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<document id="2E1839852E8F5C7C24A04EC191A2F50C" ID-DOI="10.11646/zootaxa.4018.4.1" ID-GBIF-Dataset="06511db8-7621-4067-b0c0-3786623ada21" ID-ISSN="1175-5326" ID-Zenodo-Dep="234261" ID-ZooBank="81CA1EED-5B91-4654-8BA5-9D179A7593B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461257952142" checkinUser="plazi" docAuthor="Zonstein, Irina &amp; Wahis, Raymond" docDate="2015" docId="900D9F6DFFAD404772C3FD51C8D6FE88" docLanguage="en" docName="zt04018p505.pdf" docOrigin="Zootaxa 4018 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gonaporus setitarsus I. Zonstein &amp; Wahis, sp. nov." docType="treatment" docVersion="11" lastPageNumber="488" masterDocId="6C34E715FF8940627254FF9BC972FFED" masterDocTitle="Revision of the Palaearctic genus Gonaporus Ashmead, 1902 of spider wasps (Hymenoptera: Pompilidae)" masterLastPageNumber="505" masterPageNumber="451" pageNumber="487" updateTime="1698608636130" updateUser="plazi">
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<mods:title id="D4E446DA5FF880C6E3B4A73530F40C93">Revision of the Palaearctic genus Gonaporus Ashmead, 1902 of spider wasps (Hymenoptera: Pompilidae)</mods:title>
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<mods:namePart id="62FFC3C87789C30F89F0AB20025026A3">Zonstein, Irina</mods:namePart>
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<mods:date id="443F787096148C24416A35B24566E0A5">2015</mods:date>
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<treatment id="900D9F6DFFAD404772C3FD51C8D6FE88" ID-DOI="http://doi.org/10.5281/zenodo.5665483" ID-GBIF-Taxon="119636573" ID-Zenodo-Dep="5665483" LSID="urn:lsid:plazi:treatment:900D9F6DFFAD404772C3FD51C8D6FE88" httpUri="http://treatment.plazi.org/id/900D9F6DFFAD404772C3FD51C8D6FE88" lastPageId="37" lastPageNumber="488" pageId="36" pageNumber="487">
<subSubSection id="50BE7DF0FFAD404672C3FD51CA20FCEB" pageId="36" pageNumber="487" type="nomenclature">
<paragraph id="181B2E7BFFAD404672C3FD51CB8AFD09" blockId="36.[151,850,714,774]" box="[151,760,714,740]" pageId="36" pageNumber="487">
<heading id="43539917FFAD404672C3FD51CB8AFD09" bold="true" box="[151,760,714,740]" fontSize="11" level="1" pageId="36" pageNumber="487" reason="1">
<emphasis id="2AD0F269FFAD404672C3FD51CB8AFD09" bold="true" box="[151,760,714,740]" pageId="36" pageNumber="487">
<taxonomicName id="DFA455F8FFAD404672C3FD51CBE2FD09" authority="I. Zonstein &amp; Wahis" authorityName="I. Zonstein &amp; Wahis" box="[151,656,714,740]" class="Insecta" family="Pompilidae" genus="Gonaporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="setitarsus" status="sp. nov.">
<emphasis id="2AD0F269FFAD404672C3FD51C8E3FD09" bold="true" box="[151,401,714,740]" italics="true" pageId="36" pageNumber="487">Gonaporus setitarsus</emphasis>
I. Zonstein &amp; Wahis
</taxonomicName>
,
<taxonomicNameLabel id="31E34F12FFAD404670C9FD51CB8AFD09" box="[669,760,714,740]" pageId="36" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="181B2E7BFFAD404672C3FD76CA20FCEB" blockId="36.[151,850,714,774]" box="[151,850,749,774]" pageId="36" pageNumber="487">
(
<figureCitation id="809F32FEFFAD404672CAFD76C986FCEB" box="[158,244,749,774]" captionStart="FIGURES 96 – 99" captionStartId="29.[151,264,1668,1691]" captionTargetBox="[174,1387,202,1635]" captionTargetId="figure@29.[174,1387,195,1635]" captionTargetPageId="29" captionText="FIGURES 96 – 99. Gonaporus spp. (96 – 98, ♀; 99, ♂), head (a, anterior; b, lateral; c, dorsal view). 96. G. setitarsus sp. nov. 97. G. simulator sp. nov. 98. G. spinosissimus sp. nov. 99. G. alfierii." httpUri="https://zenodo.org/record/234277/files/figure.png" pageId="36" pageNumber="487">Figs 96</figureCitation>
,
<figureCitation id="809F32FEFFAD404672AAFD75C85BFCEB" box="[254,297,750,774]" captionStart="FIGURES 105 – 109" captionStartId="32.[151,264,1946,1969]" captionTargetBox="[216,1346,375,1919]" captionTargetId="figure@32.[216,1376,369,1919]" captionTargetPageId="32" captionText="FIGURES 105 – 109. Gonaporus spp., ♂, head (a, anterior; b, lateral; c, dorsal view). 105. G. mirabilis sp. nov. 106. G. omanicus. 107. G. setitarsus sp. nov. 108. G. simulator sp. nov. 109. G. spinosissimus sp. nov." httpUri="https://zenodo.org/record/234279/files/figure.png" pageId="36" pageNumber="487">107</figureCitation>
,
