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<mods:title id="FCC7C0674E4EA505BEC91D915C2C7041">The Bostrichidae of the Maltese Islands (Coleoptera)</mods:title>
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<mods:namePart id="3D0CCECA9C6E537BC1F9C381F2147419">Nardi, Gianluca</mods:namePart>
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<mods:namePart id="4C69A84A4049EA78D17292751B63DE4E">Mifsud, David</mods:namePart>
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<mods:classification id="B215821E51B3127408435DCAEDFCE09E">journal article</mods:classification>
<mods:identifier id="358A3053A5203EC4B09117BFE874D4BB" type="DOI">http://dx.doi.org/10.3897/zookeys.481.8294</mods:identifier>
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<treatment id="85181AECD4C919EF0878573A5F7FD671" ID-GBIF-Taxon="152058227" LSID="urn:lsid:plazi:treatment:85181AECD4C919EF0878573A5F7FD671" httpUri="http://treatment.plazi.org/id/85181AECD4C919EF0878573A5F7FD671" lastPageId="10" lastPageNumber="79" pageId="9" pageNumber="78">
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<paragraph id="AD23B5FEC7FB1440F7B84C12318F266A" pageId="9" pageNumber="78">Taxon classification Animalia Coleoptera Bostrichidae</paragraph>
</subSubSection>
<subSubSection id="CD9F7A0D4E48B37CCB8F578D9770D8E2" pageId="9" pageNumber="78" type="nomenclature">
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<taxonomicName id="1EEE3B8CECDC98E1AC1DAFF37C9F32EC" ID-CoL="3L7SL" authority="Waterhouse, 1884" authorityName="Waterhouse" authorityYear="1884" class="Insecta" family="Bostrichidae" genus="Heterobostrychus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterobostrychus aequalis" order="Coleoptera" pageId="9" pageNumber="78" phylum="Arthropoda" rank="species" species="aequalis">Heterobostrychus aequalis (Waterhouse, 1884)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="33205F21C216C95EABEC4E7D65FD3882" pageId="9" pageNumber="78" type="material examined">
<paragraph id="ACB5B331C2D0E457F9C9BC6561BC75AA" pageId="9" pageNumber="78">Material examined.</paragraph>
<paragraph id="4AEE5CAD87E6AA3473BD38882FFA4F19" pageId="9" pageNumber="78">Malta: Rabat, 21.IX.2001, PS, 1 ♂ (CMM).</paragraph>
</subSubSection>
<subSubSection id="57D8D2813E9AE0D25BC45BEFA5098B14" lastPageId="10" lastPageNumber="79" pageId="9" pageNumber="78" type="chorotype">
<paragraph id="9278D24868CA58591EE13A7FF217DE0D" pageId="9" pageNumber="78">Chorotype.</paragraph>
<paragraph id="1FF1DCF7965197A304F95DE70EE86FF2" lastPageId="10" lastPageNumber="79" pageId="9" pageNumber="78">
A cosmopolitan species of Indo-Malaysian origins. It is mainly distributed in tropical and sub-tropical regions and restricted to 40° north and south of the equator (cf.
<bibRefCitation id="575F6F557052ADB6DDEFA2EB3FFF9510" author="Borowski, J" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="22" pageNumber="91" pagination="320 - 328" title="Bostrichidae." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea" year="2007">Borowski 2007</bibRefCitation>
,
<bibRefCitation id="7D6893557B81955360EB35FCBE3CC78E" author="Borowski, J" journalOrPublisher="Wydawnictwo Mantis, Olsztyn" pageId="22" pageNumber="91" title="World Catalogue of Bostrichidae (Coleoptera)." year="2007">
Borowski and
<normalizedToken id="5D7751DB21FB9CC3D9517E3E158A1510" originalValue="Węgrzynowicz">Wegrzynowicz</normalizedToken>
2007
</bibRefCitation>
,
<bibRefCitation id="C481A730365DD79432DA2C28C4F32E75" author="Azmi, MSMd" journalOrPublisher="Journal of Entomology" pageId="20" pageNumber="89" pagination="497 - 511" title="World Distribution of Heterobostrychusaequalis Waterhouse (Coleoptera: Bostrychidae)." url="10.3923/je.2011.497.511" volume="8" year="2011">Azmi et al. 2011</bibRefCitation>
: 500, fig. 2). In the Mediterranean, it was intercepted in Israel, Italy and Spain (cf.
