376 lines
59 KiB
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376 lines
59 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/dez.68.66181" ID-GBIF-Dataset="1a313eaa-9ffe-4656-9ad7-a6f19a8f1223" ID-Pensoft-Pub="1860-1324-1-207" ID-Pensoft-UUID="FF3F5EA9976253D3A2B7D94BBC40CFB0" ID-ZooBank="0D454AFB9C3142C3A5B9CD9E256C1F41" ModsDocID="1860-1324-68-1-207" checkinTime="1622665281715" checkinUser="pensoft" docAuthor="Schmidt, Joachim, Scholz, Stephan & Maddison, David R." docDate="2021" docId="CB7D5F0C41FA52B8BF76A0954218E63B" docLanguage="en" docName="DeutEntomolZeit 68(1): 207-224" docOrigin="Deutsche Entomologische Zeitschrift 68 (1)" docPubDate="2021-06-02" docSource="http://dx.doi.org/10.3897/dez.68.66181" docTitle="Balticeler kerneggeri Schmidt & Maddison 2021, sp. nov." docType="treatment" docUuid="32A6F70E-6731-490D-B563-73F3F79695B0" docUuidSource="ZooBank" docVersion="4" id="FF3F5EA9976253D3A2B7D94BBC40CFB0" lastPageNumber="207" masterDocId="FF3F5EA9976253D3A2B7D94BBC40CFB0" masterDocTitle="Balticeler kerneggeri gen. nov., sp. nov., an enigmatic Baltic amber fossil of the ground beetle subfamily Trechinae (Coleoptera, Carabidae)" masterLastPageNumber="224" masterPageNumber="207" pageNumber="207" updateTime="1732482658392" updateUser="ExternalLinkService">
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<mods:title id="26891A2297A2093B3DD254577E2FA06E">Balticeler kerneggeri gen. nov., sp. nov., an enigmatic Baltic amber fossil of the ground beetle subfamily Trechinae (Coleoptera, Carabidae)</mods:title>
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<mods:roleTerm id="97305D1276986C1002D09CE6E241132E">Author</mods:roleTerm>
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<mods:namePart id="941E87F2676ADF24FCEA9CE6433E89F4">Schmidt, Joachim</mods:namePart>
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<mods:affiliation id="8A8FF4B8829FF9AEE3E76E427E887BF9">University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitaetsplatz 2, 18055 Rostock, Germany</mods:affiliation>
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<mods:nameIdentifier id="2EBFF5C2DA7FAF8929C15E1BC0BD222E" type="email">agonumschmidt@hotmail.com</mods:nameIdentifier>
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<mods:roleTerm id="1EADB2C05762B404C5106C93EEC924DF">Author</mods:roleTerm>
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<mods:namePart id="308C510F96007187C4324658C88C212E">Scholz, Stephan</mods:namePart>
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<mods:affiliation id="3DC9E921C9E17D6A61DE99AFFFCD0441">University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitaetsplatz 2, 18055 Rostock, Germany</mods:affiliation>
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<mods:namePart id="B1EA7D4D59DB47D06AFF092EF923D93B">Maddison, David R.</mods:namePart>
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<mods:nameIdentifier id="A1FE7208B8D0B030DDDC8B29AB4FF1E9" type="ORCID">https://orcid.org/0000-0002-7152-3824</mods:nameIdentifier>
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<mods:affiliation id="B8CF7273FB994A912A4440D3795E5E67">Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
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<mods:nameIdentifier id="4FA5C68FD3A3B10135A54C520540FC83" type="email">david.maddison@science.oregonstate.edu</mods:nameIdentifier>
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<mods:title id="D7E423F47B77C5AE3B32A5EDFE5E04D6">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="C4C27B613BC81BC5E4B9B20551EFE47D">2021</mods:date>
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<mods:number id="573A3B192E90A554EFC8597223DDD420">2021-06-02</mods:number>
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<mods:number id="0EB5A307DFBE21E01D64B49AB7FFA983">68</mods:number>
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<mods:classification id="E9A7E58B755AD0B840964ACCBBA559E2">journal article</mods:classification>
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<mods:identifier id="9D857F38F82DCC0715A290C8E4632BE5" type="DOI">http://dx.doi.org/10.3897/dez.68.66181</mods:identifier>
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<subSection id="C791F3CCAD6C93DAD10A7309A548A99E" pageId="0" pageNumber="207" type="results and taxonomy">
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<treatment id="CB7D5F0C41FA52B8BF76A0954218E63B" ID-GBIF-Taxon="182486536" LSID="urn:lsid:zoobank.org:act:32A6F70E-6731-490D-B563-73F3F79695B0" httpUri="http://treatment.plazi.org/id/CB7D5F0C41FA52B8BF76A0954218E63B" lastPageNumber="207" pageId="0" pageNumber="207" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Trechinae">
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<subSubSection id="F82CD760E38AB042768ACBE8CEE5799A" pageId="0" pageNumber="207" type="nomenclature">
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<paragraph id="133D45FE862C26A65AFE38D6FBAA87AF" pageId="0" pageNumber="207">
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<taxonomicName id="3D057452CDD392A34FC1E9CA9AD1969E" LSID="http://zoobank.org/32A6F70E-6731-490D-B563-73F3F79695B0" authority="Schmidt & Maddison" authorityName="Schmidt & Maddison" authorityYear="2021" class="Insecta" family="Carabidae" genus="Balticeler" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balticeler kerneggeri" order="Coleoptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="kerneggeri" status="sp. nov.">Balticeler kerneggeri Schmidt & Maddison</taxonomicName>
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<taxonomicNameLabel id="71E9BE46EC4C6B1B8C371C82FE30F812" pageId="0" pageNumber="207">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="332CD912723569C7C072DAFB32FFFE47" pageId="0" pageNumber="207" type="type material">
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<paragraph id="B9CC8BC47F6FD23CFDBDE7239AB3F810" pageId="0" pageNumber="207">Type material.</paragraph>
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<paragraph id="C23EFF09307E72892B8DDFF63DF753A4" pageId="0" pageNumber="207">
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<emphasis id="E54969022839A78856B5B286C6E4D054" bold="true" pageId="0" pageNumber="207">
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<emphasis id="4A8BAEB41C657600DDA4C63A92718BF7" italics="true" pageId="0" pageNumber="207">Holotype</emphasis>
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:
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</emphasis>
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Male in Baltic amber, Coll. Friedrich Kernegger in the Centrum of Natural History, Hamburg (CeNak), with collection number " GPIH 04897, coll. Kernegger 1995/235". Size of the amber piece approx. 13
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<normalizedToken id="61D6BE2225FE5FD26D0B28E50AC1EEB6" originalValue="×">x</normalizedToken>
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7
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7 mm, irregularly cut, embedded in synthetic resin (
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<bibRefCitation id="4B96E187301C40A97086FA58FC2DF88C" author="Hoffeins, HW" journalOrPublisher="Polish Journal of Entomology" pageId="0" pageNumber="207" pagination="215 - 219" refId="B8" refString="Hoffeins, HW, 2001. On the preparation and conservation of amber inclusions in artificial resin. Polish Journal of Entomology 70: 215 - 219" title="On the preparation and conservation of amber inclusions in artificial resin." volume="70" year="2001">Hoffeins 2001</bibRefCitation>
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) (Fig.
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<figureCitation id="28B1729FB9E7F993D453D25C4D17D530" captionStart="Figures 1–7" captionStartId="F1" captionText="Figures 1 - 7. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the holotype (coll. Kernegger 235; 1 - 3), paratype 1 (OSAC 000 - 2900006; 4), paratype 2 (coll. Groehn 8234; 5 - 7); the enclosed photographs in 3 - 5 show the general view of the respective amber pieces, with arrows pointing to the respective location of the carabid fossil. 1. Frontolateral view of the fossil specimen. 2. Posterolateral view of the fossil specimen (the arrows point to the two setae on left side of abdominal segment VII). 3 - 5. Dorsal (3, 4) respectively left dorsolateral view (5) of the fossil specimens. 6. Head, left dorsolateral view (the white arrows point to the insertions of the four clypeal setae). 7. Right protarsomeres and anterior part of tibia with antenna cleaner, anterior surface. Abbreviations: a 1 - a 4 - antennomeres 1 - 4; ant - antenna; as - protibial anterior spur; cs - clip setae of antenna cleaner; el-l - left elytron; el-r - right elytron; hw - hind wing; ls - pronotal lateral seta; pr - pronotum; ps - protibial posterior spur; s 1, s 8 - 1 st resp. 8 th elytral stria; sas - subapical seta of elytra; sos - supraorbital seta." figureDoi="10.3897/dez.68.66181.figures1-7" httpUri="https://binary.pensoft.net/fig/551014" pageId="0" pageNumber="207">3</figureCitation>
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).