<figureCitation id="809F32FEFFAD40467361FD75C82DFCEB" box="[309,351,750,774]" captionStart="FIGURES 110 – 120" captionStartId="33.[151,264,1899,1922]" captionTargetBox="[255,1336,212,1865]" captionTargetId="figure@33.[241,1346,193,1878]" captionTargetPageId="33" captionText="FIGURES 110 – 120. Gonaporus spp., ♀, antenna. 110. G. alfierii. 111. G. ecbatanus. 112. G. emiratus sp. nov. 113. G. gracilis. 114. G. israelicus. 115. G. maureanus. 116. G. mirabilis sp. nov. 117. G. omanicus. 118. G. setitarsus sp. nov. 119. G. simulator sp. nov. 120. G. spinosissimus sp. nov." httpUri="https://zenodo.org/record/234280/files/figure.png" pageId="36" pageNumber="487">118</figureCitation>
,
<figureCitation id="809F32FEFFAD4046733FFD75C8E4FCEB" box="[363,406,750,774]" captionStart="FIGURES 121 – 130" captionStartId="34.[151,264,1838,1861]" captionTargetBox="[166,1418,207,1792]" captionTargetId="figure@34.[165,1421,193,1792]" captionTargetPageId="34" captionText="FIGURES 121 – 130. Gonaporus spp., ♂, antenna. 121. G. alfierii. 122. G. ecbatanus. 123. G. emiratus sp. nov. 124. G. gracilis. 125. G. israelicus. 126. G. jaziratensis sp. nov. 127. G. mirabilis sp. nov. 128. G. setitarsus sp. nov. 129. G. simulator sp. nov. 130. G. spinosissimus sp. nov." httpUri="https://zenodo.org/record/234281/files/figure.png" pageId="36" pageNumber="487">128</figureCitation>
,
<figureCitation id="809F32FEFFAD404673F5FD75C8BFFCEB" box="[417,461,750,774]" captionStart="FIGURES 131 – 141" captionStartId="37.[151,264,1931,1954]" captionTargetBox="[197,1414,417,1890]" captionTargetId="figure@37.[164,1428,404,1893]" captionTargetPageId="37" captionText="FIGURES 131 – 141. Gonaporus spp., ♀, protarsus (posterior spines of tarsal comb are not shadowed; anteroventral spines are shadowed). 131. G. alfierii. 132. G. ecbatanus. 133. G. emiratus sp. nov. 134. G. gracilis. 135. G. israelicus. 136. G. maureanus. 137. G. mirabilis sp. nov. 138. G. omanicus. 139. G. setitarsus sp. nov. 140. G. simulator sp. nov. 141. G. spinosissimus sp. nov." httpUri="https://zenodo.org/record/234282/files/figure.png" pageId="36" pageNumber="487">139</figureCitation>
,
<figureCitation id="809F32FEFFAD4046738CFD76CB71FCEB" box="[472,515,749,774]" captionStart="FIGURES 142 – 152" captionStartId="38.[151,264,1652,1675]" captionTargetBox="[184,1414,196,1643]" captionTargetId="figure@38.[153,1433,193,1649]" captionTargetPageId="38" captionText="FIGURES 142 – 152. Gonaporus spp., ♂, protarsus. 142. G. alfierii. 143. G. ecbatanus 144. G. emiratus sp. nov. 145. G. gracilis. 146. G. israelicus. 147. G. jaziratensis sp. nov. 148. G. mirabilis sp. nov. 149. G. omanicus. 150. G. setitarsus sp. nov. 151. G. simulator sp. nov. 152. G. spinosissimus sp. nov." httpUri="https://zenodo.org/record/234283/files/figure.png" pageId="36" pageNumber="487">150</figureCitation>
,
<figureCitation id="809F32FEFFAD4046705AFD76CB4BFCEB" box="[526,569,749,774]" captionStart="FIGURES 160 – 166" captionStartId="40.[151,264,1876,1899]" captionTargetBox="[167,1420,203,1741]" captionTargetId="figure@40.[166,1420,193,1742]" captionTargetPageId="40" captionText="FIGURES 160 – 166. Gonaporus spp., (160 – 163, ♀; 164 – 166, ♂), fore wing. 160. G. omanicus. 161. G. setitarsus sp. nov. 162. G. simulator sp. nov. 163. G. spinosissimus sp. nov. 164. G. alfierii. 165. G. ecbatanus. 166. G. emiratus sp. nov." httpUri="https://zenodo.org/record/234285/files/figure.png" pageId="36" pageNumber="487">161</figureCitation>
,
<figureCitation id="809F32FEFFAD40467011FD75CB02FCEB" box="[581,624,750,774]" captionStart="FIGURES 167 – 174" captionStartId="41.[151,264,1759,1782]" captionTargetBox="[160,1424,202,1736]" captionTargetId="figure@41.[160,1427,193,1737]" captionTargetPageId="41" captionText="FIGURES 167 – 174. Gonaporus spp., ♂, fore wing. 167. G. gracilis. 168. G. israelicus. 169. G. jaziratensis sp. nov. 170. G. mirabilis sp. nov. 171. G. omanicus. 172. G. setitarsus sp. nov. 173. G. simulator sp. nov. 174. G. spinosissimus sp. nov." httpUri="https://zenodo.org/record/234286/files/figure.png" pageId="36" pageNumber="487">172</figureCitation>
,
<figureCitation id="809F32FEFFAD40467028FD75CBD5FCEB" box="[636,679,750,774]" captionStart="FIGURES 182 – 188" captionStartId="43.[151,264,1877,1900]" captionTargetBox="[167,1417,401,1856]" captionTargetId="figure@43.[166,1420,370,1856]" captionTargetPageId="43" captionText="FIGURES 182 – 188. Gonaporus spp. (182 – 185, ♀; 186 – 188, ♂), hind wing. 182. G. omanicus. 183. G. setitarsus sp. nov. 184. G. simulator sp. nov. 185. G. spinosissimus sp. nov. 186. G. alfierii. 187. G. ecbatanus. 188. G. gracilis." httpUri="https://zenodo.org/record/234288/files/figure.png" pageId="36" pageNumber="487">183</figureCitation>
