Hemiptera: Deltocephalinae: Opsiini). Biologia, 79, 183 - 189. https: // doi. org / 10.1007 / s 11756 - 023 - 01518 - z" type="journal article" year="2024">Sunil &amp; Meshram 2024: 186</bibRefCitation>
</taxonomicName>
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, figs 2a–k, 3a–i.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BECEE982766C381DFF1EF976F965FC5C" pageId="11" pageNumber="115" type="diagnosis">
<paragraph id="F66BBA09766C381DFF1EF976FB1AFDC8" blockId="11.[151,1437,1728,2043]" pageId="11" pageNumber="115">
<emphasis id="C4A0661B766C381DFF1EF976F935FDEC" box="[151,265,1729,1754]" italics="true" pageId="11" pageNumber="115">Diagnosis</emphasis>
. Externally resembling
<taxonomicName id="31D4C18A766C381DFD94F976FAC0FDEC" authorityName="Malhotra &amp; Sharma" authorityYear="1977" box="[541,764,1728,1754]" class="Insecta" family="Cicadellidae" genus="Litura" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="115" phylum="Arthropoda" rank="species" species="bharatiensis" subGenus="Nandiana">
<emphasis id="C4A0661B766C381DFD94F976FA0FFDEC" box="[541,563,1729,1754]" italics="true" pageId="11" pageNumber="115">L.</emphasis>
(
<emphasis id="C4A0661B766C381DFDCFF976FA5CFDEC" box="[582,608,1729,1754]" italics="true" pageId="11" pageNumber="115">N.</emphasis>
)
<emphasis id="C4A0661B766C381DFDFBF977FAC0FDEC" box="[626,764,1728,1754]" italics="true" pageId="11" pageNumber="115">bharatiensis</emphasis>
</taxonomicName>
but differs in having the aedeagal shaft with paired apical proesses and both male and female witout ivory markings.
</paragraph>
<paragraph id="F66BBA09766C381DFF4EF8BFF965FC5C" blockId="11.[151,1437,1728,2043]" pageId="11" pageNumber="115">
<emphasis id="C4A0661B766C381DFF4EF8BFF9A1FC14" box="[199,413,1800,1826]" italics="true" pageId="11" pageNumber="115">Material examined</emphasis>
:
<materialsCitation id="46BCB054766C381DFE26F8BFFD59FC14" box="[431,1381,1800,1827]" collectingDate="1979-07-15" collectionCode="UASB" collectorName="C. A. Viraktamath" country="India" location="Karnataka" pageId="11" pageNumber="115" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Karnataka">
<collectingCountry id="8EC3FA99766C381DFE26F8BFFA3FFC14" box="[431,515,1800,1826]" name="India" pageId="11" pageNumber="115">INDIA</collectingCountry>
:
<collectingRegion id="341074EB766C381DFD86F8BFFABBFC14" box="[527,647,1800,1826]" country="India" name="Karnataka" pageId="11" pageNumber="115">Karnataka</collectingRegion>
:
<specimenCount id="E0D27180766C381DFD1AF8BFFA8BFC14" box="[659,695,1800,1826]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
,
<specimenCount id="E0D27180766C381DFD4FF8BFFAD5FC14" box="[710,745,1800,1826]" count="1" pageId="11" pageNumber="115" type="female">1♀</specimenCount>
, Channapatna,
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<collectingDate id="922E6521766C381DFC12F8BFFC22FC15" box="[923,1054,1800,1827]" pageId="11" pageNumber="115" value="1979-07-15">15.vii.1979</collectingDate>
</date>
,
<collectorName id="5B21DFDF766C381DFBA2F8BFFCCBFC14" box="[1067,1271,1800,1826]" pageId="11" pageNumber="115">C.A. Viraktamath</collectorName>
; (
<collectionCode id="90C522CC766C381DFA85F8BFFD61FC14" box="[1292,1373,1800,1826]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="11" pageNumber="115">UASB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="46BCB054766C381DFAFBF8BFFA88FC70" collectingDate="1985-01-20" collectionCode="BMNH" collectorName="S. A. Viraktamath" country="India" pageId="11" pageNumber="115" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="E0D27180766C381DFAFBF8BFFDABFC14" box="[1394,1431,1800,1826]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
, Bandipur,
<date id="826A9CC9766C381DFE82F89BF94BFC71" box="[267,375,1836,1863]" pageId="11" pageNumber="115" value="1985-01-20">
<collectingDate id="922E6521766C381DFE82F89BF94BFC71" box="[267,375,1836,1863]" pageId="11" pageNumber="115" value="1985-01-20">20.i.1985</collectingDate>
</date>
,
<collectorName id="5B21DFDF766C381DFE0BF89BFA78FC70" box="[386,580,1836,1862]" pageId="11" pageNumber="115">S.A. Viraktamath</collectorName>
(
<collectionCode id="90C522CC766C381DFDDBF89BFA90FC70" box="[594,684,1836,1862]" pageId="11" pageNumber="115">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="46BCB054766C381DFD37F89BF969FC5C" collectingDate="2014-04-25" collectionCode="NPC" collectorName="Hg. Light, N." country="India" elevation="1269" location="Largi" pageId="11" pageNumber="115" specimenCount="1" specimenCount-male="1" stateProvince="Himachal Pradesh">
