408 lines
45 KiB
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408 lines
45 KiB
XML
<document ID-CLB-Dataset="51924" ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-GBIF-Dataset="c2e00414-9633-4890-ab72-cb2aa3e9d174" ID-PMC="PMC10208988" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-PubMed="37250203" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak & Sutcharit, Chirasak" docDate="2023" docId="9B03C93A9D3C54BD9C53245AE285A411" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Rhiostoma tigrina Tongkerd & Tumpeesuwan 2023, sp. nov." docType="treatment" docUuid="94D81E56-7DAB-469E-B93E-0C18C59B7999" docUuidSource="ZooBank" docVersion="5" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1732652259105" updateUser="ExternalLinkService">
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<mods:titleInfo id="DA18C7447708E063201991EA76786328">
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<mods:title id="B5C194AEB6EA2E51E2846F8E0F725BE5">Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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<mods:namePart id="97BB0AA044E1B3A3EB7EDB02DFD2F9A4">Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation id="00F0813F40BBEE78285158E76D4DAA3D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="0DCAFE4E21A2A7CEC579B6D7DC705636" type="personal">
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<mods:roleTerm id="31220567B9932872976D64944E41E846">Author</mods:roleTerm>
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<mods:namePart id="40DCA1F34C6CF180F045DE79D34EA0BB">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="CE514BFFA58CCC7A013C58B07F61DC5E">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="887BDBCC05EB44D35F4B803C78CC6F06" type="personal">
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<mods:namePart id="C7AB81B60A35549DB6B6322971F92834">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="72A96E2573E19233E3349A28BF4DDD8F">Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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</mods:name>
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<mods:name id="248B6B640CE1E1C81B648E0D98B14AFB" type="personal">
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<mods:roleTerm id="136DE3094868F070690CE1050AFA417B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7AA90E9DDC1B3C30F67594D1CB070426">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="A6B755234A7898B1B01C2A1AF33D3F0F">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="92DD56B708311C0A1D30CD3EC6A08B22" type="personal">
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<mods:namePart id="48942BAD87D3E365846191B6614137CF">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation id="72D5D204357A7086881E0C88ABE8269C">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="FCF5A1819677D7271416150BA797840F" type="personal">
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<mods:namePart id="997C9BA3F587ABF9DA6D17AA8545F7D6">Panha, Somsak</mods:namePart>
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<mods:affiliation id="0A4964CEC66848DB6F175895AD988725">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="D5C4BE21F2884618A3DC3E8A1C10033F" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:namePart id="E7A57D83C27EF2EC7BBD42EADA0480DB">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="49989DF34F48B5EF4B72130ADF94600D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="AF3B84DAC1A27271C57E211EC4EDA9CF" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:typeOfResource id="75319513831691EC51DB337A5EBF4704">text</mods:typeOfResource>
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<mods:title id="C0171A93595EA71A3A27F4F85259DCFE">ZooKeys</mods:title>
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<mods:date id="83F520FB0D779DAA3A9A5F9A1D03B037">2023</mods:date>
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<mods:number id="6F9DD84186D15C02B5B41F64B2EA0D0C">2023-01-24</mods:number>
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<mods:number id="2BD89E50A9FC6B3B23287F466CBFC865">1142</mods:number>
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<mods:start id="150A5172227257905FEBADD239947F52">1</mods:start>
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<mods:end id="953CC2E9EF6AE51EB3E867D938A8DF80">144</mods:end>
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<mods:url id="B2ACAD48C4FA1DABA018DFFE59C43AE5">http://dx.doi.org/10.3897/zookeys.1142.90097</mods:url>
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<mods:classification id="7E0AAFD990E9EF501B5959EDF7470AC2">journal article</mods:classification>
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<mods:identifier id="B12415ADBCE3F9691FF6E068CBD40186" type="DOI">http://dx.doi.org/10.3897/zookeys.1142.90097</mods:identifier>
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<mods:identifier id="DBCBD5F483AF46D82A640F1AB530F608" type="Pensoft-Pub">1313-2970-1142-1</mods:identifier>
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<mods:identifier id="CCC358A73B454B297C4342F28AAAF78A" type="ZooBank">A1129EE50F9941CFB73AE771B66E2486</mods:identifier>
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<treatment id="9B03C93A9D3C54BD9C53245AE285A411" ID-GBIF-Taxon="209102249" LSID="urn:lsid:zoobank.org:act:94D81E56-7DAB-469E-B93E-0C18C59B7999" httpUri="http://treatment.plazi.org/id/9B03C93A9D3C54BD9C53245AE285A411" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Cyclophoridae" scope_subClass="Caenogastropoda" scope_subFamily="Cyclophorinae">
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<subSubSection id="5FE36BA06B1D33655595F9754294F32F" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="86A256C9172BD7AD58BFE9084A6AF501" pageId="0" pageNumber="1">
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13.