,
<figureCitation id="809F32FEFFAD404670E6FD75CBAFFCEB" box="[690,733,750,774]" captionStart="FIGURES 189 – 195" captionStartId="45.[151,264,1615,1638]" captionTargetBox="[156,1427,221,1594]" captionTargetId="figure@45.[156,1431,193,1595]" captionTargetPageId="45" captionText="FIGURES 189 – 195. Gonaporus spp., ♂, hind wing. 189. G. israelicus. 190. G. jaziratensis sp. nov. 191. G. mirabilis sp. nov. 192. G. omanicus. 193. G. setitarsus sp. nov. 194. G. simulator sp. nov. 195. G. spinosissimus sp. nov." httpUri="https://zenodo.org/record/234289/files/figure.png" pageId="36" pageNumber="487">193</figureCitation>
,
<figureCitation id="809F32FEFFAD404670BDFD76CA66FCEB" box="[745,788,749,774]" captionStart="FIGURES 200 – 207" captionStartId="48.[151,264,1819,1842]" captionTargetBox="[210,1378,228,1761]" captionTargetId="figure@48.[196,1389,214,1767]" captionTargetPageId="48" captionText="FIGURES 200 – 207. Gonaporus and Xenaporus spp., ♂, subgenital plate (a, ventral; b, lateral; c, dorsal view). 200. G. alfierii. 201. G. ecbatanus. 202. G. emiratus sp. nov. 203. G. gracilis. 204. G. israelicus. 205. G. jaziratensis sp. nov. 206. G. setitarsus sp. nov. 207. X. ignoteremus." httpUri="https://zenodo.org/record/234290/files/figure.png" pageId="36" pageNumber="487">206</figureCitation>
,
<figureCitation id="809F32FEFFAD4046714BFD75CA38FCEB" box="[799,842,750,774]" captionStart="FIGURES 212 – 215" captionStartId="51.[151,264,1914,1937]" captionTargetBox="[321,1274,197,1825]" captionTargetId="figure@51.[297,1290,195,1839]" captionTargetPageId="51" captionText="FIGURES 212 – 215. Gonaporus and Xenaporus spp., ♂, genitalia (a, dorsal; b, ventral; c, lateral view). 212. G. israelicus. 213. G. jaziratensis sp. nov. 214. G. setitarsus sp. nov. 215. X. ignoteremus." httpUri="https://zenodo.org/record/234293/files/figure.png" pageId="36" pageNumber="487">214</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="50BE7DF0FFAD404672C3FCAFC86AFAA8" pageId="36" pageNumber="487" type="diagnosis">
<paragraph id="181B2E7BFFAD404672C3FCAFC86AFAA8" blockId="36.[151,1437,820,2033]" pageId="36" pageNumber="487">
<emphasis id="2AD0F269FFAD404672C3FCAFC863FCA0" bold="true" box="[151,273,820,845]" pageId="36" pageNumber="487">Diagnosis.</emphasis>
<taxonomicName id="DFA455F8FFAD4046734DFCAECB78FCA1" box="[281,522,821,844]" class="Insecta" family="Pompilidae" genus="Gonaporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="setitarsus" status="sp. nov.">
<emphasis id="2AD0F269FFAD4046734DFCAECB78FCA1" box="[281,522,821,844]" italics="true" pageId="36" pageNumber="487">Gonaporus setitarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31E34F12FFAD40467047FCAFCB1CFCA0" box="[531,622,820,845]" pageId="36" pageNumber="487" rank="species">
<emphasis id="2AD0F269FFAD40467047FCAFCB1CFCA0" bold="true" box="[531,622,820,845]" pageId="36" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
is most similar to
<taxonomicName id="DFA455F8FFAD40467110FCAECAA3FCA0" box="[836,977,820,845]" class="Insecta" family="Pompilidae" genus="Micraporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="israelicus">
<emphasis id="2AD0F269FFAD40467110FCAECAA3FCA0" box="[836,977,820,845]" italics="true" pageId="36" pageNumber="487">G. israelicus</emphasis>
</taxonomicName>
. The females of both species have more than one long anteroventral spine on the protarsomere 1 (
<figureCitation id="809F32FEFFAD40467147FCC3CA18FC9C" box="[787,874,856,881]" captionStart="FIGURES 17 – 28. 17" captionStartId="8.[151,264,1704,1727]" captionTargetBox="[177,1420,192,1678]" captionTargetId="figure@8.[158,1428,183,1684]" captionTargetPageId="8" captionText="FIGURES 17 – 28. 17. Stigmaporus lystracantha, wing venation. 18, 19. Gonaporus spinosissimus sp. nov., metafemur (dorsal view of apical part). 20. Micraporus priesneri, wing venation. 21 – 23. Protarsus, dorsal view. 21. G. alfierii. 22. G. ecbatanus. 23. G. israelicus. 24 – 28. Protarsus, ventral view. 24. G. alfierii. 25. G. ecbatanus. 26. G. israelicus. 27. G. simulator sp. nov. 28. G. spinosissimus sp. nov. 17, 18, 20, 24 – 28, ♀; 19, 21 – 23, ♂. Scale bar 1 mm for 17 and 0.5 mm for 18 – 28." httpUri="https://zenodo.org/record/234266/files/figure.png" pageId="36" pageNumber="487">Figs 26</figureCitation>
,