<collectingRegion id="341074EB766C381DFD37F89BFBB2FC70" box="[702,910,1836,1862]" country="India" name="Himachal Pradesh" pageId="11" pageNumber="115">Himachal Pradesh</collectingRegion>
:
<specimenCount id="E0D27180766C381DFC11F89BFB80FC70" box="[920,956,1836,1862]" count="1" pageId="11" pageNumber="115" type="male">1♂</specimenCount>
,
<location id="F30BECD2766C381DFC41F89BFC3BFC70" LSID="urn:lsid:plazi:treatment:7E7D0B1F766C381DFF1EF9A9FB4FFCCC:F30BECD2766C381DFC41F89BFC3BFC70" box="[968,1031,1836,1862]" country="India" name="Largi" pageId="11" pageNumber="115" stateProvince="Himachal Pradesh">Largi</location>
,
<quantity id="312C17EC766C381DFB98F89BFC5DFC70" box="[1041,1121,1836,1862]" metricMagnitude="3" metricUnit="m" metricValue="1.269" pageId="11" pageNumber="115" unit="m" value="1269.0">
<elevation id="7DF95D3A766C381DFB98F89BFC5DFC70" box="[1041,1121,1836,1862]" metricMagnitude="3" metricUnit="m" metricValue="1.269" pageId="11" pageNumber="115" unit="m" value="1269.0">1269m</elevation>
</quantity>
,
<date id="826A9CC9766C381DFBE5F89BFCD9FC71" box="[1132,1253,1836,1863]" pageId="11" pageNumber="115" value="2014-04-25">
<collectingDate id="922E6521766C381DFBE5F89BFCD9FC71" box="[1132,1253,1836,1863]" pageId="11" pageNumber="115" value="2014-04-25">25.iv.2014</collectingDate>
</date>
,
<collectorName id="5B21DFDF766C381DFB66F89AFDBEFC71" box="[1263,1410,1836,1863]" pageId="11" pageNumber="115">Hg. Light, N.</collectorName>
M Meshram (
<collectionCode id="90C522CC766C381DFE99F8E7F971FC5C" box="[272,333,1872,1898]" country="India" httpUri="http://grbio.org/cool/nwky-2b63" name="National Pusa Collection" pageId="11" pageNumber="115">NPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BECEE982766C381DFF4EF8C3FB4FFCCC" pageId="11" pageNumber="115" type="discussion">
<paragraph id="F66BBA09766C381DFF4EF8C3FB4FFCCC" blockId="11.[151,1437,1728,2043]" pageId="11" pageNumber="115">
<emphasis id="C4A0661B766C381DFF4EF8C3F907FCB8" bold="true" box="[199,315,1908,1934]" pageId="11" pageNumber="115">Remarks.</emphasis>
<bibRefCitation id="9245C7F8766C381DFECBF8C3FA67FCB8" author="Sunil, S. &amp; Meshram, N. M." box="[322,603,1908,1934]" pageId="11" pageNumber="115" pagination="183 - 189" refId="ref8863" refString="Sunil, S. &amp; Meshram, N. M. (2024) First record of the leafhopper genus Libengaia from India, with description of new species (Hemiptera: Deltocephalinae: Opsiini). Biologia, 79, 183 - 189. https: // doi. org / 10.1007 / s 11756 - 023 - 01518 - z" type="journal article" year="2024">Sunil &amp; Meshram (2024)</bibRefCitation>
adequately described this species based on specimens collected from New
<collectingRegion id="341074EB766C381DFF1EF82FF8E9FC84" box="[151,213,1944,1970]" country="India" name="Delhi" pageId="11" pageNumber="115">Delhi</collectingRegion>
and illustrated both male and female genitalia. Here it is illustrated based on specimens collected from south
<collectingCountry id="8EC3FA99766C381DFF1EF80BF8E8FCE0" box="[151,212,1980,2006]" name="India" pageId="11" pageNumber="115">India</collectingCountry>
.
<taxonomicName id="31D4C18A766C381DFF57F80BF9A5FCE0" authorityName="Sunil &amp; Meshram" authorityYear="2023" box="[222,409,1980,2006]" class="Insecta" family="Cicadellidae" genus="Libengaia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="115" phylum="Arthropoda" rank="species" species="lamerallis" subGenus="Nandiana">
<emphasis id="C4A0661B766C381DFF57F80BF9A5FCE0" box="[222,409,1980,2006]" italics="true" pageId="11" pageNumber="115">L.(N.) lamerallis</emphasis>
</taxonomicName>
resembles
<taxonomicName id="31D4C18A766C381DFD9DF80BFADBFCE0" authorityName="Malhotra &amp; Sharma" authorityYear="1977" box="[532,743,1980,2006]" class="Insecta" family="Cicadellidae" genus="Litura" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="115" phylum="Arthropoda" rank="species" species="bharatiensis" subGenus="Nandiana">
<emphasis id="C4A0661B766C381DFD9DF80BFADBFCE0" box="[532,743,1980,2006]" italics="true" pageId="11" pageNumber="115">L.(N.) bharatiensis</emphasis>
</taxonomicName>
but differs in having bifid aedeagal processes, lamellate shafts and in the absence of processes on the preatrium of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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