<bibRefCitation id="26E87765785C5AA6C91D755E8B703F53" author="Gambetta, A" journalOrPublisher="The International Research Group on Wood Preservation" pageId="25" pageNumber="94" pagination="1 - 3" title="List of insects intercepted in imported timbers in Italy." volume="1217" year="1983">Gambetta 1983</bibRefCitation>
,
<bibRefCitation id="EF4B0884370D89919D762503EEF5476D" author="Geis, K-U" journalOrPublisher="Mitteilungen des Internationalen Entomologischen Vereins, Supplement" pageId="25" pageNumber="94" pagination="1 - 100" title="Gebietsfremde Splintholz- und Bohrkaefer, nach Mitteleuropa mit Importholz und anderen Guetern eingeschleppt. - Eine Bestandsaufnahme (Coleoptera: Lyctidae, Bostrichidae)." volume="10" year="2002">Geis 2002</bibRefCitation>
,
<bibRefCitation id="052DADCF162A4214681C4DDED226E987" author="Ratti, E" journalOrPublisher="Bollettino del Museo civico di Storia naturale di Venezia" pageId="34" pageNumber="103" title="Biodiversita della laguna di Venezia e della costa nordadriatica veneta. Segnalazioni. 72 - Heterobostrychusaequalis (Waterhouse, 1884) (InsectaColeopteraBostrichidae)." volume="53" year="2002">Ratti 2002</bibRefCitation>
,
<bibRefCitation id="51F949AB67A7359B550AEE5576F0DD40" author="Ratti, E" journalOrPublisher="Bollettino del Museo civico di Storia naturale di Venezia" pageId="35" pageNumber="104" pagination="121 - 125" title="ColeopteraLyctidae e Bostrichidae intercettati nel porto e negli ambienti urbani di Venezia." volume="55" year="2004">2004</bibRefCitation>
b,
<bibRefCitation id="D3CBD4F180ACAA416987AA0157DF7D02" author="Bahillo de la Puebla, P" journalOrPublisher="Heteropterus Revista de Entomologia" pageId="21" pageNumber="90" pagination="147 - 227" title="Los Bostrichidae Latreille, 1802 de la fauna de ibero-balear (Coleoptera)." volume="7" year="2007">Bahillo de la Puebla et al. 2007</bibRefCitation>
,
<bibRefCitation id="32EA8A038886C8EC8ED44D9745E5EF87" author="Ratti, E" journalOrPublisher="Bollettino del Museo civico di Storia naturale di Venezia" pageId="35" pageNumber="104" title="Coleotteri alieni in Italia / Alien Coleoptera in Italy. Vers. 2007 - 05 - 25" url="http://www.visitmuve.it" year="2007">Ratti 2007</bibRefCitation>
,
<bibRefCitation id="EEFB50C0268DE57BF71E5A1FCFA3F9EE" author="Azmi, MSMd" journalOrPublisher="Journal of Entomology" pageId="20" pageNumber="89" pagination="497 - 511" title="World Distribution of Heterobostrychusaequalis Waterhouse (Coleoptera: Bostrychidae)." url="10.3923/je.2011.497.511" volume="8" year="2011">Azmi et al. 2011</bibRefCitation>
). Ireland (
<bibRefCitation id="78E34849B3FE6E04D19614853712C768" author="O'Mahony, E" journalOrPublisher="Entomologist's Monthly Magazine" pageId="33" pageNumber="102" pagination=": 48" title="Lyctusbrunneus Steph (Col., Lyctidae) and Heterobostrichus [sic!] aequalis Waterh. (Col., Bostrichidae) in Ireland." volume="85" year="1949">
<normalizedToken id="E7BF38F12FFCEC9EB2D4B4AF5A6628FF" originalValue="OMahony">O'Mahony</normalizedToken>
1949
</bibRefCitation>
, as
<taxonomicName id="CE618B99D6C76E2F82C7D67A7AB1337A" genus="Heterobostrichus" lsidName="Heterobostrichus" pageId="10" pageNumber="79" rank="genus">
<pageBreakToken id="ACDDAC23DB0FE069DD9B165A6622BE98" pageId="10" pageNumber="79" start="start">Heterobostrichus</pageBreakToken>
</taxonomicName>
[sic!]
