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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.205.76790" ID-Pensoft-Pub="1314-2003-205-99" ID-Pensoft-UUID="4024A478048757B197E5CF8734331A9E" ModsDocID="1314-2003-205-99" checkinTime="1661230211668" checkinUser="pensoft" docAuthor="O'Donnell, Shawn A., Ringelberg, Jens J. &amp; Lewis, Gwilym P." docDate="2022" docId="A06F1033675A535D8FB25D9AFA0431E2" docLanguage="en" docName="PhytoKeys 205: 99-145" docOrigin="PhytoKeys 205" docPubDate="2022-08-22" docSource="http://dx.doi.org/10.3897/phytokeys.205.76790" docTitle="Entada louvelii (R. Vig.) Brenan, Kew Bull. 20: 365. 1966." docType="treatment" docVersion="1" id="4024A478048757B197E5CF8734331A9E" lastPageNumber="99" masterDocId="4024A478048757B197E5CF8734331A9E" masterDocTitle="Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade)" masterLastPageNumber="145" masterPageNumber="99" pageNumber="99" updateTime="1661230211668" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>O'Donnell, Shawn A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0731-7425</mods:nameIdentifier>
<mods:affiliation>Department of Geography and Environmental Sciences, Northumbria University, Newcastle upon Tyne, NE 1 8 ST, UK</mods:affiliation>
<mods:nameIdentifier type="email">shawn.odonnell@cantab.net</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ringelberg, Jens J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0567-5210</mods:nameIdentifier>
<mods:affiliation>Department of Systematic and Evolutionary Botany, University of Zurich, 8008 Zurich, Switzerland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lewis, Gwilym P.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2599-4577</mods:nameIdentifier>
<mods:affiliation>Accelerated Taxonomy Department, Royal Botanic Gardens, Kew, Richmond, TW 9 3 AE, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>205</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>99</mods:start>
<mods:end>145</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.205.76790</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.205.76790</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-205-99</mods:identifier>
<mods:identifier type="Pensoft-UUID">4024A478048757B197E5CF8734331A9E</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:A06F1033675A535D8FB25D9AFA0431E2" httpUri="http://treatment.plazi.org/id/A06F1033675A535D8FB25D9AFA0431E2" lastPageNumber="99" pageId="0" pageNumber="99">
<subSubSection pageId="0" pageNumber="99" type="nomenclature">
<paragraph pageId="0" pageNumber="99">
<taxonomicName LSID="A06F1033-675A-535D-8FB2-5D9AFA0431E2" authority="(R. Vig.) Brenan, Kew Bull. 20: 365. 1966." authorityName="(R. Vig.) Brenan, Kew Bull. 20: 365. 1966." class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada louvelii" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="louvelii">Entada louvelii (R. Vig.) Brenan, Kew Bull. 20: 365. 1966.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="materials_examined">
<paragraph pageId="0" pageNumber="99">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Moramanga, M, J. - F. Villiers" country="MADAGASCAR" location="Moramanga" specimenCount="1" typeStatus="lectotype">
<collectingCountry name="Madagascar">MADAGASCAR</collectingCountry>
. Analamazoatra, south of
<collectorName>
<location LSID="urn:lsid:plazi:treatment:A06F1033675A535D8FB25D9AFA0431E2:3F8B5B5FB50D3061C730C57C64E94C1A" country="MADAGASCAR" name="Moramanga">Moramanga</location>
,
<emphasis italics="true" pageId="0" pageNumber="99">M. Louvel 16</emphasis>
</collectorName>
(
<typeStatus>lectotype</typeStatus>
: P [P00452896], designated by
<collectorName>J.-F. Villiers</collectorName>
in
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="family">Leguminosae</taxonomicName>
of Madagascar, 2002: 165)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="basionym">
<paragraph pageId="0" pageNumber="99">Basionym.</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada pervillei" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="pervillei">Entada pervillei</taxonomicName>
(Vatke) R. Vig. var.
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada pervillei var. louvelii" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="pervillei" variety="louvelii">Entada pervillei louvelii</taxonomicName>
R. Vig., Notul. Syst. (Paris) 13: 347. 1949.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="description">
<paragraph pageId="0" pageNumber="99">Description.</paragraph>
<paragraph pageId="0" pageNumber="99">
Tree 10-15 m tall, with elevated nectaries at nodes (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Putative extrafloral nectaries on herbarium specimens of five of the six native Madagascan species of Entada A E. leptostachya, Du Puy et al. M 235 B E. louvelii, Reserves Naturelles RN 1447 C E. pervillei, Service Forestier 10525 _ SF D E. pervillei, Service Forestier 11481 _ SF E E. rheedei, Gautier LG 3153 F E. tuberosa, Jongkind et al. 3264. Scale bars: 2 mm (A-F). Photos: S O'Donnell." figureDoi="10.3897/phytokeys.205.76790.figure4" httpUri="https://binary.pensoft.net/fig/732699" pageId="0" pageNumber="99">4B</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Leaves</emphasis>
: petiole 2-4 cm long, grooved above; rachis 9-18 cm long, winged, no tendril; pinnae 11-20 pairs per leaf, 3-9 cm long, with 24-46 pairs of leaflets; leaflets 3-7
<normalizedToken originalValue="×">x</normalizedToken>
1-1.75 mm, oblong to oblong-elliptic, apex rounded-obtuse to sub-acute and mucronate, base asymmetric and sub-truncate; lamina glabrous (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Entada louvelii vegetative and reproductive structures. A branch and shoots bearing leaves and terminal panicles of spikes, Madagascar (photo: P Antilahimena CC BY-NC-ND 3.0, MBG (2021)) B terminal panicle of flower spikes, Madagascar (photo: D Du Puy) C spikes bearing open, sessile flowers and closed flower buds, Madagascar (photo: D Du Puy) D leaves and fruits nearing maturity, Madagascar (photo: D Du Puy)." figureDoi="10.3897/phytokeys.205.76790.figure16" httpUri="https://binary.pensoft.net/fig/732711" pageId="0" pageNumber="99">16A, B, D</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Inflorescence</emphasis>
: a terminal panicle of spikes, each spike 5-19 cm long, rachis pubescent (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Entada louvelii vegetative and reproductive structures. A branch and shoots bearing leaves and terminal panicles of spikes, Madagascar (photo: P Antilahimena CC BY-NC-ND 3.0, MBG (2021)) B terminal panicle of flower spikes, Madagascar (photo: D Du Puy) C spikes bearing open, sessile flowers and closed flower buds, Madagascar (photo: D Du Puy) D leaves and fruits nearing maturity, Madagascar (photo: D Du Puy)." figureDoi="10.3897/phytokeys.205.76790.figure16" httpUri="https://binary.pensoft.net/fig/732711" pageId="0" pageNumber="99">16A, B</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Flowers</emphasis>
: white, 4-5 mm long, sessile to sub-sessile; calyx cream-coloured, obconical, 1.5-2 mm long, shallowly toothed, glabrous; petals 3.5-4 mm long; stamen filaments 5-7.25 mm long (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Entada louvelii vegetative and reproductive structures. A branch and shoots bearing leaves and terminal panicles of spikes, Madagascar (photo: P Antilahimena CC BY-NC-ND 3.0, MBG (2021)) B terminal panicle of flower spikes, Madagascar (photo: D Du Puy) C spikes bearing open, sessile flowers and closed flower buds, Madagascar (photo: D Du Puy) D leaves and fruits nearing maturity, Madagascar (photo: D Du Puy)." figureDoi="10.3897/phytokeys.205.76790.figure16" httpUri="https://binary.pensoft.net/fig/732711" pageId="0" pageNumber="99">16C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Fruit</emphasis>
: a torulose, laterally compressed craspedium, 15-20
<normalizedToken originalValue="×">x</normalizedToken>
3-6.5 cm, with transverse septa between seeds dividing the fruit into one-seeded segments which, upon ripening, fall from the persistent replum (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Entada louvelii vegetative and reproductive structures. A branch and shoots bearing leaves and terminal panicles of spikes, Madagascar (photo: P Antilahimena CC BY-NC-ND 3.0, MBG (2021)) B terminal panicle of flower spikes, Madagascar (photo: D Du Puy) C spikes bearing open, sessile flowers and closed flower buds, Madagascar (photo: D Du Puy) D leaves and fruits nearing maturity, Madagascar (photo: D Du Puy)." figureDoi="10.3897/phytokeys.205.76790.figure16" httpUri="https://binary.pensoft.net/fig/732711" pageId="0" pageNumber="99">16D</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Seeds</emphasis>
: elliptic, 1.5-2.4
<normalizedToken originalValue="×">x</normalizedToken>
0.6-1.2 cm, light brown, pleurogram lacking.
</paragraph>
<caption doi="10.3897/phytokeys.205.76790.figure16" httpUri="https://binary.pensoft.net/fig/732711" pageId="0" pageNumber="99" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Figure 16.</emphasis>
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada louvelii" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="louvelii">
<emphasis italics="true" pageId="0" pageNumber="99">Entada louvelii</emphasis>
</taxonomicName>
vegetative and reproductive structures.
<emphasis bold="true" pageId="0" pageNumber="99">A</emphasis>
branch and shoots bearing leaves and terminal panicles of spikes, Madagascar (photo: P Antilahimena
<emphasis italics="true" pageId="0" pageNumber="99">CC BY-NC-ND 3.0</emphasis>
,
<bibRefCitation author="Maurin, O" journalOrPublisher="The New Phytologist" pageId="0" pageNumber="99" publicationUrl="Tropicos.org" refId="B37" refString="2021. . Tropicos.org" url="Tropicos.org" year="2021">MBG (2021)</bibRefCitation>
)
<emphasis bold="true" pageId="0" pageNumber="99">B</emphasis>
terminal panicle of flower spikes, Madagascar (photo: D Du Puy)
<emphasis bold="true" pageId="0" pageNumber="99">C</emphasis>
spikes bearing open, sessile flowers and closed flower buds, Madagascar (photo: D Du Puy)
<emphasis bold="true" pageId="0" pageNumber="99">D</emphasis>
leaves and fruits nearing maturity, Madagascar (photo: D Du Puy).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="distribution">
<paragraph pageId="0" pageNumber="99">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="99">Madagascar (east).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="habitat">
<paragraph pageId="0" pageNumber="99">Habitat and ecology.</paragraph>
<paragraph pageId="0" pageNumber="99">
Moist forest, up to 1000 m alt. (
<bibRefCitation author="Villiers, J-F" editor="Du Puy, DJ" journalOrPublisher="Kew Publishing, Richmond" pageId="0" pageNumber="99" pagination="163 - 169" refId="B61" refString="Villiers, J-F, 2002. 3. Entada. In: Du Puy, DJ, Ed., The Leguminosae of Madagascar. Kew Publishing, Richmond: 163 - 169" title="3. Entada." volumeTitle="The Leguminosae of Madagascar." year="2002">Villiers 2002</bibRefCitation>
, p. 167); disturbed or dry forest (
<bibRefCitation author="Lungu, S" journalOrPublisher="Kew Publishing, Richmond" pageId="0" pageNumber="99" refId="B34" refString="Lungu, S, 1995. Systematics and economic botany of the genus Entada Adans. PhD Thesis, University of Reading, United Kingdom." title="Systematics and economic botany of the genus Entada Adans. PhD Thesis, University of Reading, United Kingdom." year="1995">Lungu 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>