<figureCitation id="809F32FEFFAD40467121FCC3CAA9FC9D" box="[885,987,856,881]" captionStart="FIGURES 131 – 141" captionStartId="37.[151,264,1931,1954]" captionTargetBox="[197,1414,417,1890]" captionTargetId="figure@37.[164,1428,404,1893]" captionTargetPageId="37" captionText="FIGURES 131 – 141. Gonaporus spp., ♀, protarsus (posterior spines of tarsal comb are not shadowed; anteroventral spines are shadowed). 131. G. alfierii. 132. G. ecbatanus. 133. G. emiratus sp. nov. 134. G. gracilis. 135. G. israelicus. 136. G. maureanus. 137. G. mirabilis sp. nov. 138. G. omanicus. 139. G. setitarsus sp. nov. 140. G. simulator sp. nov. 141. G. spinosissimus sp. nov." httpUri="https://zenodo.org/record/234282/files/figure.png" pageId="36" pageNumber="487">135, 139</figureCitation>
); and both sexes have dense pubescence on the metapostnotum (
<figureCitation id="809F32FEFFAD404673EEFCE7CB34FC78" box="[442,582,892,917]" captionStart="FIGURES 29 – 37" captionStartId="10.[151,264,1740,1763]" captionTargetBox="[170,1421,195,1713]" captionTargetId="figure@10.[151,1436,193,1719]" captionTargetPageId="10" captionText="FIGURES 29 – 37. Gonaporus and Xenaporus spp. (29, 30, 32 – 37, ♀; 31, ♂), metapostnotum (dorsal view). 29. G. alfierii. 30 – 32. G. israelicus from Israel (30, 31) and Mauritania (32). 33. G. ecbatanus. 34. G. simulator sp. nov. 35. G. spinosissimus sp. nov. 36. X. eremocanus. 37. X. tamerlan. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/234267/files/figure.png" pageId="36" pageNumber="487">Figs 30–32</figureCitation>
). The other congeners have mostly short anteroventral spines on the protarsomere 1 (sometimes only one spine is long).
<taxonomicName id="DFA455F8FFAD404670BDFC3ACA06FC54" box="[745,884,928,953]" class="Insecta" family="Pompilidae" genus="Gonaporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="simulator" status="sp. nov.">
<emphasis id="2AD0F269FFAD404670BDFC3ACA06FC54" box="[745,884,928,953]" italics="true" pageId="36" pageNumber="487">G. simulator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31E34F12FFAD40467129FC3BCAAAFC54" box="[893,984,928,953]" pageId="36" pageNumber="487" rank="species">
<emphasis id="2AD0F269FFAD40467129FC3BCAAAFC54" bold="true" box="[893,984,928,953]" pageId="36" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
has pubescence on the metapostnotum, but the pubescence is very sparse (
<figureCitation id="809F32FEFFAD40467063FC5FCBE2FC30" box="[567,656,964,989]" captionStart="FIGURES 29 – 37" captionStartId="10.[151,264,1740,1763]" captionTargetBox="[170,1421,195,1713]" captionTargetId="figure@10.[151,1436,193,1719]" captionTargetPageId="10" captionText="FIGURES 29 – 37. Gonaporus and Xenaporus spp. (29, 30, 32 – 37, ♀; 31, ♂), metapostnotum (dorsal view). 29. G. alfierii. 30 – 32. G. israelicus from Israel (30, 31) and Mauritania (32). 33. G. ecbatanus. 34. G. simulator sp. nov. 35. G. spinosissimus sp. nov. 36. X. eremocanus. 37. X. tamerlan. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/234267/files/figure.png" pageId="36" pageNumber="487">Fig. 34</figureCitation>
). Males of
<taxonomicName id="DFA455F8FFAD40467148FC5ECADEFC30" box="[796,940,964,989]" class="Insecta" family="Pompilidae" genus="Micraporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="israelicus">
<emphasis id="2AD0F269FFAD40467148FC5ECADEFC30" box="[796,940,964,989]" italics="true" pageId="36" pageNumber="487">G. israelicus</emphasis>
</taxonomicName>
and
<taxonomicName id="DFA455F8FFAD404671B8FC5ECD09FC31" box="[1004,1147,965,988]" class="Insecta" family="Pompilidae" genus="Gonaporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="setitarsus" status="sp. nov.">
<emphasis id="2AD0F269FFAD404671B8FC5ECD09FC31" box="[1004,1147,965,988]" italics="true" pageId="36" pageNumber="487">G. setitarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31E34F12FFAD404676D2FC5FCD97FC30" box="[1158,1253,964,989]" pageId="36" pageNumber="487" rank="species">
<emphasis id="2AD0F269FFAD404676D2FC5FCD97FC30" bold="true" box="[1158,1253,964,989]" pageId="36" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
share a unique character among
<taxonomicName id="DFA455F8FFAD4046733EFC72C89AFBED" box="[362,488,1001,1024]" class="Insecta" family="Pompilidae" genus="Gonaporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="genus">
<emphasis id="2AD0F269FFAD4046733EFC72C89AFBED" box="[362,488,1001,1024]" italics="true" pageId="36" pageNumber="487">Gonaporus</emphasis>
</taxonomicName>
species—the asymmetrical and widened protarsomere 5, and similarly to
<taxonomicName id="DFA455F8FFAD404677D0FC72C86BFBC9" class="Insecta" family="Pompilidae" genus="Gonaporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="jaziratensis" status="sp. nov.">