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</paragraph>
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<paragraph id="4E77973C3B8850955A3FA3D322D8DC9C" pageId="0" pageNumber="207">
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Preservation status: The amber is clear but pervaded by few flowlines dorsally of the embedded beetle fossil. The fossil is partly covered by milky coating on the ventral side of the body. The exoskeleton of the specimen and its negative imprint on the inclusion wall is well preserved and could therefore be visualized in detail using micro-CT (Figs
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<figureCitation id="91D8E26A41DD2BA1071E75ADD8CB1AC4" captionStart="Figures 11–14" captionStartId="F3" captionText="Figures 11 - 14. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the holotype. 11. Dorsal aspect (the black arrows point to the positions of the discal seta on left elytron). 12. Ventral aspect. 13. Right lateral aspect. 14. Left lateral aspect (the black arrows point to the positions of the seta of the umbilicate series on left elytron)." figureDoi="10.3897/dez.68.66181.figures11-14" httpUri="https://binary.pensoft.net/fig/551016" pageId="0" pageNumber="207">11-14</figureCitation>
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). In contrast, the aedeagus is poorly preserved inside the fossilized beetle body: it is strongly shrunken and deformed and therefore, its original shape and structure could not be determined.
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</paragraph>
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<paragraph id="364389A99B5CA94178AC86457882C25C" pageId="0" pageNumber="207">Syninclusions: None.</paragraph>
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<paragraph id="87EE2357D3FD088DB0D11AA3FBBFEA53" pageId="0" pageNumber="207">
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<emphasis id="528DEF0A2DAC154E114A4E9A3610D282" bold="true" pageId="0" pageNumber="207">
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<emphasis id="82F6E1235C902245FC4E1AD890842E1F" italics="true" pageId="0" pageNumber="207">Paratype 1</emphasis>
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:
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</emphasis>
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Male in Baltic amber, deposited in the Oregon State University Collection, with specimen number OSAC_0002900006. Size of the amber piece approx. 21
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<normalizedToken id="06C6D9478FA0F637ADD7B6D988CB8799" originalValue="×">x</normalizedToken>
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8
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<normalizedToken id="06A3A30C5E4ECA520F8C4E4330A27445" originalValue="×">x</normalizedToken>
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4 mm, irregularly cut (Fig.
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<figureCitation id="D06ABC3B26B82EA2C124AB07DE2679C8" captionStart="Figures 1–7" captionStartId="F1" captionText="Figures 1 - 7. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the holotype (coll. Kernegger 235; 1 - 3), paratype 1 (OSAC 000 - 2900006; 4), paratype 2 (coll. Groehn 8234; 5 - 7); the enclosed photographs in 3 - 5 show the general view of the respective amber pieces, with arrows pointing to the respective location of the carabid fossil. 1. Frontolateral view of the fossil specimen. 2. Posterolateral view of the fossil specimen (the arrows point to the two setae on left side of abdominal segment VII). 3 - 5. Dorsal (3, 4) respectively left dorsolateral view (5) of the fossil specimens. 6. Head, left dorsolateral view (the white arrows point to the insertions of the four clypeal setae). 7. Right protarsomeres and anterior part of tibia with antenna cleaner, anterior surface. Abbreviations: a 1 - a 4 - antennomeres 1 - 4; ant - antenna; as - protibial anterior spur; cs - clip setae of antenna cleaner; el-l - left elytron; el-r - right elytron; hw - hind wing; ls - pronotal lateral seta; pr - pronotum; ps - protibial posterior spur; s 1, s 8 - 1 st resp. 8 th elytral stria; sas - subapical seta of elytra; sos - supraorbital seta." figureDoi="10.3897/dez.68.66181.figures1-7" httpUri="https://binary.pensoft.net/fig/551014" pageId="0" pageNumber="207">4</figureCitation>
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).
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</paragraph>
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<paragraph id="608F7C42A6FE8F38B5B66B95F42E921B" pageId="0" pageNumber="207">
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Preservation status: The embedded beetle fossil is almost completely surrounded by flowlines. Head and prothorax are additionally covered by milky coating. Therefore, only small parts of the beetle body are visible using light microscopy (Fig.
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<figureCitation id="F76138705B3956FCA093EBC05D29973F" captionStart="Figures 1–7" captionStartId="F1" captionText="Figures 1 - 7. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the holotype (coll. Kernegger 235; 1 - 3), paratype 1 (OSAC 000 - 2900006; 4), paratype 2 (coll. Groehn 8234; 5 - 7); the enclosed photographs in 3 - 5 show the general view of the respective amber pieces, with arrows pointing to the respective location of the carabid fossil. 1. Frontolateral view of the fossil specimen. 2. Posterolateral view of the fossil specimen (the arrows point to the two setae on left side of abdominal segment VII). 3 - 5. Dorsal (3, 4) respectively left dorsolateral view (5) of the fossil specimens. 6. Head, left dorsolateral view (the white arrows point to the insertions of the four clypeal setae). 7. Right protarsomeres and anterior part of tibia with antenna cleaner, anterior surface. Abbreviations: a 1 - a 4 - antennomeres 1 - 4; ant - antenna; as - protibial anterior spur; cs - clip setae of antenna cleaner; el-l - left elytron; el-r - right elytron; hw - hind wing; ls - pronotal lateral seta; pr - pronotum; ps - protibial posterior spur; s 1, s 8 - 1 st resp. 8 th elytral stria; sas - subapical seta of elytra; sos - supraorbital seta." figureDoi="10.3897/dez.68.66181.figures1-7" httpUri="https://binary.pensoft.net/fig/551014" pageId="0" pageNumber="207">4</figureCitation>
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). The exoskeleton of the fossil and its negative imprint on the inclusion wall are moderately well preserved and could therefore be imaged using micro-CT, but the head capsule is partly shrunken and displaced into the prothorax (Figs
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<figureCitation id="6A8A6A5A560CC5C3E0FCCDF97270A8C6" captionStart="Figures 26–28" captionStartId="F6" captionText="Figures 26 - 28. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the paratype 1 (OSAC 000 - 2900006) using different grey scales. 26, 27. Dorsal aspect. 28. Right lateral aspect. The aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software in 27 and 28. Abbreviations: bb - air bubble attached to the left posterior part of the beetle body; ph - shrunken and posteriorly displaced head capsule of the fossil specimen." figureDoi="10.3897/dez.68.66181.figures26-28" httpUri="https://binary.pensoft.net/fig/551019" pageId="0" pageNumber="207">26-28</figureCitation>
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). The aedeagus is rather well preserved inside the fossilized beetle body and could therefore be imaged using micro-CT in most details (Figs
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<figureCitation id="BDFB39C9E4E2377714DE85C863BE2CDC" captionStart="Figures 26–28" captionStartId="F6" captionText="Figures 26 - 28. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the paratype 1 (OSAC 000 - 2900006) using different grey scales. 26, 27. Dorsal aspect. 28. Right lateral aspect. The aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software in 27 and 28. Abbreviations: bb - air bubble attached to the left posterior part of the beetle body; ph - shrunken and posteriorly displaced head capsule of the fossil specimen." figureDoi="10.3897/dez.68.66181.figures26-28" httpUri="https://binary.pensoft.net/fig/551019" pageId="0" pageNumber="207">27</figureCitation>
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,
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<figureCitation id="85B3062ABAEC690D56DA8FB138BF24A7" captionStart="Figures 26–28" captionStartId="F6" captionText="Figures 26 - 28. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the paratype 1 (OSAC 000 - 2900006) using different grey scales. 26, 27. Dorsal aspect. 28. Right lateral aspect. The aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software in 27 and 28. Abbreviations: bb - air bubble attached to the left posterior part of the beetle body; ph - shrunken and posteriorly displaced head capsule of the fossil specimen." figureDoi="10.3897/dez.68.66181.figures26-28" httpUri="https://binary.pensoft.net/fig/551019" pageId="0" pageNumber="207">28</figureCitation>
