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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1150.93760" ID-GBIF-Dataset="ef137bea-e9d6-406b-b016-95acd7758286" ID-PMC="PMC10208737" ID-Pensoft-Pub="1313-2970-1150-1" ID-Pensoft-UUID="2D0F861C78665B9BABB241437CA5ED53" ID-PubMed="37234100" ID-ZooBank="A38C5FB69F664F858788AAA53D21704D" ModsDocID="1313-2970-1150-1" checkinTime="1677664009133" checkinUser="pensoft" docAuthor="Crews, Sarah C." docDate="2023" docId="5ADC6A85461C521E9FD2E36D7D84A334" docLanguage="en" docName="ZooKeys 1150: 1-189" docOrigin="ZooKeys 1150" docPubDate="2023-02-27" docSource="http://dx.doi.org/10.3897/zookeys.1150.93760" docTitle="Karaops markharveyi Crews 2023, sp. nov." docType="treatment" docUuid="01C8D958-A493-45B8-9C3F-6F6A83AE44EC" docUuidSource="ZooBank" docVersion="4" id="2D0F861C78665B9BABB241437CA5ED53" lastPageNumber="1" masterDocId="2D0F861C78665B9BABB241437CA5ED53" masterDocTitle="But wait, there's more! Descriptions of new species and undescribed sexes of flattie spiders (Araneae, Selenopidae, Karaops) from Australia" masterLastPageNumber="189" masterPageNumber="1" pageNumber="1" updateTime="1732488240156" updateUser="ExternalLinkService">
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<mods:title id="D40DAF9BA4C260148EDFA97DA5E465D2">But wait, there's more! Descriptions of new species and undescribed sexes of flattie spiders (Araneae, Selenopidae, Karaops) from Australia</mods:title>
</mods:titleInfo>
<mods:name id="35A38F8556747CDC9AF80A8BD1B32518" type="personal">
<mods:role id="752D3CBEF45B82A3BA170737339F4BC5">
<mods:roleTerm id="635C972FB3271600B492100CF0EBFD09">Author</mods:roleTerm>
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<mods:namePart id="3C7D99043981E80636AD180DE2EDF71B">Crews, Sarah C.</mods:namePart>
<mods:nameIdentifier id="AB461280A1A1125474309661298EEC79" type="ORCID">https://orcid.org/0000-0001-9360-6236</mods:nameIdentifier>
<mods:affiliation id="F11B783555AFA5CABB8C5069CA223D24">California Academy of Sciences, Department of Entomology, 55 Music Concourse Drive, San Francisco, CA, 94118, USA</mods:affiliation>
<mods:nameIdentifier id="A919637C19A426C5310D60CEF556405C" type="email">screwsemail@gmail.com</mods:nameIdentifier>
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<mods:title id="EB461B79078C36AEDDE1EF9FC4EF7AA1">ZooKeys</mods:title>
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<mods:part id="220DCE1825533E5618E08FBE97D465A3">
<mods:date id="B0E8B53AA6173C8F67718D5F8415D55B">2023</mods:date>
<mods:detail id="B567944E2889B4D31AE47C6A70A8418C" type="pubDate">
<mods:number id="B3BCDFB8CAA642C0249EBA0B48F9FF30">2023-02-27</mods:number>
</mods:detail>
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<mods:number id="197E3667A9A6F2E93348BE126B0B1F7D">1150</mods:number>
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<mods:start id="40929D287AB233D98BD67709D7B4E0F8">1</mods:start>
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<mods:url id="87FE2D2DA025A957BCD5BCA79A925B93">http://dx.doi.org/10.3897/zookeys.1150.93760</mods:url>
</mods:location>
<mods:classification id="90AB933CD54D89E9BACC15C7AC56D75E">journal article</mods:classification>
<mods:identifier id="C471B06C94D11C93C2F320238A2F5C33" type="DOI">http://dx.doi.org/10.3897/zookeys.1150.93760</mods:identifier>
<mods:identifier id="4B86061806FCA2464E88BC34DBB3F7BA" type="Pensoft-Pub">1313-2970-1150-1</mods:identifier>
<mods:identifier id="608F605A7D38D7186E97F1FDA6CF7DC2" type="ZooBank">A38C5FB69F664F858788AAA53D21704D</mods:identifier>
<mods:identifier id="214C3719B2D22C313E2A696F88139815" type="Pensoft-UUID">2D0F861C78665B9BABB241437CA5ED53</mods:identifier>
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<treatment id="5ADC6A85461C521E9FD2E36D7D84A334" ID-GBIF-Taxon="209383670" LSID="urn:lsid:zoobank.org:act:01C8D958-A493-45B8-9C3F-6F6A83AE44EC" httpUri="http://treatment.plazi.org/id/5ADC6A85461C521E9FD2E36D7D84A334" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Selenopidae" scope_order="Araneae">
<subSubSection id="522939AD647B779A8415695D71F0EDF0" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="7A5BB8D2A10EA7C047A9C3478780FD05" pageId="0" pageNumber="1">
<taxonomicName id="27DA21DF32C9950700302436EF62E637" ID-CoL="C7KP3" LSID="https://zoobank.org/01C8D958-A493-45B8-9C3F-6F6A83AE44EC" authority="Crews, 2023" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi" status="sp. nov.">Karaops markharveyi</taxonomicName>
<taxonomicNameLabel id="E5A7043EC520EDBBCE9510544BC5ED19" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C922EA733170304B628B39A743E75920" pageId="0" pageNumber="1" type="description">
<paragraph id="103FBE3190E4FED9FA687ADF3664512C" pageId="0" pageNumber="1">