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<taxonomicName id="786475728985A58763F27572200E32CC" ID-CoL="B3CKP" LSID="https://zoobank.org/94D81E56-7DAB-469E-B93E-0C18C59B7999" authority="Tongkerd & Tumpeesuwan" authorityName="Tongkerd & Tumpeesuwan" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma tigrina" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tigrina" status="sp. nov.">Rhiostoma tigrina Tongkerd & Tumpeesuwan</taxonomicName>
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<taxonomicNameLabel id="FA9C97F95BAF1184E43F0D9B6545F295" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="544C64FD50379862547EBD4CF3595D07" pageId="0" pageNumber="1" type="description">
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<paragraph id="56D1C6BA03FC9E4AABE908D18B8BB5E6" pageId="0" pageNumber="1">
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<figureCitation id="1919D985E551FDC9E9BBF729E81E0549" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">Figs 18</figureCitation>
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<figureCitation id="8D3D7496C448BF984BB9A157493D868A" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Shell of Rhiostoma tigrina sp. nov. A holotype CUMZ 4497 / 1 from Tham Pum Tham Pla, Mae Sai, Chiang Rai B paratype CUMZ 4497 / 2 from type locality C paratype CUMZ 4806 from type locality D specimen CUMZ 4795 from Tham Phra, Chiang Rai E, F specimens CUMZ 4807 Mae Suai, Chiang Rai" figureDoi="10.3897/zookeys.1142.90097.figure31" httpUri="https://binary.pensoft.net/fig/799889" pageId="0" pageNumber="1">, 31</figureCitation>
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<figureCitation id="A47AFAE645EBFDC3F1F7A538D9CF4DAC" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">, 32F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D143908B341285884F9FBB0BAFD9EF52" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="95D9AE14BAE06644204469515A813AF7" pageId="0" pageNumber="1">
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<taxonomicName id="09A9A305BADC2077A9D6B1AD893A3C47" authority="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit, 2023" authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus" status="sp. nov.">Rhiostoma</taxonomicName>
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sp. 1-
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<bibRefCitation id="A6559D7EE3805D8C5A98275EC3AA936F" author="Tumpeesuwan, S" journalOrPublisher="Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" pageId="0" pageNumber="1" refId="B175" refString="Tumpeesuwan, S, 2001. Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand. Master Thesis, Department of Biology, Faculty of Science, Chulalongkorn University" title="Taxonomy of snorkel snails genus Rhiostoma Benson, 1860 in Thailand." year="2001">Tumpeesuwan 2001</bibRefCitation>
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: 59-64, figs 4.19-4.21 (in part).
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</paragraph>
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</subSubSection>
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<subSubSection id="88E80BD30DA5CAFF880A085AE44CA792" pageId="0" pageNumber="1" type="type material">
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<paragraph id="45FE9D8C1684B8F30CC6EBE98A7952DD" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="043C7285FB32ADC343B9A25EB9377278" pageId="0" pageNumber="1">
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<emphasis id="2A131E83F67100F36BD4B59A05528DC5" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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CUMZ 4497/1 (cW 19.5 mm, cH 10.5 mm, dL 20.8 mm; Fig.
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<figureCitation id="A74214E9C37128F61CA70D38CF1044CC" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Shell of Rhiostoma tigrina sp. nov. A holotype CUMZ 4497 / 1 from Tham Pum Tham Pla, Mae Sai, Chiang Rai B paratype CUMZ 4497 / 2 from type locality C paratype CUMZ 4806 from type locality D specimen CUMZ 4795 from Tham Phra, Chiang Rai E, F specimens CUMZ 4807 Mae Suai, Chiang Rai" figureDoi="10.3897/zookeys.1142.90097.figure31" httpUri="https://binary.pensoft.net/fig/799889" pageId="0" pageNumber="1">31A</figureCitation>
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).
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<emphasis id="216341476FD20C54045D087B49FCCF90" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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CUMZ 3909 (29 adults + 10 juveniles), CUMZ 4497/2 (2 shells; Figs
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<figureCitation id="82CD906D2E1B701EFBD0EBA9BB2E8B91" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Shell of Rhiostoma tigrina sp. nov. A holotype CUMZ 4497 / 1 from Tham Pum Tham Pla, Mae Sai, Chiang Rai B paratype CUMZ 4497 / 2 from type locality C paratype CUMZ 4806 from type locality D specimen CUMZ 4795 from Tham Phra, Chiang Rai E, F specimens CUMZ 4807 Mae Suai, Chiang Rai" figureDoi="10.3897/zookeys.1142.90097.figure31" httpUri="https://binary.pensoft.net/fig/799889" pageId="0" pageNumber="1">31B</figureCitation>
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,
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<figureCitation id="736DC39FC9E94A39779A3AD15029254B" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32F</figureCitation>
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), CUMZ 4806 (13 adults + 5 juveniles; Fig.