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,
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<figureCitation id="E6FEE7B83A336C0790C424D4A4D18737" captionStart="Figures 32–35" captionStartId="F8" captionText="Figures 32 - 35. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the aedeagus of paratype 1 (OSAC 000 - 2900006) (32, 33) and paratype 2 (coll. Groehn 8234) (34), and transverse sections through the abdomen with aedeagus of paratype 1 (35). 32 a, 33, 34 a. Left lateral view. 32 b. Left dorsolateral view. 32 c. Dorsal view. 32 d, 34 b. Ventral view. The parameres were separated using the segmentation function of Amira software and highlighted by green colour in 32 and 34. The position of the selected slice in 35 a is shown as a red line in 35 b which illustrates the aedeagus in right lateral view. Abbreviations: do - dorsal opening (= dorsally connected basal and apical ostia); la - apical lamella of aedeagus; lbl, lbr - left resp. right lobe of median lobe bulb; md - membranous dorsal surface of the median lobe; mv - ventral surface of the median lobe; pma-l - apical apophysis of the left paramere; pma-r - preserved remain of the apical apophysis of the right paramere; pmb-l, pmb-r - basal part of the left respectively right paramere; pmm-l, pmm-r - discoidal middle portion of the left respectively right paramere; st-IV - sternit IV." figureDoi="10.3897/dez.68.66181.figures32-35" httpUri="https://binary.pensoft.net/fig/551021" pageId="0" pageNumber="207">32</figureCitation>
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,
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<figureCitation id="D9B84BB0C40C4C5BC92B29419979BB8B" captionStart="Figures 32–35" captionStartId="F8" captionText="Figures 32 - 35. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the aedeagus of paratype 1 (OSAC 000 - 2900006) (32, 33) and paratype 2 (coll. Groehn 8234) (34), and transverse sections through the abdomen with aedeagus of paratype 1 (35). 32 a, 33, 34 a. Left lateral view. 32 b. Left dorsolateral view. 32 c. Dorsal view. 32 d, 34 b. Ventral view. The parameres were separated using the segmentation function of Amira software and highlighted by green colour in 32 and 34. The position of the selected slice in 35 a is shown as a red line in 35 b which illustrates the aedeagus in right lateral view. Abbreviations: do - dorsal opening (= dorsally connected basal and apical ostia); la - apical lamella of aedeagus; lbl, lbr - left resp. right lobe of median lobe bulb; md - membranous dorsal surface of the median lobe; mv - ventral surface of the median lobe; pma-l - apical apophysis of the left paramere; pma-r - preserved remain of the apical apophysis of the right paramere; pmb-l, pmb-r - basal part of the left respectively right paramere; pmm-l, pmm-r - discoidal middle portion of the left respectively right paramere; st-IV - sternit IV." figureDoi="10.3897/dez.68.66181.figures32-35" httpUri="https://binary.pensoft.net/fig/551021" pageId="0" pageNumber="207">33</figureCitation>
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,
|
||
<figureCitation id="1841203512873FE4C4E4393DE01C1177" captionStart="Figures 32–35" captionStartId="F8" captionText="Figures 32 - 35. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the aedeagus of paratype 1 (OSAC 000 - 2900006) (32, 33) and paratype 2 (coll. Groehn 8234) (34), and transverse sections through the abdomen with aedeagus of paratype 1 (35). 32 a, 33, 34 a. Left lateral view. 32 b. Left dorsolateral view. 32 c. Dorsal view. 32 d, 34 b. Ventral view. The parameres were separated using the segmentation function of Amira software and highlighted by green colour in 32 and 34. The position of the selected slice in 35 a is shown as a red line in 35 b which illustrates the aedeagus in right lateral view. Abbreviations: do - dorsal opening (= dorsally connected basal and apical ostia); la - apical lamella of aedeagus; lbl, lbr - left resp. right lobe of median lobe bulb; md - membranous dorsal surface of the median lobe; mv - ventral surface of the median lobe; pma-l - apical apophysis of the left paramere; pma-r - preserved remain of the apical apophysis of the right paramere; pmb-l, pmb-r - basal part of the left respectively right paramere; pmm-l, pmm-r - discoidal middle portion of the left respectively right paramere; st-IV - sternit IV." figureDoi="10.3897/dez.68.66181.figures32-35" httpUri="https://binary.pensoft.net/fig/551021" pageId="0" pageNumber="207">35</figureCitation>
|
||
); its external shape particularly near base, including the lateral lobes of the median lobe basal bulb, seem somewhat deformed (Fig.
|
||
<figureCitation id="83373DAF7CA431DACFA446E8A814886F" captionStart="Figures 32–35" captionStartId="F8" captionText="Figures 32 - 35. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the aedeagus of paratype 1 (OSAC 000 - 2900006) (32, 33) and paratype 2 (coll. Groehn 8234) (34), and transverse sections through the abdomen with aedeagus of paratype 1 (35). 32 a, 33, 34 a. Left lateral view. 32 b. Left dorsolateral view. 32 c. Dorsal view. 32 d, 34 b. Ventral view. The parameres were separated using the segmentation function of Amira software and highlighted by green colour in 32 and 34. The position of the selected slice in 35 a is shown as a red line in 35 b which illustrates the aedeagus in right lateral view. Abbreviations: do - dorsal opening (= dorsally connected basal and apical ostia); la - apical lamella of aedeagus; lbl, lbr - left resp. right lobe of median lobe bulb; md - membranous dorsal surface of the median lobe; mv - ventral surface of the median lobe; pma-l - apical apophysis of the left paramere; pma-r - preserved remain of the apical apophysis of the right paramere; pmb-l, pmb-r - basal part of the left respectively right paramere; pmm-l, pmm-r - discoidal middle portion of the left respectively right paramere; st-IV - sternit IV." figureDoi="10.3897/dez.68.66181.figures32-35" httpUri="https://binary.pensoft.net/fig/551021" pageId="0" pageNumber="207">33</figureCitation>
|
||
), probably because the fossil specimen was embedded during an immature stage of its development; the apical apophysis of the right paramere is only partly preserved (Fig.
|
||
<figureCitation id="9C2EDCCFAF659FE901122F0FE2364A83" captionStart="Figures 32–35" captionStartId="F8" captionText="Figures 32 - 35. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the aedeagus of paratype 1 (OSAC 000 - 2900006) (32, 33) and paratype 2 (coll. Groehn 8234) (34), and transverse sections through the abdomen with aedeagus of paratype 1 (35). 32 a, 33, 34 a. Left lateral view. 32 b. Left dorsolateral view. 32 c. Dorsal view. 32 d, 34 b. Ventral view. The parameres were separated using the segmentation function of Amira software and highlighted by green colour in 32 and 34. The position of the selected slice in 35 a is shown as a red line in 35 b which illustrates the aedeagus in right lateral view. Abbreviations: do - dorsal opening (= dorsally connected basal and apical ostia); la - apical lamella of aedeagus; lbl, lbr - left resp. right lobe of median lobe bulb; md - membranous dorsal surface of the median lobe; mv - ventral surface of the median lobe; pma-l - apical apophysis of the left paramere; pma-r - preserved remain of the apical apophysis of the right paramere; pmb-l, pmb-r - basal part of the left respectively right paramere; pmm-l, pmm-r - discoidal middle portion of the left respectively right paramere; st-IV - sternit IV." figureDoi="10.3897/dez.68.66181.figures32-35" httpUri="https://binary.pensoft.net/fig/551021" pageId="0" pageNumber="207">32d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="BEE3348C6421A7EFC782BB20E3269051" doi="10.3897/dez.68.66181.figures26-28" httpUri="https://binary.pensoft.net/fig/551019" pageId="0" pageNumber="207" start="Figures 26–28" startId="F6">
|
||
<paragraph id="20EBF185D60F09D9DDFC0910774B0120" pageId="0" pageNumber="207">
|
||
<emphasis id="4D027E59B3E7B776446989568251D490" bold="true" pageId="0" pageNumber="207">Figures 26-28.</emphasis>
|
||
<taxonomicName id="53934C9573E39FC69B91F12A0A0A46E0" authorityName="Schmidt & Maddison" authorityYear="2021" class="Insecta" family="Carabidae" genus="Balticeler" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balticeler kerneggeri" order="Coleoptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="kerneggeri">
|
||
<emphasis id="F7F7D79C74AE7011D2E6EB49EC76B67B" italics="true" pageId="0" pageNumber="207">Balticeler kerneggeri</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov., volume rendering of the paratype 1 (OSAC 000-2900006) using different grey scales.