<figureCitation id="A5104646C901FED426D531183EB9D708" captionStart="Figure 42" captionStartId="F50" captionText="Figure 42. Karaops mareeba sp. nov. and Karaops markharveyi sp. nov. A Karaops mareeba sp. nov., paratype female, Desailly Range, Queensland (QMS 120908) B Karaops mareeba sp. nov., holotype female, Desailly Range, Queensland (QMS 110872) C same, epigyne D Karaops markharveyi sp. nov., 4.6 km from Drysdale River Road, Gibb River Road, Western Australia, holotype female (sel _ 1249, WAM T 155625) E Karaops mareeba sp. nov., holotype female, endogyne, Desailly Range, Queensland (QMS 110872) F sandstone ridge habitat of Karaops markharveyi sp. nov. G Karaops mareeba sp. nov., holotype female, epigyne, caudal, Desailly Range, Queensland (QMS 110872). Scale bars: 0.5 mm (C, E, G); 2 mm (A, B, D)." figureDoi="10.3897/zookeys.1150.93760.figure42" httpUri="https://binary.pensoft.net/fig/816430" pageId="0" pageNumber="1">Figs 42D, F</figureCitation>
<figureCitation id="787633203A860283EF93D73DAE4A343E" captionStart="Figure 43" captionStartId="F51" captionText="Figure 43. Karaops markharveyi sp. nov. A paratype female, Gibb River Road, Western Australia (sel _ 1253, WAM T 155629) B paratype male, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1250, WAM T 155626)." figureDoi="10.3897/zookeys.1150.93760.figure43" httpUri="https://binary.pensoft.net/fig/816431" pageId="0" pageNumber="1">, 43A, B</figureCitation>
<figureCitation id="C75116AE2AC857F5719BACD600F1474E" captionStart="Figure 44" captionStartId="F52" captionText="Figure 44. Karaops markharveyi sp. nov. A paratype male, 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1250, WAM T 155626) B adult male, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) C holotype female, epigyne, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1249, WAM T 155625) D same, endogyne E paratype female, epigyne, Gibb River Road, Western Australia (sel _ 1253, WAM T 155629) F same, endogyne G adult male, Judbarra National Park, Northern Territory (sel _ 1330, MAGNT A 004886) H adult female, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888). Scale bars: 0.5 mm (C-F); 1 mm (B, G, H)." figureDoi="10.3897/zookeys.1150.93760.figure44" httpUri="https://binary.pensoft.net/fig/816432" pageId="0" pageNumber="1">, 44 A-H</figureCitation>
<figureCitation id="81C971D3D84100F8027A48ACE97E671D" captionStart="Figure 45" captionStartId="F53" captionText="Figure 45. Karaops markharveyi sp. nov. from Judbarra National Park, Northern Territory A adult male (sel _ 1330, MAGNT A 004886) B adult male (sel _ 1329, MAGNT A 004885)." figureDoi="10.3897/zookeys.1150.93760.figure45" httpUri="https://binary.pensoft.net/fig/816433" pageId="0" pageNumber="1">, 45 A, B</figureCitation>
<figureCitation id="1F71318B4674718486E77478B114D6F0" captionStart="Map 1" captionStartId="F3" captionText="Map 1. Map of Australia showing areas that are expanded in subsequent figures. CD = Central Desert group, D = Karaops dawara group, F = Karaops francesae group, K = Kimberley group, UA = species that are not assigned to a group, PG = Pilbara-Gascoyne group." figureDoi="10.3897/zookeys.1150.93760.map1" httpUri="https://binary.pensoft.net/fig/816383" pageId="0" pageNumber="1">, Maps 1</figureCitation>
<figureCitation id="9ED30A3F82F5C7D3DFD968F511CF6DDA" captionStart="Map 8" captionStartId="F44" captionText="Map 8. Species that are not currently placed in a species group. Diamonds (juveniles): blue = Little Mertens, red = Ruby Plains, pink = Nackeroo Lookout; Circles: yellow = Karaops yumbubaarnji sp. nov., green = Karaops markharveyi sp. nov., purple = Karaops kennerleyorum sp. nov., blue = Karaops madhawundu sp. nov., red = Karaops mareeba sp. nov. Juveniles are treated as distinct species based on molecular data." figureDoi="10.3897/zookeys.1150.93760.map8" httpUri="https://binary.pensoft.net/fig/816424" pageId="0" pageNumber="1">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B957D48225D77032CC9CA6C297E2B87" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="D7E8A58EE78A0AB3C7E3185BEBFBFF86" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="4818CD8DA3EF3174F7213C131D09C37E" pageId="0" pageNumber="1">
<materialsCitation id="9922BA876E2E175618B4178CB91F60CC" collectingDate="2016-05-22" collectorName="S. Crews, J. DeJong" county="Gibb River Road" elevation="491" latitude="-16.103525" location="River Road" longLatPrecision="1" longitude="126.579056" municipality="Gibb River" specimenCount="♀" stateProvince="Western Australia" typeStatus="Holotype">
<emphasis id="E9651E2EC3D96B0EEB99C35E41B34BF9" bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus id="F15C6F434FE4E9D5E0D964BA2CB58F32">Holotype</typeStatus>
</emphasis>
:
<collectingRegion id="F27332807D3AD4D8D1CF7B119649A82B" country="Australia" name="Western Australia">Western Australia</collectingRegion>
<specimenCount id="8FAE589685240DDF4081818DB2EB0ABB"></specimenCount>
(reared in captivity);
<collectingCounty id="367FDDF3A8F824B06C9919AEA23340FF">
<collectingMunicipality id="B733A0820401A3CD2BA4A011E36C4B43">
Gibb
<location id="339A084BEA0F208DA1FBCA1D0CAB147F" LSID="urn:lsid:plazi:treatment:5ADC6A85461C521E9FD2E36D7D84A334:339A084BEA0F208DA1FBCA1D0CAB147F" county="Gibb River Road" latitude="-16.103525" longLatPrecision="1" longitude="126.579056" municipality="Gibb River" name="River Road" stateProvince="Western Australia">River Road</location>
</collectingMunicipality>
</collectingCounty>
, ~
<quantity id="71424D9F1750B25228468A6B28D46CD2" metricMagnitude="3" metricUnit="m" metricValue="4.6" unit="km" value="4.6">4.6 km</quantity>