<emphasis id="2AD0F269FFAD404677D0FC72C86BFBC9" italics="true" pageId="36" pageNumber="487">G. jaziratensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31E34F12FFAD4046734BFB97C808FBC8" box="[287,378,1036,1061]" pageId="36" pageNumber="487" rank="species">
<emphasis id="2AD0F269FFAD4046734BFB97C808FBC8" bold="true" box="[287,378,1036,1061]" pageId="36" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
, they have long hyaline spines on the protarsus (
<figureCitation id="809F32FEFFAD404671F5FB97CA8AFBC8" box="[929,1016,1036,1061]" captionStart="FIGURES 17 – 28. 17" captionStartId="8.[151,264,1704,1727]" captionTargetBox="[177,1420,192,1678]" captionTargetId="figure@8.[158,1428,183,1684]" captionTargetPageId="8" captionText="FIGURES 17 – 28. 17. Stigmaporus lystracantha, wing venation. 18, 19. Gonaporus spinosissimus sp. nov., metafemur (dorsal view of apical part). 20. Micraporus priesneri, wing venation. 21 – 23. Protarsus, dorsal view. 21. G. alfierii. 22. G. ecbatanus. 23. G. israelicus. 24 – 28. Protarsus, ventral view. 24. G. alfierii. 25. G. ecbatanus. 26. G. israelicus. 27. G. simulator sp. nov. 28. G. spinosissimus sp. nov. 17, 18, 20, 24 – 28, ♀; 19, 21 – 23, ♂. Scale bar 1 mm for 17 and 0.5 mm for 18 – 28." httpUri="https://zenodo.org/record/234266/files/figure.png" pageId="36" pageNumber="487">Figs 23</figureCitation>
,
<figureCitation id="809F32FEFFAD40467650FB97CD19FBC8" box="[1028,1131,1036,1061]" captionStart="FIGURES 142 – 152" captionStartId="38.[151,264,1652,1675]" captionTargetBox="[184,1414,196,1643]" captionTargetId="figure@38.[153,1433,193,1649]" captionTargetPageId="38" captionText="FIGURES 142 – 152. Gonaporus spp., ♂, protarsus. 142. G. alfierii. 143. G. ecbatanus 144. G. emiratus sp. nov. 145. G. gracilis. 146. G. israelicus. 147. G. jaziratensis sp. nov. 148. G. mirabilis sp. nov. 149. G. omanicus. 150. G. setitarsus sp. nov. 151. G. simulator sp. nov. 152. G. spinosissimus sp. nov." httpUri="https://zenodo.org/record/234283/files/figure.png" pageId="36" pageNumber="487">146, 150</figureCitation>
). However, the males of
<taxonomicName id="DFA455F8FFAD404677D0FB96C876FBA4" class="Insecta" family="Pompilidae" genus="Micraporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="israelicus">
<emphasis id="2AD0F269FFAD404677D0FB96C876FBA4" italics="true" pageId="36" pageNumber="487">G. israelicus</emphasis>
</taxonomicName>
and
<taxonomicName id="DFA455F8FFAD4046736AFBAAC8B8FBA5" box="[318,458,1073,1096]" class="Insecta" family="Pompilidae" genus="Gonaporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="setitarsus" status="sp. nov.">
<emphasis id="2AD0F269FFAD4046736AFBAAC8B8FBA5" box="[318,458,1073,1096]" italics="true" pageId="36" pageNumber="487">G. setitarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31E34F12FFAD40467387FBABCB5CFBA4" box="[467,558,1072,1097]" pageId="36" pageNumber="487" rank="species">
<emphasis id="2AD0F269FFAD40467387FBABCB5CFBA4" bold="true" box="[467,558,1072,1097]" pageId="36" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
have very convex subgenital plate with two longitudinal keels on its ventral side (
<figureCitation id="809F32FEFFAD40467283FBCFC80EFB80" box="[215,380,1108,1133]" captionStart="FIGURES 200 – 207" captionStartId="48.[151,264,1819,1842]" captionTargetBox="[210,1378,228,1761]" captionTargetId="figure@48.[196,1389,214,1767]" captionTargetPageId="48" captionText="FIGURES 200 – 207. Gonaporus and Xenaporus spp., ♂, subgenital plate (a, ventral; b, lateral; c, dorsal view). 200. G. alfierii. 201. G. ecbatanus. 202. G. emiratus sp. nov. 203. G. gracilis. 204. G. israelicus. 205. G. jaziratensis sp. nov. 206. G. setitarsus sp. nov. 207. X. ignoteremus." httpUri="https://zenodo.org/record/234290/files/figure.png" pageId="36" pageNumber="487">Figs 204, 206</figureCitation>
) and genitalia with widened paramere and characteristic shape of volsella, that is pointed apically (
<figureCitation id="809F32FEFFAD404672ABFBE3C8D2FB7D" box="[255,416,1144,1169]" captionStart="FIGURES 212 – 215" captionStartId="51.[151,264,1914,1937]" captionTargetBox="[321,1274,197,1825]" captionTargetId="figure@51.[297,1290,195,1839]" captionTargetPageId="51" captionText="FIGURES 212 – 215. Gonaporus and Xenaporus spp., ♂, genitalia (a, dorsal; b, ventral; c, lateral view). 212. G. israelicus. 213. G. jaziratensis sp. nov. 214. G. setitarsus sp. nov. 215. X. ignoteremus." httpUri="https://zenodo.org/record/234293/files/figure.png" pageId="36" pageNumber="487">Figs 212, 214</figureCitation>