<taxonomicName id="E5CACF8F5A7278B274CF9E19403E8A9E" lsidName="aequalis" pageId="10" pageNumber="79" rank="species" species="aequalis">aequalis</taxonomicName>
), France (cf.
<bibRefCitation id="68E70212283C7D682032CBD3B36BFEFF" author="Brustel, H" journalOrPublisher="R. A. R. E." pageId="22" pageNumber="91" pagination="65 - 68" title="Les Bostrichidae Latreille, 1802 de la faune de France: especes autochtones, interceptees, introduites ou susceptibles de l'etre (Coleoptera)." volume="23" year="2014">Brustel and Aberlenc 2014</bibRefCitation>
) and Oregon (
<bibRefCitation id="4C555C5DF5BCD57F465BD679EAE6417F" author="Westcott, RL" journalOrPublisher="Zootaxa" pageId="38" pageNumber="107" pagination="1 - 33" title="New records and other notes for Oregon Coleoptera." volume="1142" year="2006">Westcott et al. 2006</bibRefCitation>
) must be added to the countries in which this species was intercepted (cf.
<bibRefCitation id="AD00653E6F0E79176C77D9938E7F5C86" author="Azmi, MSMd" journalOrPublisher="Journal of Entomology" pageId="20" pageNumber="89" pagination="497 - 511" title="World Distribution of Heterobostrychusaequalis Waterhouse (Coleoptera: Bostrychidae)." url="10.3923/je.2011.497.511" volume="8" year="2011">Azmi et al. 2011</bibRefCitation>
), eventhough the former record was based on collection of death specimens (larvae and adults), which was later ignored (cf.
<bibRefCitation id="AF3DB162C8ECC3C6D7CC764B5AB91683" pageId="10" pageNumber="79">Anderson et al. 1997</bibRefCitation>
,
<bibRefCitation id="123A82F3C1DFD53B2404474EFF641777" author="Geis, K-U" journalOrPublisher="Mitteilungen des Internationalen Entomologischen Vereins, Supplement" pageId="25" pageNumber="94" pagination="1 - 100" title="Gebietsfremde Splintholz- und Bohrkaefer, nach Mitteleuropa mit Importholz und anderen Guetern eingeschleppt. - Eine Bestandsaufnahme (Coleoptera: Lyctidae, Bostrichidae)." volume="10" year="2002">Geis 2002</bibRefCitation>
,
<bibRefCitation id="DF9F15798991C85B088559F97E459803" author="Nardi, G" editor="Audisio, P" journalOrPublisher="Comune di Roma, Dipartimento X Area Risorsa Suolo e Tutela Ambientale, Quaderni dell'Ambiente" pageId="32" pageNumber="101" title="Fauna Europaea: Coleoptera 2, Beetles. Fauna Europaea version 1.0" url="http://www.faunaeur.org" year="2004 b">Nardi 2004b</bibRefCitation>
,
<bibRefCitation id="83DFE38E681A2D7268FF87D5D43F21FD" author="Borowski, J" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="22" pageNumber="91" pagination="320 - 328" title="Bostrichidae." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea" year="2007">Borowski 2007</bibRefCitation>
,
<bibRefCitation id="E445036D0A3AB47AE62F6D6B71EBEFA4" author="Denux, O" editor="Roques, A" journalOrPublisher="BioRisk" pageId="24" pageNumber="93" pagination="315 - 406" title="Coleoptera families other than Cerambycidae, Curculionidae sensu lato, Chrysomelidae sensu lato and Coccinelidae. Chapter 8.5." volume="4" volumeTitle="Alien terrestrial arthropods of Europe." year="2010">Denux and Zagatti 2010</bibRefCitation>
,
<bibRefCitation id="47A9F743978E347463D6FB6C489CB663" author="Alexander, KNA" journalOrPublisher="National Parks and Wildlife Service, Department of the Arts, Heritage and the Gaeltacht, Dublin, Ireland" pageId="19" pageNumber="88" title="The beetles of decaying wood in Ireland. A provisional annotated checklist of saproxylic Coleoptera. Irish Wildlife Manuals, No. 65." year="2012">Alexander and Anderson 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="26EA8E6DF7B07CD638E0DF2970E7EA28" pageId="10" pageNumber="79" type="ecology">
<paragraph id="3A09B474EB569E0EFEC93F64AF90C651" pageId="10" pageNumber="79">Ecology.</paragraph>
<paragraph id="57084AE7DE5A961535268A745DF7A92A" pageId="10" pageNumber="79">
Polyphagous species attacking some 36 unrelated host-plant genera; this species breeds not only in logs, but also in planks, furniture, plywood and roots of manioc (cf.
<bibRefCitation id="F7726B0B97C18E1C2E7652B97AEE8F04" author="Fisher, WS" journalOrPublisher="United States Department of Agriculture Miscellaneous Publications" pageId="25" pageNumber="94" pagination="1 - 157" title="A revision of the North American Species of Beetles belonging to the family Bostrichidae." volume="698" year="1950">Fisher 1950</bibRefCitation>
,
<bibRefCitation id="799FAC5B2CAC1648FB127666E909B9D8" author="Kalshoven, LGE" journalOrPublisher="Entomologische Berichten" pageId="29" pageNumber="98" pagination="242 - 257" title="Notes on the biology of Indonesian Bostrychidae (Coleoptera)." volume="23" year="1963 a">Kalshoven 1963a</bibRefCitation>
,
<bibRefCitation id="72A54CD9DB400C8447620FA51F65D287" author="Kalshoven, LGE" journalOrPublisher="Beaufortia" pageId="29" pageNumber="98" pagination="232 - 240" title="Ecological data on some neotropical Scolytidae, Platypodidae, and Bostrychidae (Coleoptera), mainly of Surinam." volume="9" year="1963 b">1963b</bibRefCitation>
,
<bibRefCitation id="A602583CB28B1B66623EEA06D1DD10AA" author="Horion, A" journalOrPublisher="Entomologische Blaetter" pageId="27" pageNumber="96" pagination="9 - 42" title="Zwoelfter Nachtrag zum Verzeichnis der mitteleuropaeischen Kaefer." volume="68" year="1972">Horion 1972</bibRefCitation>
,
<bibRefCitation id="77F82AEA2C4CFE2DE0837A4355B635F2" author="Gambetta, A" journalOrPublisher="The International Research Group on Wood Preservation" pageId="25" pageNumber="94" pagination="1 - 3" title="List of insects intercepted in imported timbers in Italy." volume="1217" year="1983">Gambetta 1983</bibRefCitation>
,
<bibRefCitation id="EF9EEDC91DBFF6C792DE9562C7BA6A21" author="Wang, H" journalOrPublisher="International Network for Bamboo and Rattan (INBAR), Beijing, Eindhoven, New Delhi" pageId="38" pageNumber="107" title="Insect Pests of Bamboos in Asia - An Illustrated Manual." year="1996">Wang et al. 1996</bibRefCitation>
, as
<taxonomicName id="9DA3E7E3A45222FBC2CA641F38D4449B" genus="Heterobostrachus" lsidName="Heterobostrachus" pageId="10" pageNumber="79" rank="genus">Heterobostrachus</taxonomicName>
[sic!]
<taxonomicName id="21061F5B279C98C965ADE4DAC114985A" lsidName="aequalis" pageId="10" pageNumber="79" rank="species" species="aequalis">aequalis</taxonomicName>
,
<bibRefCitation id="30F88231CA3D636578B027BA1C56A6BB" author="Geis, K-U" journalOrPublisher="Mitteilungen des Internationalen Entomologischen Vereins, Supplement" pageId="25" pageNumber="94" pagination="1 - 100" title="Gebietsfremde Splintholz- und Bohrkaefer, nach Mitteleuropa mit Importholz und anderen Guetern eingeschleppt. - Eine Bestandsaufnahme (Coleoptera: Lyctidae, Bostrichidae)." volume="10" year="2002">Geis 2002</bibRefCitation>
, Maes 2005,
<bibRefCitation id="1D3F349969CDDB77220399365DAE9704" author="Aguilera, AO" journalOrPublisher="Mitteilungen Landesmuseum Joanneum, Zoologie" pageId="19" pageNumber="88" title="Analisis de riesgo para Heterobostrychusaequalis" url="http://www.semarnat.gob.mx/temas/gestionambiental/forestalsuelos/Paginas/saludforestal.aspx" year="2006">Aguilera 2006</bibRefCitation>
,
<bibRefCitation id="250E67EFF1FA7985F9F10E4E632CB33F" author="Bahillo de la Puebla, P" journalOrPublisher="Heteropterus Revista de Entomologia" pageId="21" pageNumber="90" pagination="147 - 227" title="Los Bostrichidae Latreille, 1802 de la fauna de ibero-balear (Coleoptera)." volume="7" year="2007">Bahillo de la Puebla et al. 2007</bibRefCitation>
,
<bibRefCitation id="1ECB17DAE3F24A6E1A7A6F6E02AA49F9" author="Sittichaya, W" journalOrPublisher="ZooKeys" pageId="36" pageNumber="105" pagination="33 - 51" title="An illustrated key to powder post beetles (Coleoptera, Bostrichidae) associated with rubberwood in Thailand, with new records and a checklist of species found in Southern Thailand." url="10.3897/zookeys.26.88" volume="26" year="2009">Sitticaya et al. 2009</bibRefCitation>
,
<bibRefCitation id="21E4151BB532D6C459B027FF2B08668D" author="Robinson, WH" journalOrPublisher="Cambridge University Press, London" pageId="35" pageNumber="104" title="Urban Insects and Arachnids: A Handbook of Urban Entomology." year="2013">Robinson 2013</bibRefCitation>
).
</paragraph>
<paragraph id="A8EEF4A6D8D4F8A4588671CAABD43DE0" pageId="10" pageNumber="79">
Notes. First record for Malta. This species become established in some countries where it was accidentally introduced. Temperatures of 17 °C and below are said to be unsuitable for the species to breed (cf.
<bibRefCitation id="AA6A3694F6CC129E2ADB7EB175648AB7" author="Ivie, MA" editor="Arnett, RHJr (†)" journalOrPublisher="CRC Press, Boca Raton, London, New York, Washington" pageId="28" pageNumber="97" pagination="233 - 244" title="69. Bostrichidae Latreille 1802." volumeTitle="American Beetles Polyphaga: Scarabaeoidea through Curculionoidea" year="2002">Ivie 2002</bibRefCitation>
,
<bibRefCitation id="BD4C41DDB62214B24DAFBDB45668E6A0" author="Azmi, MSMd" journalOrPublisher="Journal of Entomology" pageId="20" pageNumber="89" pagination="497 - 511" title="World Distribution of Heterobostrychusaequalis Waterhouse (Coleoptera: Bostrychidae)." url="10.3923/je.2011.497.511" volume="8" year="2011">Azmi et al. 2011</bibRefCitation>
). Thus, considering the warm climate of the Maltese Islands, (
<bibRefCitation id="9E2C11F0BF1BFC66E097B8587BEB6C6B" author="Chetcuti, D" journalOrPublisher="Malta Univesity Press, Msida" pageId="23" pageNumber="92" title="The climate of the Maltese Islands: a review" year="1992">Chetcuti et al. 1992</bibRefCitation>
), it is highly likely that the species is already an established one. The above specimen was collected with a light trap on a terrace, mainly surrounded by agricultural land (P. Sammut, pers. comm., 2002).
</paragraph>
</subSubSection>
</treatment>
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