from
<location id="FB75209CB496DB75ECC8FE64D79BCB94" LSID="urn:lsid:plazi:treatment:5ADC6A85461C521E9FD2E36D7D84A334:FB75209CB496DB75ECC8FE64D79BCB94" county="Gibb River Road" latitude="-16.103525" longLatPrecision="1" longitude="126.579056" municipality="Gibb River" name="Drysdale River Road" stateProvince="Western Australia">
<location id="ADB47290B6EFF64F74B87A9D50BDF7B0" LSID="urn:lsid:plazi:treatment:5ADC6A85461C521E9FD2E36D7D84A334:ADB47290B6EFF64F74B87A9D50BDF7B0" county="Gibb River Road" latitude="-16.103525" longLatPrecision="1" longitude="126.579056" municipality="Gibb River" name="Drysdale River" stateProvince="Western Australia">
Drysdale
<location id="D8902781176E9612196C3304C5218E73" LSID="urn:lsid:plazi:treatment:5ADC6A85461C521E9FD2E36D7D84A334:D8902781176E9612196C3304C5218E73" county="Gibb River Road" latitude="-16.103525" longLatPrecision="1" longitude="126.579056" municipality="Gibb River" name="River Road" stateProvince="Western Australia">River Road</location>
</location>
</location>
, heading east;
<geoCoordinate id="3382E5828AC3790988D6485F3E025574" degrees="16" direction="south" minutes="6" orientation="latitude" precision="1" seconds="12.69" value="-16.103525">16°6'12.69&quot;S</geoCoordinate>
,
<geoCoordinate id="83A8ED97FDE28B0CE95AFA43362275BA" degrees="126" direction="east" minutes="34" orientation="longitude" precision="1" seconds="44.60" value="126.579056">126°34'44.60&quot;E</geoCoordinate>
; ~
<quantity id="CE0D0CF919657F822A94DB758BF82DC5" metricMagnitude="2" metricUnit="m" metricValue="4.91" unit="m" value="491.0">
<elevation id="DEBDA0778675D398A467E727A3D7B0D0" metricMagnitude="2" metricUnit="m" metricValue="4.91" unit="m" value="491.0">491 m</elevation>
</quantity>
;
<collectingDate id="F427F02B78368EFA90A493FD1E192E18" value="2016-05-22">22 May 2016</collectingDate>
;
<collectorName id="717B16CFF4EFFF217F94D84BD0285F1C">S. Crews</collectorName>
,
<collectorName id="ABAD9F8AD5F0FED1D87DA70089479F33">J. DeJong</collectorName>
leg.; sandstone outcrop on south side of road; sel_1249; SCC16_050; (WAM T155625)
</materialsCitation>
.
<materialsCitation id="685CFD30AD88EB3F9CFF5083D22A40E2" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="S. Crews &amp; J. DeJong" county="Gibb River Road" elevation="491" latitude="-16.103525" location="River Road" longLatPrecision="1" longitude="126.579056" municipality="Gibb River" specimenCount="♂" stateProvince="Western Australia" typeStatus="Paratypes">
<emphasis id="F894CE6459077C682F21C4D9D93BE8A8" bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus id="5094608E66BD63239C32D482702ED805">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="89C41A7374AB2A06D6AE9FAB9F7AEDB5"></specimenCount>
(reared in captivity); same data as previous; sel_1250; (WAM T155626)
</materialsCitation>
<materialsCitation id="AB09409A8D9FB7E7060693CF523A7FB3" collectingDate="2016-05-12" collectorName="S. Crews, J. DeJong" county="Gibb River Road" elevation="410" latitude="-15.960447" location="River Road" longLatPrecision="1" longitude="126.88259" municipality="Gibb River" specimenCount="♀" stateProvince="Northern Territory" typeStatus="paratype">
<specimenCount id="1503F9E84D95E50E519BC740BC4A2795"></specimenCount>
(reared in captivity);
<collectingCounty id="64BD789C236894FE41E2AA6BF8F33C24">
<collectingMunicipality id="F646D6EBEBFAEB7A676B75EE5495E2B2">
Gibb
<location id="0F30B2E831E1C0314A1F95E7D879A64B" LSID="urn:lsid:plazi:treatment:5ADC6A85461C521E9FD2E36D7D84A334:0F30B2E831E1C0314A1F95E7D879A64B" county="Gibb River Road" latitude="-15.960447" longLatPrecision="1" longitude="126.88259" municipality="Gibb River" name="River Road" stateProvince="Northern Territory">River Road</location>
</collectingMunicipality>
</collectingCounty>
, heading east;
<geoCoordinate id="7904B8E01A433BC882BCECEF60B21AC1" degrees="15" direction="south" minutes="57" orientation="latitude" precision="1" seconds="37.61" value="-15.960447">15°57'37.61&quot;S</geoCoordinate>
,
<geoCoordinate id="57C253E7B3DF4BE0449AD465880C9BDA" degrees="126" direction="east" minutes="52" orientation="longitude" precision="1" seconds="57.32" value="126.88259">126°52'57.32&quot;E</geoCoordinate>
; ~
<quantity id="E70EF845BA627BAB9EC773565A8E9E29" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">
<elevation id="AA02A760D1B8CBD0370557988A6C2694" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">410 m</elevation>
</quantity>
;
<collectingDate id="97BECC0CEE6E4CABF04F40D9E1061AAC" value="2016-05-12">12 May 2016</collectingDate>
;
<collectorName id="39A57824F8FE7456799FF119BC7B157A">S. Crews</collectorName>
,
<collectorName id="67E550F0444ED52B5771EF808642A7C8">J. DeJong</collectorName>
leg.; sandstone outcrop on south side of road; sel_1253; SCC16_051; (WAM T155629) •
<collectingRegion id="720047EFEC003EDD275D320F865B8A1C" country="Australia" name="Northern Territory">Northern Territory</collectingRegion>
</materialsCitation>
<materialsCitation id="C1731CB167FFBBA614CAB138201FA041" collectingDate="2016-06-01" collectorName="S. Crews, J. DeJong" county="Judbarra National Park" latitude="-15.605731" location="Joe Creek" longLatPrecision="1" longitude="131.07918" municipality="Joe Creek Walk" specimenCount="♀" stateProvince="Northern Territory" typeStatus="paratype">