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<figureCitation id="1703BD4AA86A83C3C1C7FE4E9C020281" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Shell of Rhiostoma tigrina sp. nov. A holotype CUMZ 4497 / 1 from Tham Pum Tham Pla, Mae Sai, Chiang Rai B paratype CUMZ 4497 / 2 from type locality C paratype CUMZ 4806 from type locality D specimen CUMZ 4795 from Tham Phra, Chiang Rai E, F specimens CUMZ 4807 Mae Suai, Chiang Rai" figureDoi="10.3897/zookeys.1142.90097.figure31" httpUri="https://binary.pensoft.net/fig/799889" pageId="0" pageNumber="1">31C</figureCitation>
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), CUMZ 10191 (6 adults + 4 juveniles), NHMUK 20220441 (5 shells), and SMF 368676 (5 shells). All paratypes are from the type locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="25A187AA320A92A8C6AF9940A7DF0EDF" pageId="0" pageNumber="1" type="type locality">
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<paragraph id="2B7EDED00AB26930B2B7934CACF16F6A" pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph id="90E6C451EEA01F33A76123F0917553D5" pageId="0" pageNumber="1">
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Tham Pum Tham Pla, Mae Sai District, Chiang Rai Province, Thailand (
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<geoCoordinate id="2016513AF8293036D35CB8667EAC30D9" degrees="20" direction="north" minutes="21" orientation="latitude" precision="1" seconds="00.4" value="20.350111">20°21'00.4"N</geoCoordinate>
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,
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<geoCoordinate id="1EFC5EF9F352B848514D98C3B9150703" degrees="99" direction="east" minutes="51" orientation="longitude" precision="1" seconds="25.2" value="99.857">99°51'25.2"E</geoCoordinate>
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). Limestone hill covered by dry deciduous forest.
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</paragraph>
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</subSubSection>
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<subSubSection id="72614DEA7E6926DCBD7E1CE0AB40FABD" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="89B5B4A9D36EB430A06D1CFDBED5B8B0" pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph id="2F0A1D1DE4027F009004A8D1559CBCB2" pageId="0" pageNumber="1">
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<materialsCitation id="D4AA34B9E122ADAA4B00E6ECC8655315" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Chiang Dao, B. Degerbol" country="Thailand" county="North Thailand" elevation="400" location="Muang" municipality="Tham Samta" specimenCount="1" stateProvince="Chiang Rai">
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<emphasis id="A1E2D1EB73566D9BDC6D33DFBBB9181F" bold="true" pageId="0" pageNumber="1">
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<collectingCountry id="01C42FA9F18E9004729E56C19B0DE9A5" name="Thailand">Thailand</collectingCountry>
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</emphasis>
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:
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<collectorName id="81EF9CFB349FDB9096251A49403383BF">Chiang Dao</collectorName>
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,
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<collectingCounty id="23D4D3D80A932013E173CA5D130023BA">North Thailand</collectingCounty>
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: ZMUC ex.
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<collectorName id="96639696A0E447CCEAC256A9F7C5E2A1">
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B.
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<normalizedToken id="088B105F48F869336BC6D703B12F0A6F" originalValue="Degerbøl">Degerbol</normalizedToken>
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</collectorName>
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collection (1 alcohol specimen at
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<quantity id="B07B6ACF262843BA5F007BDE8182C821" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
|
|
<elevation id="2E88013C08DDC69C548DADD7DB934A28" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</elevation>
|
|
</quantity>
|
|
, and 2 alcohol specimens at
|
|
<quantity id="679025B607976B313A3C7D37017CC282" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
|
|
<elevation id="32CEA8071424600FFDAB5D8FAB8F8DAF" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</elevation>
|
|
</quantity>
|
|
).
|
|
<collectingMunicipality id="41475A68D6727BFDA5713B7243DBF7FA">Tham Samta</collectingMunicipality>
|
|
,
|
|
<location id="054EF91F16D4EB1CC72C02DA5DB9B95C" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:054EF91F16D4EB1CC72C02DA5DB9B95C" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Muang" stateProvince="Chiang Rai">Muang</location>
|
|
,
|
|
<location id="9A66061390801BFE9F072840F3F49F3C" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:9A66061390801BFE9F072840F3F49F3C" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Maehongsorn" stateProvince="Chiang Rai">Maehongsorn</location>
|
|
: CUMZ 4441.
|
|
<location id="5C542BB9FEB1D9FB8D816B63D488F11B" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:5C542BB9FEB1D9FB8D816B63D488F11B" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Wat Pa Tham Wua" stateProvince="Chiang Rai">Wat Pa Tham Wua</location>
|
|
,
|
|
<location id="6F9D479D8C8A742EFA7E1BE516556F65" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:6F9D479D8C8A742EFA7E1BE516556F65" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Pang Mapha" stateProvince="Chiang Rai">Pang Mapha</location>
|
|
,
|
|
<location id="AD88526579E12A04DB4592A62600E818" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:AD88526579E12A04DB4592A62600E818" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Maehongsorn" stateProvince="Chiang Rai">Maehongsorn</location>
|
|
: CUMZ 10028.