). The male of
<taxonomicName id="DFA455F8FFAD40467010FBE2CB97FB7D" box="[580,741,1145,1168]" class="Insecta" family="Pompilidae" genus="Gonaporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="jaziratensis" status="sp. nov.">
<emphasis id="2AD0F269FFAD40467010FBE2CB97FB7D" box="[580,741,1145,1168]" italics="true" pageId="36" pageNumber="487">G. jaziratensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31E34F12FFAD404670B9FBE3CA3AFB7C" box="[749,840,1144,1169]" pageId="36" pageNumber="487" rank="species">
<emphasis id="2AD0F269FFAD404670B9FBE3CA3AFB7C" bold="true" box="[749,840,1144,1169]" pageId="36" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
has convex subgenital plate but lacking longitudinal keels (
<figureCitation id="809F32FEFFAD4046728BFB07C833FB58" box="[223,321,1180,1205]" captionStart="FIGURES 200 – 207" captionStartId="48.[151,264,1819,1842]" captionTargetBox="[210,1378,228,1761]" captionTargetId="figure@48.[196,1389,214,1767]" captionTargetPageId="48" captionText="FIGURES 200 – 207. Gonaporus and Xenaporus spp., ♂, subgenital plate (a, ventral; b, lateral; c, dorsal view). 200. G. alfierii. 201. G. ecbatanus. 202. G. emiratus sp. nov. 203. G. gracilis. 204. G. israelicus. 205. G. jaziratensis sp. nov. 206. G. setitarsus sp. nov. 207. X. ignoteremus." httpUri="https://zenodo.org/record/234290/files/figure.png" pageId="36" pageNumber="487">Fig. 205</figureCitation>
), the parameres are not widened apically and the volsella is oval and not pointed apically (
<figureCitation id="809F32FEFFAD4046777AFB07CCE2FB58" box="[1326,1424,1180,1205]" captionStart="FIGURES 212 – 215" captionStartId="51.[151,264,1914,1937]" captionTargetBox="[321,1274,197,1825]" captionTargetId="figure@51.[297,1290,195,1839]" captionTargetPageId="51" captionText="FIGURES 212 – 215. Gonaporus and Xenaporus spp., ♂, genitalia (a, dorsal; b, ventral; c, lateral view). 212. G. israelicus. 213. G. jaziratensis sp. nov. 214. G. setitarsus sp. nov. 215. X. ignoteremus." httpUri="https://zenodo.org/record/234293/files/figure.png" pageId="36" pageNumber="487">Fig. 213</figureCitation>
). The female of
<taxonomicName id="DFA455F8FFAD40467316FB5AC8BCFB35" box="[322,462,1217,1240]" class="Insecta" family="Pompilidae" genus="Gonaporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="setitarsus" status="sp. nov.">
<emphasis id="2AD0F269FFAD40467316FB5AC8BCFB35" box="[322,462,1217,1240]" italics="true" pageId="36" pageNumber="487">G. setitarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31E34F12FFAD4046738CFB5BCB46FB34" box="[472,564,1216,1241]" pageId="36" pageNumber="487" rank="species">
<emphasis id="2AD0F269FFAD4046738CFB5BCB46FB34" bold="true" box="[472,564,1216,1241]" pageId="36" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from the females of
<taxonomicName id="DFA455F8FFAD40467128FB5ACD78FB34" box="[892,1034,1216,1241]" class="Insecta" family="Pompilidae" genus="Micraporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="israelicus">
<emphasis id="2AD0F269FFAD40467128FB5ACD78FB34" box="[892,1034,1216,1241]" italics="true" pageId="36" pageNumber="487">G. israelicus</emphasis>
</taxonomicName>
by the presence of three (not two) spines in the anteroventral row of the tarsal comb. The male of
<taxonomicName id="DFA455F8FFAD40467125FB7ECA8CFB11" box="[881,1022,1253,1276]" class="Insecta" family="Pompilidae" genus="Gonaporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="setitarsus" status="sp. nov.">
<emphasis id="2AD0F269FFAD40467125FB7ECA8CFB11" box="[881,1022,1253,1276]" italics="true" pageId="36" pageNumber="487">G. setitarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="31E34F12FFAD40467652FB7FCD13FB10" box="[1030,1121,1252,1277]" pageId="36" pageNumber="487" rank="species">
<emphasis id="2AD0F269FFAD40467652FB7FCD13FB10" bold="true" box="[1030,1121,1252,1277]" pageId="36" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from the male of
<taxonomicName id="DFA455F8FFAD404677D0FB7EC876FACC" class="Insecta" family="Pompilidae" genus="Micraporus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="israelicus">
<emphasis id="2AD0F269FFAD404677D0FB7EC876FACC" italics="true" pageId="36" pageNumber="487">G. israelicus</emphasis>
</taxonomicName>
by the presence of six (not three) posterior spines on the protarsomere 1 and by the longer protarsomere 1 (Table 2).