<specimenCount id="719AEB67CDEFF7F3E015CEBDE8CF923B"></specimenCount>
,
<specimenCount id="6E24FD7D71C0EC33EF255FACD7CCF723"></specimenCount>
(reared in captivity);
<collectingCounty id="A52EB71EB113CB09B4164C698933E5B2">Judbarra National Park</collectingCounty>
,
<collectingMunicipality id="7BEF6647C139C2EFEFAE47CFE0422B2E">
<location id="62CD11002D277F689BDE75C3BD3C893A" LSID="urn:lsid:plazi:treatment:5ADC6A85461C521E9FD2E36D7D84A334:62CD11002D277F689BDE75C3BD3C893A" county="Judbarra National Park" latitude="-15.605731" longLatPrecision="1" longitude="131.07918" municipality="Joe Creek Walk" name="Joe Creek" stateProvince="Northern Territory">Joe Creek</location>
Walk
</collectingMunicipality>
;
<geoCoordinate id="5E349B1A7422844458FF8D5DBD9991BF" degrees="15" direction="south" minutes="36" orientation="latitude" precision="1" seconds="20.63" value="-15.605731">15°36'20.63&quot;S</geoCoordinate>
,
<geoCoordinate id="DF051941A078184CC668236F23DE4A0A" degrees="131" direction="east" minutes="4" orientation="longitude" precision="1" seconds="45.02" value="131.07918">131°4'45.02&quot;E</geoCoordinate>
;
<collectingDate id="5A5CD8DEB6132E1A986A99CFC9A65487" value="2016-06-01">1 Jun. 2016</collectingDate>
;
<collectorName id="5C03DE762D5087A10E0E08A930F40940">S. Crews</collectorName>
,
<collectorName id="B63CE6D131AA71FCB38AC58240A05343">J. DeJong</collectorName>
leg.; under pink sandstone rocks beneath escarpment; sel_1330, 1332; SCC16_070; (MAGNT A004886, A004888)
</materialsCitation>
.
<materialsCitation id="2A06685E0BB4610FB41177588E08E238" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="S. Crews &amp; J. DeJong" county="Gibb River Road" elevation="491" latitude="-16.103525" location="Western Australia" longLatPrecision="1" longitude="126.579056" municipality="Gibb River" specimenCount="2" stateProvince="Western Australia" typeStatus="Other material examined">
<emphasis id="032015041A89F7C3AAD88BE07F33BF91" bold="true" pageId="0" pageNumber="1">
<typeStatus id="D3E6289D43D26A2E5B3379206D4F9D14" isOtherTypeStatus="true">Other material examined</typeStatus>
</emphasis>
:
<collectingRegion id="CCEDE0C0FD1B8094B4E4DD0B7DF12768" country="Australia" name="Western Australia">Western Australia</collectingRegion>
<specimenCount id="F8C79D30640AF579BA130AF0D486851E" type="generic">2 imm.</specimenCount>
, same data as holotype; sel_1248, 1251; (WAM T155624, T155627)
</materialsCitation>
<materialsCitation id="37CE35CB78127E38AD10ACEBD3E7B7C7" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="S. Crews &amp; J. DeJong" county="Gibb River Road" elevation="491" latitude="-16.103525" location="River Road" longLatPrecision="1" longitude="126.579056" municipality="Gibb River" specimenCount="1" stateProvince="Western Australia">
<specimenCount id="E785B33A27C901740642CF957B1B7155" type="generic">1 imm.</specimenCount>
; same data as sel_1253; sel_1251; (WAM T155627)
</materialsCitation>
<materialsCitation id="C2F271CB7ADF495B87CFD6E140B1B4B3" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="S. Crews &amp; J. DeJong" county="Gibb River Road" elevation="491" latitude="-16.103525" location="River Road" longLatPrecision="1" longitude="126.579056" municipality="Gibb River" specimenCount="♂" stateProvince="Western Australia">
<specimenCount id="5B43E8517D7572DAB7AF386D5E4C5FCD"></specimenCount>
(reared in captivity); same data as previous; sel_1252; (WAM T155628)
</materialsCitation>
<materialsCitation id="12A1E9282FF7BF0494A3D094B618FA21" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="S. Crews &amp; J. DeJong" county="Gibb River Road" elevation="491" latitude="-16.103525" location="River Road" longLatPrecision="1" longitude="126.579056" municipality="Gibb River" specimenCount="2" specimenCount-male="2" stateProvince="Western Australia">
<specimenCount id="FCC8B8B7965B99F0D851A92CCFE7DD02" type="male">2♂</specimenCount>
(reared in captivity); same data as sel_1330, 1332; sel_1329, 1331; (MAGNT A004885, A004887)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90BA894159807394569F6B4BBB0D8B1E" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="61E5609DA6DD64E9FE0C6F5175A5F0E9" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="DA19BF5118099CF690A06E29C764A552" pageId="0" pageNumber="1">
Females of
<taxonomicName id="5FF01A64AD1DA3BBAFB03E8B4A220965" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi">
<emphasis id="D9C864B3BB6AE39561CDA17AA974082E" italics="true" pageId="0" pageNumber="1">Karaops markharveyi</emphasis>
</taxonomicName>
sp. nov. are most similar to
<taxonomicName id="E40796C81245C9193BD4D3A71E77ED86" lsidName="K. madhawundu" pageId="0" pageNumber="1" rank="species" species="madhawundu">
<emphasis id="6E875D866EBB9A9F475D9151228FFFF6" italics="true" pageId="0" pageNumber="1">K. madhawundu</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="541CE34C515DC21C59F8DA479AB192D5" lsidName="K. mareeba" pageId="0" pageNumber="1" rank="species" species="mareeba">
<emphasis id="CC6B3BD7EA99DF6013FF5F7F052E3587" italics="true" pageId="0" pageNumber="1">K. mareeba</emphasis>
