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<mods:title id="0E0D7D88D2E764FCE29DC60736667A98">Review of the Chinese Stomis, with description of nine new taxa and a new key (Carabidae: Pterostichini)</mods:title>
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<mods:date id="CEA5EC758B8C036A1EBE0FC155ADBADE">2024</mods:date>
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<treatment id="9D4DF639FFFDF973A5BBDECBA4B2AD6C" ID-DOI="http://doi.org/10.5281/zenodo.13933886" ID-Zenodo-Dep="13933886" LSID="urn:lsid:zoobank.org:act:E873CCD5-D763-43E2-8933-60AC0AE9D838" httpUri="http://treatment.plazi.org/id/9D4DF639FFFDF973A5BBDECBA4B2AD6C" lastPageNumber="9" pageId="8" pageNumber="9">
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<taxonomicName id="D2E43CACFFFDF973A5BBDECBA4EFA8A4" authorityName="Sciaky" authorityYear="2024" box="[151,427,151,178]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="exilis" status="subsp. nov." subSpecies="pauli">
<emphasis id="27909B3DFFFDF973A5BBDECBA474A8A4" bold="true" box="[151,304,151,178]" italics="true" pageId="8" pageNumber="9">Stomis exilis</emphasis>
ssp.
<emphasis id="27909B3DFFFDF973A441DECBA4EFA8A4" bold="true" box="[365,427,151,178]" italics="true" pageId="8" pageNumber="9">pauli</emphasis>
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<taxonomicNameLabel id="3CA32646FFFDF973A49EDEC4A75FA8A4" box="[434,539,152,178]" pageId="8" pageNumber="9" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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<paragraph id="155B472FFFFDF973A5BBDEE7A614A8C0" blockId="8.[151,848,151,214]" box="[151,848,187,214]" pageId="8" pageNumber="9">https://zoobank.org/e873ccd5-d763-43e2-8933-60ac0ae9d838</paragraph>
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<subSubSection id="5DFE14A4FFFDF973A5BBDF5FA453A9C4" pageId="8" pageNumber="9" type="diagnosis">
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<emphasis id="27909B3DFFFDF973A5BBDF5FA44EA90B" bold="true" box="[151,266,259,285]" pageId="8" pageNumber="9">Diagnosis</emphasis>
. A
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<emphasis id="27909B3DFFFDF973A41EDF58A439A908" box="[306,381,260,286]" italics="true" pageId="8" pageNumber="9">Stomis</emphasis>
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of medium length (
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), black with evident bronze or green hue on head, pronotum and elytra, with head narrow, but with eyes rather convex; pronotum with maximum width at the level of anterior third, margins almost rectilinearly constricted towards hind angles; elytra without shoulder denticle, markedly elongate and moderately flat on disc, with striae rather shallow; median lobe of aedeagus in lateral view narrow and very elongate, with inferior margin sinuate; in dorsal view with apical lamella thicker than in the two other subspecies.
</paragraph>
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<subSubSection id="5DFE14A4FFFDF973A5EBDF87A1EEACFC" pageId="8" pageNumber="9" type="description">
<paragraph id="155B472FFFFDF973A5EBDF87A0D2ABDF" blockId="8.[151,1437,259,1402]" pageId="8" pageNumber="9">
<emphasis id="27909B3DFFFDF973A5EBDF87A41DA9E3" bold="true" box="[199,345,475,501]" pageId="8" pageNumber="9">Description.</emphasis>
Habitus as in
<figureCitation id="8DDF5BAAFFFDF973A4DDDF80A770A9E0" box="[497,564,476,502]" captionStart="FIGURES 912" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[335,1243,181,1915]" captionTargetId="figure-18@20.[323,1263,181,1919]" captionTargetPageId="20" captionText="FIGURES 912. Habitus of: (9) S. exilis ssp. pauli HT (10.9 mm); (10) S. collucens from Yunnan, Yulongshan (11.5 mm); (11) S. politus from Yunnan, Zhongdian (11.9 mm); (12) S. habashanensis from Yunnan, Habashan (14.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13933854" httpUri="https://zenodo.org/record/13933854/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
. Length