|
|
<location id="CBE57A5F2FB494531C70E6552ED5C59B" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:CBE57A5F2FB494531C70E6552ED5C59B" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Tham Luang" stateProvince="Chiang Rai">Tham Luang</location>
|
|
,
|
|
<location id="B6212B66610414F1FF18DA55D9246D2D" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:B6212B66610414F1FF18DA55D9246D2D" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Khun Nam Nang Non" stateProvince="Chiang Rai">Khun Nam Nang Non</location>
|
|
,
|
|
<location id="936767E66504423222DE0432ABF2A1A4" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:936767E66504423222DE0432ABF2A1A4" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Mae Sai" stateProvince="Chiang Rai">Mae Sai</location>
|
|
,
|
|
<collectingRegion id="1752F6D3C2F4BA5BDE34517504622906" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
: CUMZ 10186, 10188.
|
|
<location id="F3B79839BFC106E798B4B9226C0FD671" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:F3B79839BFC106E798B4B9226C0FD671" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Tham Pha Chom" stateProvince="Chiang Rai">Tham Pha Chom</location>
|
|
,
|
|
<location id="144DCEB6CEBECAA4F485827296D26E25" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:144DCEB6CEBECAA4F485827296D26E25" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Mae Sai" stateProvince="Chiang Rai">Mae Sai</location>
|
|
,
|
|
<collectingRegion id="0977CAC346A4C4703F54A2BFA43D2A27" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
: CUMZ 4347.
|
|
<location id="19C17021C728696EE6109CC9931C46E4" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:19C17021C728696EE6109CC9931C46E4" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Tham Pla" stateProvince="Chiang Rai">Tham Pla</location>
|
|
,
|
|
<location id="84AA395CA38F0D2B6B9E63986757B688" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:84AA395CA38F0D2B6B9E63986757B688" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Mae Sai" stateProvince="Chiang Rai">Mae Sai</location>
|
|
,
|
|
<collectingRegion id="4385BB846A7A184FD02331EA8371EC4E" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
: CUMZ 4346, 4803.
|
|
<location id="03B6BC84475E1E5A336F7F38AF992F58" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:03B6BC84475E1E5A336F7F38AF992F58" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Tham Saohin Prayanak" stateProvince="Chiang Rai">Tham Saohin Prayanak</location>
|
|
,
|
|
<location id="1C88A8DBFE717CFA3AF8397C332935A0" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:1C88A8DBFE717CFA3AF8397C332935A0" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Mae Sai" stateProvince="Chiang Rai">Mae Sai</location>
|
|
,
|
|
<collectingRegion id="46C2AB2BEC57CEA8948557A834CCCA02" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
: CUMZ 3906, 4442, 10190, 10193.
|
|
<location id="F2C78223978405AB2D5281C0D4F51589" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:F2C78223978405AB2D5281C0D4F51589" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Wat Tham Pla School" stateProvince="Chiang Rai">Wat Tham Pla School</location>
|
|
,
|
|
<location id="BB21D564A85C12666C7DE6B1AECFE7FA" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:BB21D564A85C12666C7DE6B1AECFE7FA" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Mae Sai" stateProvince="Chiang Rai">Mae Sai</location>
|
|
,
|
|
<collectingRegion id="79B30268646E5B27FC88706EA40F769C" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
: CUMZ 3913, 4844, 10189.
|
|
<location id="264D7A1C12E9295D134B2163F378EC6C" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:264D7A1C12E9295D134B2163F378EC6C" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Tham Mae Suai" stateProvince="Chiang Rai">Tham Mae Suai</location>
|
|
,
|
|
<location id="4634086DA2B867D8981ABA72B7C60CE7" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:4634086DA2B867D8981ABA72B7C60CE7" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Mae Suai" stateProvince="Chiang Rai">Mae Suai</location>
|
|
,
|
|
<collectingRegion id="956A446460608090AADED723485D2D16" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
: CUMZ 3907, 4345, 4807 (Fig.
|
|
<figureCitation id="88B2FB28F860CA1512300CCAAA592817" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Shell of Rhiostoma tigrina sp. nov. A holotype CUMZ 4497 / 1 from Tham Pum Tham Pla, Mae Sai, Chiang Rai B paratype CUMZ 4497 / 2 from type locality C paratype CUMZ 4806 from type locality D specimen CUMZ 4795 from Tham Phra, Chiang Rai E, F specimens CUMZ 4807 Mae Suai, Chiang Rai" figureDoi="10.3897/zookeys.1142.90097.figure31" httpUri="https://binary.pensoft.net/fig/799889" pageId="0" pageNumber="1">31E, F</figureCitation>
|
|
).
|
|
<location id="1DD05DC7927D6A310AB7D730FF7D88AE" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:1DD05DC7927D6A310AB7D730FF7D88AE" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Khao Tham Phra" stateProvince="Chiang Rai">Khao Tham Phra</location>
|
|
,
|
|
<location id="327ED311F98E7D2E32075A325E9B4135" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:327ED311F98E7D2E32075A325E9B4135" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Muang" stateProvince="Chiang Rai">Muang</location>
|
|
,
|
|
<collectingRegion id="338D55C10AD2CC27E6E2A7585029B59E" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
: CUMZ 3915, 4795 (Fig.
|
|
<figureCitation id="0D7585745951672A244237BA39B47660" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Shell of Rhiostoma tigrina sp. nov. A holotype CUMZ 4497 / 1 from Tham Pum Tham Pla, Mae Sai, Chiang Rai B paratype CUMZ 4497 / 2 from type locality C paratype CUMZ 4806 from type locality D specimen CUMZ 4795 from Tham Phra, Chiang Rai E, F specimens CUMZ 4807 Mae Suai, Chiang Rai" figureDoi="10.3897/zookeys.1142.90097.figure31" httpUri="https://binary.pensoft.net/fig/799889" pageId="0" pageNumber="1">31D</figureCitation>
|
|
), 4797, 4839, 4890, 10187.
|
|
<location id="13B3A8668988C869105A37B62C31FC70" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:13B3A8668988C869105A37B62C31FC70" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Wat Pha Cha Lui" stateProvince="Chiang Rai">Wat Pha Cha Lui</location>
|
|
,
|
|
<location id="E46123CFA487DC8B8F792BB3A8101EA1" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:E46123CFA487DC8B8F792BB3A8101EA1" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Pa Dad" stateProvince="Chiang Rai">Pa Dad</location>
|
|
,
|
|
<collectingRegion id="C75160CD689CA11C268B78AC0ED4EB2F" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
: CUMZ 4435.
|
|
<location id="7F87F1A8ACA5A37347E2E48A90D7E527" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:7F87F1A8ACA5A37347E2E48A90D7E527" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Wat Tham Phra Bumpenboon" stateProvince="Chiang Rai">Wat Tham Phra Bumpenboon</location>
|
|
,
|
|
<location id="D7A27E8F7DA51BA580B432B67C3C0AF9" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:D7A27E8F7DA51BA580B432B67C3C0AF9" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Phan" stateProvince="Chiang Rai">Phan</location>
|
|
,
|
|
<collectingRegion id="91F5AA4684DD87D986FD0272087889C7" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
: CUMZ 3914, 4495, 4496, 4706, 4846, 10027, 10034.
|
|
<location id="AF0444004A038B8FA849E2E0CF8BF6C4" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:AF0444004A038B8FA849E2E0CF8BF6C4" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Ban Pang Ma Yao" stateProvince="Chiang Rai">Ban Pang Ma Yao</location>
|
|
,
|
|
<location id="BD525A0C94C8F4809F8095427AB921F3" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:BD525A0C94C8F4809F8095427AB921F3" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Chiang Dao" stateProvince="Chiang Rai">Chiang Dao</location>
|
|
,
|
|
<collectingRegion id="F178FB44FD8048697C86BBBB5483D4F2" country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
|
|
: CUMZ 10185.
|
|
<location id="C49E11CFB9E5806344222A3D948F4EFD" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:C49E11CFB9E5806344222A3D948F4EFD" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Tham Chiang Dao" stateProvince="Chiang Rai">Tham Chiang Dao</location>
|
|
,
|
|
<location id="FEDEE58466EE709BB107397DAAC864BE" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:FEDEE58466EE709BB107397DAAC864BE" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Chiang Dao" stateProvince="Chiang Rai">Chiang Dao</location>
|
|
,
|
|
<collectingRegion id="BE8A7E2B421252547D331953B42515C5" country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
|
|
: CUMZ 3911, 4438, 4499, 4881, 10181, 10182, 10183, 10192.
|
|
<location id="AAC6AC668E6FB788ACC27E3305195BD7" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:AAC6AC668E6FB788ACC27E3305195BD7" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Tham Brichinda" stateProvince="Chiang Rai">Tham Brichinda</location>
|
|
,
|
|
<location id="5538B9C1C045E19BBA1130133E8A9A3A" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:5538B9C1C045E19BBA1130133E8A9A3A" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Chom Thong" stateProvince="Chiang Rai">Chom Thong</location>
|
|
,
|
|
<collectingRegion id="4D030DB57EED17A2EB670051A5B40687" country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
|
|
: CUMZ 10195.
|
|
<location id="CACC5FD569AD46072F51E63E74A26062" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:CACC5FD569AD46072F51E63E74A26062" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Huai Mae Muk Waterfall" stateProvince="Chiang Rai">Huai Mae Muk Waterfall</location>
|
|
,
|
|
<location id="633138784FF1A260A4A9DE3BE0308AF2" LSID="urn:lsid:plazi:treatment:9B03C93A9D3C54BD9C53245AE285A411:633138784FF1A260A4A9DE3BE0308AF2" country="Thailand" county="North Thailand" municipality="Tham Samta" name="Fang" stateProvince="Chiang Rai">Fang</location>
|
|
,
|
|
<collectingRegion id="0857FE769F154BF510066154F27F019B" country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
|
|
: CUMZ 10184
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
<caption id="52344A87867470C803E5C48ED1A50D9B" doi="10.3897/zookeys.1142.90097.figure31" httpUri="https://binary.pensoft.net/fig/799889" pageId="0" pageNumber="1" start="Figure 31" startId="F31">
|
|
<paragraph id="D7A8FFF28C8CFE4CD77586A773B32681" pageId="0" pageNumber="1">
|
|
<emphasis id="3A01FEF1B3A229307357ADBD0BCB1ADC" bold="true" pageId="0" pageNumber="1">Figure 31.</emphasis>
|
|
Shell of
|
|
<taxonomicName id="698A3FEECE19953BC62D88984DA70CC1" authorityName="Tongkerd & Tumpeesuwan" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma tigrina" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tigrina">
|
|
<emphasis id="883D788DDF5195725D93BCC1058DE581" italics="true" pageId="0" pageNumber="1">Rhiostoma tigrina</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis id="EA4E4A4A4E6AA74A3B234843CCB95966" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
<typeStatus id="983A5554957ED7925A7037CA9D8291FB">holotype</typeStatus>
|
|
CUMZ 4497/1 from Tham Pum Tham Pla, Mae Sai,
|
|
<collectingRegion id="93D45C74074E6BD1861BB58EB0C2306D" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
<emphasis id="7BF07BB67637825720D2BDB967B7875D" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
<typeStatus id="8ECB75921A3A2934ECB4618729515404">paratype</typeStatus>
|
|
CUMZ 4497/2 from type locality
|
|
<emphasis id="38AB24EE1592F9EACFE6CE81E3E11558" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
<typeStatus id="1DEDE0C260C1184709A7F7D59DD4018C">paratype</typeStatus>
|
|
CUMZ 4806 from type locality
|
|
<emphasis id="1348F3D2F2A91A6AE4903FBDDC857985" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
specimen CUMZ 4795 from Tham Phra,
|
|
<collectingRegion id="CCA66FA7AD488B93142B441B77AC0675" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
<emphasis id="C8EDC212DAAFCE93E387B725A01D10FF" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
|
|
specimens CUMZ 4807 Mae Suai,
|
|
<collectingRegion id="09D91B7BF5EC30AB9177AB6C8FA80B8E" country="Thailand" name="Chiang Rai">Chiang Rai</collectingRegion>
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="AB197CD2AF78EA767AD83402A0E1BF34" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="2B277E4FED3E9E09CFEBA03A59488C0A" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph id="3F19C30FFB27A3716C71A2BC4E256D8C" pageId="0" pageNumber="1">Shell small to medium, slightly thin and depressed. Detached whorl medium in length, curved and descending. Breathing device tubular and usually attached to preceding whorl. Shell colour with dark brown zigzag patterns on both sides.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="2DA23D7B995D1871F67F09B49C5C7920" pageId="0" pageNumber="1" type="differential diagnosis">
|
|
<paragraph id="4DBE4C0487C4B5A78222B8508C546FA6" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
|
<paragraph id="CC54FF2210AC017CF51D0107F9A7F6EA" pageId="0" pageNumber="1">
|
|
This new species differs from
|
|
<taxonomicName id="2E7DD06087EC9BF8275F2B6010095BCC" lsidName="R. marioni" pageId="0" pageNumber="1" rank="species" species="marioni">
|
|
<emphasis id="98A26DD37B042FFDA7AABA9EF9B3DA54" italics="true" pageId="0" pageNumber="1">R. marioni</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="CF577A00842145D5B06D1F7CA2D87C7C" lsidName="R. jalorensis" pageId="0" pageNumber="1" rank="species" species="jalorensis">
|
|
<emphasis id="F35DBCD36932179685441472F391AD4F" italics="true" pageId="0" pageNumber="1">R. jalorensis</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="AC13EF13B522C0F71F4512EEFDD1216F" lsidName="R. thachi" pageId="0" pageNumber="1" rank="species" species="thachi">
|
|
<emphasis id="F665AAC162279027752FB9FD9C4F6B23" italics="true" pageId="0" pageNumber="1">R. thachi</emphasis>
|
|
</taxonomicName>
|
|
in having a brown zigzag colour pattern and short detached whorl. In contrast, these three species have uniformly brownish to dark brown shells (rarely with a blotched pattern) and long and twisted detached whorls. In addition,
|
|
<taxonomicName id="EBE378E6E586FC3DFA38D2F1748309EE" lsidName="R. thachi" pageId="0" pageNumber="1" rank="species" species="thachi">
|
|
<emphasis id="317D222459AE10628B60B32D5D9F0EBA" italics="true" pageId="0" pageNumber="1">R. thachi</emphasis>
|
|
</taxonomicName>
|
|
has a broader expanded aperture on the palatal side, while
|
|
<taxonomicName id="FD8469764E67CD1ABEB328CD2F335199" lsidName="R. trigrina" pageId="0" pageNumber="1" rank="species" species="trigrina">
|
|
<emphasis id="DB0CB0349DB82F32DBF5CC96C846354A" italics="true" pageId="0" pageNumber="1">R. trigrina</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. has a thickened, slightly expanded, multi-layered lip.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B721ACB6AA30801D905722ED930E175F" pageId="0" pageNumber="1" type="description">
|
|
<paragraph id="9C88A48853752F45B89C9A8F5610C0B0" pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph id="8348C7737B38FE2CC278BB262E9387F8" pageId="0" pageNumber="1">
|
|
<emphasis id="30802617C4CE889CB10DF52FCC845076" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="A77756CF4DCDECB4355CF37FFAB9CEE3" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
|
.
|
|
</emphasis>
|
|
Shell small to medium, cW 15.7-20.5 mm, cH 8.5-11.7 mm, slightly thin, and sub-discoidal to discoidal; detached-whorl length 4.5-13.5 mm. Apex acute and dark; spire convex to nearly flat. Whorls 4 to 5, convex, increasing regularly; suture wide and shallow; last whorl rounded and stout. Shell surface with fine growth lines. Periostracum thin, corneous, and transparent. Shell with brown to dark brown zigzag patterns and faded on ventral shell surface, with narrow, dark, spiral band on periphery. Detached whorl medium in length, approximately the same length as aperture width, curved and descending. Peristome circular and double; lip thickened, expanded and multi-layered. Aperture opened sub-ventrally. Breathing device tubular and curved or straight, and its tip usually attached to preceding whorl; outer lip forming a long and closed tube, and located just behind apertural lip; inner lip with hole inside aperture. Umbilicus widely opened and deep. Operculum calcareous, cup-shaped, and multispiral (Fig.
|
|
<figureCitation id="9F91BDC6B006A15E32B810C4BD660960" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Shell of Rhiostoma tigrina sp. nov. A holotype CUMZ 4497 / 1 from Tham Pum Tham Pla, Mae Sai, Chiang Rai B paratype CUMZ 4497 / 2 from type locality C paratype CUMZ 4806 from type locality D specimen CUMZ 4795 from Tham Phra, Chiang Rai E, F specimens CUMZ 4807 Mae Suai, Chiang Rai" figureDoi="10.3897/zookeys.1142.90097.figure31" httpUri="https://binary.pensoft.net/fig/799889" pageId="0" pageNumber="1">31</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="5E8145E6A7AA3CA8A0F42C612150CDD5" pageId="0" pageNumber="1">
|
|
<emphasis id="683CAB07FFC9F2EC93F26AA7F1304C50" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="647466E79FDF0AFAC26731B002F26A2D" italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
|
.
|
|
</emphasis>
|
|
Teeth arrangement and shape are similar to those of
|
|
<taxonomicName id="0B9C9913BAEEC22DC16F59452E5B7BCA" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
|
<emphasis id="0F982BD4952D1E9C8008E13692CE7207" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
|
</taxonomicName>
|
|
. Central tooth with triangular central cusp, flanked by two lateral cusps on each side, each pair with pointed tips. Lateral teeth composed of four cusps; central cusp large with dull tip and flanked by one outer and two inner cusps. Marginal teeth each consisting of three pointed cusps (Fig.
|
|
<figureCitation id="082C107138C435745E32BE9F8B616C83" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="03B4D833C022D3F00B1E90EEF8C7AB2D" doi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1" start="Figure 32" startId="F32">
|
|
<paragraph id="8496BFA8D25EF9E308323C89BCB019EF" pageId="0" pageNumber="1">
|
|
<emphasis id="CC6A96FF91B10456C0B736B292F458B4" bold="true" pageId="0" pageNumber="1">Figure 32.</emphasis>
|
|
Radula morphology
|
|
<emphasis id="E0B415895587BB4530DC0D6B12E89A7B" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
<taxonomicName id="30D0F6D245395F8DC9CC3EFF4D5323CF" baseAuthorityName="Haines" baseAuthorityYear="1855" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma housei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="housei">
|
|
<emphasis id="23FEA612D4D9CE4BE74B22857CF59D3F" italics="true" pageId="0" pageNumber="1">Rhiostoma housei</emphasis>
|
|
</taxonomicName>
|
|
specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi
|
|
<emphasis id="DD7401D5B41F252A5D2DA0322DE77820" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
<taxonomicName id="04FAEA6FD2ABABFC2F97B1ABED90B463" authorityName="Pfeiffer" authorityYear="1862" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma hainesi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">
|
|
<emphasis id="2B994C04086EAB07E82854A03F45781E" italics="true" pageId="0" pageNumber="1">Rhiostoma hainesi</emphasis>
|
|
</taxonomicName>
|
|
specimen from Chanthaburi
|
|
<emphasis id="77A6FE557984845C1FBE22EA232F0E49" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
<taxonomicName id="F434F50874874CD79DA2F6C3E8ACBC99" authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma jalorensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jalorensis">
|
|
<emphasis id="AD5ED1A4EDCCC781566C10C101A21EC2" italics="true" pageId="0" pageNumber="1">Rhiostoma jalorensis</emphasis>
|
|
</taxonomicName>
|
|
specimen CUMZ 3994 from Wat Tham Sua, Krabi
|
|
<emphasis id="B765E10FDC24AB608CC32A121056DFC8" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
<taxonomicName id="41507F56CABCC1AC914588DCE4F2AEE4" authorityName="Tongkerd & Panha" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma ebenozostera" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ebenozostera">
|
|
<emphasis id="FF9395BF5CFC56B1B57D8651D3AB32C7" italics="true" pageId="0" pageNumber="1">Rhiostoma ebenozostera</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei
|
|
<emphasis id="BBA99C0695AEA044E2B319FCD6D5358D" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
<taxonomicName id="09B34B5782DB4B03AE87013E5FE08116" authorityName="Tongkerd & Tumpeesuwan" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma lannaense" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lannaense">
|
|
<emphasis id="8E9E13090E0006A592E0BF92F9754772" italics="true" pageId="0" pageNumber="1">Rhiostoma lannaense</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai
|
|
<emphasis id="200CEB811DDEA30E439EDA4E6D599DD3" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
<taxonomicName id="8054653A9B0F36B27D2C73FFDBB3B5B0" authorityName="Tongkerd & Tumpeesuwan" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma tigrina" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tigrina">
|
|
<emphasis id="2D90640117553453394473AFE1D58FC7" italics="true" pageId="0" pageNumber="1">Rhiostoma tigrina</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. paratype CUMZ 4497/2 from Mae Sai, Chiang Rai.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="2DA828DA6BAF395F3287CBDC95CEAB94" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="30A416D6AB00170B01835331598B2D07" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="39C1D2FABDFF2B9B29F2B5D728678EF6" pageId="0" pageNumber="1">
|
|
The specific name
|
|
<emphasis id="42ED09556E071B4901D1DE191F193A64" italics="true" pageId="0" pageNumber="1">tigrina</emphasis>
|
|
comes from the Latin word
|
|
<emphasis id="0EB136FB0A255943EA76B8DFA24EB388" italics="true" pageId="0" pageNumber="1">tigris</emphasis>
|
|
. It refers to the brownish streak or zigzag colour pattern on the shell surface of this new species, which is similar to the colour pattern of the Bengal tiger,
|
|
<taxonomicName id="27893B2A4C83FECC5B1043DF326988D0" authority="tigris" class="Mammalia" family="Felidae" genus="Panthera" kingdom="Animalia" lsidName="Panthera tigris subsp. tigris" order="Carnivora" pageId="0" pageNumber="1" phylum="Chordata" rank="subSpecies" species="tigris" subSpecies="tigris">
|
|
<emphasis id="73B24B827C0DE03260EFD1A0BFC8834D" italics="true" pageId="0" pageNumber="1">Panthera tigris tigris</emphasis>
|
|
</taxonomicName>
|
|
(Linnaeus, 1758).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="92A453065584B7DA2B1D42C09B736950" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="C38FACB4FA5C9BBE7F8D6E2B99B0756A" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="E583893DF7EF3AAD05F5A2D62D49D15C" pageId="0" pageNumber="1">
|
|
This species is mainly distributed in northern Thailand in Chiang Rai, Maehongsorn, and Chiang Mai provinces (Fig.
|
|
<figureCitation id="CD2C055F336E9A79750F0E6A04FC48F2" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">18</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="15D1954A0BE46FFB5D26E75771D640CC" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="DB1C0E17A07AF9901BCFA8C90541752B" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="D4173F20EE0DE88DC77873B0C53465FD" pageId="0" pageNumber="1">
|
|
This species has high variation in shell colour, from dark brown zigzag patterns to whitish shells without patterns (Fig.
|
|
<figureCitation id="2D1B52DDCCBE2BE04A9E6C08B62CFAD2" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Shell of Rhiostoma tigrina sp. nov. A holotype CUMZ 4497 / 1 from Tham Pum Tham Pla, Mae Sai, Chiang Rai B paratype CUMZ 4497 / 2 from type locality C paratype CUMZ 4806 from type locality D specimen CUMZ 4795 from Tham Phra, Chiang Rai E, F specimens CUMZ 4807 Mae Suai, Chiang Rai" figureDoi="10.3897/zookeys.1142.90097.figure31" httpUri="https://binary.pensoft.net/fig/799889" pageId="0" pageNumber="1">31F</figureCitation>
|
|
) among specimens from the same collection localities.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |