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<mods:title id="C112C1EB8A3BE0D49CCF813AE1A13CED">A taxonomic revision of the catshark genus Haploblepharus Garman 1913 (Chondrichthyes: Carcharhiniformes: Scyliorhinidae).</mods:title>
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<mods:namePart id="43CF82B3BAAB7A44B19A535DF4D490D5">Brett A. Human</mods:namePart>
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<treatment id="A4BB123BD41E9989A9B0379559F46E92" ID-DOI="http://doi.org/10.5281/zenodo.6248469" ID-GBIF-Taxon="100117391" ID-Zenodo-Dep="6248469" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92" httpUri="http://treatment.plazi.org/id/A4BB123BD41E9989A9B0379559F46E92" lastPageNumber="20" pageNumber="14">
<subSubSection id="266506081C8FC59716DA8AB822E5C20C" type="nomenclature">
<paragraph id="612EBB323101C0C1490AC64F2C4E2175" pageNumber="14">
<taxonomicName id="237E79F49EEB95A135651FBA9AC71E45" ID-CoL="3JL89" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="pictus">
Haploblepharus pictus (
<normalizedToken id="1D879BFAE85A9FA64B1EEBAE403D1737" originalValue="Müller">Mueller</normalizedToken>
&amp; Henle, 1838)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C871572249F30D34109AD0813D9BD094" type="description">
<paragraph id="BEC3FE925174695CA456654F8F59F28D" pageNumber="14">(Fig. 5, Table 2)</paragraph>
</subSubSection>
<subSubSection id="3AFFF72349A13A1B76771AEC2A0A1AB3" type="reference_group">
<paragraph id="B16DDC20C0A60040D0158BAB8BD86D77" pageNumber="14">
<taxonomicName id="BA21915CADF025D4FD378F01E7F1D4C9" LSID-ZBK="urn:lsid:zoobank.org:act:A528ABC8-509D-47A7-9129-3B9249AA0B02" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Scyllium pictum Müller &amp; Henle 1838:4" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="pictum">
Scyllium pictum
<normalizedToken id="C02B3F91FC79A8212E82ADEE50F558C0" originalValue="Müller">Mueller</normalizedToken>
&amp; Henle, 1838
</taxonomicName>
: 4.
</paragraph>
<paragraph id="2B0A78257E858A4A679ED54787463F60" pageNumber="14">
<taxonomicName id="F744FD2F8FD75D760F4F62CE505CE071" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="edwardsii">Haploblepharus edwardsii</taxonomicName>
:
<normalizedToken id="F339EAE0EB576B92F642EA1D7C055A63" originalValue="Günther">Guenther</normalizedToken>
, 1870: 401 (in part); Regan, 1908: 463 (in part); Garman, 1913: 102 (in part); Thompson, 1914: 137 (in part); Gilchrist &amp; Thompson, 1916: 283 (in part); Fowler, 1941: 64 (in part); von Bonde, 1945a: 1 (in part); von Bonde, 1945b: 220 (in part); Smith, 1949: 54 (in part); Smith, 1950: 879 (in part).
</paragraph>
</subSubSection>
<subSubSection id="152C3F3A792F519FE9894E69FC969552" type="nomenclature">
<paragraph id="9F7BFD161A5A16A225D60E15B71B4113" pageNumber="14">
Type series and locality. Probable
<typeStatus id="44DD1C37B3196865FE2D7EDD14FEEED2">syntypes</typeStatus>
(see Remarks below), five specimens in four lots.
<specimenCode id="4BAE958BAAB40EA78CF0CC24A5570B55">
<collectionCode id="7FDA7E34A8F4683EC331A9E088DFE3D2">RMNH</collectionCode>
4161
</specimenCode>
, two specimens, mature male 522 mm TL, and mature female 490 mm TL, both are dried specimens. The female appears to be in poor condition and the male in very poor condition.
<specimenCode id="0EF3593E5581F4D84329CC13DDDEB646">
<collectionCode id="60174573F473058813C383FF45B35BB3">RMNH</collectionCode>
4162
</specimenCode>
, immature female 135 mm TL, in reasonable condition.
<specimenCode id="3D648061BFABFEE28800918CE3A53A01">
<collectionCode id="8D18956519B281BE9FD71ACD318152F6">RMNH</collectionCode>
4163
</specimenCode>
, immature male 199 mm TL, in reasonable condition.
<specimenCode id="CE2C5FCDDEAB19E4BA31606F8BF2758F">
<collectionCode id="1689C65ED641778F509B578455849DB3">RMNH</collectionCode>
4164
</specimenCode>
, immature female 143 mm TL, in good condition.
</paragraph>
<paragraph id="080E6788B6A60C1A405DDD1EF06EBB5F" pageNumber="14">
<materialsCitation id="4222A60C6DF157785F19CFF33E036C3C" ID-GBIF-Occurrence="923889893" collectingDate="1834" collectionCode="RMNH" collectorName="H.B. van Horstok" country="South Africa" location="“Vom Cap der guten Hoffnung” (from Cape of Good Hope)" specimenCode="RMNH 4161 - 4164" typeStatus="Syntypes">
The entire syntypic series collected
<location id="22D96D3FA683B5099F8531DA32C13888" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:22D96D3FA683B5099F8531DA32C13888">“Vom Cap der guten Hoffnung” (from Cape of Good Hope)</location>
,
<collectingCountry id="BCE2A69AC337C63A9120D0A2CD403B86">South Africa</collectingCountry>
,
<collectingDate id="04C1376F6A9B96E7295F47DF950F26F9">1826-1834</collectingDate>
(Leg. &amp; don.:
<collectorName id="1779B7FCBE80E9E3AC3A4A82881EF339">H.B. van Horstok</collectorName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D7FB128D122B050F5A74EB517C4D0B06" type="diagnosis">
<paragraph id="F858C6172F39B0BCB4E27366135E5D44" pageNumber="14">
Diagnosis. The author was not able to personally measure the putative type series (see remarks below), and given the highly contorted condition of those specimens, the measurements obtainable from those specimens would likely be inaccurate. The following is based on the mean measurements of
<taxonomicName id="214A706541B525C2F7E56FD97527B21E" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
specimens examined by the author (see study material).
<taxonomicName id="6041A4AF78EDEAF2C4E792AE3AFBA4D9" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
is distinguished from other
<taxonomicName id="EA6A633024E10AB366EDDCF680DE7D84" LSID-ZBK="urn:lsid:zoobank.org:act:87028B01-5D5B-4004-918B-492D3059F90E" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus Garman 1913:101" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="genus">Haploblepharus</taxonomicName>
sharks in having a slender body in juvenile individuals but becoming stockier in mature individuals, abdomen width 11.2% TL; snout broadly rounded, not coming to a point; head width at the posterior margin of the orbit 13.0% TL; head strongly depressed, head height at the posterior margin of the orbit 6.7% TL; trunk somewhat depressed, trunk height 10.2% TL and trunk width 12.3% TL; claspers of mature males stout, inner length 6.2 times the base.
<taxonomicName id="B0552775FBFAFDF2B6B87333C959F897" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
specimens have 45-83 rows of teeth in the upper jaw (mean 72.0) and 47-75 rows of teeth in the lower jaw (mean 62.8). The number of vertebral centra in
<taxonomicName id="67D37A0E863C7BA987958B8DC14371A3" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
specimens ranges from 93-133 (mean 124.5) total.
<taxonomicName id="00FF828AF2ED79FC31EC92DAC87C5E17" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
has a highly variable colour pattern that can superficially resemble
<taxonomicName id="BB8556C7DDA3EDEA1074E57964A5451D" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
or
<taxonomicName id="511404A1C7DB168F052DB90902BEA187" LSID-ZBK="urn:lsid:zoobank.org:act:5B016902-D9E6-47E1-9F91-E23E5AA226C2" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus fuscus Smith 1950:883" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="fuscus">H. fuscus</taxonomicName>
in some individuals, although
<taxonomicName id="EA0C92E34BEC60FE5AFA13E6ACC25ACE" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
more commonly has 6-8 distinct, dark saddles.
</paragraph>
</subSubSection>
<subSubSection id="D15B769D4E8531A99903EF869838C22A" type="description">
<paragraph id="1C8D39C5B38F20641CB9E2A6B2AC40BC" lastPageNumber="15" pageNumber="14">
Description. Morphometric and meristic data are given in Table 2. The following is based on the mean measurements of
<taxonomicName id="BD18502A5B8773F98F9CF04A912F23CD" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
specimens examined by the author (see Study material):
<taxonomicName id="AFCE57C46E52EA4E8DAE589167B5341A" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
is a slender to moderately stocky bodied
<taxonomicName id="3F964152387C1A21119A8860ED7C0543" LSID-ZBK="urn:lsid:zoobank.org:act:87028B01-5D5B-4004-918B-492D3059F90E" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus Garman 1913:101" order="Carcharhiniformes" pageNumber="14" phylum="Chordata" rank="genus">Haploblepharus</taxonomicName>
shark with a relatively broad head, head width at the pectoral origin3.63 times the preoral length; head length 1.26 times distance from snout tip to first gill slit; height of first gill slit 1.76 times the height of the fifth gill slit; eye length 4.29 times longer than spiracle length; basimandibular cartilage found at the symphysis of the Meckels cartilage in the lower jaw; mouth length 1.23 times the prenarial length; mouth width 6.6 times the upper labial furrow length; labial cartilages present; nasal lobes fused into a nasal flap that covers the excurrent apertures and extends to the mouth; interorbital width 1.19 times the nasal flap width; head very strongly depressed, head width at the posterior margin of the orbit 1.95 times its height; head width 1.48 times its height; trunk depressed, trunk width 1.21 times its height; abdomen not depressed, abdomen width 0.99 times its height; tail not depressed, tail width 1.04 times its height; caudal peduncle strongly compressed, caudal peduncle width 0.61 times its height; precaudal length 1.76 times the distance from snout to first dorsal fin; dorsal fins rounded; height of first dorsal fin 1.04 times that of the second dorsal fin; first dorsal fin length equal to the length of its anterior margin; second dorsal fin length equal to the length of its anterior margin; pectoral fin to pelvic fin space 1.33 times the interdorsal space; pectoral and pelvic fins rounded; pectoral fin height 1.8 times the height of the pelvic fin; pectoral fin length 0.94 times the length of its anterior margin; pelvic fin length 1.46 times the length of its anterior margin; claspers of mature males stout, clasper inner length 6.17 times the base; anal fin to caudal fin space 1.1 times the head height at the origin of the pectoral fin; length of anal fin base 1.21 the length of the second dorsal fin base; anal fin length 1.4 times the length of its anterior margin; distance from pectoral fin insertion to the midpoint of the first dorsal fin length 1.51 times the caudal dorsal margin length. Vertebral counts: total 124.5 (93-133), 35 (30-38) monospondylous, 52.2 (35-57) precaudal diplospondylous and 37.7 (28-45) caudal diplospondylous vertebrae. Dental formula: upper jaw (left) 35.6 (23-40), (right) 36.4 (22-43); lower jaw (left) 31.6 (23-37), (right) 31.2 (24-38).
</paragraph>
<paragraph id="90421E4A1FF3FB597B9084DD53582A3C" pageNumber="16">
Size and sexual maturity. In this study,
<taxonomicName id="E6B19120752E62E947D86FBFB0F9BBE0" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="16" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
males were found to be embryonic at 103 mm TL, juvenile at 165 mm TL to 369 mm TL, adolescent at 282 mm TL to 407 mm TL, and mature at 400 mm TL to 570 mm TL. Females were found to be juvenile at 110 mm TL to 306 mm TL, adolescent at 430 mm TL to 533 mm TL, and mature at 359 mm TL to 597 mm TL. There is no sexual dimorphism apparent in this species.
</paragraph>
<paragraph id="63101CE425D07F49CA1301768E25DE91" pageNumber="16">
Colouration.
<taxonomicName id="2260D13195BAA27F823616216CE633F0" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="16" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
has a background dorsal colouration pale to very dark brown, almost black, pale grey brown to dark grey brown, red brown, or grey; white spots variably present (Figs. 5A, C-E) or absent (Fig. 5F), occurring on the saddles when spots present, between and on the saddles, but never just between the saddles; number of saddles, when present, 6 to 8, variably placed; centre of saddles range from orange, brown, dark brown to almost black, margins variably present, when margins present may be indistinct; background colouration extending to dorsal surface of pectoral and pelvic fins, including spots when present; saddles not extending to dorsal surface of pectoral and pelvic fins. Ventral colouration abruptly uniform white (Fig. 5B), cream or red brown (latter probably a preservation artefact); ventral surfaces of pectoral and pelvic fins, as well as anal fin, same colouration as the dorsal background colouration, occasionally dark spots and blotches present.
</paragraph>
</subSubSection>
<subSubSection id="FC220F692393E6C3E037AA5B79A965A2" type="discussion">
<paragraph id="514066541DC17AAE8B67E4DEC6563634" pageNumber="17">
Comparison with other species.
<taxonomicName id="A128C1072CDBA205829009F5B559C508" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
has a highly variable colour pattern and is easily misidentified for other members of the genus when using colour pattern alone.
<taxonomicName id="DA1B6622788BB816A9B575C8799BF25B" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
is a relatively large member of the genus, second to
<taxonomicName id="99AA84660BE6C0A49A91FA75D07A1064" LSID-ZBK="urn:lsid:zoobank.org:act:5B016902-D9E6-47E1-9F91-E23E5AA226C2" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus fuscus Smith 1950:883" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="fuscus">H. fuscus</taxonomicName>
in overall size, and is more robust than
<taxonomicName id="D27549993E36546CCCCC3F035AAB4E7B" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
or
<taxonomicName id="8C0CC0C0253BF63F6AD6FC2839108B60" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="kistnasamyi">H. kistnasamyi</taxonomicName>
. The head is more bluntly rounded than in
<taxonomicName id="B214F4B355DD1DAD688833DFD28D7B8D" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
and
<taxonomicName id="279913F878558441B06CC0D8E7C250A2" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="kistnasamyi">H. kistnasamyi</taxonomicName>
, and about equally so with
<taxonomicName id="938E930374C5825D7CBCDAB559C54EE4" LSID-ZBK="urn:lsid:zoobank.org:act:5B016902-D9E6-47E1-9F91-E23E5AA226C2" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus fuscus Smith 1950:883" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="fuscus">H. fuscus</taxonomicName>
. Head and body is depressed more so than in
<taxonomicName id="9B8F89B73D4B8897311C6C2148C8FDCB" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="kistnasamyi">H. kistnasamyi</taxonomicName>
and
<taxonomicName id="81CCC94E7A8302BF49FC57BA6FBAE0AA" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
, but less so than
<taxonomicName id="5B2A67859F7D316836B808CCE1D3E46C" LSID-ZBK="urn:lsid:zoobank.org:act:5B016902-D9E6-47E1-9F91-E23E5AA226C2" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus fuscus Smith 1950:883" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="fuscus">H. fuscus</taxonomicName>
. Claspers of mature males are equivalent in size to
<taxonomicName id="529ABC262A29C70F3367ECE38F21EF5A" LSID-ZBK="urn:lsid:zoobank.org:act:5B016902-D9E6-47E1-9F91-E23E5AA226C2" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus fuscus Smith 1950:883" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="fuscus">H. fuscus</taxonomicName>
and
<taxonomicName id="AEA70DE3153A55D37760B32C34F84005" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="kistnasamyi">H. kistnasamyi</taxonomicName>
, although longer and stouter than in
<taxonomicName id="8E2F6B83C071DABD4D736EB59F698836" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
.
<taxonomicName id="91586E65D46EF3A7EAD1F873609AF0FC" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
has a lower total tooth count than
<taxonomicName id="E3B7E35AE6E38CA719B32E26632374C6" LSID-ZBK="urn:lsid:zoobank.org:act:5B016902-D9E6-47E1-9F91-E23E5AA226C2" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus fuscus Smith 1950:883" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="fuscus">H. fuscus</taxonomicName>
, but higher than either
<taxonomicName id="5CA796CCE449121E0CA9EB0FFC533C99" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
or
<taxonomicName id="AADBC13F84017450EFA5D775636EED50" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="kistnasamyi">H. kistnasamyi</taxonomicName>
.
<taxonomicName id="C020D1214BEC89950BB76C05B2DF11FD" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
is most similar to
<taxonomicName id="F2508EF5E8739C1E927CC601D85D1050" LSID-ZBK="urn:lsid:zoobank.org:act:5B016902-D9E6-47E1-9F91-E23E5AA226C2" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus fuscus Smith 1950:883" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="fuscus">H. fuscus</taxonomicName>
in overall morphology, and is distinguished from that species by having distinct saddles and spots, less vertebrae, and more tooth rows. The variable colour pattern of
<taxonomicName id="8680D8CCDE0E3049664229481906541E" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
can superficially resemble all other
<taxonomicName id="9B5C7212D92F509273E44D290E689EF5" LSID-ZBK="urn:lsid:zoobank.org:act:87028B01-5D5B-4004-918B-492D3059F90E" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus Garman 1913:101" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="genus">Haploblepharus</taxonomicName>
species, particularly
<taxonomicName id="A7BAA4B321ADEC0E7353D1154186A8C4" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
(Fig. 5D).
</paragraph>
</subSubSection>
<subSubSection id="0262B091F6A1148223D6761FC8CC7E9A" type="discussion">
<paragraph id="2805464FD7B1290A5900FA4E8DFB4C7B" lastPageNumber="18" pageNumber="17">
Remarks.
<normalizedToken id="B46D8A7000D93A556D58B2C921007F3F" originalValue="Müller">Mueller</normalizedToken>
&amp; Henle (1838-1841) refer to specimens in the Museum in Leyden (now the Rijksmuuseum van Natuurlijke Historie, Leiden, RMNH) and in the British Museum (BMNH), but give no further details of the type material. RMNH 4161 and RMNH 4164 bear the name
<taxonomicName id="43E15DC89D9C3571EFB541D13CCD6B95" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyllium" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="edwardsii">Scyllium edwardsii</taxonomicName>
, while RMNH 4162 and RMNH 4163, have two names written on the label,
<taxonomicName id="B1B8E84778B3A238A04E05956317E706" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyllium" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="edwardsii">Scyllium edwardsii</taxonomicName>
and
<taxonomicName id="6926A3102D2E023C127F1CD434168020" LSID-ZBK="urn:lsid:zoobank.org:act:A528ABC8-509D-47A7-9129-3B9249AA0B02" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Scyllium pictum Müller &amp; Henle 1838:4" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="pictum">Scyllium pictum</taxonomicName>
. All four lots were in the museum during the time of
<normalizedToken id="BD3A5EE91E021D5E88401E57672D42A7" originalValue="Müller">Mueller</normalizedToken>
and Henles visit to the museum (M. van Oijen, pers. comm.). Photographs of these specimens (supplied by M. van Oijen) were examined by the author. The specimens are in poor condition, and although the colour patterning on most of the specimens are considerably faded, saddles are visible on all specimens, and all specimens appear to be
<taxonomicName id="7D0D03FCB915C8E5DC54E03E898F34E1" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
. Although there is no label identifying them as such, the RMNH specimens are probably the syntypes for
<taxonomicName id="392D07F3CAB7F6379027EC21E7D1FAF1" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
. Of the scyliorhinid specimens in the BMNH that have been assigned a species name, none are referable to
<taxonomicName id="EF35D75EE693162052617D0347774D4D" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
. As there are many lots in the BMNH under the name
<taxonomicName id="109CAF722B0F1DAA9333241D556F2F40" LSID-ZBK="urn:lsid:zoobank.org:act:BE653649-A8A7-4C6E-910E-F69CE41A83A3" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Scyllium Cuvier 1816:124" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="genus">Scyllium</taxonomicName>
sp., it was not practical to examine all of these specimens remotely, and the status of types for
<taxonomicName id="19B5A51EE429B994FB4F717BEE811285" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="17" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
in the BMNH remains unresolved.
</paragraph>
<paragraph id="2DE9D672D2BC7D4FDA993D69A6B854F7" pageNumber="18">
Eschmeyer (1998) refers to two possible syntypes in the Naturhistorisches Museum, Vienna, Lot NMW 78529, represented by an immature female (NMW 78529:1 approx. 200 mm TL) and an immature male (NMW 78529:2, approx. 110 mm TL). These specimens were purchased in 1842 from Ludwig Parreyss by the Naturhistorisches Museum, and may have been acquired from the museums in Leiden or London, however there are no labels accompanying the specimens indicating from where the specimens were procured, or whether they were used as type material. If these specimens did come from either the Leiden or London museums, then the specimens were probably available at the time
<normalizedToken id="6E2003571D421A9AE3906FBE34BAC54F" originalValue="Müller">Mueller</normalizedToken>
and Henle visited those museums. The only label accompanying the specimens currently, is a label dated at the time Steindachner was at the museum, which was some time after the specimens were deposited there (H. Wellendorf, pers. comm.). Photos taken of these specimens (supplied by H. Wellendorf) were examined by the author and identified as
<taxonomicName id="FE5C0AA02F0AEEAAC85D61EFA6111BFD" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
, however, NMW 78529 probably does not represent type material. Springer (1979) made no reference to type material for
<taxonomicName id="EF91086D0ADC71EF8EA938FF8174950F" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
, and Compagno (1984b, 1988) did not locate type material for
<taxonomicName id="82A712E1D6135763AF18D43F3BF61832" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
.
</paragraph>
<paragraph id="79B6D22BA1316875557D2DE3A6339BC1" pageNumber="18">
Since the description of
<taxonomicName id="6C7470EAB2DF6A8356C54A6B366BAFED" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
by
<normalizedToken id="54D8A1DADA649AEBB19113DD54935EF5" originalValue="Müller">Mueller</normalizedToken>
&amp; Henle (1838-1841), many authors have chosen to synonymise it with
<taxonomicName id="C8EEB7DE8E948EACAE8ABDDD0D5FCF76" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
(e.g., Fowler, 1941; Smith, 1950), and this was due, in the most part, to the loss of the syntypes of
<taxonomicName id="96E50D5CDE2982A3F5608F88D13555C9" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
. Bass et al. (1975) revived
<taxonomicName id="B84B37E7769D69522B853036241EE17D" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
, which has been followed by subsequent authors. Bass et al. (1975) report that
<taxonomicName id="D17EC547466D4342F19BD3FF63A6F3DE" LSID-ZBK="urn:lsid:zoobank.org:act:E79C1C0C-404E-4BA5-98F9-1E9B9E158492" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Squalus pictus Lichtenstein 1823:111" order="Squaliformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">Squalus pictus</taxonomicName>
described by Lichtenstein in 1823 is probably not
<taxonomicName id="B2567ECC7FD68EF75491E52407B38C96" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
and may or may not be referable to
<taxonomicName id="8CFB6FFC0104DD77077A073B034DB95B" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
. There is no type material for this species and the description is insufficient to unambiguously identify this species.
</paragraph>
<paragraph id="D7F82D9066B6D866FB13913F2A59B5B6" pageNumber="18">
<taxonomicName id="107919C64E5246B52801DB69C14FC7DE" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
is regularly misidentified, mostly for
<taxonomicName id="D1C4A30A8C77DCE8A0C3FDE29BDE47A5" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
(Fig. 5D), but also for
<taxonomicName id="F6E3B9F05CF3E5ED4050DD466F827C7A" LSID-ZBK="urn:lsid:zoobank.org:act:5B016902-D9E6-47E1-9F91-E23E5AA226C2" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus fuscus Smith 1950:883" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="fuscus">H. fuscus</taxonomicName>
(Fig. 5F). This misidentification has led to an easterly range extension in its distribution (Fig. 6, and see below). The problems with misidentification may also underestimate the abundance of this species and overestimate the abundance of
<taxonomicName id="BB81FB80D14111858A78A2013F143F7F" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
. It is hoped that the current revision will aid in the identification of these species in the field and allow for a more accurate representation of distribution and abundance for both
<taxonomicName id="10FD4CDE0266F54992E7F778044C8EB3" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
and
<taxonomicName id="9CB0E53DCB550D38FFEEB00BFC357E2E" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
.
</paragraph>
<paragraph id="146A032B10B748E47DF2AAEA3B973474" pageNumber="18">
The morphometric proportions given here agree with those given by Bass et al. (1975) and Springer (1979). The dentition and denticles have been described and illustrated by Bass et al. (1975) and Compagno (1984b). Bass et al. (1975) found that there was strong sexual heterodonty in
<taxonomicName id="EE1CCA31DB36722C389618BD7053DEDE" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
. Vertebral counts given here agree with Bass et al. (1975) and Springer (1979).
</paragraph>
<paragraph id="97BD05C2C92BFA882E957C23F4B5CB0C" pageNumber="18">
<taxonomicName id="AC5FCE879CD3FBF4755902FF99BB127D" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
has a highly variable colour pattern (Fig. 5), some of these stunning, with no identifiable forms, as opposed to
<taxonomicName id="DB7B4EF6EBE77A62C13515A2CD4C8526" class="Elasmobranchii" family="Scyliorhinidae" genus="Poroderma" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pantherinum">Poroderma pantherinum</taxonomicName>
for example, which also has a highly variable colour pattern, but with identifiable forms (Compagno et al., 1989; Human, 2006a). Saddles are rarely absent, and some specimens resembling
<taxonomicName id="776E244135B232C9C76638BE0165C032" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
has led to the misidentification of
<taxonomicName id="E02C189CE49A84C24EBE9F38793F9DC5" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
as that species.
</paragraph>
<paragraph id="3F2433CF001843BF03FF08C6E15C6ADA" pageNumber="18">
Yeld &amp; Smit (2006) have identified a blood parasite in this species (along with
<taxonomicName id="49605F27942A7CD166F4CA625799903A" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
) that constitutes the first record of a blood parasite from a South African chondrichthyan, and
<taxonomicName id="FCCB19AD13357E03653A1D99C8269B7C" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
is the type host for
<taxonomicName id="8DBBF9C1375CAC4C77B913A2C478D524" class="Kinetoplastea" family="Trypanosomatidae" genus="Trypanosoma" higherTaxonomySource="CoL" kingdom="Protozoa" order="Trypanosomatida" pageNumber="18" phylum="Euglenozoa" rank="species" species="haploblephari">Trypanosoma haploblephari Yeld &amp; Smit 2006</taxonomicName>
(Mastigophora: Trypanosomatidae).
</paragraph>
</subSubSection>
<subSubSection id="F9A4F8FB284DF51257D5C9503930D0CA" type="distribution">
<paragraph id="F401C767EA7EE840A155B8D9A7F4A465" pageNumber="18">
Distribution. Figure 6 shows the distribution of
<taxonomicName id="2276A3155CE1C7F5F5F9466CCA3AB397" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
as verified by this study.
<taxonomicName id="DD1D70FD4A0F1AC177C5E4A2C20C3C1F" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="18" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
occurs north of
<normalizedToken id="E811143251DE8D7AAF076DB83DBC1051" originalValue="Lüderitz">Luederitz</normalizedToken>
, Namibia, extending south into South Africa to Cape of Good Hope, then east to into the Indian Ocean, east of the Storms River mouth, Eastern Cape.
</paragraph>
</subSubSection>
<subSubSection id="DB294070B07031A531028EAC338D2CAC" type="distribution">
<paragraph id="2CE144F927BD656E9C61611E1A53157D" pageNumber="20">
The present revision of
<taxonomicName id="15808CCA9085F23FC04B9113C4002999" LSID-ZBK="urn:lsid:zoobank.org:act:87028B01-5D5B-4004-918B-492D3059F90E" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus Garman 1913:101" order="Carcharhiniformes" pageNumber="20" phylum="Chordata" rank="genus">Haploblepharus</taxonomicName>
has led to a major range extension for
<taxonomicName id="BF6ED11ED7BA2EDA0336AB823F2D572E" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="20" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
, which was previously thought to occur only as far east as Cape Agulhas (Bass et al., 1975; Compagno, 1988; Compagno et al., 1989). The previous distribution of
<taxonomicName id="79BF3B4E6D6532185E6E72BB6490DFA0" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="20" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
was due to misidentification of this species with
<taxonomicName id="E1EFF2FB69F363358349559FF56D1783" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="20" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
and
<taxonomicName id="39D5D593EF01F4ABFF1666AB6FF64FA2" LSID-ZBK="urn:lsid:zoobank.org:act:5B016902-D9E6-47E1-9F91-E23E5AA226C2" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus fuscus Smith 1950:883" order="Carcharhiniformes" pageNumber="20" phylum="Chordata" rank="species" species="fuscus">H. fuscus</taxonomicName>
, the former in particular, east of Cape Agulhas.
</paragraph>
<paragraph id="5ECC5429889616A972BD6FDBFE9D541E" pageNumber="20">
<taxonomicName id="170AB49D93D1EE1B3312006127632EA0" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="20" phylum="Chordata" rank="species" species="pictus">Haploblepharus pictus</taxonomicName>
seems to be more abundant west of Cape Agulhas (which makes it somewhat surprising that it was not recorded by Clark (1997)), although it appears not to be uncommon east of Cape Agulhas. Dr. P. D. Cowley (SAIAB) has shown the author photographs of a number of live
<taxonomicName id="AA30331DF00D9D44E6DF91282010189E" LSID-ZBK="urn:lsid:zoobank.org:act:87028B01-5D5B-4004-918B-492D3059F90E" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Haploblepharus Garman 1913:101" order="Carcharhiniformes" pageNumber="20" phylum="Chordata" rank="genus">Haploblepharus</taxonomicName>
specimens from Titsikama National Park, Eastern Cape, with colour patterns that resemble
<taxonomicName id="FA0BC0F67F7AEAEFF79E1D36EC69CAB6" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="20" phylum="Chordata" rank="species" species="edwardsii">H. edwardsii</taxonomicName>
, however, these specimens proved to be
<taxonomicName id="908C5DBF3C265535E9EDCAD9AB52961B" class="Elasmobranchii" family="Scyliorhinidae" genus="Haploblepharus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="20" phylum="Chordata" rank="species" species="pictus">H. pictus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="57F147CBE67E9CD0F2B082B0B7E8F517" type="etymology">
<paragraph id="F2258104923D160A5DE6917E838E7F9F" pageNumber="20">
Etymology. Not provided by
<normalizedToken id="E450650396DD4C3B9C3EB1DD4DF27101" originalValue="Müller">Mueller</normalizedToken>
&amp; Henle in their original description, however, the Latin word, pictus, means painted or coloured and is a clear reference to the variegated colour pattern of this species.
</paragraph>
<paragraph id="0B9D4D8CA866C4E10C4F1ABC1FFC2B9E" pageNumber="20">Common name. The common name, pretty happy, was introduced in Compagno &amp; Human (2003). It is otherwise known as the dark shyshark, or skaamoog in Afrikaans.</paragraph>
</subSubSection>
<subSubSection id="103A82581DC7EF43D48E6FB2AEBC67BE" type="materials_examined">
<paragraph id="CB7EF135A9968DAD62FCB02B0B46FA74" pageNumber="20">
Study material.
<materialsCitation id="9611F49D89963A66CBDE9A523528C490" ID-GBIF-Occurrence="923889897" collectionCode="BAH" country="South Africa" latitude="-34.233333" location="Millers Point, False Bay" longitude="18.476667" specimenCode="BAH 20000617.08" stateProvince="Western Cape">
<specimenCode id="D3F6590C0FDAE777FF8F8E2941534483">
<collectionCode id="A22C8095606DD52CCF8BB60FD15409A0">BAH</collectionCode>
20000617.08
</specimenCode>
, male 560 mm TL,
<location id="798306EB2F559070F5E22E41913545E7" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:798306EB2F559070F5E22E41913545E7" country="South Africa" latitude="-18.476667" name="Millers Point, False Bay" stateProvince="Western Cape">Millers Point, False Bay</location>
,
<collectingRegion id="3FD40703316B3E1BEF56863F8BC1FD39">Western Cape</collectingRegion>
,
<collectingCountry id="8C6DD7BAF6ED9AE2A9D094D750B55100">South Africa</collectingCountry>
,
<geoCoordinate id="0D35FC6533E0E59ECFB2C81D979656AB" direction="south" orientation="latitude" value="-34.233334">34°14'S</geoCoordinate>
<geoCoordinate id="7CEA52CA57F768BE9BF3606B8FF7FC4B" direction="south" orientation="latitude" value="-18.476667">18°28.6'S</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="B924FFBBDCE156DCFC44AAF89C00E03B" ID-GBIF-Occurrence="923889902" collectionCode="BAH" country="South Africa" location="Millers Point, False Bay" specimenCode="BAH 20000617.09" stateProvince="Western Cape">
<specimenCode id="92CF03DBF3502DB0BA2F450CFE3C97A5">
<collectionCode id="2EBA34C15F6E012918665EEAC9ADD05F">BAH</collectionCode>
20000617.09
</specimenCode>
, male 700 mm TL,
<location id="0238C98BC7C7006A7441D8DCD3A7E4E9" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:0238C98BC7C7006A7441D8DCD3A7E4E9" country="South Africa" name="Millers Point, False Bay" stateProvince="Western Cape">Millers Point, False Bay</location>
,
<collectingRegion id="A7943CEFD82E4A7070330506F3089EF6">Western Cape</collectingRegion>
,
<collectingCountry id="C10B3F2313292B7D8F1F5FB58A814A72">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="FF4E858EBA312E28ECADC6639475A5D9" ID-GBIF-Occurrence="923889898" collectionCode="BAH" country="South Africa" location="Millers Point, False Bay" specimenCode="BAH 20001007.03" stateProvince="Western Cape">
<specimenCode id="E1A736563F73F9CBF24F6B4A7CA1A974">
<collectionCode id="53C7C0FE5D1BEF5E2E308E2F2614CCE7">BAH</collectionCode>
20001007.03
</specimenCode>
, male 545 mm TL,
<location id="829AD305987390D1E77A516B8E85BC39" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:829AD305987390D1E77A516B8E85BC39" country="South Africa" name="Millers Point, False Bay" stateProvince="Western Cape">Millers Point, False Bay</location>
,
<collectingRegion id="FDFB423FBE73FC6947E72844060C1E90">Western Cape</collectingRegion>
,
<collectingCountry id="D99509AC559D869768D8B826DB76CBF5">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="194C0901B3F0E3B2365B502EA38B548A" ID-GBIF-Occurrence="923889900" collectionCode="BAH" country="South Africa" latitude="-34.585" location="Klein Bay, Gans Bay" longitude="19.345" specimenCode="BAH 20010308.04" stateProvince="Western Cape">
<specimenCode id="E7C94726C0F5EC3D119BE6E31D0F6F81">
<collectionCode id="85D3B2414B1BA45CAA822B71E0544868">BAH</collectionCode>
20010308.04
</specimenCode>
, 1 specimen,
<location id="FCA7A28C6737C829192ADE79447B476F" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:FCA7A28C6737C829192ADE79447B476F" country="South Africa" latitude="-34.585" longitude="19.345" name="Klein Bay, Gans Bay" stateProvince="Western Cape">Klein Bay, Gans Bay</location>
,
<collectingRegion id="292375A09076AC7B645A9DB406F7C188">Western Cape</collectingRegion>
,
<collectingCountry id="B4548060E5B18C55FA926E79ED9F40B4">South Africa</collectingCountry>
,
<geoCoordinate id="6FE2A81ACBDD5E648FA752F35A04A592" direction="south" orientation="latitude" value="-34.585">34°35.1'S</geoCoordinate>
<geoCoordinate id="AE475E4A5673D7AC4E41C26D2CF68C58" direction="east" orientation="longitude" value="19.345">19°20.7'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="838CBFA128BABCC4393B7EC3F090099F" ID-GBIF-Occurrence="923889899" collectionCode="BAH" country="South Africa" latitude="-33.9" location="Granger Bay, Table Bay" longitude="18.4" specimenCode="BAH 20010405.06" stateProvince="Western Cape">
<specimenCode id="F993BC297ED22A6BED0AF37EDE886707">
<collectionCode id="D0722D0B11AE89E4781D74082A133F91">BAH</collectionCode>
20010405.06
</specimenCode>
, male 595 mm TL,
<location id="20DDD997D7897687CA2CCF925BC5A0CE" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:20DDD997D7897687CA2CCF925BC5A0CE" country="South Africa" latitude="-33.9" longitude="18.4" name="Granger Bay, Table Bay" stateProvince="Western Cape">Granger Bay, Table Bay</location>
,
<collectingRegion id="26D960B3F7F28A82D193CAC1C239A53B">Western Cape</collectingRegion>
,
<collectingCountry id="DAB906F341BDD815BC1DC79E91F896FD">South Africa</collectingCountry>
,
<geoCoordinate id="49C065309558496D1608C33A629E9360" direction="south" orientation="latitude" value="-33.9">33°54'S</geoCoordinate>
<geoCoordinate id="5C21CC8338D3CDB9677143712DF2BD6E" direction="east" orientation="longitude" value="18.4">18°24'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="9B5E4A437C82631DB2D134DCCCCC8EF3" ID-GBIF-Occurrence="923889896" collectionCode="BAH" country="South Africa" location="Granger Bay, Table Bay" specimenCode="BAH 20010405.07" stateProvince="Western Cape">
<specimenCode id="8B24EC2C166ABF02D27F6968BF47B064">
<collectionCode id="E12E4EFB5D012655F704E342A4418E5D">BAH</collectionCode>
20010405.07
</specimenCode>
, male 489 mm TL,
<location id="3DB595A66D70F5D67E3C288E992CD8A7" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:3DB595A66D70F5D67E3C288E992CD8A7" country="South Africa" name="Granger Bay, Table Bay" stateProvince="Western Cape">Granger Bay, Table Bay</location>
,
<collectingRegion id="0CCFF72F4D4CC32201BE8233ABA975C8">Western Cape</collectingRegion>
,
<collectingCountry id="A01B78F1BC3D5312ED39552AC54B1F86">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="AA20296CEB87AE5348FD3A59213714CC" ID-GBIF-Occurrence="923889901" collectionCode="BAH" country="South Africa" location="Granger Bay, Table Bay" specimenCode="BAH 20010405.08" stateProvince="Western Cape">
<specimenCode id="04B301597BAE29631503C582DEAD4510">
<collectionCode id="356BDAFABBF37104109F85626521D6C4">BAH</collectionCode>
20010405.08
</specimenCode>
, male 580 mm TL,
<location id="15F80D9F8D7C78391D92CCB599CDA68B" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:15F80D9F8D7C78391D92CCB599CDA68B" country="South Africa" name="Granger Bay, Table Bay" stateProvince="Western Cape">Granger Bay, Table Bay</location>
,
<collectingRegion id="A6DA507458C447576DD7A75FFE9FDA62">Western Cape</collectingRegion>
,
<collectingCountry id="940F6AA541F17B41E98E95FF6F5E5FAA">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="029D5DF9A74F7755D6A5C384170BD655" ID-GBIF-Occurrence="923889907" collectionCode="BAH" country="South Africa" location="Granger Bay, Table Bay" specimenCode="BAH 20010405.09" stateProvince="Western Cape">
<specimenCode id="00E3F9C0DF240D085FEC9148013E12FF">
<collectionCode id="C8BEBDE9B6161656C5308587964EE102">BAH</collectionCode>
20010405.09
</specimenCode>
, male 641 mm TL,
<location id="BD28C2821F80F48659C11EB3104404A2" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:BD28C2821F80F48659C11EB3104404A2" country="South Africa" name="Granger Bay, Table Bay" stateProvince="Western Cape">Granger Bay, Table Bay</location>
,
<collectingRegion id="BAAC65D548DDEFB8D1C63D80EA77AD2D">Western Cape</collectingRegion>
,
<collectingCountry id="AE3B0B51CC79F8AF329743A937E0AE00">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="EB4C13B1277E1C6D04866BF4C91C2949" ID-GBIF-Occurrence="923889905" collectionCode="BAH" country="South Africa" location="Granger Bay, Table Bay" specimenCode="BAH 20010405.10" stateProvince="Western Cape">
<specimenCode id="8A98170EABE2E319945BB6268085F94A">
<collectionCode id="9894A282E36A443FC407601CC077C4A7">BAH</collectionCode>
20010405.10
</specimenCode>
, female 420 mm TL,
<location id="B343334990186695091777D6169A7E7C" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:B343334990186695091777D6169A7E7C" country="South Africa" name="Granger Bay, Table Bay" stateProvince="Western Cape">Granger Bay, Table Bay</location>
,
<collectingRegion id="AF7AFB8C7E7A7873B1BB0EEB12B985A0">Western Cape</collectingRegion>
,
<collectingCountry id="B5A199FD1D7C7855941766B8910A06A6">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="8F00F48949A5BBAF859440BEAEBCBD88" ID-GBIF-Occurrence="923889903" collectionCode="BAH" country="South Africa" location="Granger Bay, Table Bay" specimenCode="BAH 20010405.11" stateProvince="Western Cape">
<specimenCode id="84F6E5DBFA4AA7E9570B7213B581DDFC">
<collectionCode id="B339CF3383B08F2893B9EE378E00ABD8">BAH</collectionCode>
20010405.11
</specimenCode>
, male 580 mm TL,
<location id="3DA9DB720A06DFE9DC9E866C5F516311" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:3DA9DB720A06DFE9DC9E866C5F516311" country="South Africa" name="Granger Bay, Table Bay" stateProvince="Western Cape">Granger Bay, Table Bay</location>
,
<collectingRegion id="714669DF5C072604DA9079C8CB68BBAE">Western Cape</collectingRegion>
,
<collectingCountry id="A0BBA823170AD347202679C8FD95D4F4">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="36FA6336EF2E0535598CDF9F00C5C388" ID-GBIF-Occurrence="923889910" collectionCode="BAH" country="South Africa" latitude="-34.185833" location="Long Beach, False Bay" longitude="18.426666" specimenCode="BAH 20011119.17" stateProvince="Western Cape">
<specimenCode id="BDBE7AAAB77DE2DB61BAD3B105D28CB9">
<collectionCode id="35CDC3F95321E75B9DDB0265FD7E0ED4">BAH</collectionCode>
20011119.17
</specimenCode>
, 1 specimen,
<location id="0ED3F77FB4429A4B1D614BCE1797D093" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:0ED3F77FB4429A4B1D614BCE1797D093" country="South Africa" latitude="-34.185833" longitude="18.426666" name="Long Beach, False Bay" stateProvince="Western Cape">Long Beach, False Bay</location>
,
<collectingRegion id="AAE7AE0A671CF78327DC55143D4D55D2">Western Cape</collectingRegion>
,
<collectingCountry id="9EDA7C10CDDDB5B0DB1FDA772420B8B8">South Africa</collectingCountry>
,
<geoCoordinate id="A09173FA4D25EBD27BB558EAA91ABDC7" direction="south" orientation="latitude" value="-34.185833">34°11.15'S</geoCoordinate>
<geoCoordinate id="4C548634BFC735D191B5322CDC3B0962" direction="east" orientation="longitude" value="18.426666">18°25.6'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="ABA879AA63ED7AE8B7C78F3D55EE3D4F" ID-GBIF-Occurrence="923889915" collectionCode="BAH" country="South Africa" latitude="-34.204334" location="Froggy Pond, False Bay" longitude="18.459333" specimenCode="BAH 20021014.01" stateProvince="Western Cape">
<specimenCode id="6EB344709F04727E69250370F7ADBD1F">
<collectionCode id="B56F753980953DDBCFE0096F43DD2B94">BAH</collectionCode>
20021014.01
</specimenCode>
, gravid female 433 mm PCL,
<location id="81201585D16D485C97A0612970F39A77" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:81201585D16D485C97A0612970F39A77" country="South Africa" latitude="-34.204334" longitude="18.459333" name="Froggy Pond, False Bay" stateProvince="Western Cape">Froggy Pond, False Bay</location>
,
<collectingRegion id="E389A0C3B2E9014D12B3713921766A42">Western Cape</collectingRegion>
,
<collectingCountry id="F2647DEFF0D7811C088A7DE0319603C2">South Africa</collectingCountry>
,
<geoCoordinate id="8378ABE9072B52FA2A1BE274FA246A87" direction="south" orientation="latitude" value="-34.204334">34°12.26'S</geoCoordinate>
<geoCoordinate id="82CC890C8D2A954A3B3C10A9145C644C" direction="east" orientation="longitude" value="18.459333">18°27.56'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="F826D19F6EF3A9CA17BEBC82AB287BB3" ID-GBIF-Occurrence="923889906" collectionCode="BAH" country="South Africa" location="Froggy Pond, False Bay" specimenCode="BAH 20021014.02" stateProvince="Western Cape">
<specimenCode id="9E08757FCCBE8B2776901FE3E6B170EC">
<collectionCode id="672DA122D5855C439ECF379B11CA757B">BAH</collectionCode>
20021014.02
</specimenCode>
, adolescent female 493 mm TL,
<location id="732F55DF78EFE8EFF3DB86C38B225937" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:732F55DF78EFE8EFF3DB86C38B225937" country="South Africa" name="Froggy Pond, False Bay" stateProvince="Western Cape">Froggy Pond, False Bay</location>
,
<collectingRegion id="EF9B3C2F4DDD6AED2B31950923BFEE55">Western Cape</collectingRegion>
,
<collectingCountry id="C6159B29D467BCE5EAB8F70C5337CED1">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="5B44EB04F44E43B5C8526DB1223CB039" ID-GBIF-Occurrence="923889911" collectionCode="BAH" country="South Africa" location="Froggy Pond, False Bay" specimenCode="BAH 20021014.04" stateProvince="Western Cape">
<specimenCode id="5B34B1C1A74A9CDAA1DB67346561C430">
<collectionCode id="CF46A50BECBF283C27FA31E108B55CF8">BAH</collectionCode>
20021014.04
</specimenCode>
, mature male 559 mm TL,
<location id="167788ADE7EC4959D1599590933BD16B" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:167788ADE7EC4959D1599590933BD16B" country="South Africa" name="Froggy Pond, False Bay" stateProvince="Western Cape">Froggy Pond, False Bay</location>
,
<collectingRegion id="8FB3AFF25542250C1E3A338FA3C4C192">Western Cape</collectingRegion>
,
<collectingCountry id="62CB016144388B53B496E97CD203A500">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="EE76534F37C94A2BAB6D827558FC6D6B" ID-GBIF-Occurrence="923889904" collectionCode="BAH" country="South Africa" location="Froggy Pond, False Bay" specimenCode="BAH 20021014.05" stateProvince="Western Cape">
<specimenCode id="696E9CA1F8E8C4A4E2F4E36B84DCE071">
<collectionCode id="AD82C44181FAA0D002E26BC9B4CF9F79">BAH</collectionCode>
20021014.05
</specimenCode>
, mature male 540 mm TL,
<location id="CC116AF7C737E5BF43DFF4BDB1728304" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:CC116AF7C737E5BF43DFF4BDB1728304" country="South Africa" name="Froggy Pond, False Bay" stateProvince="Western Cape">Froggy Pond, False Bay</location>
,
<collectingRegion id="B7957BFA5B7AC4B1A8AC6F7B0EE47EC0">Western Cape</collectingRegion>
,
<collectingCountry id="C3B88AD5B4D51B85FA00BF4845AEC21E">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="DBF2A2C7E9C8F12846F469ED52E0DC95" ID-GBIF-Occurrence="923889912" collectionCode="BAH" country="South Africa" location="Froggy Pond, False Bay" specimenCode="BAH 20021014.06" stateProvince="Western Cape">
<specimenCode id="953487E475F217D3E279AD7C35A65793">
<collectionCode id="5C84C49D77D1D49D70884A72D1984FDF">BAH</collectionCode>
20021014.06
</specimenCode>
, mature female 506 mm TL,
<location id="A52106F2BD9760E0CEB3462C470D8D64" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:A52106F2BD9760E0CEB3462C470D8D64" country="South Africa" name="Froggy Pond, False Bay" stateProvince="Western Cape">Froggy Pond, False Bay</location>
,
<collectingRegion id="E3D23D6FBFEFA6FBFF5E8CC855133EF8">Western Cape</collectingRegion>
,
<collectingCountry id="811B2AB7312C0FBE3C345ECED9F4698E">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="56561B54755725ACAC5C8128E8C21D42" ID-GBIF-Occurrence="923889914" collectionCode="BAH" country="South Africa" location="Millers Point, False Bay" specimenCode="BAH 20021015.03" stateProvince="Western Cape">
<specimenCode id="7DADBF6834DCAB9453D56E94A83B5581">
<collectionCode id="F8A13FDE552927901929CAD4CDFF22DC">BAH</collectionCode>
20021015.03
</specimenCode>
, adolescent female 496 mm TL,
<location id="97EFFD4683ED4A550BEEC2A574E61463" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:97EFFD4683ED4A550BEEC2A574E61463" country="South Africa" name="Millers Point, False Bay" stateProvince="Western Cape">Millers Point, False Bay</location>
,
<collectingRegion id="69E4D129D59D7A05BFBCB26F037641DE">Western Cape</collectingRegion>
,
<collectingCountry id="BA26CB0F33C24783F59243D51C6B0FF0">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="9880FBBA749DD765EE87347EC9FCE497" ID-GBIF-Occurrence="923889908" collectionCode="BAH" country="South Africa" location="Millers Point, False Bay" specimenCode="BAH 20021015.04" stateProvince="Western Cape">
<specimenCode id="77D061D36742C74BE4F4C16311450AC1">
<collectionCode id="4FA05B9D56F4A9F79BC2946E2A9740FB">BAH</collectionCode>
20021015.04
</specimenCode>
, mature male 514 mm TL,
<location id="C36A28322FCCCE9C3300C0148FB5AB12" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:C36A28322FCCCE9C3300C0148FB5AB12" country="South Africa" name="Millers Point, False Bay" stateProvince="Western Cape">Millers Point, False Bay</location>
,
<collectingRegion id="D904464D4C493FA26A26BA8C95C5CD95">Western Cape</collectingRegion>
,
<collectingCountry id="902353C5FB764A09A4CCFEB997EF654E">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="B809A01AFE351FC79ACFD2A10EEBA6B1" ID-GBIF-Occurrence="923889913" collectionCode="BAH" country="South Africa" location="Millers Point, False Bay" specimenCode="BAH 20021015.05" stateProvince="Western Cape">
<specimenCode id="3533DF85ECD05D6442203E2A202F2A00">
<collectionCode id="0B5167061D9A43F092E44DCF33AA18BB">BAH</collectionCode>
20021015.05
</specimenCode>
, mature male 570 mm TL,
<location id="A749FDF398F856A4045D1B8CA7B59A85" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:A749FDF398F856A4045D1B8CA7B59A85" country="South Africa" name="Millers Point, False Bay" stateProvince="Western Cape">Millers Point, False Bay</location>
,
<collectingRegion id="C0A3E924358D289DD2725F5E7C02F3BA">Western Cape</collectingRegion>
,
<collectingCountry id="B84462F23D28B2299F3AC95005748D4B">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="7F662CB5506CD3C10A2A6E8381D27708" ID-GBIF-Occurrence="923889918" collectionCode="BAH" country="South Africa" location="Millers Point, False Bay" specimenCode="BAH 20021015.06" stateProvince="Western Cape">
<specimenCode id="A37F1CD4851597B742FACE8B8D1EE5AD">
<collectionCode id="F844FE537A0C37A322834612ABA7E7BA">BAH</collectionCode>
20021015.06
</specimenCode>
, mature male 484 mm TL,
<location id="05D2EF70DECF23DC6F556409C8B4B198" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:05D2EF70DECF23DC6F556409C8B4B198" country="South Africa" name="Millers Point, False Bay" stateProvince="Western Cape">Millers Point, False Bay</location>
,
<collectingRegion id="D17CE33BEAD4957E1B8EE1F9B99E2862">Western Cape</collectingRegion>
,
<collectingCountry id="590E7371B74928D97F578E98373DB256">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="C4A5BFF19F5C4CDE9F796CF503270CD4" ID-GBIF-Occurrence="923889909" collectionCode="BAH" country="South Africa" location="Froggy Pond, False Bay" specimenCode="BAH 20021017.01" stateProvince="Western Cape">
<specimenCode id="B986747FA5318467959B261E63541A91">
<collectionCode id="0471AE73047871B65A18E41FFA60E68E">BAH</collectionCode>
20021017.01
</specimenCode>
, mature male 517 mm TL,
<location id="64434028528B35CA86FFE2FBD7868AC1" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:64434028528B35CA86FFE2FBD7868AC1" country="South Africa" name="Froggy Pond, False Bay" stateProvince="Western Cape">Froggy Pond, False Bay</location>
,
<collectingRegion id="8E5C2DC95937C9D6134B87C74300D81F">Western Cape</collectingRegion>
,
<collectingCountry id="A86E6EBC6C43BB2B4456022C237460DC">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="F1DA68A80AF276FE47F99B06AE419878" ID-GBIF-Occurrence="923889916" collectionCode="BAH" country="South Africa" location="Millers Point, False Bay" specimenCode="BAH 20021020.01" stateProvince="Western Cape">
<specimenCode id="26FE1777367630407CDD7CCCECE99470">
<collectionCode id="389C2B2319BA46C8CFA50FD608D5F69E">BAH</collectionCode>
20021020.01
</specimenCode>
, mature male 460 mm TL,
<location id="5EE514684B7206219B58DF9531DA74E1" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:5EE514684B7206219B58DF9531DA74E1" country="South Africa" name="Millers Point, False Bay" stateProvince="Western Cape">Millers Point, False Bay</location>
,
<collectingRegion id="0181D35178391F0203478BC87E6B6A02">Western Cape</collectingRegion>
,
<collectingCountry id="FF6DE6FF88D798C777EDA473B32A5788">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="B4285DB501D86F86D2D5EA37A36456F0" ID-GBIF-Occurrence="923889921" collectionCode="BAH" country="South Africa" location="Millers Point, False Bay" specimenCode="BAH 20021020.02" stateProvince="Western Cape">
<specimenCode id="1BE186D9C2DBBBA776BDE7EC64FA99A8">
<collectionCode id="740E124E61774DD5E291AF43DECDED6F">BAH</collectionCode>
20021020.02
</specimenCode>
, adolescent female 455 mm TL,
<location id="45BBB2BEA9E4E85C8B7FB9DA75726C1A" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:45BBB2BEA9E4E85C8B7FB9DA75726C1A" country="South Africa" name="Millers Point, False Bay" stateProvince="Western Cape">Millers Point, False Bay</location>
,
<collectingRegion id="2BA0BD94F06EC3864C0360998D505AA9">Western Cape</collectingRegion>
,
<collectingCountry id="588E767E3B4A87F5EF2FD09F3AB6B97D">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="066D33C6963D4317F4E22F609C411046" ID-GBIF-Occurrence="923889919" collectionCode="SAM" country="South Africa" latitude="-32.766666" location="Berg River Mouth, St Helena Bay" longitude="18.15" specimenCode="BGR. 1. H" stateProvince="Western Cape">
<specimenCode id="9C6C253DD4623702DFEDEF24D52E2AD7">BGR.1.H</specimenCode>
, deposited at
<collectionCode id="B6DC35BA0E4E19E3A614BA7713C3CFAB" collectionName="Australia, South Australia, Adelaide, South Australian Museum">SAM</collectionCode>
, embryonic male 103 mm TL,
<location id="21918CBC370053D85F12603A42FD5D4B" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:21918CBC370053D85F12603A42FD5D4B" country="South Africa" latitude="-32.766666" longitude="18.15" name="Berg River Mouth, St Helena Bay" stateProvince="Western Cape">Berg River Mouth, St Helena Bay</location>
,
<collectingRegion id="6DCA0136008A9C24FB9F3167C07ECF9A">Western Cape</collectingRegion>
,
<collectingCountry id="0C970E77898996D00D7C8164AFC5EE02">South Africa</collectingCountry>
,
<geoCoordinate id="16CB1C3D8C5653ABE9843A0DC3A2A38A" direction="south" orientation="latitude" value="-32.766666">32°46'S</geoCoordinate>
<geoCoordinate id="2B692AE70904CFACACBEB45996F6C1CC" direction="east" orientation="longitude" value="18.15">18°09'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="0D9E34CFA31ED9C0ED75469FEDB3AFCD" ID-GBIF-Occurrence="923889917" collectionCode="MJS" country="South Africa" latitude="-34.406666" location="Rocky Bank, False Bay" longitude="18.591667" specimenCode="MJS 960526" stateProvince="Western Cape">
<specimenCode id="EB36D5C42C29FBF521533989771FC5C1">
<collectionCode id="9D86B45DB5628AC211B153EDEA3C1388">MJS</collectionCode>
960526
</specimenCode>
, adolescent female 430 mm TL,
<location id="749E09AB13EC4A08018314A7EA5F56AC" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:749E09AB13EC4A08018314A7EA5F56AC" country="South Africa" latitude="-34.406666" longitude="18.591667" name="Rocky Bank, False Bay" stateProvince="Western Cape">Rocky Bank, False Bay</location>
,
<collectingRegion id="2F611345B778366075D1739E9641A8E8">Western Cape</collectingRegion>
,
<collectingCountry id="AC33DC14C7FACE62CFD4D50CBD4F5E90">South Africa</collectingCountry>
,
<geoCoordinate id="89755E684249FF305F1813FD57210ED3" direction="south" orientation="latitude" value="-34.406666">34°24.4'S</geoCoordinate>
<geoCoordinate id="6B47EA8FB040C3CBA74F7D120FF91120" direction="east" orientation="longitude" value="18.591667">18°35.5'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="5217582BACDA5D96C186B0469F9CBB44" ID-GBIF-Occurrence="923889925" collectionCode="MJS" country="South Africa" latitude="-34.182835" location="Slangkop" longitude="18.307667" specimenCode="MJS 970124" stateProvince="Western Cape">
<specimenCode id="F8B674A8E40BF841B156C059D0739362">
<collectionCode id="C03FF4CECAE379657F8C67A88E1308BB">MJS</collectionCode>
970124
</specimenCode>
, gravid female 517 mm TL,
<location id="22F53FD707685A15B72940DCCFF436D1" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:22F53FD707685A15B72940DCCFF436D1" country="South Africa" latitude="-34.182835" longitude="18.307667" name="Slangkop" stateProvince="Western Cape">Slangkop</location>
,
<collectingRegion id="05C0C7B2708A81A5E87133E4A143F6E0">Western Cape</collectingRegion>
,
<collectingCountry id="31BEB43C6A3B51E296F10861CB266C80">South Africa</collectingCountry>
,
<geoCoordinate id="A97AD48B5964AD9950031A179A3283C9" direction="south" orientation="latitude" value="-34.182835">34°10.97'S</geoCoordinate>
<geoCoordinate id="0C0C3DF43EBEA3497CE3643B903DC06F" direction="east" orientation="longitude" value="18.307667">18°18.46'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="FD44CE5C4A498A68ECFE57BADA1DBC11" ID-GBIF-Occurrence="923889926" collectionCode="MJS" country="South Africa" latitude="-34.512665" location="Die Kelders, Gans Bay" longitude="19.337334" specimenCode="MJS 970128" stateProvince="Western Cape">
<specimenCode id="345D22A97989330AD3470616E49F1F73">
<collectionCode id="00D961B0725DC23AE9898E80340C1B02">MJS</collectionCode>
970128
</specimenCode>
, 3 specimens, mature male 554 mm TL, two mature females 547 mm TL and 597 mm TL,
<location id="DA35525C798288E5FF8F11FC69CB9796" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:DA35525C798288E5FF8F11FC69CB9796" country="South Africa" latitude="-34.512665" longitude="19.337334" name="Die Kelders, Gans Bay" stateProvince="Western Cape">Die Kelders, Gans Bay</location>
,
<collectingRegion id="2002014F457A4AC25CDE16658F6587BB">Western Cape</collectingRegion>
,
<collectingCountry id="379498923B85B74CFDD77574EFB97941">South Africa</collectingCountry>
,
<geoCoordinate id="1611B7709FC650464015BFE4A5A78A71" direction="south" orientation="latitude" value="-34.512665">34°30.76'S</geoCoordinate>
<geoCoordinate id="AA422D97C7871525CF982625B4F15D26" direction="east" orientation="longitude" value="19.337334">19°20.24'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="6F570F666506D60A9330A41B67D3465D" ID-GBIF-Occurrence="923889924" collectionCode="NMW" country="South Africa" location="“ Cap ” [Cape]" specimenCode="NMW 78529">
<specimenCode id="8535B559AD0E8027CAEBD87C7AD9BFA9">
<collectionCode id="8D3AA55E7143FF689620FAFC02B22A36">NMW</collectionCode>
78529
</specimenCode>
, 2 specimens, immature male ~110 mm TL, immature female ~200 mm TL,
<location id="B74AB7A04F508481E986B8DE2FFD5B88" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:B74AB7A04F508481E986B8DE2FFD5B88" country="South Africa" name="“ Cap ”">“Cap”</location>
,
<collectingCountry id="1AE036D97A81BA19F1EC9357A206BD9E">South Africa</collectingCountry>
</materialsCitation>
; RMNH 4161, probable syntypes, see under Type Series and Locality for details; RMNH 4162, probable syntype, see under Type Series and Locality for details; RMNH 4163, probable syntype, see under Type Series and Locality for details; RMNH 4164, probable syntype, see under Type Series and Locality for details;
<materialsCitation id="E19603E95EC5438AC6184CF91AA2BDCA" ID-GBIF-Occurrence="923889920" collectionCode="RUSI" country="South Africa" latitude="-33.916668" location="Woodstock Power Station (site no longer in existence, reclaimed land), Cape Town" longitude="18.433332" specimenCode="RUSI 4115" stateProvince="Western Cape">
<specimenCode id="67D2DD1FD7B367D1C236F63683FBC1D8">
<collectionCode id="05A4B671495912B9BE056E85D177C893">RUSI</collectionCode>
4115
</specimenCode>
, juvenile male 165 mm TL,
<location id="052C5538E3F24198358C9514CDB35856" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:052C5538E3F24198358C9514CDB35856" country="South Africa" latitude="-33.916668" longitude="18.433332" name="Woodstock Power Station (site no longer in existence, reclaimed land), Cape Town" stateProvince="Western Cape">Woodstock Power Station (site no longer in existence, reclaimed land), Cape Town</location>
,
<collectingRegion id="41C19CF03C8E8671C4FE869F6C74B352">Western Cape</collectingRegion>
,
<collectingCountry id="CFC18039CD213AEF7085CA3405136427">South Africa</collectingCountry>
,
<geoCoordinate id="63EE195D6C983DBB47C85297619331A3" direction="south" orientation="latitude" value="-33.916668">33°55'S</geoCoordinate>
<geoCoordinate id="E89BCC4294BCF905C5C4FFA8F974FD0D" direction="east" orientation="longitude" value="18.433332">18°26'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="B436DAD451839B619C73B1BB6643370D" ID-GBIF-Occurrence="923889923" collectionCode="RUSI" country="South Africa" latitude="-33.8" location="Robben Island, Table Bay" longitude="18.366667" specimenCode="RUSI 4117" stateProvince="Western Cape">
<specimenCode id="AC4B238EE1207900F976279B5D6CA583">
<collectionCode id="97E92FD0194E16CBD461B52C7DB38B4F">RUSI</collectionCode>
4117
</specimenCode>
, adolescent female 483 mm TL,
<location id="7E18DB2ABDDE76538CC5982508587B31" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:7E18DB2ABDDE76538CC5982508587B31" country="South Africa" latitude="-33.8" longitude="18.366667" name="Robben Island, Table Bay" stateProvince="Western Cape">Robben Island, Table Bay</location>
,
<collectingRegion id="5599527A0AB69D12DA653895D3C60257">Western Cape</collectingRegion>
,
<collectingCountry id="BC2319599BB03CA1A01C206C0F6A5B2F">South Africa</collectingCountry>
,
<geoCoordinate id="87780FCD130336E1582D3BFEF6D50F48" direction="south" orientation="latitude" value="-33.8">33°48'S</geoCoordinate>
<geoCoordinate id="026B3642CBFF5A5D221B3EA8FC9E7ACC" direction="east" orientation="longitude" value="18.366667">18°22'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="B7FA4DF48CB6EAE49A68C5A070C94CA7" ID-GBIF-Occurrence="923889922" collectionCode="RUSI ( previously ORI)" country="South Africa" latitude="-34.35125" location="Cape Point" longitude="18.485695" specimenCode="RUSI 6083, previously ORI 6083" stateProvince="Western Cape">
<specimenCode id="89FB9A71376BB97E6A621C73DB7D9E90">
<collectionCode id="8898C111BF241FA0259EB6CE531EF653">RUSI</collectionCode>
6083, previously
<collectionCode id="72B1B163716058E2D66C8F21A95B29CD">ORI</collectionCode>
6083
</specimenCode>
, mature male 565 mm TL,
<location id="FC3F5549F4C26CC998E85425A4F965FC" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:FC3F5549F4C26CC998E85425A4F965FC" country="South Africa" latitude="-34.35125" longitude="18.485695" name="Cape Point" stateProvince="Western Cape">Cape Point</location>
,
<collectingRegion id="DD46A4DE8FBBDABFE53F9DEDE3C73EEB">Western Cape</collectingRegion>
,
<collectingCountry id="B5692753FC6994D4B4E21AA16A3ED1A6">South Africa</collectingCountry>
,
<geoCoordinate id="A59762AD427207855D5C8FBB6C405455" direction="south" orientation="latitude" value="-34.35125">34°21'4.5&quot;S</geoCoordinate>
<geoCoordinate id="9E8CAB074451CE42D323B30B563FE97F" direction="east" orientation="longitude" value="18.485695">18°29'8.5&quot;E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="198A4FF51C5E1CC00676FE09AE87DC6F" ID-GBIF-Occurrence="923889927" collectionCode="RUSI ( previously ORI)" country="South Africa" location="Cape Point" specimenCode="RUSI 6084, previously ORI 2874" stateProvince="Western Cape">
<specimenCode id="BBA2220908965EAC774339C819C7FC7A">
<collectionCode id="FA9B7C33D16F293B3D784BDCB38B667D">RUSI</collectionCode>
6084, previously
<collectionCode id="6DE4A0C7C0734511CEA345CD74612AFF">ORI</collectionCode>
2874
</specimenCode>
, adolescent female 533 mm TL,
<location id="8C75A17C1E2C6EC9C90C3491DBE5982A" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:8C75A17C1E2C6EC9C90C3491DBE5982A" country="South Africa" name="Cape Point" stateProvince="Western Cape">Cape Point</location>
,
<collectingRegion id="A97D66F43DDA06675FD452246E12D94B">Western Cape</collectingRegion>
,
<collectingCountry id="9EDF977D01F946494F10E796032182E3">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="CEB874969F5A79E8239793DB13265203" ID-GBIF-Occurrence="923889944" collectionCode="RUSI ( previously ORI)" country="South Africa" latitude="-33.916668" location="Woodstock (presently reclaimed land), Cape Town" longitude="18.433332" specimenCode="RUSI 6160, previously ORI 2932" stateProvince="Western Cape">
<specimenCode id="8FF3596AF1E6761B519E05EB47518288">
<collectionCode id="03577E8193DE53BA5FD986C7658C130A">RUSI</collectionCode>
6160, previously
<collectionCode id="C56A12DBAD6327C790C095160BBED92F">ORI</collectionCode>
2932
</specimenCode>
, juvenile male 199 mm TL,
<location id="EDA8B6322DFE34264FAEE1975F1C38B5" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:EDA8B6322DFE34264FAEE1975F1C38B5" country="South Africa" latitude="-33.916668" longitude="18.433332" name="Woodstock (presently reclaimed land), Cape Town" stateProvince="Western Cape">Woodstock (presently reclaimed land), Cape Town</location>
,
<collectingRegion id="2FF0DAACEFA78A29A7A8B3F4E7ABD0BB">Western Cape</collectingRegion>
,
<collectingCountry id="9750A0C6A06D21F886C92876DE7C551A">South Africa</collectingCountry>
,
<geoCoordinate id="F39948646D170F5295AEEFA698599CF1" direction="south" orientation="latitude" value="-33.916668">33°55'S</geoCoordinate>
<geoCoordinate id="FF54154B441AA67E216008858587D32C" direction="east" orientation="longitude" value="18.433332">18°26'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="A8AC803295CA744CDE685D8BAC486797" ID-GBIF-Occurrence="923889931" collectionCode="RUSI" country="South Africa" latitude="-34.336666" location="Platboom, Cape Point" longitude="18.444166" specimenCode="RUSI 39993" stateProvince="Western Cape">
<specimenCode id="B0E60BF6CF5C787CCEDEE64BF28DAAD2">
<collectionCode id="5ABC11F86A5DA9EEAA371471358B1670">RUSI</collectionCode>
39993
</specimenCode>
, 3 specimens, two juvenile males 320 mm TL and 369 mm TL, mature male 537 mm TL,
<location id="4B500FF3C99E03851481BEC38FFF9631" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:4B500FF3C99E03851481BEC38FFF9631" country="South Africa" latitude="-34.336666" longitude="18.444166" name="Platboom, Cape Point" stateProvince="Western Cape">Platboom, Cape Point</location>
,
<collectingRegion id="A4BE087CA47ED8C2B7D6E543413E8079">Western Cape</collectingRegion>
,
<collectingCountry id="7C873E82C84E7604AD439006156DC080">South Africa</collectingCountry>
,
<geoCoordinate id="3808D75205D0A188A1364B63EEFB7EBA" direction="south" orientation="latitude" value="-34.336666">34°20.2'S</geoCoordinate>
<geoCoordinate id="19A49D7E0E039F4E544B51A910BA36A0" direction="east" orientation="longitude" value="18.444166">18°26.65'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="845713EEBAB7AED69078CD9D502BA8FE" ID-GBIF-Occurrence="923889933" collectionCode="RUSI" country="South Africa" location="Berg River Mouth, St Helena Bay" specimenCode="RUSI 44494" stateProvince="Western Cape">
<specimenCode id="74B9A40AE0E6B244003BF89BC61E402B">
<collectionCode id="A1C789833A8C15B2D55BB1E5D579A8B1">RUSI</collectionCode>
44494
</specimenCode>
, mature male 432 mm TL,
<location id="9898F810A5C61FE270D1455E1689271B" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:9898F810A5C61FE270D1455E1689271B" country="South Africa" name="Berg River Mouth, St Helena Bay" stateProvince="Western Cape">Berg River Mouth, St Helena Bay</location>
,
<collectingRegion id="AE09D28D8FB39F09486B0869A8AE945D">Western Cape</collectingRegion>
,
<collectingCountry id="1303F3F9CBC9F172396A4F14687909FA">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="AE7DDF77614C09E17B9DD641EA308B35" ID-GBIF-Occurrence="923889937" collectionCode="RUSI" country="Namibia" latitude="-26.116667" location="Hottentot Bay" longitude="14.95" specimenCode="RUSI 48499">
<specimenCode id="9C0E8FB84BD87140148951AF09590937">
<collectionCode id="254B80A12308CB7E32B4645C7FDCAF3C">RUSI</collectionCode>
48499
</specimenCode>
, gravid female 359 mm TL,
<location id="43146D03B5485B15F7AC74F55BEF8795" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:43146D03B5485B15F7AC74F55BEF8795" country="Namibia" latitude="-26.116667" longitude="14.95" name="Hottentot Bay">Hottentot Bay</location>
,
<collectingCountry id="C25D1563C86E0A5D07CC6F2112D4B9C4">Namibia</collectingCountry>
,
<geoCoordinate id="D955C7F6D58667D34242C9F920891A67" direction="south" orientation="latitude" value="-26.116667">26°07'S</geoCoordinate>
<geoCoordinate id="B7B6845FC175EA8E9D53DDE3BF30BD8A" direction="east" orientation="longitude" value="14.95">14°57'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="5CC02F63F8C1341262854B6AAEC45EBD" ID-GBIF-Occurrence="923889939" collectionCode="RUSI" country="Namibia" latitude="-26.65" location="Luederitz" longitude="15.15" specimenCode="RUSI 48500">
<specimenCode id="6EF24DA05F5B00B40DFA2B73C4A66C0E">
<collectionCode id="F6B39EF5F9D13D5204113C5C2E103F69">RUSI</collectionCode>
48500
</specimenCode>
, mature male,
<normalizedToken id="D6506B770A452543677A82604483619A" originalValue="Lüderitz">
<location id="B544256FF61F4D73FBCFE70AA55F0EF3" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:B544256FF61F4D73FBCFE70AA55F0EF3" country="Namibia" latitude="-26.65" longitude="15.15" name="Luederitz">Luederitz</location>
</normalizedToken>
,
<collectingCountry id="9AEF58E92F24D4286C83E7DAC1B0740D">Namibia</collectingCountry>
,
<geoCoordinate id="66AC490E29C63F23A95B9248EFE3D2D3" direction="south" orientation="latitude" value="-26.65">26°39'S</geoCoordinate>
<geoCoordinate id="0E83D3114F7CB52351CE3A2D3A5CF624" direction="east" orientation="longitude" value="15.15">15°09'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="D11438CA0A6C9EBD1AF8033EBC7746B6" ID-GBIF-Occurrence="923889928" collectionCode="SAM" country="South Africa" latitude="-33.8975" location="Table Bay, Cape Town" longitude="18.435" specimenCode="SAM 3225" stateProvince="Western Cape">
<specimenCode id="FD6C45468EC7C61C8A08DE0DA69E9870">
<collectionCode id="3D045D71DF244B4095E51182BC3FC5E2">SAM</collectionCode>
3225
</specimenCode>
, mature male,
<location id="925AFE83EFE8322D72BD12B6C2B03BA3" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:925AFE83EFE8322D72BD12B6C2B03BA3" country="South Africa" latitude="-33.8975" longitude="18.435" name="Table Bay, Cape Town" stateProvince="Western Cape">Table Bay, Cape Town</location>
,
<collectingRegion id="9E467549FA355E2B42DD0A2C11A2132D">Western Cape</collectingRegion>
,
<collectingCountry id="8891842F8AC7AE9063B9472A5B66B077">South Africa</collectingCountry>
,
<geoCoordinate id="AD778EB75DD5446C3AACBFCBCA1DB65E" direction="south" orientation="latitude" value="-33.8975">33°53.85'S</geoCoordinate>
<geoCoordinate id="F572919F8CDE901891A708C4E9830401" direction="east" orientation="longitude" value="18.435">18°26.1'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="5126154635A3445E70B80340A3A3686A" ID-GBIF-Occurrence="923889943" collectionCode="SAM" country="South Africa" latitude="-32.975" location="Saldanha Bay" longitude="17.85" specimenCode="SAM 10142" stateProvince="Western Cape">
<specimenCode id="42CED34102D88AEDB19DFE6413D4E028">
<collectionCode id="563DDD59E1E3BE9D87BC0160CDB60C94">SAM</collectionCode>
10142
</specimenCode>
, adolescent female,
<location id="DE2C200A8F9F9B49D6587D7648F39DBF" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:DE2C200A8F9F9B49D6587D7648F39DBF" country="South Africa" latitude="-32.975" longitude="17.85" name="Saldanha Bay" stateProvince="Western Cape">Saldanha Bay</location>
,
<collectingRegion id="E80069DEDECB150EEBC213A34FE15E0A">Western Cape</collectingRegion>
,
<collectingCountry id="FE025DB55EDBBE9E812C06711F4CC976">South Africa</collectingCountry>
,
<geoCoordinate id="F05F836333CA8935FDFBA7CE3DB7E323" direction="south" orientation="latitude" value="-32.975">32°58.5'S</geoCoordinate>
<geoCoordinate id="B5BA61DBBF8FFB5ADE4E98CB3AC2C6F2" direction="east" orientation="longitude" value="17.85">17°51'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="A2B27C9BA27662BA9FA365EF54C13768" ID-GBIF-Occurrence="923889930" collectionCode="SAM" country="South Africa" latitude="-34.185" location="Simonstown Naval Dockyard, False Bay" longitude="18.438334" specimenCode="SAM 21941" stateProvince="Western Cape">
<specimenCode id="CDECBB32AC2B6E6466832E3CE38BBC85">
<collectionCode id="F291BB77000732742E89A8E748D7D307">SAM</collectionCode>
21941
</specimenCode>
, adolescent male 282 mm TL,
<location id="C99F0BF77692126FA63645CC6196546F" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:C99F0BF77692126FA63645CC6196546F" country="South Africa" latitude="-34.185" longitude="18.438334" name="Simonstown Naval Dockyard, False Bay" stateProvince="Western Cape">Simonstown Naval Dockyard, False Bay</location>
,
<collectingRegion id="1BA852D5A366D5F6536F05192C52F711">Western Cape</collectingRegion>
,
<collectingCountry id="1D1DF54CF592EC8A6B0A35E854CB6D8E">South Africa</collectingCountry>
,
<geoCoordinate id="2A6A9486BF57BBF8D5C4CED6198B3EA8" direction="south" orientation="latitude" value="-34.185">34°11.1'S</geoCoordinate>
<geoCoordinate id="4CEBC69B5E2FAF81F1FDEE89E2C7B0FD" direction="east" orientation="longitude" value="18.438334">18°26.3'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="C70EDFF1BD7E79674C774788D9C52431" ID-GBIF-Occurrence="923889942" collectionCode="SAM" country="South Africa" location="Saldanha Bay" specimenCode="SAM 22996" stateProvince="Western Cape">
<specimenCode id="2BFA1DC918F823EF51266DB5749441F8">
<collectionCode id="37CCA2191A4F8E70C844A9CF2ADC92DD">SAM</collectionCode>
22996
</specimenCode>
, 3 specimens,
<location id="EBBAC2EA77DD92B5FCF1B6688EFBC808" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:EBBAC2EA77DD92B5FCF1B6688EFBC808" country="South Africa" name="Saldanha Bay" stateProvince="Western Cape">Saldanha Bay</location>
,
<collectingRegion id="7CE6F76D99E65575D98A9C37E9B2EBE5">Western Cape</collectingRegion>
,
<collectingCountry id="8CD97CB54F07ED2ACC3FD75528E399A0">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="DF64868672EBE49B80CF9BA2AAE60E6F" ID-GBIF-Occurrence="923889929" collectionCode="SAM" country="South Africa" latitude="-33.916668" location="Salt River Power station filter screens, Table Bay" longitude="18.433332" specimenCode="SAM 23096" stateProvince="Western Cape">
<specimenCode id="C7094A4305335D487948AD54D0479029">
<collectionCode id="6600433AC10A5D2496A360995A150BB1">SAM</collectionCode>
23096
</specimenCode>
, juvenile male 335 mm TL,
<location id="5EB715491785F1B60BE73D4F8E0F42AD" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:5EB715491785F1B60BE73D4F8E0F42AD" country="South Africa" latitude="-33.916668" longitude="18.433332" name="Salt River Power station filter screens, Table Bay" stateProvince="Western Cape">Salt River Power station filter screens, Table Bay</location>
,
<collectingRegion id="EE0CFB778A4D0E2BD7FBBE1463505C29">Western Cape</collectingRegion>
,
<collectingCountry id="211D6ED7D15320C47ACB22620AF73943">South Africa</collectingCountry>
,
<geoCoordinate id="FCBC5166390841F08D84367082DD1229" direction="south" orientation="latitude" value="-33.916668">33°55'S</geoCoordinate>
<geoCoordinate id="B848520865A23653352604CE41255184" direction="east" orientation="longitude" value="18.433332">18°26'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="E5FCA59A04F1EB0DCD7C77EFB0D2CE4E" ID-GBIF-Occurrence="923889935" collectionCode="SAM" country="Namibia" location="North of Orange River Mouth" specimenCode="SAM 23296">
<specimenCode id="92C96D2BEC03CD61780CA9504EAF6991">
<collectionCode id="708FDBED778DC2BA278695DE73D84CF4">SAM</collectionCode>
23296
</specimenCode>
, 2 specimens, juvenile male 218mm TL, juvenile female 242 mm TL,
<location id="46AB5A5619829EBF981C8F49FF6CD101" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:46AB5A5619829EBF981C8F49FF6CD101" country="Namibia" name="North of Orange River Mouth">North of Orange River Mouth</location>
,
<collectingCountry id="0E8862C455DCA63C84A9FA79780C6782">Namibia</collectingCountry>
(no coordinates)
</materialsCitation>
;
<materialsCitation id="F89493D16A2F6062F96F13E5D190948D" ID-GBIF-Occurrence="923889941" collectionCode="SAM" country="Namibia" latitude="-28.033333" location="70 miles (112 km) North of Orange River Mouth" longitude="15.75" specimenCode="SAM 23307">
<specimenCode id="AC32575326F70E6A0FF1CCABA0E6FB7F">
<collectionCode id="5A7403C18C12F45412E82B687F885E8E">SAM</collectionCode>
23307
</specimenCode>
, juvenile male 135 mm TL,
<location id="0662BA83B37FC5D9C984F0622C215BC4" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:0662BA83B37FC5D9C984F0622C215BC4" country="Namibia" latitude="-28.033333" longitude="15.75" name="70 miles (112 km) North of Orange River Mouth">70 miles (112 km) North of Orange River Mouth</location>
,
<collectingCountry id="E3AC8812DF19E184735ACB1C95EC1AEB">Namibia</collectingCountry>
, approx.
<geoCoordinate id="8091465C6D18F4EF6D9E940E184FB99E" direction="south" orientation="latitude" value="-28.033333">28°02'S</geoCoordinate>
<geoCoordinate id="19856ABC9490A31C1E850DD8A7A5B47E" direction="east" orientation="longitude" value="15.75">15°45'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="B362530D2CD6F9B51A6A0D7496EA91DF" ID-GBIF-Occurrence="923889934" collectionCode="SAM" country="South Africa" latitude="-33.093334" location="Schaapen Island, Langebaan Lagoon" longitude="18.025" specimenCode="SAM 23578" stateProvince="Western Cape">
<specimenCode id="3569D6483FC8319FC9F4D8C6163F1D6D">
<collectionCode id="4A25BC9EF03652BD55BD9D8ED777203F">SAM</collectionCode>
23578
</specimenCode>
, embryonic male 83 mm TL,
<location id="3DC77D48FAD8F5EB172F8DD6A4B612B3" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:3DC77D48FAD8F5EB172F8DD6A4B612B3" country="South Africa" latitude="-33.093334" longitude="18.025" name="Schaapen Island, Langebaan Lagoon" stateProvince="Western Cape">Schaapen Island, Langebaan Lagoon</location>
,
<collectingRegion id="5994B4B237DB9234EABC03A5CE65B788">Western Cape</collectingRegion>
,
<collectingCountry id="895F89021697A95B81FCD5B0CFB82758">South Africa</collectingCountry>
,
<geoCoordinate id="3B3EBDB8602155A4E73A8B10F70D505E" direction="south" orientation="latitude" value="-33.093334">33°05.6'S</geoCoordinate>
<geoCoordinate id="235D50B3ACF3D34014E20A3FDD8DE3B8" direction="east" orientation="longitude" value="18.025">18°01.5E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="9C14BA9F86C916348CBCCA82D04EB961" ID-GBIF-Occurrence="923889932" collectionCode="SAM" country="South Africa" latitude="-34.05" location="Hout Bay" longitude="18.35" specimenCode="SAM 23598" stateProvince="Western Cape">
<specimenCode id="76827D9CFF49ECA56C2173FC02B86912">
<collectionCode id="C9A9C39D11FFC3524FB22C8D86AD0D96">SAM</collectionCode>
23598
</specimenCode>
, 4 specimens, 2 juvenile males 128 mm TL and 276 mm TL, 2 juvenile females, 110 mm TL and 128 mm TL,
<location id="64A8D2F474648E8D463EDF57632E1903" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:64A8D2F474648E8D463EDF57632E1903" country="South Africa" latitude="-34.05" longitude="18.35" name="Hout Bay" stateProvince="Western Cape">Hout Bay</location>
,
<collectingRegion id="660D89077B83BBCA4D58A6232AF9C395">Western Cape</collectingRegion>
,
<collectingCountry id="DFE1624D2C78C99C9413B4CA827974DA">South Africa</collectingCountry>
,
<geoCoordinate id="49A98686452C40816FBD8F33D29A4108" direction="south" orientation="latitude" value="-34.05">34°03'S</geoCoordinate>
<geoCoordinate id="976DD70BDF17EB75449D38FE9C8DF4C6" direction="east" orientation="longitude" value="18.35">18°21'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="CA67373DAF7D79964549F271F0C3C069" ID-GBIF-Occurrence="923889938" collectionCode="SAM" country="South Africa" latitude="-32.791668" location="South of Cape Columbine" longitude="17.78" specimenCode="SAM 23811" stateProvince="Western Cape">
<specimenCode id="51381A200B6B1332EDFC3AFBDE305F0C">
<collectionCode id="8B9BB3F65E7863CEB2BBDC9B2926F761">SAM</collectionCode>
23811
</specimenCode>
, 1 specimen,
<location id="20A6E6192877832E44F1CA6872CD7A90" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:20A6E6192877832E44F1CA6872CD7A90" country="South Africa" latitude="-32.791668" longitude="17.78" name="South of Cape Columbine" stateProvince="Western Cape">South of Cape Columbine</location>
,
<collectingRegion id="ADD6FFCB534BD5D928C171B026624D7E">Western Cape</collectingRegion>
,
<collectingCountry id="330E8D1C14ED8E2635CE146EC893C3D0">South Africa</collectingCountry>
,
<geoCoordinate id="E7FC415E7FCED23D0BA02BF94B2D9B9A" direction="south" orientation="latitude" value="-32.791668">32°47.5'S</geoCoordinate>
<geoCoordinate id="167EC39A1D9B307CFB59852401C1469A" direction="east" orientation="longitude" value="17.78">17°46.8'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="34EF800036D9DDDB6820299D2F788DAC" ID-GBIF-Occurrence="923889945" collectionCode="SAM" country="Namibia" location="Luederitz" specimenCode="SAM 24345">
<specimenCode id="6EFCF57B6D378EAC6298A5F7FC2F371F">
<collectionCode id="DE5C0BFA2B4916FA9A9718342FC06AD2">SAM</collectionCode>
24345
</specimenCode>
, 5 specimens, 2 juvenile males 235 mm TL and 268 mm TL, adolescent male 407 mm TL, mature male 516 mm TL, adolescent female 411 mm TL,
<normalizedToken id="B4C065BE76D141655DE1772C5FF781DC" originalValue="Lüderitz">
<location id="45F5D06BE1B54AC548521D5229D930F2" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:45F5D06BE1B54AC548521D5229D930F2" country="Namibia" name="Luederitz">Luederitz</location>
</normalizedToken>
,
<collectingCountry id="31AB637F5F81DE11704FDBA62327FFF1">Namibia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="6AC94FB444A0C67C2D7C57BC3C6874BB" ID-GBIF-Occurrence="923889940" collectionCode="SAM" country="South Africa" location="Bredasdorp (erroneous, true locality probably Arniston, Struis Bay or Cape Agulhas)" specimenCode="SAM 24545" stateProvince="Western Cape">
<specimenCode id="47BB6C7D2157C04CB037D5BE3FC0E7A7">
<collectionCode id="029C84FD4C557675F54F4286E7B4B3D9">SAM</collectionCode>
24545
</specimenCode>
, 1 specimen,
<location id="D50D4108475D51D78FD31B759D8F7C8D" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:D50D4108475D51D78FD31B759D8F7C8D" country="South Africa" name="Bredasdorp (erroneous, true locality probably Arniston, Struis Bay or Cape Agulhas)" stateProvince="Western Cape">Bredasdorp (erroneous, true locality probably Arniston, Struis Bay or Cape Agulhas)</location>
,
<collectingRegion id="925B9FA9C1C8F7A20529E723DA2D2F7A">Western Cape</collectingRegion>
,
<collectingCountry id="7CBA34706EC9D1E11F241A0911DDAB41">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="CFAC17D43423F45E7A56C78BAC0E2AF0" ID-GBIF-Occurrence="923889936" collectionCode="SAM" country="South Africa" latitude="-32.075" location="Lamberts Bay" longitude="18.308332" specimenCode="SAM 26385" stateProvince="Western Cape">
<specimenCode id="2594B694876BB9C0448E99FB96D9E72C">
<collectionCode id="0D996E2424FB39D7412927EC759DEB44">SAM</collectionCode>
26385
</specimenCode>
, juvenile female 306 mm TL,
<location id="E0B19FACC30ABB3795773E07A3C5D074" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:E0B19FACC30ABB3795773E07A3C5D074" country="South Africa" latitude="-32.075" longitude="18.308332" name="Lamberts Bay" stateProvince="Western Cape">Lamberts Bay</location>
,
<collectingRegion id="059B097A01064D916531C5C8DED1973D">Western Cape</collectingRegion>
,
<collectingCountry id="F0441F007487A4ABA9FD347DF8C342FF">South Africa</collectingCountry>
,
<geoCoordinate id="8C7E664D7B2CFE4F65DEA3E31DB1D91E" direction="south" orientation="latitude" value="-32.075">32°04.5'S</geoCoordinate>
<geoCoordinate id="FB04D105DF18756AAD47B432D0DFD4B7" direction="east" orientation="longitude" value="18.308332">18°18.5'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="CC0A481802CF5EB3EFCB37C1A6D07058" ID-GBIF-Occurrence="923889955" collectionCode="SAM" country="South Africa" latitude="-34.35125" location="Cape Peninsula" longitude="18.485695" specimenCode="SAM 29303" stateProvince="Western Cape">
<specimenCode id="384CC391EFC94FAF0BD04B48F6A60293">
<collectionCode id="495E1152480DAC86D1B301C8D6E85F6F">SAM</collectionCode>
29303
</specimenCode>
, 13 specimens,
<location id="01A23F8F80D53117F6464BD1DBC3EE78" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:01A23F8F80D53117F6464BD1DBC3EE78" country="South Africa" latitude="-34.35125" longitude="18.485695" name="Cape Peninsula" stateProvince="Western Cape">Cape Peninsula</location>
,
<collectingRegion id="E41F1F63213CFDA9FF645B3F20ACE8F8">Western Cape</collectingRegion>
,
<collectingCountry id="C0AACFCDD9A60E55FD656484EC339428">South Africa</collectingCountry>
,
<geoCoordinate id="EA20B962A3C614D8E64FD07849E1B163" direction="south" orientation="latitude" value="-34.35125">34°21'4.5&quot;S</geoCoordinate>
<geoCoordinate id="3DFDD4153DC425ADA1B95C4EE6AA6B5B" direction="east" orientation="longitude" value="18.485695">18°29'8.5&quot;E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="F1816CE1F67772721B94C2B99ADE7904" ID-GBIF-Occurrence="923889954" collectionCode="SAM" country="Namibia" location="Mowe Point" specimenCode="SAM 29337">
<specimenCode id="3C60EE9A2B5695A3E0FF28A89A671A6A">
<collectionCode id="B5338D8B3008559A84BEE7E7DD05096B">SAM</collectionCode>
29337
</specimenCode>
, mature female,
<location id="49F3BEDF713490C5D0177EAE643D3064" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:49F3BEDF713490C5D0177EAE643D3064" country="Namibia" name="Mowe Point">Mowe Point</location>
,
<collectingCountry id="19F08038A0BFDDE7A3D3480ECA30F034">Namibia</collectingCountry>
(no coordinates)
</materialsCitation>
;
<materialsCitation id="399D4A4A3FC3359664D97867A9E6F403" ID-GBIF-Occurrence="923889948" collectionCode="SAM" country="South Africa" location="Cape Point" specimenCode="SAM 32526" stateProvince="Western Cape">
<specimenCode id="7E97A93F002FEA2E086359073026A946">
<collectionCode id="A11C60CCF2526EEFEB36EB06FDB8CA2B">SAM</collectionCode>
32526
</specimenCode>
, mature female,
<location id="2C0A75424DE1F989C621A6E1BF645E01" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:2C0A75424DE1F989C621A6E1BF645E01" country="South Africa" name="Cape Point" stateProvince="Western Cape">Cape Point</location>
,
<collectingRegion id="2B7ED32E2434361DEE1F5A25D2936DF8">Western Cape</collectingRegion>
,
<collectingCountry id="421CEF57A452287A47B3C882753650B8">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="6B7E437A254C8A26426A130529BD8990" ID-GBIF-Occurrence="923889947" collectionCode="SAM" country="South Africa" location="Cape Point" specimenCode="SAM 32556" stateProvince="Western Cape">
<specimenCode id="C2FF78D1B6F47AE8820010A05720BA63">
<collectionCode id="5BD8590FFAAEA19316C89F96AF6C313A">SAM</collectionCode>
32556
</specimenCode>
, mature female,
<location id="A492374C5079D16C8E52265396BD5A66" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:A492374C5079D16C8E52265396BD5A66" country="South Africa" name="Cape Point" stateProvince="Western Cape">Cape Point</location>
,
<collectingRegion id="EE1A72ED23CF2852B05BBDFD7B7FD1BE">Western Cape</collectingRegion>
,
<collectingCountry id="8A480847CED088A245394CCAAB48FB04">South Africa</collectingCountry>
</materialsCitation>
;
<materialsCitation id="AE00B923632AD29A10271A4E639A4BAB" ID-GBIF-Occurrence="923889946" collectionCode="SAM" country="South Africa" latitude="-34.135" location="Titsikama" longitude="24.44" specimenCode="SAM 32617" stateProvince="Eastern Cape">
<specimenCode id="7138DA467E0D45F0548E393AC28B244A">
<collectionCode id="5E400597C8468DA4CBA60C9A7200382D">SAM</collectionCode>
32617
</specimenCode>
, mature female 553 mm TL,
<location id="AC664362E89CCA67C10B62BD6CF409CE" LSID="urn:lsid:plazi:treatment:A4BB123BD41E9989A9B0379559F46E92:AC664362E89CCA67C10B62BD6CF409CE" country="South Africa" latitude="-34.135" longitude="24.44" name="Titsikama" stateProvince="Eastern Cape">Titsikama</location>
,
<collectingRegion id="436D52217A25D9D6747EA5856AF3C88B">Eastern Cape</collectingRegion>
,
<collectingCountry id="470F115B351C2C88A6231966539EC234">South Africa</collectingCountry>
,
<geoCoordinate id="95E1D56ED7D78909B9A308A42108E7B6" direction="south" orientation="latitude" value="-34.135">34°08.1'S</geoCoordinate>
<geoCoordinate id="E5C5DEBA5A155853C8353FE045881FF6" direction="east" orientation="longitude" value="24.44">24°26.4'E</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>