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; body black with bronze or green hue on elytra; antennae, tibiae and tarsi dark reddish-brown, femora black; mandibles dark reddish-brown, other buccal parts reddish-brown; head smooth, with evident but feeble ocular convexity; mandibles very long, longer than rest of the head, the right one without notch; eyes rather small, not protruding from ocular convexity. Pronotum narrow, longer than wide, widest at the anterior fourth; lateral margins rectilinearly constricted towards base, only with a hint of sinuation just before base; anterior margin feebly advanced at middle; anterior angles scarcely distinct; base rectilinear at middle, advanced forward at sides; all basal surface with scattered punctuation; basal foveae rather narrow, starting from the basal angles, longer than one third of the pronotal length; basal angles almost completely rounded, scarcely distinguishable; basal seta not in the basal angle but moved forward; lateral gutter narrow all along its length. Elytra elongate oval, rather convex, widest almost at middle, with evident humeral tooth; striae feebly but evidently punctate, well distinct until apex, although more superficial in the apical fourth; intervals very slightly convex, humeri very rounded in shape; lateral gutter well marked and rather wide, mainly in anterior and posterior fourth. Median lobe of aedeagus in lateral view narrow and very elongate, with inferior margin sinuate; in dorsal view with apical lamella thicker than in the two other subspecies (
<figureCitation id="8DDF5BAAFFFDF973A7D3DDECA616ABDF" box="[767,850,943,970]" captionStart="FIGURES 3236" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[199,1389,187,1913]" captionTargetId="figure-18@26.[193,1395,181,1919]" captionTargetPageId="26" captionText="FIGURES 3236. Aedeagus in lateral and dorsal view of: (32) S. deuvei ssp. deuvei from Sichuan, pass Zhangla-Nanping; (33) S. romani from Sichuan, mts. 15 km W Keluodong; (34) S. exilis ssp. exilis from Shaanxi, Taibai Shan; (35) S. exilis ssp. brivioi from (36) S. exilis ssp. pauli HT." figureDoi="http://doi.org/10.5281/zenodo.13933870" httpUri="https://zenodo.org/record/13933870/files/figure.png" pageId="8" pageNumber="9">Fig. 36</figureCitation>
). Ratios: El/Pl = 2.17; Ew/Pw = 1.35; El/Ew = 1.61.
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<emphasis id="27909B3DFFFDF973A5EBDD8FA426ABFB" bold="true" box="[199,354,979,1005]" pageId="8" pageNumber="9">Type-locality</emphasis>
:
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W
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,
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,
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.
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<emphasis id="27909B3DFFFDF973A5EBDDABA40AAC07" bold="true" box="[199,334,1015,1041]" pageId="8" pageNumber="9">Type-series</emphasis>
.
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,
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leg. (
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.2010,
<collectorName id="B81122F9FFFDF973A722DA1CA7F5AC4C" box="[526,689,1088,1114]" pageId="8" pageNumber="9">Jaroslav Turna</collectorName>
leg.:
<specimenCount id="03E28CA6FFFDF973A7C1DA1CA651AC4C" box="[749,789,1088,1114]" count="4" pageId="8" pageNumber="9" type="male">4 ♂</specimenCount>
(
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); W
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<collectingRegion id="D72089CDFFFDF973A6A9DA1CA695AC4C" box="[901,977,1088,1114]" country="China" name="Henan" pageId="8" pageNumber="9">Henan</collectingRegion>
,
<location id="103B11F4FFFDF973A6F7DA1CA119AC4C" LSID="urn:lsid:plazi:treatment:9D4DF639FFFDF973A5BBDECBA4B2AD6C:103B11F4FFFDF973A6F7DA1CA119AC4C" box="[987,1117,1088,1114]" latitude="33.516666" longLatPrecision="1204" longitude="111.933334" name="Funiu Shan" pageId="8" pageNumber="9" stateProvince="Henan">Funiu Shan</location>
, Baotianman,
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<geoCoordinate id="70D021E8FFFDF973A045DA1CA590AC68" degrees="111" direction="east" minutes="56" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="111.933334">111° 56 E</geoCoordinate>
,
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<elevation id="9EC9A01CFFFDF973A5F3DA3FA438AC68" box="[223,380,1123,1150]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.75" metricValueMin="1.5" pageId="8" pageNumber="9" unit="m" value="1625.0" valueMax="1750.0" valueMin="1500.0">15001750 m</elevation>
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,
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10.