</paragraph>
</subSubSection>
<subSubSection id="50BE7DF0FFAD40467293FACBCB13F81D" pageId="36" pageNumber="487" type="description">
<paragraph id="181B2E7BFFAD40467293FACBCDA6F965" blockId="36.[151,1437,820,2033]" pageId="36" pageNumber="487">
<emphasis id="2AD0F269FFAD40467293FACBC82AFA84" bold="true" box="[199,344,1360,1385]" pageId="36" pageNumber="487">Description.</emphasis>
<emphasis id="2AD0F269FFAD4046730BFACBC8C1FA84" box="[351,435,1360,1385]" italics="true" pageId="36" pageNumber="487">Female</emphasis>
. Body length
<quantity id="DF5C839EFFAD40467004FACBCBDAFA85" box="[592,680,1360,1385]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="36" pageNumber="487" unit="mm" value="5.7">5.7 mm</quantity>
. Structure: see in Table 1. Coloration: Head mostly black; inner and outer orbits with narrow orange stripes along eye margin; clypeus black on basal half and orange on apical; mandible mostly yellowish-orange, dark orange on apical 0.33; scape orange dorsally, yellowish-orange ventrally; flagellum brown dorsally, orange-brown ventrally. Mesosoma mostly black except of orange posterior edge of pronotum; tegula orange. Legs mostly orange; coxae black basally, orange apically; tarsi orange-brown; spur of fore leg hyaline basally, blackish apically; other spurs mostly hyaline, slightly blackish apically; spines of tarsal comb mostly hyaline, slightly orange basally; other spines orange. Metasoma brownish-orange; pygidium black, shiny. Setae: Head and propleuron with numerous long fine whitish setae; clypeus with moderately long brownish setae. Protarsomere 1 with 3 long spines posteriorly and 3 long spines anteroventrally (
<figureCitation id="809F32FEFFAD40467631F9EBCDB5F965" box="[1125,1223,1648,1673]" captionStart="FIGURES 131 – 141" captionStartId="37.[151,264,1931,1954]" captionTargetBox="[197,1414,417,1890]" captionTargetId="figure@37.[164,1428,404,1893]" captionTargetPageId="37" captionText="FIGURES 131 – 141. Gonaporus spp., ♀, protarsus (posterior spines of tarsal comb are not shadowed; anteroventral spines are shadowed). 131. G. alfierii. 132. G. ecbatanus. 133. G. emiratus sp. nov. 134. G. gracilis. 135. G. israelicus. 136. G. maureanus. 137. G. mirabilis sp. nov. 138. G. omanicus. 139. G. setitarsus sp. nov. 140. G. simulator sp. nov. 141. G. spinosissimus sp. nov." httpUri="https://zenodo.org/record/234282/files/figure.png" pageId="36" pageNumber="487">Fig. 139</figureCitation>
).
</paragraph>
<paragraph id="181B2E7BFFAD40467293F90FCB13F81D" blockId="36.[151,1437,820,2033]" pageId="36" pageNumber="487">
<emphasis id="2AD0F269FFAD40467293F90FC873F940" box="[199,257,1684,1709]" italics="true" pageId="36" pageNumber="487">Male</emphasis>
. Body length
<quantity id="DF5C839EFFAD404673EDF90FCB39F941" box="[441,587,1684,1709]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.8" metricValueMin="3.9" pageId="36" pageNumber="487" unit="mm" value="4.85" valueMax="5.8" valueMin="3.9">3.9–5.8 mm</quantity>
. Structure: see Table 2. Subgenital plate (
<figureCitation id="809F32FEFFAD40467603F90FCDB1F940" box="[1111,1219,1684,1709]" captionStart="FIGURES 200 – 207" captionStartId="48.[151,264,1819,1842]" captionTargetBox="[210,1378,228,1761]" captionTargetId="figure@48.[196,1389,214,1767]" captionTargetPageId="48" captionText="FIGURES 200 – 207. Gonaporus and Xenaporus spp., ♂, subgenital plate (a, ventral; b, lateral; c, dorsal view). 200. G. alfierii. 201. G. ecbatanus. 202. G. emiratus sp. nov. 203. G. gracilis. 204. G. israelicus. 205. G. jaziratensis sp. nov. 206. G. setitarsus sp. nov. 207. X. ignoteremus." httpUri="https://zenodo.org/record/234290/files/figure.png" pageId="36" pageNumber="487">Fig. 206</figureCitation>
): hirsute, reversetrapeziform, trapeziform in cross-section. Genitalia as in
<figureCitation id="809F32FEFFAD4046714DF923CA08F93D" box="[793,890,1720,1745]" captionStart="FIGURES 212 – 215" captionStartId="51.[151,264,1914,1937]" captionTargetBox="[321,1274,197,1825]" captionTargetId="figure@51.[297,1290,195,1839]" captionTargetPageId="51" captionText="FIGURES 212 – 215. Gonaporus and Xenaporus spp., ♂, genitalia (a, dorsal; b, ventral; c, lateral view). 212. G. israelicus. 213. G. jaziratensis sp. nov. 214. G. setitarsus sp. nov. 215. X. ignoteremus." httpUri="https://zenodo.org/record/234293/files/figure.png" pageId="36" pageNumber="487">Fig. 214</figureCitation>
. Coloration: Head mostly black; inner orbit with narrow orange stripe along eye margin; clypeus black basally, orange-brown apically; mandible mostly yellow, orange-brown apically; antenna yellowish-orange ventrally, brown dorsally. Mesosoma mostly black except of orange lateral and posterior edges of pronotum; tegula light yellow. Legs mostly orange; coxae mostly orange, narrowly black basally; protarsus mostly orange, meso- and metatarsus mostly orange to mostly brown, apical tarsomeres of all legs brown; spur of fore leg whitish basally, blackish apically; other spurs brownish-orange; leg spines hyaline to light orange. Metasoma: Segment 1–2 orange, segment 3 orange basally, blackish-orange apically; segments 4–6 mostly black, yellowish-orange apically; T7 with large white spot medially. Setae: Posterior part of head with long fine whitish setae.