|
||
<emphasis id="C01E215E105585D24E768AF577173C41" bold="true" pageId="0" pageNumber="207">26, 27.</emphasis>
|
||
Dorsal aspect.
|
||
<emphasis id="54E68C8E0D3E8A90A7EE96F7D304829A" bold="true" pageId="0" pageNumber="207">28.</emphasis>
|
||
Right lateral aspect. The aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software in
|
||
<emphasis id="54B12D3D26094BE7A1B353FF170CDB17" bold="true" pageId="0" pageNumber="207">27</emphasis>
|
||
and
|
||
<emphasis id="379EF988C4221F8BF732FC7085AB3E49" bold="true" pageId="0" pageNumber="207">28</emphasis>
|
||
. Abbreviations:
|
||
<emphasis id="86AAEB02DD5BED47C06F3A1D0D726449" bold="true" pageId="0" pageNumber="207">bb</emphasis>
|
||
- air bubble attached to the left posterior part of the beetle body;
|
||
<emphasis id="6DA72FAE9A4A8BB0C2DB10B90E5F755B" bold="true" pageId="0" pageNumber="207">ph</emphasis>
|
||
- shrunken and posteriorly displaced head capsule of the fossil specimen.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA0ADE4980B0D309D64674CB33B8100F" pageId="0" pageNumber="207">Syninclusions: None.</paragraph>
|
||
<paragraph id="6FA2AFAE47D22CC414FE14AFD34427EB" pageId="0" pageNumber="207">
|
||
<emphasis id="7A65E07DA4CF094590924E56AF29D6E2" bold="true" pageId="0" pageNumber="207">
|
||
<emphasis id="EAE2E75A4D3A70D94EC729C8ABD17B36" italics="true" pageId="0" pageNumber="207">Paratype 2</emphasis>
|
||
:
|
||
</emphasis>
|
||
Male in Baltic amber, Coll. Carsten
|
||
<normalizedToken id="4C7B4883D6D8867AF1A4965A596E7FEC" originalValue="Gröhn">Groehn</normalizedToken>
|
||
in the collection of the Geological-Palaeontological Institute of the University of Hamburg (GPIH), now in the Centrum of Natural History, Hamburg (CeNak), with collection number " GPIH 5044, coll.
|
||
<normalizedToken id="E4555D6760CEF882BC95FB707EE131D7" originalValue="Gröhn">Groehn</normalizedToken>
|
||
8234". The amber piece was nearly triangularly cut with an edge length of about 33, 26, and 16 mm, and with width of 5 mm (Fig.
|
||
<figureCitation id="19201D56269ED80864B575960797626E" captionStart="Figures 1–7" captionStartId="F1" captionText="Figures 1 - 7. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the holotype (coll. Kernegger 235; 1 - 3), paratype 1 (OSAC 000 - 2900006; 4), paratype 2 (coll. Groehn 8234; 5 - 7); the enclosed photographs in 3 - 5 show the general view of the respective amber pieces, with arrows pointing to the respective location of the carabid fossil. 1. Frontolateral view of the fossil specimen. 2. Posterolateral view of the fossil specimen (the arrows point to the two setae on left side of abdominal segment VII). 3 - 5. Dorsal (3, 4) respectively left dorsolateral view (5) of the fossil specimens. 6. Head, left dorsolateral view (the white arrows point to the insertions of the four clypeal setae). 7. Right protarsomeres and anterior part of tibia with antenna cleaner, anterior surface. Abbreviations: a 1 - a 4 - antennomeres 1 - 4; ant - antenna; as - protibial anterior spur; cs - clip setae of antenna cleaner; el-l - left elytron; el-r - right elytron; hw - hind wing; ls - pronotal lateral seta; pr - pronotum; ps - protibial posterior spur; s 1, s 8 - 1 st resp. 8 th elytral stria; sas - subapical seta of elytra; sos - supraorbital seta." figureDoi="10.3897/dez.68.66181.figures1-7" httpUri="https://binary.pensoft.net/fig/551014" pageId="0" pageNumber="207">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FFD7CF6F19A77A9F201BCFBFE620D190" pageId="0" pageNumber="207">
|
||
Preservation status: The amber is clear; large flowlines are attached to the left lateral side of the embedded beetle. The ventral surface of the beetle body is partly covered by milky coating. Antenna, legs, and dorsal surface of head are clear and thus well visible using light microscopy (Figs
|
||
<figureCitation id="A05D712C481BC43D51587AC18D9971C6" captionStart="Figures 1–7" captionStartId="F1" captionText="Figures 1 - 7. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the holotype (coll. Kernegger 235; 1 - 3), paratype 1 (OSAC 000 - 2900006; 4), paratype 2 (coll. Groehn 8234; 5 - 7); the enclosed photographs in 3 - 5 show the general view of the respective amber pieces, with arrows pointing to the respective location of the carabid fossil. 1. Frontolateral view of the fossil specimen. 2. Posterolateral view of the fossil specimen (the arrows point to the two setae on left side of abdominal segment VII). 3 - 5. Dorsal (3, 4) respectively left dorsolateral view (5) of the fossil specimens. 6. Head, left dorsolateral view (the white arrows point to the insertions of the four clypeal setae). 7. Right protarsomeres and anterior part of tibia with antenna cleaner, anterior surface. Abbreviations: a 1 - a 4 - antennomeres 1 - 4; ant - antenna; as - protibial anterior spur; cs - clip setae of antenna cleaner; el-l - left elytron; el-r - right elytron; hw - hind wing; ls - pronotal lateral seta; pr - pronotum; ps - protibial posterior spur; s 1, s 8 - 1 st resp. 8 th elytral stria; sas - subapical seta of elytra; sos - supraorbital seta." figureDoi="10.3897/dez.68.66181.figures1-7" httpUri="https://binary.pensoft.net/fig/551014" pageId="0" pageNumber="207">6</figureCitation>
|
||
,
|
||
<figureCitation id="45B2D1B974DCB822F2A65BCC884C0810" captionStart="Figures 1–7" captionStartId="F1" captionText="Figures 1 - 7. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the holotype (coll. Kernegger 235; 1 - 3), paratype 1 (OSAC 000 - 2900006; 4), paratype 2 (coll. Groehn 8234; 5 - 7); the enclosed photographs in 3 - 5 show the general view of the respective amber pieces, with arrows pointing to the respective location of the carabid fossil. 1. Frontolateral view of the fossil specimen. 2. Posterolateral view of the fossil specimen (the arrows point to the two setae on left side of abdominal segment VII). 3 - 5. Dorsal (3, 4) respectively left dorsolateral view (5) of the fossil specimens. 6. Head, left dorsolateral view (the white arrows point to the insertions of the four clypeal setae). 7. Right protarsomeres and anterior part of tibia with antenna cleaner, anterior surface. Abbreviations: a 1 - a 4 - antennomeres 1 - 4; ant - antenna; as - protibial anterior spur; cs - clip setae of antenna cleaner; el-l - left elytron; el-r - right elytron; hw - hind wing; ls - pronotal lateral seta; pr - pronotum; ps - protibial posterior spur; s 1, s 8 - 1 st resp. 8 th elytral stria; sas - subapical seta of elytra; sos - supraorbital seta." figureDoi="10.3897/dez.68.66181.figures1-7" httpUri="https://binary.pensoft.net/fig/551014" pageId="0" pageNumber="207">7</figureCitation>
|
||
). The skeleton of the fossil beetle specimen together with its aedeagus is moderately well preserved and could therefore be imaged using micro-CT (Figs
|
||
<figureCitation id="CB844622459DDD8E9B806452F93DE5C3" captionStart="Figures 29–31" captionStartId="F7" captionText="Figures 29 - 31. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the paratype 2 (coll. Groehn 8234) using different grey scales. 29. Dorsal aspect. 30. Left lateral aspect. 31. Right lateral aspect (downsized visualization with respect to 30) to show position of aedeagus (separated using the segmentation function of Amira software and highlighted by red colour) in the beetle body." figureDoi="10.3897/dez.68.66181.figures29-31" httpUri="https://binary.pensoft.net/fig/551020" pageId="0" pageNumber="207">29-31</figureCitation>
|
||
); however, the basal portion of the aedeagus is partly decayed so that the parameres are preserved only with the more strongly sclerotized median parts (Fig.
|
||
<figureCitation id="2418B47EA602F1056947F675D6C4AEE4" captionStart="Figures 32–35" captionStartId="F8" captionText="Figures 32 - 35. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the aedeagus of paratype 1 (OSAC 000 - 2900006) (32, 33) and paratype 2 (coll. Groehn 8234) (34), and transverse sections through the abdomen with aedeagus of paratype 1 (35). 32 a, 33, 34 a. Left lateral view. 32 b. Left dorsolateral view. 32 c. Dorsal view. 32 d, 34 b. Ventral view. The parameres were separated using the segmentation function of Amira software and highlighted by green colour in 32 and 34. The position of the selected slice in 35 a is shown as a red line in 35 b which illustrates the aedeagus in right lateral view. Abbreviations: do - dorsal opening (= dorsally connected basal and apical ostia); la - apical lamella of aedeagus; lbl, lbr - left resp. right lobe of median lobe bulb; md - membranous dorsal surface of the median lobe; mv - ventral surface of the median lobe; pma-l - apical apophysis of the left paramere; pma-r - preserved remain of the apical apophysis of the right paramere; pmb-l, pmb-r - basal part of the left respectively right paramere; pmm-l, pmm-r - discoidal middle portion of the left respectively right paramere; st-IV - sternit IV." figureDoi="10.3897/dez.68.66181.figures32-35" httpUri="https://binary.pensoft.net/fig/551021" pageId="0" pageNumber="207">34 a, b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="5404884E102A912BD4DBECF468B45169" doi="10.3897/dez.68.66181.figures29-31" httpUri="https://binary.pensoft.net/fig/551020" pageId="0" pageNumber="207" start="Figures 29–31" startId="F7">
|
||
<paragraph id="F66B4D25FC5A7E2D489489016A4CEA9D" pageId="0" pageNumber="207">
|
||
<emphasis id="3432AFB67B2FE1A456361BD13305F3B3" bold="true" pageId="0" pageNumber="207">Figures 29-31.</emphasis>
|
||
<taxonomicName id="FEAF979896AB3785F114173416687E8A" authorityName="Schmidt & Maddison" authorityYear="2021" class="Insecta" family="Carabidae" genus="Balticeler" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balticeler kerneggeri" order="Coleoptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="kerneggeri">
|
||
<emphasis id="98CED5163D9B34DABF7C632C535F22EE" italics="true" pageId="0" pageNumber="207">Balticeler kerneggeri</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov., volume rendering of the paratype 2 (coll. Groehn 8234) using different grey scales.
|
||
<emphasis id="493B4016229164B38052077E2BFC1D65" bold="true" pageId="0" pageNumber="207">29.</emphasis>
|
||
Dorsal aspect.
|
||
<emphasis id="34144E26697F7911F544DAF293252EB7" bold="true" pageId="0" pageNumber="207">30.</emphasis>
|
||
Left lateral aspect.
|
||
<emphasis id="167053935AFFC0BCFB24FC7FF7BAED65" bold="true" pageId="0" pageNumber="207">31.</emphasis>
|
||
Right lateral aspect (downsized visualization with respect to
|
||
<emphasis id="AC6FEBE8BC9F9BFB548F389A5BE325A0" bold="true" pageId="0" pageNumber="207">30</emphasis>
|
||
) to show position of aedeagus (separated using the segmentation function of Amira software and highlighted by red colour) in the beetle body.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="46B03017867B727781605702DFB324F2" pageId="0" pageNumber="207">Syninclusions: 1 stellate hair, few dust particles.</paragraph>
|
||
<paragraph id="E7D3653C32ED78E922D0E941533C09DB" pageId="0" pageNumber="207">
|
||
<emphasis id="7B432D7F577B0EBE78FD1BFC1617E12A" bold="true" pageId="0" pageNumber="207">
|
||
<emphasis id="6E1281B6F281ACE1CCB26132970BB2B0" italics="true" pageId="0" pageNumber="207">Paratype 3</emphasis>
|
||
:
|
||
</emphasis>
|
||
Specimen of unknown sex in Baltic amber, deposited in the Oregon State University Collection, with amber piece number OSAC_AMB0000387. According to the dealer (Marius Veta, Palanga) from whom the amber was purchased, this is most likely Baltic amber from the Yantarni mine, but there is a slight chance that it is Rovno amber. Size of the amber piece approx. 14
|
||
<normalizedToken id="FED5753157D02F1D297537E3338CD6DA" originalValue="×">x</normalizedToken>
|
||
8
|
||
<normalizedToken id="CB496AC03B631B81B9DFBC4199029C2A" originalValue="×">x</normalizedToken>
|
||
2 mm, irregularly cut (Fig.
|
||
<figureCitation id="88D9338EF97E9A0C529D9700AADC4E7A" captionStart="Figures 8–10" captionStartId="F2" captionText="Figures 8 - 10. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the paratype 3 (OSAC 000 - 2900387; 8), paratype 4 (OSAC 000 - 2900600; 9), paratype 5 (coll. Sciaky; 10); the enclosed photographs show the general view of the respective amber pieces, with arrow in 8 pointing to the location of the carabid fossil. 8. Dorsal view. 9. Left lateral view. 10. Right dorsolateral view. Abbreviations: bs - pronotal basal seta; ds-a, ds-m, ds-p - anterior, medial respectively posterior discal seta on right elytron; ls - pronotal lateral seta." figureDoi="10.3897/dez.68.66181.figures8-10" httpUri="https://binary.pensoft.net/fig/551015" pageId="0" pageNumber="207">8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="07F4196C9C05FC52D74583941D5D02C7" pageId="0" pageNumber="207">
|
||
Preservation status: The amber is clear but the ventral surface of the embedded beetle is completely covered by milky coating. Flowlines within the amber extend to each lateral side of the fossil. Most parts of the legs and the tip of the abdomen were abraded during polishing process and are thus lacking. Therefore, only the dorsal side of the beetle body is visible using light microscopy (Fig.
|
||
<figureCitation id="958A4E515FB16AC84D7B1C44543F624D" captionStart="Figures 8–10" captionStartId="F2" captionText="Figures 8 - 10. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the paratype 3 (OSAC 000 - 2900387; 8), paratype 4 (OSAC 000 - 2900600; 9), paratype 5 (coll. Sciaky; 10); the enclosed photographs show the general view of the respective amber pieces, with arrow in 8 pointing to the location of the carabid fossil. 8. Dorsal view. 9. Left lateral view. 10. Right dorsolateral view. Abbreviations: bs - pronotal basal seta; ds-a, ds-m, ds-p - anterior, medial respectively posterior discal seta on right elytron; ls - pronotal lateral seta." figureDoi="10.3897/dez.68.66181.figures8-10" httpUri="https://binary.pensoft.net/fig/551015" pageId="0" pageNumber="207">8</figureCitation>
|
||
). The negative imprint of the fossil on the inclusion wall has low contrast in the X-ray analyses and therefore, certain details of external morphology could not be imaged and the sex of the specimen could not be assessed (Figs
|
||
<figureCitation id="F727215FB1A0D5772B03131A9757E993" captionStart="Figures 36–42" captionStartId="F9" captionText="Figures 36 - 42. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the paratype 3 (OSAC 000 - 2900387) (36, 37), paratype 4 (OSAC 000 - 2900600) (38 - 40), paratype 5 (coll. Sciaky) (41, 42). 36, 38, 41. Dorsal aspect. 37, 39. Left lateral aspect. 40. Dorso-frontal aspect. 42. Right lateral aspect." figureDoi="10.3897/dez.68.66181.figures36-42" httpUri="https://binary.pensoft.net/fig/551022" pageId="0" pageNumber="207">36</figureCitation>
|
||
,
|
||
<figureCitation id="F6F969DDCA5A750359C393339642A170" captionStart="Figures 36–42" captionStartId="F9" captionText="Figures 36 - 42. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the paratype 3 (OSAC 000 - 2900387) (36, 37), paratype 4 (OSAC 000 - 2900600) (38 - 40), paratype 5 (coll. Sciaky) (41, 42). 36, 38, 41. Dorsal aspect. 37, 39. Left lateral aspect. 40. Dorso-frontal aspect. 42. Right lateral aspect." figureDoi="10.3897/dez.68.66181.figures36-42" httpUri="https://binary.pensoft.net/fig/551022" pageId="0" pageNumber="207">37</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="219DB25DBC0844AE7A4D169D1239882C" doi="10.3897/dez.68.66181.figures32-35" httpUri="https://binary.pensoft.net/fig/551021" pageId="0" pageNumber="207" start="Figures 32–35" startId="F8">
|
||
<paragraph id="4ED3F5C7C92076B165B5E9C10F9AD3EF" pageId="0" pageNumber="207">
|
||
<emphasis id="DDD4FF27E26B9DD5B440E4A92D2BC0F1" bold="true" pageId="0" pageNumber="207">Figures 32-35.</emphasis>
|
||
<taxonomicName id="5B9B0F3A7B35E56DF555B0FE43E16FF2" authorityName="Schmidt & Maddison" authorityYear="2021" class="Insecta" family="Carabidae" genus="Balticeler" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balticeler kerneggeri" order="Coleoptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="kerneggeri">
|
||
<emphasis id="3A2D5573D7822B3F46B4377907526914" italics="true" pageId="0" pageNumber="207">Balticeler kerneggeri</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov., volume rendering of the aedeagus of paratype 1 (OSAC 000-2900006) (
|
||
<emphasis id="D8B2C2E3512640A160D68E955150F2C6" bold="true" pageId="0" pageNumber="207">32, 33</emphasis>
|
||
) and paratype 2 (coll. Groehn 8234) (
|
||
<emphasis id="33CFF4E44225E8DF59231FD995697F29" bold="true" pageId="0" pageNumber="207">34</emphasis>
|
||
), and transverse sections through the abdomen with aedeagus of paratype 1 (
|
||
<emphasis id="281792EE52437D347227364BE0FB2721" bold="true" pageId="0" pageNumber="207">35</emphasis>
|
||
).
|
||
<emphasis id="54C5D1FB9FC497F9BAD1F4409D190267" bold="true" pageId="0" pageNumber="207">32a, 33, 34a.</emphasis>
|
||
Left lateral view.
|
||
<emphasis id="1A204501EC4E7FAF2D79A5E8C788394B" bold="true" pageId="0" pageNumber="207">32b.</emphasis>
|
||
Left dorsolateral view.
|
||
<emphasis id="ADCAD56F40169A3960890F3407246A2C" bold="true" pageId="0" pageNumber="207">32c.</emphasis>
|
||
Dorsal view.
|
||
<emphasis id="1B03DF5FCD3608CEA5A8EE8D8B27B1D3" bold="true" pageId="0" pageNumber="207">32d, 34b.</emphasis>
|
||
Ventral view. The parameres were separated using the segmentation function of Amira software and highlighted by green colour in
|
||
<emphasis id="249F60F773AEB6DFE75DC3894CF6D991" bold="true" pageId="0" pageNumber="207">32</emphasis>
|
||
and
|
||
<emphasis id="34B4C5F706E48144EF39D626D07EFD19" bold="true" pageId="0" pageNumber="207">34</emphasis>
|
||
. The position of the selected slice in
|
||
<emphasis id="443F5972FDBC9E7020F8D03D4B34715D" bold="true" pageId="0" pageNumber="207">35a</emphasis>
|
||
is shown as a red line in
|
||
<emphasis id="EB49D116868F1B8C2133F82E2E4ED50F" bold="true" pageId="0" pageNumber="207">35b</emphasis>
|
||
which illustrates the aedeagus in right lateral view. Abbreviations:
|
||
<emphasis id="983DB6A5109E159E508940F06CC4FA72" bold="true" pageId="0" pageNumber="207">do</emphasis>
|
||
- dorsal opening (= dorsally connected basal and apical ostia);
|
||
<emphasis id="438AAD663CCC51327E5EF1157A23C9A0" bold="true" pageId="0" pageNumber="207">la</emphasis>
|
||
- apical lamella of aedeagus;
|
||
<emphasis id="D2BDB2C1CFB2F629B8939769EB5B5177" bold="true" pageId="0" pageNumber="207">lbl, lbr</emphasis>
|
||
- left resp. right lobe of median lobe bulb;
|
||
<emphasis id="C49F8247AF64808FA7DE6131D6FCFCD0" bold="true" pageId="0" pageNumber="207">md</emphasis>
|
||
- membranous dorsal surface of the median lobe;
|
||
<emphasis id="AE1B36CC2F706AB77A9BAB3B8E8F48B0" bold="true" pageId="0" pageNumber="207">mv</emphasis>
|
||
- ventral surface of the median lobe;
|
||
<emphasis id="85E51B79EDF1AFF4FA3FD4460F066794" bold="true" pageId="0" pageNumber="207">pma-l</emphasis>
|
||
- apical apophysis of the left paramere;
|
||
<emphasis id="BB2F60CE4CC70A9E09C9EC6E926CF510" bold="true" pageId="0" pageNumber="207">pma-r</emphasis>
|
||
- preserved remain of the apical apophysis of the right paramere;
|
||
<emphasis id="99E5131C223CBB5375DBDAA1014F3C58" bold="true" pageId="0" pageNumber="207">pmb-l, pmb-r</emphasis>
|
||
- basal part of the left respectively right paramere;
|
||
<emphasis id="8D3640DF94BCBA9BB158F64E9E8F9B76" bold="true" pageId="0" pageNumber="207">pmm-l, pmm-r</emphasis>
|
||
- discoidal middle portion of the left respectively right paramere;
|
||
<emphasis id="6770B902355D7152D3A7E0CC2C853C39" bold="true" pageId="0" pageNumber="207">st-IV</emphasis>
|
||
- sternit IV.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C1E0B45642A464D8A76292CC9FB213FC" doi="10.3897/dez.68.66181.figures36-42" httpUri="https://binary.pensoft.net/fig/551022" pageId="0" pageNumber="207" start="Figures 36–42" startId="F9">
|
||
<paragraph id="F4622A005552778CEFFE9F3CF6C0B328" pageId="0" pageNumber="207">
|
||
<emphasis id="380A33D090AE700EA928F2A44E18B27B" bold="true" pageId="0" pageNumber="207">Figures 36-42.</emphasis>
|
||
<taxonomicName id="8183CA0D2B85121C6D27C1F9E5A59AB2" authorityName="Schmidt & Maddison" authorityYear="2021" class="Insecta" family="Carabidae" genus="Balticeler" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balticeler kerneggeri" order="Coleoptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="species" species="kerneggeri">
|
||
<emphasis id="D17B9C96D143780DB02FD7FBDA0115AA" italics="true" pageId="0" pageNumber="207">Balticeler kerneggeri</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov., volume rendering of the paratype 3 (OSAC 000-2900387) (
|
||
<emphasis id="4546C4A57C1ACE8BA6AB9D1E210DE426" bold="true" pageId="0" pageNumber="207">36, 37</emphasis>
|
||
), paratype 4 (OSAC 000-2900600) (
|
||
<emphasis id="D2A1D5551CBFACCB507127F9EFD14B1F" bold="true" pageId="0" pageNumber="207">38-40</emphasis>
|
||
), paratype 5 (coll. Sciaky) (
|
||
<emphasis id="441E507CDBD68DB59E390B22C0E08C10" bold="true" pageId="0" pageNumber="207">41, 42</emphasis>
|
||
).
|
||
<emphasis id="955A97C3FCEA33E58E99DFDFD35C1028" bold="true" pageId="0" pageNumber="207">36, 38, 41.</emphasis>
|
||
Dorsal aspect.
|
||
<emphasis id="3F39A69236A75C7D54815C32A6A3E056" bold="true" pageId="0" pageNumber="207">37, 39.</emphasis>
|
||
Left lateral aspect.
|
||
<emphasis id="8E32513C3BD91859D5DDE9B936D6D9E6" bold="true" pageId="0" pageNumber="207">40.</emphasis>
|
||
Dorso-frontal aspect.
|
||
<emphasis id="4046053652F3071CF7F34C37422BB8A7" bold="true" pageId="0" pageNumber="207">42.</emphasis>
|
||
Right lateral aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1F624CE3B3C0C940B86AE9FDF5D43461" pageId="0" pageNumber="207">Syninclusions: One spider.</paragraph>
|
||
<paragraph id="632E52D10315B98D4C1495A2C5559E9B" pageId="0" pageNumber="207">
|
||
<emphasis id="43958182AC82E1ACBEEA6DD9DB91450E" bold="true" pageId="0" pageNumber="207">
|
||
<emphasis id="4B7226280A6628B5DACC854E88EA5C3E" italics="true" pageId="0" pageNumber="207">Paratype 4</emphasis>
|
||
:
|
||
</emphasis>
|
||
Specimen of unknown sex in Baltic amber, deposited in the Oregon State University Collection, with amber piece number OSAC_AMB0000600. Size of the amber piece approx. 13
|
||
<normalizedToken id="0A8E4D17861586CDE9DED76B6BE2A24C" originalValue="×">x</normalizedToken>
|
||
8
|
||
<normalizedToken id="6BC677C6199C46BB11F06FF453587EB1" originalValue="×">x</normalizedToken>
|
||
5 mm, cut into an approximately trapezoidal form (Fig.
|
||
<figureCitation id="A93EADD144F8EDEB6CEF89C34E375D10" captionStart="Figures 8–10" captionStartId="F2" captionText="Figures 8 - 10. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the paratype 3 (OSAC 000 - 2900387; 8), paratype 4 (OSAC 000 - 2900600; 9), paratype 5 (coll. Sciaky; 10); the enclosed photographs show the general view of the respective amber pieces, with arrow in 8 pointing to the location of the carabid fossil. 8. Dorsal view. 9. Left lateral view. 10. Right dorsolateral view. Abbreviations: bs - pronotal basal seta; ds-a, ds-m, ds-p - anterior, medial respectively posterior discal seta on right elytron; ls - pronotal lateral seta." figureDoi="10.3897/dez.68.66181.figures8-10" httpUri="https://binary.pensoft.net/fig/551015" pageId="0" pageNumber="207">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F1AFADFD91049D055C1F8EFCA87E8F58" pageId="0" pageNumber="207">
|
||
Preservation status: The amber is pervaded by numerous air bubbles on the right side to the embedded beetle fossil; the latter is well visible from dorsal, left lateral and ventral sides using light microscope (Fig.
|
||
<figureCitation id="868E6AA1FF854CC0643177168D9EF2BC" captionStart="Figures 8–10" captionStartId="F2" captionText="Figures 8 - 10. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the paratype 3 (OSAC 000 - 2900387; 8), paratype 4 (OSAC 000 - 2900600; 9), paratype 5 (coll. Sciaky; 10); the enclosed photographs show the general view of the respective amber pieces, with arrow in 8 pointing to the location of the carabid fossil. 8. Dorsal view. 9. Left lateral view. 10. Right dorsolateral view. Abbreviations: bs - pronotal basal seta; ds-a, ds-m, ds-p - anterior, medial respectively posterior discal seta on right elytron; ls - pronotal lateral seta." figureDoi="10.3897/dez.68.66181.figures8-10" httpUri="https://binary.pensoft.net/fig/551015" pageId="0" pageNumber="207">9</figureCitation>
|
||
), but the head and thorax are covered by milky coating ventrally. The stone was very likely altered by autoclaving in order to reduce milky coating; this is evident from the blackened appendages of the beetle which are slightly deformed (particularly tibiae and tarsi), and from one of the synincluded
|
||
<taxonomicName id="1A4BF4FB5F61238BB5432F5E9E5ED662" authorityName="Schmidt & Maddison" authorityYear="2021" kingdom="Animalia" lsidName="" order="Collembola" pageId="0" pageNumber="207" rank="order">Collembola</taxonomicName>
|
||
which has a roasted appearance (for the effect of autoclaving on amber fossils see
|
||
<bibRefCitation id="063A3EF85A996A7B301DA8B2E8756C37" DOI="https://doi.org/10.2478/v10200-012-0005-z" author="Hoffeins, C" journalOrPublisher="Polish Journal of Entomology" pageId="0" pageNumber="207" pagination="165 - 181" refId="B9" refString="Hoffeins, C, 2012. On Baltic amber inclusions treated in an autoclave. Polish Journal of Entomology 81: 165 - 181, DOI: https://doi.org/10.2478/v10200-012-0005-z" title="On Baltic amber inclusions treated in an autoclave." url="https://doi.org/10.2478/v10200-012-0005-z" volume="81" year="2012">Hoffeins 2012</bibRefCitation>
|
||
). Very probably as a consequence of autoclaving, the negative imprint of the fossil on the inclusion wall has low contrast in the X-ray analyses; certain details of external morphology could thus not be imaged, and the genital armatures are not preserved (Figs
|
||
<figureCitation id="F3D1892320240C8E4006B744BBC7BDB6" captionStart="Figures 36–42" captionStartId="F9" captionText="Figures 36 - 42. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the paratype 3 (OSAC 000 - 2900387) (36, 37), paratype 4 (OSAC 000 - 2900600) (38 - 40), paratype 5 (coll. Sciaky) (41, 42). 36, 38, 41. Dorsal aspect. 37, 39. Left lateral aspect. 40. Dorso-frontal aspect. 42. Right lateral aspect." figureDoi="10.3897/dez.68.66181.figures36-42" httpUri="https://binary.pensoft.net/fig/551022" pageId="0" pageNumber="207">38-40</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2EF17C8CFA850E2ACC2466FEFF3F4770" pageId="0" pageNumber="207">
|
||
Syninclusions: One rove beetle (
|
||
<taxonomicName id="B77277CD0F3C5DB7C0D5F724A4DCE7E8" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="207" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
|
||
), one springtail (
|
||
<taxonomicName id="9E7C3DFE526937F672206747E57C0EF4" authorityName="Schmidt & Maddison" authorityYear="2021" kingdom="Animalia" lsidName="" order="Collembola" pageId="0" pageNumber="207" rank="order">Collembola</taxonomicName>
|
||
), remains of a second springtail.
|
||
</paragraph>
|
||
<paragraph id="15A8511EC22DB9151CC9E3C58C969BA2" pageId="0" pageNumber="207">
|
||
<emphasis id="6AA630BF4369C62D52D48AB1313E4A3D" bold="true" pageId="0" pageNumber="207">
|
||
<emphasis id="669F6FEEC271FBACBC37640D8D3E8425" italics="true" pageId="0" pageNumber="207">Paratype 5</emphasis>
|
||
:
|
||
</emphasis>
|
||
Specimen of unknown sex in Baltic amber, ex collectio Riccardo Sciaky (Milano), now preserved in the Museum
|
||
<normalizedToken id="58423EE39432E70FD9BF5569B432A17A" originalValue="für">fuer</normalizedToken>
|
||
Naturkunde Berlin, with specimen number MB.I.8614. Size of the amber piece approx. 10
|
||
<normalizedToken id="DF4058BACEA5247E5888EC99242629CF" originalValue="×">x</normalizedToken>
|
||
5
|
||
<normalizedToken id="351D22E92465EB0EC74BF93473873688" originalValue="×">x</normalizedToken>
|
||
5 mm, polished into a somewhat bean-shaped piece (Fig.