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</date>
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<materialsCitation id="A58C4D72FFFDF973A490DA3FA688AC68" box="[444,972,1123,1150]" collectionCode="CM" collectorName="Jaroslav Turna" pageId="8" pageNumber="9" specimenCount="2" specimenCount-female="2" stateProvince="Henan" typeStatus="paratype">
6.VII.2006,
<collectorName id="B81122F9FFFDF973A779DA38A7BCAC68" box="[597,760,1124,1150]" pageId="8" pageNumber="9">Jaroslav Turna</collectorName>
leg.:
<specimenCount id="03E28CA6FFFDF973A61ADA38A624AC68" box="[822,864,1124,1150]" count="2" pageId="8" pageNumber="9" type="female">2 ♀</specimenCount>
(
<collectionCode id="73F5DFEAFFFDF973A643DA38A6E4AC68" box="[879,928,1124,1150]" country="China" name="Chongqing Museum" pageId="8" pageNumber="9">CM</collectionCode>
); W
</materialsCitation>
<materialsCitation id="A58C4D72FFFDF973A6FFDA38A75CACB4" collectionCode="V" elevation="1625" latitude="33.516666" location="Funiu Shan" longLatPrecision="1204" longitude="111.933334" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Henan" typeStatus="paratype">
<collectingRegion id="D72089CDFFFDF973A6FFDA38A15BAC68" box="[979,1055,1124,1150]" country="China" name="Henan" pageId="8" pageNumber="9">Henan</collectingRegion>
,
<location id="103B11F4FFFDF973A105DA38A1E8AC68" LSID="urn:lsid:plazi:treatment:9D4DF639FFFDF973A5BBDECBA4B2AD6C:103B11F4FFFDF973A105DA38A1E8AC68" box="[1065,1196,1124,1150]" latitude="33.516666" longLatPrecision="1204" longitude="111.933334" name="Funiu Shan" pageId="8" pageNumber="9" stateProvince="Henan">Funiu Shan</location>
, Baotianman,
<geoCoordinate id="70D021E8FFFDF973A067DA38A5EFACB4" degrees="33" direction="north" minutes="31" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="33.516666">33° 31 N</geoCoordinate>
<geoCoordinate id="70D021E8FFFDF973A59FDAD4A468ACB4" box="[179,300,1159,1186]" degrees="111" direction="east" minutes="56" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="111.933334">111° 56 E</geoCoordinate>
,
<quantity id="D21CEACAFFFDF973A415DADBA493ACB4" box="[313,471,1159,1186]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.75" metricValueMin="1.5" pageId="8" pageNumber="9" unit="m" value="1625.0" valueMax="1750.0" valueMin="1500.0">
<elevation id="9EC9A01CFFFDF973A415DADBA493ACB4" box="[313,471,1159,1186]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.75" metricValueMin="1.5" pageId="8" pageNumber="9" unit="m" value="1625.0" valueMax="1750.0" valueMin="1500.0">15001750 m</elevation>
</quantity>
,
<date id="615A61EFFFFDF973A4C8DADBA75CACB4" box="[484,536,1159,1186]" pageId="8" pageNumber="9" value="2009-05-15" valueMax="2009-06-05" valueMin="2009-05-15">
15.
<collectionCode id="73F5DFEAFFFDF973A72EDADBA75CACB4" box="[514,536,1159,1186]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="9" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="A58C4D72FFFDF973A734DADBA71FACB4" box="[536,603,1159,1186]" collectionCode="VI" elevation="1625" isTailEnumeration="true" latitude="33.516666" location="Funiu Shan" longLatPrecision="1204" longitude="111.933334" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Henan" typeStatus="paratype">
5.