</paragraph>
</subSubSection>
<subSubSection id="50BE7DF0FFAC40477293FF0CCDF2FF42" box="[199,1152,151,176]" pageId="37" pageNumber="488" type="etymology">
<paragraph id="181B2E7BFFAC40477293FF0CCDF2FF42" blockId="37.[151,1437,151,357]" box="[199,1152,151,176]" pageId="37" pageNumber="488">
<emphasis id="2AD0F269FFAC40477293FF0CC83CFF5D" bold="true" box="[199,334,151,176]" pageId="37" pageNumber="488">Etymology.</emphasis>
The specific epithet refers to the strongly spined protarsus of both sexes.
</paragraph>
</subSubSection>
<subSubSection id="50BE7DF0FFAC40477293FF27CA11FF15" pageId="37" pageNumber="488" type="materials_examined">
<paragraph id="181B2E7BFFAC40477293FF27CA11FF15" blockId="37.[151,1437,151,357]" pageId="37" pageNumber="488">
<emphasis id="2AD0F269FFAC40477293FF27C8C0FF38" bold="true" box="[199,434,188,213]" pageId="37" pageNumber="488">Material examined.</emphasis>
<typeStatus id="C71F90D9FFAC404773EFFF27CB55FF38" box="[443,551,188,213]" pageId="37" pageNumber="488" type="holotype">Holotype</typeStatus>
: ♂,
<collectingCountry id="60B36EEBFFAC4047700DFF27CB91FF39" box="[601,739,188,212]" name="Pakistan" pageId="37" pageNumber="488">PAKISTAN</collectingCountry>
: Karachi, 1911, T. R. Bell (
<collectionCode id="7EB5B6BEFFAC40477676FF27CD0EFF39" box="[1058,1148,188,212]" pageId="37" pageNumber="488">BMNH</collectionCode>
).
<typeStatus id="C71F90D9FFAC404776C4FF26CC70FF38" box="[1168,1282,189,213]" pageId="37" pageNumber="488" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="60B36EEBFFAC4047775AFF27CCEBFF39" box="[1294,1433,188,212]" name="Pakistan" pageId="37" pageNumber="488">PAKISTAN</collectingCountry>
: Karachi, T. R. Bell,
<specimenCount id="0EA2E5F2FFAC4047732CFF7BC8ECFF15" box="[376,414,224,248]" pageId="37" pageNumber="488" type="generic">2♂</specimenCount>
; Karachi, T. R. Bell, 1♀ (all in
<collectionCode id="7EB5B6BEFFAC404770AAFF7BCA25FF15" box="[766,855,224,248]" pageId="37" pageNumber="488">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="50BE7DF0FFAC40477293FE9FC8D5FEAD" pageId="37" pageNumber="488" type="distribution">
<paragraph id="181B2E7BFFAC40477293FE9FCB36FEF1" blockId="37.[151,1437,151,357]" box="[199,580,260,285]" pageId="37" pageNumber="488">
<emphasis id="2AD0F269FFAC40477293FE9FC812FEF0" bold="true" box="[199,352,260,285]" pageId="37" pageNumber="488">Distribution.</emphasis>
<collectingCountry id="60B36EEBFFAC40477333FE9FC8B4FEF0" box="[359,454,260,285]" name="Pakistan" pageId="37" pageNumber="488">Pakistan</collectingCountry>
(
<figureCitation id="809F32FEFFAC40477381FE9FCB4AFEF1" box="[469,568,260,285]" captionStart="FIGURE 217" captionStartId="53.[151,250,1215,1237]" captionTargetBox="[151,1433,529,1168]" captionTargetId="figure@53.[151,1436,526,1173]" captionTargetPageId="53" captionText="FIGURE 217. Distribution of Gonaporus spp., G. gracilis;, G. israelicus;, G. omanicus;, G. setitarsus sp. nov.;, G. simulator sp. nov." httpUri="https://zenodo.org/record/234295/files/figure.png" pageId="37" pageNumber="488">Fig. 217</figureCitation>
).
</paragraph>
<paragraph id="181B2E7BFFAC40477293FEBCC8D5FEAD" blockId="37.[151,1437,151,357]" box="[199,423,295,320]" pageId="37" pageNumber="488">
<emphasis id="2AD0F269FFAC40477293FEBCC858FEAD" bold="true" box="[199,298,295,320]" pageId="37" pageNumber="488">Habitat.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="50BE7DF0FFAC40477293FED7C8D6FE88" box="[199,420,332,357]" pageId="37" pageNumber="488" type="biology_ecology">
<paragraph id="181B2E7BFFAC40477293FED7C8D6FE88" blockId="37.[151,1437,151,357]" box="[199,420,332,357]" pageId="37" pageNumber="488">
<emphasis id="2AD0F269FFAC40477293FED7C855FE88" bold="true" box="[199,295,332,357]" pageId="37" pageNumber="488">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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