</taxonomicName>
sp. nov. by the large, round accessory bulbs (Figs
<figureCitation id="E9B5DCC7EC227620D7841AA94D2E7955" captionStart="Figure 44" captionStartId="F52" captionText="Figure 44. Karaops markharveyi sp. nov. A paratype male, 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1250, WAM T 155626) B adult male, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) C holotype female, epigyne, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1249, WAM T 155625) D same, endogyne E paratype female, epigyne, Gibb River Road, Western Australia (sel _ 1253, WAM T 155629) F same, endogyne G adult male, Judbarra National Park, Northern Territory (sel _ 1330, MAGNT A 004886) H adult female, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888). Scale bars: 0.5 mm (C-F); 1 mm (B, G, H)." figureDoi="10.3897/zookeys.1150.93760.figure44" httpUri="https://binary.pensoft.net/fig/816432" pageId="0" pageNumber="1">44C-F</figureCitation>
;
<figureCitation id="9EF2A47878ED1C86135EA36EBF918227" captionStart="Figure 47" captionStartId="F55" captionText="Figure 47. Karaops markharveyi sp. nov. A holotype female, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1249, WAM T 155625) B habitat, Judbarra National Park, Northern Territory C holotype female, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1249, WAM T 155625) D epigyne, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888) E same, endogyne. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure47" httpUri="https://binary.pensoft.net/fig/816435" pageId="0" pageNumber="1">47D, E</figureCitation>
) but can be differentiated by the much smaller size: females of the new species average 3.58 (
<emphasis id="E6081B43EFB331AB66961DB484BB97CA" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 3), whereas
<taxonomicName id="330087AE9C648BFAD45B831B59AB6B4D" lsidName="K. madhawundu" pageId="0" pageNumber="1" rank="species" species="madhawundu">
<emphasis id="98C2C2EC7D01D786BD273643D9B0F899" italics="true" pageId="0" pageNumber="1">K. madhawundu</emphasis>
</taxonomicName>
sp. nov. have a median size of 7.31 (
<emphasis id="AF57776A7BE54F560549EFF5B033AA4E" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 3) and
<taxonomicName id="BF743289827A7BEB42973F355BBB1850" lsidName="K. mareeba" pageId="0" pageNumber="1" rank="species" species="mareeba">
<emphasis id="4F5CFC9B56516AE323CF46DEE0182588" italics="true" pageId="0" pageNumber="1">K. mareeba</emphasis>
</taxonomicName>
sp. nov. a median size of 7.63 (
<emphasis id="9D7747D04DAFFAB7A34E52D5D04A83ED" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 2). Additionally, the spermathecae of
<taxonomicName id="E24D637941C9D82A20D18D5CE3460704" lsidName="K. mareeba" pageId="0" pageNumber="1" rank="species" species="mareeba">
<emphasis id="BE51B451BFFD9AA0791F96C4C4F585AC" italics="true" pageId="0" pageNumber="1">K. mareeba</emphasis>
</taxonomicName>
sp. nov. are directed medially at ~ 45°angle, and the copulatory openings are beneath a lobe (42C, E).
</paragraph>
<paragraph id="C2170AFAC469EBC1DC53246B74572CEC" pageId="0" pageNumber="1">
The male palps do not resemble any other known Australian selenopid (Fig.
<figureCitation id="DC4ABF4E5E07DCE026767593DD9A8031" captionStart="Figure 46" captionStartId="F54" captionText="Figure 46. Karaops markharveyi sp. nov. and Karaops sp. A paratype male, palp, ventral, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1250, WAM T 155626) B same, retrolateral C same, dorsal D habitat of Karaops sp., Timber Creek, Nackeroo Lookout, Northern Territory E Karaops markharveyi sp. nov., palp, dorsal, tilted forward to see chemosensory brush, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) F habitat of Karaops markharveyi sp. nov., Judbarra National Park, Northern Territory. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure46" httpUri="https://binary.pensoft.net/fig/816434" pageId="0" pageNumber="1">46A-C, E</figureCitation>
); however, the males are unknown for
<taxonomicName id="DD6B66D510B583460B5AAAC2C0C05EE5" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops madhawundu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madhawundu">
<emphasis id="7471C5B678CD13AF4D104BC2A6F4048B" italics="true" pageId="0" pageNumber="1">Karaops madhawundu</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="A6D4055EAC962092D8B80AA5F74E7759" lsidName="K. mareeba" pageId="0" pageNumber="1" rank="species" species="mareeba">
<emphasis id="258684094EDA0EF508816E31A7163562" italics="true" pageId="0" pageNumber="1">K. mareeba</emphasis>
</taxonomicName>
sp. nov., the two species that most closely resemble this new species, and molecular data suggest they are closely related to one another (Suppl. material 1). The cymbium has a superficial resemblance to that of
<taxonomicName id="5A5F19919B4154FA79490FCB635A460A" lsidName="K. yumbubaarnji" pageId="0" pageNumber="1" rank="species" species="yumbubaarnji">
<emphasis id="A0EEC9B7EE83F799708C3E5390C11910" italics="true" pageId="0" pageNumber="1">K. yumbubaarnji</emphasis>
</taxonomicName>
sp. nov. by the chemosensory setae on the tip (Fig.