<collectionCode id="73F5DFEAFFFDF973A716DADBA71FACB4" box="[570,603,1159,1186]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="9">VI</collectionCode>
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<materialsCitation id="A58C4D72FFFDF973A776DADBA16DACB4" box="[602,1065,1159,1186]" collectionCode="CM" collectorName="Jaroslav Turna" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Henan" typeStatus="paratype">
.2009,
<collectorName id="B81122F9FFFDF973A785DAD4A60AACB4" box="[681,846,1160,1186]" pageId="8" pageNumber="9">Jaroslav Turna</collectorName>
leg.:
<specimenCount id="03E28CA6FFFDF973A6A3DAD4A6FEACB4" box="[911,954,1160,1186]" count="1" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
(
<collectionCode id="73F5DFEAFFFDF973A6E6DAD4A6BFACB4" box="[970,1019,1160,1186]" country="China" name="Chongqing Museum" pageId="8" pageNumber="9">CM</collectionCode>
); W
</materialsCitation>
<materialsCitation id="A58C4D72FFFDF973A11DDAD4A75BACD3" collectionCode="V" elevation="2000" latitude="33.7" location="Funiu Shan" longLatPrecision="1204" longitude="112.25" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Henan" typeStatus="paratype">
<collectingRegion id="D72089CDFFFDF973A11DDAD4A139ACB4" box="[1073,1149,1160,1186]" country="China" name="Henan" pageId="8" pageNumber="9">Henan</collectingRegion>
,
<location id="103B11F4FFFDF973A1A5DAD4A049ACB4" LSID="urn:lsid:plazi:treatment:9D4DF639FFFDF973A5BBDECBA4B2AD6C:103B11F4FFFDF973A1A5DAD4A049ACB4" box="[1161,1293,1160,1186]" latitude="33.7" longLatPrecision="1204" longitude="112.25" name="Funiu Shan" pageId="8" pageNumber="9" stateProvince="Henan">Funiu Shan</location>
, Shirenshan,
<geoCoordinate id="70D021E8FFFDF973A5BBDAF0A441ACD0" box="[151,261,1196,1222]" degrees="33" direction="north" minutes="42" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="33.7">33° 42 N</geoCoordinate>
<geoCoordinate id="70D021E8FFFDF973A421DAF0A4C3ACD0" box="[269,391,1195,1222]" degrees="112" direction="east" minutes="15" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="112.25">112° 15 E</geoCoordinate>
,
<quantity id="D21CEACAFFFDF973A4BFDAF0A4AFACD0" box="[403,491,1196,1222]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="m" value="2000.0">
<elevation id="9EC9A01CFFFDF973A4BFDAF0A4AFACD0" box="[403,491,1196,1222]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="m" value="2000.0">2000 m</elevation>
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,
<date id="615A61EFFFFDF973A4D4DAF7A75BACD3" box="[504,543,1195,1221]" pageId="8" pageNumber="9" value="2006-05-09" valueMax="2006-07-09" valueMin="2006-05-09">
9.
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</date>
</materialsCitation>
<materialsCitation id="A58C4D72FFFDF973A733DAF7A173ACD0" box="[543,1079,1195,1222]" collectionCode="CM" collectorName="Jaroslav Turna" pageId="8" pageNumber="9" specimenCount="2" specimenCount-female="2" stateProvince="Henan" typeStatus="paratype">
9.VII.2006,
<collectorName id="B81122F9FFFDF973A794DAF0A619ACD0" box="[696,861,1196,1222]" pageId="8" pageNumber="9">Jaroslav Turna</collectorName>
leg.:
<specimenCount id="03E28CA6FFFDF973A6B1DAF0A68CACD0" box="[925,968,1196,1222]" count="2" pageId="8" pageNumber="9" type="female">2 ♀</specimenCount>
(
<collectionCode id="73F5DFEAFFFDF973A6F4DAF0A14DACD0" box="[984,1033,1196,1222]" country="China" name="Chongqing Museum" pageId="8" pageNumber="9">CM</collectionCode>
); W
</materialsCitation>
<materialsCitation id="A58C4D72FFFDF973A113DAF0A7F0ACFC" collectionCode="V" elevation="1625" latitude="34.283333" location="Shenlingzhai" longLatPrecision="1200" longitude="111.71667" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Henan" typeStatus="paratype">
<collectingRegion id="D72089CDFFFDF973A113DAF0A1CFACD0" box="[1087,1163,1196,1222]" country="China" name="Henan" pageId="8" pageNumber="9">Henan</collectingRegion>
,
<location id="103B11F4FFFDF973A1BADAF0A06FACD0" LSID="urn:lsid:plazi:treatment:9D4DF639FFFDF973A5BBDECBA4B2AD6C:103B11F4FFFDF973A1BADAF0A06FACD0" box="[1174,1323,1196,1222]" latitude="34.283333" longLatPrecision="1200" longitude="111.71667" name="Shenlingzhai" pageId="8" pageNumber="9" stateProvince="Henan">Shenlingzhai</location>
, Xionger Shan,
<geoCoordinate id="70D021E8FFFDF973A5F1DA8CA40EACFC" box="[221,330,1232,1258]" degrees="34" direction="north" minutes="17" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="34.283333">34° 17 N</geoCoordinate>
<geoCoordinate id="70D021E8FFFDF973A47DDA8CA48CACFC" box="[337,456,1232,1258]" degrees="111" direction="east" minutes="43" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="111.71667">111° 43 E</geoCoordinate>
,
<quantity id="D21CEACAFFFDF973A4F8DA93A735ACFC" box="[468,625,1231,1258]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.75" metricValueMin="1.5" pageId="8" pageNumber="9" unit="m" value="1625.0" valueMax="1750.0" valueMin="1500.0">
<elevation id="9EC9A01CFFFDF973A4F8DA93A735ACFC" box="[468,625,1231,1258]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.75" metricValueMin="1.5" pageId="8" pageNumber="9" unit="m" value="1625.0" valueMax="1750.0" valueMin="1500.0">15001750 m</elevation>
</quantity>
,
<date id="615A61EFFFFDF973A751DA93A7F0ACFC" box="[637,692,1231,1258]" pageId="8" pageNumber="9" value="2010-05-25" valueMax="2010-06-17" valueMin="2010-05-25">
25.
<collectionCode id="73F5DFEAFFFDF973A7B2DA93A7F0ACFC" box="[670,692,1231,1258]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="9" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="A58C4D72FFFDF973A798DA93A646ACFC" box="[692,770,1231,1258]" collectionCode="VI" elevation="1625" isTailEnumeration="true" latitude="34.283333" location="Shenlingzhai" longLatPrecision="1200" longitude="111.71667" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Henan" typeStatus="paratype">
17.
<collectionCode id="73F5DFEAFFFDF973A7CEDA93A646ACFC" box="[738,770,1231,1258]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="9">VI</collectionCode>
</materialsCitation>
<materialsCitation id="A58C4D72FFFDF973A62EDA93A1E1ACFC" box="[770,1189,1231,1258]" collectionCode="CM" collectorName="Jaroslav Turna" pageId="8" pageNumber="9" specimenCount="2" specimenCount-female="2" stateProvince="Henan" typeStatus="paratype">
.2010,
<collectorName id="B81122F9FFFDF973A67CDA8CA6B0ACFC" box="[848,1012,1232,1258]" pageId="8" pageNumber="9">Jaroslav Turna</collectorName>
leg.:
<specimenCount id="03E28CA6FFFDF973A11FDA8CA119ACFC" box="[1075,1117,1232,1258]" count="2" pageId="8" pageNumber="9" type="female">2 ♀</specimenCount>
(
<collectionCode id="73F5DFEAFFFDF973A140DA8CA1D9ACFC" box="[1132,1181,1232,1258]" country="China" name="Chongqing Museum" pageId="8" pageNumber="9">CM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5DFE14A4FFFDF973A5EBDAA8A7F7AD24" pageId="8" pageNumber="9" type="etymology">
<paragraph id="155B472FFFFDF973A5EBDAA8A7F7AD24" blockId="8.[151,1437,259,1402]" pageId="8" pageNumber="9">
<emphasis id="27909B3DFFFDF973A5EBDAA8A40CAD18" bold="true" box="[199,328,1268,1294]" pageId="8" pageNumber="9">Etymology</emphasis>
. This specific epithet is from the Latinized first name of my friend Paolo Magrini, an excellent specialist of endogeous coleopterological fauna.
</paragraph>
</subSubSection>
<subSubSection id="5DFE14A4FFFDF973A5EBDB67A4B2AD6C" pageId="8" pageNumber="9" type="distribution">
<paragraph id="155B472FFFFDF973A5EBDB67A4B2AD6C" blockId="8.[151,1437,259,1402]" pageId="8" pageNumber="9">
<emphasis id="27909B3DFFFDF973A5EBDB67A41DAD43" bold="true" box="[199,345,1339,1365]" pageId="8" pageNumber="9">Distribution</emphasis>
. Known from few localities in western
<collectingRegion id="D72089CDFFFDF973A60CDB60A629AD40" box="[800,877,1340,1366]" country="China" name="Henan" pageId="8" pageNumber="9">Henan</collectingRegion>
; until now it is the easternmost species of
<taxonomicName id="D2E43CACFFFDF973A07EDB60A0D9AD40" authorityName="Clairville" authorityYear="1806" box="[1362,1437,1340,1366]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="27909B3DFFFDF973A07EDB60A0D9AD40" box="[1362,1437,1340,1366]" italics="true" pageId="8" pageNumber="9">Stomis</emphasis>
</taxonomicName>
known from continental
<collectingCountry id="6DF307BFFFFDF973A487DB3CA4B6AD6C" box="[427,498,1376,1402]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>