|
||
<figureCitation id="64F1D5DA176F56680E2E77B76A9D8584" captionStart="Figures 8–10" captionStartId="F2" captionText="Figures 8 - 10. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the paratype 3 (OSAC 000 - 2900387; 8), paratype 4 (OSAC 000 - 2900600; 9), paratype 5 (coll. Sciaky; 10); the enclosed photographs show the general view of the respective amber pieces, with arrow in 8 pointing to the location of the carabid fossil. 8. Dorsal view. 9. Left lateral view. 10. Right dorsolateral view. Abbreviations: bs - pronotal basal seta; ds-a, ds-m, ds-p - anterior, medial respectively posterior discal seta on right elytron; ls - pronotal lateral seta." figureDoi="10.3897/dez.68.66181.figures8-10" httpUri="https://binary.pensoft.net/fig/551015" pageId="0" pageNumber="207">10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F475997581EBC80F95DF7E128211E406" pageId="0" pageNumber="207">
|
||
Preservation status: Rather poor; head and ventral surface of the beetle fossil are completely covered by milky coating; in addition, flow lines of the amber surrounding the specimen (Fig.
|
||
<figureCitation id="66932F857F380105C9A423C6176CFACA" captionStart="Figures 8–10" captionStartId="F2" captionText="Figures 8 - 10. Balticeler kerneggeri gen. nov., sp. nov., light microscopic images of the paratype 3 (OSAC 000 - 2900387; 8), paratype 4 (OSAC 000 - 2900600; 9), paratype 5 (coll. Sciaky; 10); the enclosed photographs show the general view of the respective amber pieces, with arrow in 8 pointing to the location of the carabid fossil. 8. Dorsal view. 9. Left lateral view. 10. Right dorsolateral view. Abbreviations: bs - pronotal basal seta; ds-a, ds-m, ds-p - anterior, medial respectively posterior discal seta on right elytron; ls - pronotal lateral seta." figureDoi="10.3897/dez.68.66181.figures8-10" httpUri="https://binary.pensoft.net/fig/551015" pageId="0" pageNumber="207">10</figureCitation>
|
||
). The negative imprint of the fossil on the inclusion wall has moderate low contrast in the X-ray analyses (Figs
|
||
<figureCitation id="B8B9E1CC24C8891E77A6508C4C17A33E" captionStart="Figures 36–42" captionStartId="F9" captionText="Figures 36 - 42. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the paratype 3 (OSAC 000 - 2900387) (36, 37), paratype 4 (OSAC 000 - 2900600) (38 - 40), paratype 5 (coll. Sciaky) (41, 42). 36, 38, 41. Dorsal aspect. 37, 39. Left lateral aspect. 40. Dorso-frontal aspect. 42. Right lateral aspect." figureDoi="10.3897/dez.68.66181.figures36-42" httpUri="https://binary.pensoft.net/fig/551022" pageId="0" pageNumber="207">41</figureCitation>
|
||
,
|
||
<figureCitation id="F49E1EA8C3CBD719B469BC19B012BF6C" captionStart="Figures 36–42" captionStartId="F9" captionText="Figures 36 - 42. Balticeler kerneggeri gen. nov., sp. nov., volume rendering of the paratype 3 (OSAC 000 - 2900387) (36, 37), paratype 4 (OSAC 000 - 2900600) (38 - 40), paratype 5 (coll. Sciaky) (41, 42). 36, 38, 41. Dorsal aspect. 37, 39. Left lateral aspect. 40. Dorso-frontal aspect. 42. Right lateral aspect." figureDoi="10.3897/dez.68.66181.figures36-42" httpUri="https://binary.pensoft.net/fig/551022" pageId="0" pageNumber="207">42</figureCitation>
|
||
), and the genital armatures are not preserved.
|
||
</paragraph>
|
||
<paragraph id="723241AF33A36A199E30C1507631618C" pageId="0" pageNumber="207">Syninclusions: None.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3C224F3C80FD636A04586D44D4814BD9" pageId="0" pageNumber="207" type="description">
|
||
<paragraph id="363DBDEFE7B2C25E040D3CB786061AFC" pageId="0" pageNumber="207">Description.</paragraph>
|
||
<paragraph id="8B43F286EDA52F09B5F8D812174AAC93" pageId="0" pageNumber="207">Color: All specimens appear unicolored dark throughout, markedly shiny. No metallic reflection is visible.</paragraph>
|
||
<paragraph id="55CE83AF2D4734211ACD631EA3703A9E" pageId="0" pageNumber="207">
|
||
Microsculpture: Head, pronotum and elytra with very finely impressed very small transverse meshes (visible at magnifications of> 80
|
||
<normalizedToken id="AF3C600AC59A4B84C01B997C3CAF777D" originalValue="×">x</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6E6BB49BDE94E3608EDBA2E3EBAE33E1" pageId="0" pageNumber="207">Measurements and proportions, given as mean (min-max values):</paragraph>
|
||
<paragraph id="C715D67E758523122EF651783F771078" pageId="0" pageNumber="207">Standardized body length 3.43 (3.01-3.77) mm.</paragraph>
|
||
<paragraph id="C2A16A46BD5D4EB9C36C74B17FB44139" pageId="0" pageNumber="207">PW/HW = 1.24 (1.21-1.28).</paragraph>
|
||
<paragraph id="84CCED39B06367FBF7DF71F2A3C7DFA3" pageId="0" pageNumber="207">PW/PL = 1.09 (1.02-1.13).</paragraph>
|
||
<paragraph id="4E073436EBCBD032E11DE6978B49F35E" pageId="0" pageNumber="207">PW/PWb = 1.39 (1.33-1.46).</paragraph>
|
||
<paragraph id="94221ACF07F26655147101AE93B1F9A8" pageId="0" pageNumber="207">PWb/PWa = 1.03 (0.97-1.08).</paragraph>
|
||
<paragraph id="AF7814854D32F028DDF338FE5D5DDA2B" pageId="0" pageNumber="207">EW/PW = 1.66 (1.60-1.77).</paragraph>
|
||
<paragraph id="A24BDFCB35E7E8C975C9B89267A89C05" pageId="0" pageNumber="207">EL/EW = 1.55 (1.50-1.59).</paragraph>
|
||
<paragraph id="B64F3398EE742A1B6E7DD94BB005FD1D" pageId="0" pageNumber="207">EL/FL = 2.18 (2.13-2.21).</paragraph>
|
||
<paragraph id="6ABBC1BF8C6169AAF8BD40264C4A84F0" pageId="0" pageNumber="207">EL/AedL = 2.36, 2.63 (paratypes 1, 2).</paragraph>
|
||
<paragraph id="734B514705D22CFA033205350CA6CB88" pageId="0" pageNumber="207">
|
||
For individual measurements and proportions see Tables
|
||
<tableCitation id="56FF05780B02AC65AF4C668E8DE97442" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements [µm]." httpUri="http://table.plazi.org/id/F51158147A1515BB565B411237896E8D" pageId="0" pageNumber="207" tableUuid="F51158147A1515BB565B411237896E8D">2</tableCitation>
|
||
,
|
||
<tableCitation id="939F1F63C37C16F30E26189BC91ECB85" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Body length and proportions." httpUri="http://table.plazi.org/id/DA6DEFEE7654ACB23A8DB426DB526F5F" pageId="0" pageNumber="207" tableUuid="DA6DEFEE7654ACB23A8DB426DB526F5F">3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0938A90D193BCCDF9AE63A9019980DA6" pageId="0" pageNumber="207" type="derivation of species epithet">
|
||
<paragraph id="D720828EABEB71A4723152FFE6F51C91" pageId="0" pageNumber="207">Derivation of species epithet.</paragraph>
|
||
<paragraph id="C23E3AC1BEEE29CCD8A67DB89CBCD0EE" pageId="0" pageNumber="207">The species epithet is given in honor of Friedrich Kernegger, Hamburg, who found a piece of amber bearing a fossilized specimen of this extraordinarily interesting species more than 25 years ago.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |