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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1453" ID-GBIF-Dataset="51bb4676-675d-429f-86ed-819a712454bc" ID-PMC="PMC3260773" ID-Pensoft-Pub="1313-2970-130-421" ID-PubMed="22259291" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-421" ModsDocOrigin="ZooKeys 130" ModsDocTitle="New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae)" checkinTime="1451249849988" checkinUser="pensoft" docAuthor="Perrichot, Vincent, Ortega-Blanco, Jaime, McKellar, Ryan C., Delclos, Xavier, Azar, Dany, Nel, Andre, Tafforeau, Paul &amp; Engel, Michael S." docDate="2011" docId="A4805418925C8B55EDE0F4AC4D0BBAC5" docLanguage="en" docName="ZooKeys 130: 421-453" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1453" docTitle="Iberomaimetsha nihtmara Ortega-Blanco, Delclos &amp; Engel, sp. n." docType="treatment" docVersion="4" lastPageNumber="437" masterDocId="7C0CFFB3434B4C34FFA1FFD2FFEEFFD8" masterDocTitle="New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae)" masterLastPageNumber="453" masterPageNumber="421" pageNumber="433" updateTime="1668152281281" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perrichot, Vincent</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ortega-Blanco, Jaime</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>McKellar, Ryan C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Delclos, Xavier</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Azar, Dany</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nel, Andre</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tafforeau, Paul</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Engel, Michael S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>130</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>421</mods:start>
<mods:end>453</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.130.1453</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.130.1453</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-130-421</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152030999" LSID="urn:lsid:zoobank.org:act:8E3C73D0-4CE1-4910-9DE6-45A0D3BC2F1D" httpUri="http://treatment.plazi.org/id/A4805418925C8B55EDE0F4AC4D0BBAC5" lastPageId="16" lastPageNumber="437" pageId="12" pageNumber="433">
<subSubSection pageId="12" pageNumber="433" type="nomenclature">
<paragraph pageId="12" pageNumber="433">
<taxonomicName LSID="urn:lsid:zoobank.org:act:8E3C73D0-4CE1-4910-9DE6-45A0D3BC2F1D" authority="Ortega-Blanco, Delclos &amp; Engel" class="Insecta" family="Maimetshidae" genus="Iberomaimetsha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Iberomaimetsha nihtmara" order="Hymenoptera" pageId="12" pageNumber="433" phylum="Arthropoda" rank="species" species="nihtmara">
Iberomaimetsha nihtmara Ortega-Blanco,
<normalizedToken originalValue="Delclòs">Delclos</normalizedToken>
&amp; Engel
</taxonomicName>
<taxonomicNameLabel pageId="12" pageNumber="433">sp. n.</taxonomicNameLabel>
Figs 810
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="433" type="type material">
<paragraph pageId="12" pageNumber="433">Type material.</paragraph>
<paragraph pageId="12" pageNumber="433">
Holotype MCNA 13049 (?); paratypes MCNA 8758(♀?), MCNA 8790 (♂), MCNA 9373 (?), MCNA 9918 (♀), and MCNA 10732 (?). All specimens deposited in the Museo de Ciencias Naturales de
<normalizedToken originalValue="Álava">Alava</normalizedToken>
, Vitoria-Gasteiz, Spain.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="433" type="type locality">
<paragraph pageId="12" pageNumber="433">Type locality.</paragraph>
<paragraph pageId="12" pageNumber="433">
<normalizedToken originalValue="Peñacerrada">Penacerrada</normalizedToken>
I, Moraza, Burgos Province, Basque-Cantabrian Basin, Spain.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="433" type="stratigraphic horizon">
<paragraph pageId="12" pageNumber="433">Stratigraphic horizon.</paragraph>
<paragraph pageId="12" pageNumber="433">Escucha Formation, Lower Cretaceous, Lower Albian.</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="433" type="diagnosis">
<paragraph pageId="12" pageNumber="433">Diagnosis.</paragraph>
<paragraph pageId="12" pageNumber="433">
Mandibles distinctly large, exserted, asymmetrical, right 4-toothed, left 3-toothed, arched inwards (teeth pointing toward teeth of opposite mandible); basal flagellomeres (from F1 to F3-4) distinctly larger than apical flagellomeres (F1-F4 about 2
<normalizedToken originalValue="1.5×">-1.5x</normalizedToken>
the length of F7-F14); protibial distal margin with a comb of stiff setae; cell [1M] square or rhomboidal, distinctly smaller than [2Cu] which has a short but distinct connection between 1M+Cu and M (then, [2Cu] composed by five aristae); 2Rs+M longer than 1m-cu and longer than cell [1Rs]; apicalmost sector of Cu arising from almost apical posterior margin of [2Cu]; metasoma moderately short, as long as mesosoma.
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="437" pageId="12" pageNumber="433" type="description">
<paragraph pageId="12" pageNumber="433">Description.</paragraph>
<paragraph lastPageId="15" lastPageNumber="436" pageId="12" pageNumber="433">
Body length (from paratypes, as incomplete metasoma in holotype) around 2.55-2.61 mm. Head rounded in anterior view, length 0.41 mm, width 0.68 mm with a short occipital carina and apparently some polygonal dorsal sculpture; compound eyes not bulging, slightly oval, separated dorsally by 0.39 mm; toruli about at same distance to inner margin of compound eyes and each other; antennae 2.54 mm in length, inserted on frons below midpoint of compound eyes; scape and pedicel short and globular, flagellomeres cylindrical; lengths of antennomeres as follows (all in mm): scape 0.12, pedicel 0.09, flagellomeres 0.29, 0.28, 0.27, 0.23, 0.20, 0.18, 0.15, 0.14, 0.12, 0.12, 0.11, 0.11, 0.10, 0.12; malar space short, length approximately 0.11 mm. Mesosoma (dorsal view) 0.95 mm in length; pronotum not distinct dorsally; mesoscutal length 0.27 mm, with
<pageBreakToken pageId="13" pageNumber="434" start="start">parallel</pageBreakToken>
notauli and longitudinal medial line, all three present as linear series of grooves more than well-delimited lines (Fig. 9); axillae wide and long, well distinct, separating completely mesoscutellum and mesoscutum; mesoscutellum distinctly convex, length 0.22 mm; propodeum 0.29 mm in length, areolate (seen in paratype MCNA 8790). Legs distinctly elongate; division of trochantellus not visible; tarsi with no plantar lobes visible; leg measurements (all in mm): profemur 0.82, protibia 0.46, protarsomeres
<normalizedToken originalValue="IV">I-V</normalizedToken>
0.34/0.15/0.10/0.09/0.10; mid-legs very poorly preserved but apparently with mesofemur 0.68, mesotibia?, meso
<pageBreakToken pageId="14" pageNumber="435" start="start">tarsomeres</pageBreakToken>
0.24/0.12/0.10/?/?; metafemur 1.0, metatibia 0.83, metatarsomeres 0.46/0.20/0.15/0.09/0.12. Forewing hyaline, length 2.1 mm, maximum width 0.80 mm, densely covered by microtrichia; all veins tubular except for nebulous 2rs-m and 2m-cu, thus, only cells [2Rs] and [2M] pseudo-opened apically (Figs 9, 10)
<pageBreakToken pageId="15" pageNumber="436" start="start">;</pageBreakToken>
1rs-m not aligned with r-rs; 2m-cu curved S-like; veins M and Cu reaching wing margin (although M somewhat nebulous), with apicalmost sector of Cu slightly angled posteriorly; cell [1R1] longer than both cells [1Rs]+[2Rs]; cell [1Rs] almost
<normalizedToken originalValue="¾">3/4</normalizedToken>
the surface of cell [1M] and almost half of [2Rs]; vein r-rs emerging at two-thirds from base of pterostigma; cell [2R1] short, with vein Rs slightly anteriorly bent distad 2rs-m. Hind wing difficult to observe, hyaline and densely covered by microtrichia, length approximately 1.22 mm, maximum width approximately 0.41 mm; three strong hamuli present on costal vein in distal half of wing. Metasoma (checked on paratype MCNA 8790) pedunculate, short and slightly flattened, length 0.98 mm, greatest height 0.41 mm; sterna hard, convex; no apparent metasomal armature on sternum II or III.
</paragraph>
<paragraph pageId="16" pageNumber="437">
<pageBreakToken pageId="16" pageNumber="437" start="start">Male</pageBreakToken>
: Parameres spatulate, with apparently three short stiff setae apically (Fig. 10).
</paragraph>
<caption pageId="16" pageNumber="437">
<paragraph pageId="16" pageNumber="437">
Figure 8. Photomicrographs of holotype of
<taxonomicName class="Insecta" family="Maimetshidae" genus="Iberomaimetsha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Iberomaimetsha nihtmara" order="Hymenoptera" pageId="16" pageNumber="437" phylum="Arthropoda" rank="species" species="nihtmara">Iberomaimetsha nihtmara</taxonomicName>
Ortega-Blanco,
<normalizedToken originalValue="Delclòs">Delclos</normalizedToken>
, and Engel sp. n. (MCNA 13049) A Dorsal aspect B Ventral aspect.
</paragraph>
</caption>
<caption pageId="16" pageNumber="437">
<paragraph pageId="16" pageNumber="437">
Figure 9. Habitus diagram of holotype of
<taxonomicName class="Insecta" family="Maimetshidae" genus="Iberomaimetsha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Iberomaimetsha nihtmara" order="Hymenoptera" pageId="16" pageNumber="437" phylum="Arthropoda" rank="species" species="nihtmara">Iberomaimetsha nihtmara</taxonomicName>
Ortega-Blanco,
<normalizedToken originalValue="Delclòs">Delclos</normalizedToken>
, and Engel sp. n. (MCNA 13049), with inset of facial aspect.
</paragraph>
</caption>
<caption pageId="16" pageNumber="437">
<paragraph pageId="16" pageNumber="437">
Figure 10. Habitus diagram of male paratype of
<taxonomicName class="Insecta" family="Maimetshidae" genus="Iberomaimetsha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Iberomaimetsha nihtmara" order="Hymenoptera" pageId="16" pageNumber="437" phylum="Arthropoda" rank="species" species="nihtmara">Iberomaimetsha nihtmara</taxonomicName>
Ortega-Blanco,
<normalizedToken originalValue="Delclòs">Delclos</normalizedToken>
, and Engel sp. n. (MCNA 8790), with inset of facial aspect.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="16" pageNumber="437" type="etymology">
<paragraph pageId="16" pageNumber="437">Etymology.</paragraph>
<paragraph pageId="16" pageNumber="437">The specific epithet is a noun in apposition and is the Anglo-Saxon word for nightmare, in reference to the terrifying appearance of the species in frontal view and with its enlarged mandibles.</paragraph>
</subSubSection>
</treatment>
</document>