<figureCitation id="BEE86E6E30881E8DDA8447CF838EF3CB" captionStart="Figure 36" captionStartId="F43" captionText="Figure 36. Karaops yumbubaarnji sp. nov., holotype male, Cathedral Gorge Lookout, Purnululu National Park, Western Australia (sel _ 1300, WAM T 155676) A palp, ventral, arrow indicates expanded part below B palp, retrolateral C close up of the median apophysis showing small, spine-like process. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure36" httpUri="https://binary.pensoft.net/fig/816423" pageId="0" pageNumber="1">36A, B</figureCitation>
), and the body size of both species is small, but the other features of the palp are not similar, and the abdominal pattern differs between the two (Figs
<figureCitation id="8CBC7FC4D7D6B46E810080D868C419ED" captionStart="Figure 34" captionStartId="F41" captionText="Figure 34. Karaops yumbubaarnji sp. nov., Cathedral Gorge Lookout, Purnululu National Park, Western Australia A live spider on sandstone B habitat, Bungle Bungles." figureDoi="10.3897/zookeys.1150.93760.figure34" httpUri="https://binary.pensoft.net/fig/816421" pageId="0" pageNumber="1">34A</figureCitation>
,
<figureCitation id="0245DF536F5336243F65209CD1E678B2" captionStart="Figure 35" captionStartId="F42" captionText="Figure 35. Karaops yumbubaarnji sp. nov., Cathedral Gorge Lookout, Purnululu National Park, Western Australia A holotype male (sel _ 1300, WAM T 155676) B adult female." figureDoi="10.3897/zookeys.1150.93760.figure35" httpUri="https://binary.pensoft.net/fig/816422" pageId="0" pageNumber="1">35A</figureCitation>
,
<figureCitation id="36DD184306D7C62198CD6095EC2ECA95" captionStart="Figure 44" captionStartId="F52" captionText="Figure 44. Karaops markharveyi sp. nov. A paratype male, 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1250, WAM T 155626) B adult male, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) C holotype female, epigyne, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1249, WAM T 155625) D same, endogyne E paratype female, epigyne, Gibb River Road, Western Australia (sel _ 1253, WAM T 155629) F same, endogyne G adult male, Judbarra National Park, Northern Territory (sel _ 1330, MAGNT A 004886) H adult female, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888). Scale bars: 0.5 mm (C-F); 1 mm (B, G, H)." figureDoi="10.3897/zookeys.1150.93760.figure44" httpUri="https://binary.pensoft.net/fig/816432" pageId="0" pageNumber="1">44B</figureCitation>
,
<figureCitation id="9DF76A13FDD59FEC7DBE7403FECA1B73" captionStart="Figure 45" captionStartId="F53" captionText="Figure 45. Karaops markharveyi sp. nov. from Judbarra National Park, Northern Territory A adult male (sel _ 1330, MAGNT A 004886) B adult male (sel _ 1329, MAGNT A 004885)." figureDoi="10.3897/zookeys.1150.93760.figure45" httpUri="https://binary.pensoft.net/fig/816433" pageId="0" pageNumber="1">45A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A6027E5794F54800D47802E8C33E92C1" pageId="0" pageNumber="1" type="description">
<paragraph id="8C1A6F589DE951625877F2AD130D8BB5" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="C6D20F6F779A5033B7936DC289F37D3B" pageId="0" pageNumber="1">
<emphasis id="94C6DDFA59336FC261962CDD9449C6FB" bold="true" pageId="0" pageNumber="1">Female</emphasis>
(holotype). Total length 4.09. Carapace length 1.99, width 2.39. Chelicerae: promargin with three teeth, retromargin with two teeth (1-0-1). Eyes: AER recurved; PER strongly recurved; diameters, AME 0.12, ALE 0.08, PME 0.16, PLE 0.26; interdistances AME-PME 0.04, PME-ALE 0.11, ALE-PLE 0.17, PME-PME 0.72, ALE-ALE 1.01, AME-AME 0.38, PLE-PLE 1.22. Sternum length 0.88, width 1.18. Abdomen length 2.10, width 1.52. Color (in life Figs
<figureCitation id="6DCEFC300AB3A9ED953FFDA6015C33C1" captionStart="Figure 43" captionStartId="F51" captionText="Figure 43. Karaops markharveyi sp. nov. A paratype female, Gibb River Road, Western Australia (sel _ 1253, WAM T 155629) B paratype male, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1250, WAM T 155626)." figureDoi="10.3897/zookeys.1150.93760.figure43" httpUri="https://binary.pensoft.net/fig/816431" pageId="0" pageNumber="1">43A</figureCitation>
,
<figureCitation id="5000C1528DF7CCE4D3EF63F399DCE4FA" captionStart="Figure 47" captionStartId="F55" captionText="Figure 47. Karaops markharveyi sp. nov. A holotype female, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1249, WAM T 155625) B habitat, Judbarra National Park, Northern Territory C holotype female, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1249, WAM T 155625) D epigyne, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888) E same, endogyne. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure47" httpUri="https://binary.pensoft.net/fig/816435" pageId="0" pageNumber="1">47A, C</figureCitation>
/preserved Figs
<figureCitation id="23D142ECE77153810B90FBEE6A996638" captionStart="Figure 42" captionStartId="F50" captionText="Figure 42. Karaops mareeba sp. nov. and Karaops markharveyi sp. nov. A Karaops mareeba sp. nov., paratype female, Desailly Range, Queensland (QMS 120908) B Karaops mareeba sp. nov., holotype female, Desailly Range, Queensland (QMS 110872) C same, epigyne D Karaops markharveyi sp. nov., 4.6 km from Drysdale River Road, Gibb River Road, Western Australia, holotype female (sel _ 1249, WAM T 155625) E Karaops mareeba sp. nov., holotype female, endogyne, Desailly Range, Queensland (QMS 110872) F sandstone ridge habitat of Karaops markharveyi sp. nov. G Karaops mareeba sp. nov., holotype female, epigyne, caudal, Desailly Range, Queensland (QMS 110872). Scale bars: 0.5 mm (C, E, G); 2 mm (A, B, D)." figureDoi="10.3897/zookeys.1150.93760.figure42" httpUri="https://binary.pensoft.net/fig/816430" pageId="0" pageNumber="1">42D</figureCitation>
,
<figureCitation id="BD0A431E29FD93D47471F4B66C39DAD8" captionStart="Figure 44" captionStartId="F52" captionText="Figure 44. Karaops markharveyi sp. nov. A paratype male, 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1250, WAM T 155626) B adult male, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) C holotype female, epigyne, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1249, WAM T 155625) D same, endogyne E paratype female, epigyne, Gibb River Road, Western Australia (sel _ 1253, WAM T 155629) F same, endogyne G adult male, Judbarra National Park, Northern Territory (sel _ 1330, MAGNT A 004886) H adult female, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888). Scale bars: 0.5 mm (C-F); 1 mm (B, G, H)." figureDoi="10.3897/zookeys.1150.93760.figure44" httpUri="https://binary.pensoft.net/fig/816432" pageId="0" pageNumber="1">44H</figureCitation>
): Carapace: reddish brown with two large dark patches behind PLEs extending to middle of carapace next to furrow, some dark patches laterally and posterior to furrow, white setae behind eyes, and in patches on carapace. Chelicerae: yellow-brown, anterior black except medially. Maxillae: yellow, dusky on outer edge. Labium: brownish yellow, pale distally. Sternum: pale yellowish white. Abdomen: dorsally dark brown with orangish spots anterolaterally, mediolaterally, posterolaterally, posteriorly, all but posterior patches with black dot, all orangish areas with tufts of white setae on edges, posterior dark; ventrally yellowish white. Anterior spinnerets: yellowish brown, posterior spinnerets black except ventrally yellowish brown. Legs: yellowish with black marks dorsally and ventrally on Cx I-III, dark mark anteriorly on all Tr,, annulations on all legs, only completely encircle legs at Mt, centers open but dusky on Fm, Pt, and Fm-Pt joint, dark annulations on Ti at Pt-Ti joint and medially, dark annulations on Mt at Ti-Mt joint and at Mt-Ta joint, Ta tips dark; flat, white setae enlarged distally ventrally on all Fm, Pt, Ti, annulation at Pt-Mt joint, tips of Ta dark; spination leg I Fm d 1-1-1, Ti v 2-2-2-2-2, Mt v 2-2-2; leg II Fm d 1-1-1, Ti v 2-2-2-2-2, Mt v 2-2-2; leg III Fm d 1-1-1, Ti v 1-1, Mt v 2-0, 5 pair of small spines near Mt-Ta; leg IV Fm d 1-1-1, rl 0-0-1, rl 0-1-1, Ti v 1-1-1, Mt v 1-1; leg formula 2341; measurements leg I 7.43 (2.41, 0.90, 2.00. 1.23, 0.89); leg II 9.78 (3.82, 0.88, 2.34, 1.64, 1.10); leg III 9.56 (3.40, 0.83, 2.39, 1.18, 1.76); leg IV 8.33 (2.94, 0.60, 2.03, 1.65, 1.11). Palp spination Fm d 0-1-2; 1.84 (0.59, 0.36, 0.27, 0.63); claw with five teeth; dark spots dorsolaterally and ventrally on Fm. Epigyne: EP rounded triangular; MF slightly depressed; LL conspicuous posteriorly, touching; COs laterally in depression. Endogyne: CDs very short; ABs large, round; S oblong; FDs large, directed anterolaterally; small pdf.
</paragraph>
<caption id="90182DF89C737A0AAC25377EC76CDA6C" doi="10.3897/zookeys.1150.93760.figure43" httpUri="https://binary.pensoft.net/fig/816431" pageId="0" pageNumber="1" start="Figure 43" startId="F51">
<paragraph id="506C7C352588EEA957109465F0AA9345" pageId="0" pageNumber="1">
<emphasis id="0335267BAFF4180BCF64A73959FB7765" bold="true" pageId="0" pageNumber="1">Figure 43.</emphasis>
<taxonomicName id="DCF05DC69644DAE61E1C343B7AB8A772" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi">
<emphasis id="97663D6047D20A6F4BBEF32B19F23266" italics="true" pageId="0" pageNumber="1">Karaops markharveyi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="484A93BE4744DE2F0CF8F6F741CAE652" bold="true" pageId="0" pageNumber="1">A</emphasis>
paratype female, Gibb River Road, Western Australia (sel_1253, WAM T155629)
<emphasis id="FC60E236C302E6A3A648371C7334AFE1" bold="true" pageId="0" pageNumber="1">B</emphasis>
paratype male, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel_1250, WAM T155626).
</paragraph>
</caption>
<caption id="1E761DCF7E39EF24FB8EF6C2FBA7DEE4" doi="10.3897/zookeys.1150.93760.figure44" httpUri="https://binary.pensoft.net/fig/816432" pageId="0" pageNumber="1" start="Figure 44" startId="F52">
<paragraph id="39ECD8BF962BD10191D00BD5254602DE" pageId="0" pageNumber="1">
<emphasis id="EDD6CD8B878DE6C9FFAA6CD08FE61008" bold="true" pageId="0" pageNumber="1">Figure 44.</emphasis>
<taxonomicName id="D9A13C18E186FBAE3B1847006DCECECE" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi">
<emphasis id="61D1774919168D2F736F1BF3D6F33C4D" italics="true" pageId="0" pageNumber="1">Karaops markharveyi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="B38BC6A640026A3A2AA6D3BBFC0BD037" bold="true" pageId="0" pageNumber="1">A</emphasis>
paratype male, 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel_1250, WAM T155626)
<emphasis id="5F3C2901C8F2D54D0ECCD0E33D3689A2" bold="true" pageId="0" pageNumber="1">B</emphasis>
adult male, Judbarra National Park, Northern Territory (sel_1329, MAGNT A004885)
<emphasis id="44A5C9E7BDE6FF24FEB2F2FF3DE8947E" bold="true" pageId="0" pageNumber="1">C</emphasis>
holotype female, epigyne, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel_1249, WAM T155625)
<emphasis id="2221FCD955F6369D4CECF36F76702DAD" bold="true" pageId="0" pageNumber="1">D</emphasis>
same, endogyne
<emphasis id="ED2B0A59AD68A3DCB47CA39B32E0A394" bold="true" pageId="0" pageNumber="1">E</emphasis>
paratype female, epigyne, Gibb River Road, Western Australia (sel_1253, WAM T155629)
<emphasis id="B7E4D95687C48017707087E9598ED8DC" bold="true" pageId="0" pageNumber="1">F</emphasis>
same, endogyne
<emphasis id="21F9251CB94205EC90DAE30638B34427" bold="true" pageId="0" pageNumber="1">G</emphasis>
adult male, Judbarra National Park, Northern Territory (sel_1330, MAGNT A004886)
<emphasis id="30834FC2A10C6BD940EB0723A9950CDC" bold="true" pageId="0" pageNumber="1">H</emphasis>
adult female, Judbarra National Park, Northern Territory (sel_1332, MAGNT A004888). Scale bars: 0.5 mm (
<emphasis id="7AC4264A19FB219DA83E1C0058360078" bold="true" pageId="0" pageNumber="1">C-F</emphasis>
); 1 mm (
<emphasis id="B3E398F78BC618808F83C9D246765F0F" bold="true" pageId="0" pageNumber="1">B, G, H</emphasis>
).
</paragraph>
</caption>
<caption id="8D97FE5E894AEE8C5C3F33735D0E2A2A" doi="10.3897/zookeys.1150.93760.figure45" httpUri="https://binary.pensoft.net/fig/816433" pageId="0" pageNumber="1" start="Figure 45" startId="F53">
<paragraph id="BC0E7F8952DC15E3F7146276B213E8DE" pageId="0" pageNumber="1">
<emphasis id="8D0306B4577010828037A4B3D848BDBD" bold="true" pageId="0" pageNumber="1">Figure 45.</emphasis>
<taxonomicName id="9A3829934277C7614295CEE5CD72C5F0" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi">
<emphasis id="FD10B97F576CF08321F6A90AD86C21C6" italics="true" pageId="0" pageNumber="1">Karaops markharveyi</emphasis>
</taxonomicName>
sp. nov. from Judbarra National Park, Northern Territory
<emphasis id="E62BEAFF650137EE87B6D707D9212C72" bold="true" pageId="0" pageNumber="1">A</emphasis>
adult male (sel_1330, MAGNT A004886)
<emphasis id="4977430CC7A386A6C6D21ECCEE2851D1" bold="true" pageId="0" pageNumber="1">B</emphasis>
adult male (sel_1329, MAGNT A004885).
</paragraph>
</caption>
<caption id="60E70974B3958474ED8116AA90301FAF" doi="10.3897/zookeys.1150.93760.figure46" httpUri="https://binary.pensoft.net/fig/816434" pageId="0" pageNumber="1" start="Figure 46" startId="F54">
<paragraph id="050BA125B6B68F52DFC437502CAB929E" pageId="0" pageNumber="1">
<emphasis id="934279BA8ADE91A6D72BC39D3CC9F051" bold="true" pageId="0" pageNumber="1">Figure 46.</emphasis>
<taxonomicName id="25E0BC812FE049C4F024299F90CEDD7A" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi">
<emphasis id="6F1B362DA230C86A7445D226850EE944" italics="true" pageId="0" pageNumber="1">Karaops markharveyi</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="C511676A988F2EFF8340B2C3B1E53FB5" authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="D46237912BA24B62BC2C38080F0E5F88" italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp.
<emphasis id="4169A236253641D7C74C7EF30949D01D" bold="true" pageId="0" pageNumber="1">A</emphasis>
paratype male, palp, ventral, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel_1250, WAM T155626)
<emphasis id="EB1A4B2C11A1E6CD7D4E893AC4B9EB7E" bold="true" pageId="0" pageNumber="1">B</emphasis>
same, retrolateral
<emphasis id="7DAC687A3D24311E2A7C9A809A4643FE" bold="true" pageId="0" pageNumber="1">C</emphasis>
same, dorsal
<emphasis id="41E999C829DFAA997BC28860E3DD1DA9" bold="true" pageId="0" pageNumber="1">D</emphasis>
habitat of
<taxonomicName id="4B474E49316A65FC79BC57D37AD5F1D6" authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="EA8892F7A65E533C53C2E570FD0180C4" italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp., Timber Creek, Nackeroo Lookout, Northern Territory
<emphasis id="F444B4E1C263EDC9F16EB42B85247450" bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName id="A9C3867F1442F0DDD594D3D133F12C39" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi">
<emphasis id="67854D66A88C92591987EBB6CB10FEE5" italics="true" pageId="0" pageNumber="1">Karaops markharveyi</emphasis>
</taxonomicName>
sp. nov., palp, dorsal, tilted forward to see chemosensory brush, Judbarra National Park, Northern Territory (sel_1329, MAGNT A004885)
<emphasis id="7BAE179E504107ED560BCECA2F0D2E7A" bold="true" pageId="0" pageNumber="1">F</emphasis>
habitat of
<taxonomicName id="616E32F94245A58A1AF7B1537F9433B3" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi">
<emphasis id="C7DD35B08CEB81871EBF64B5E2F5AD73" italics="true" pageId="0" pageNumber="1">Karaops markharveyi</emphasis>
</taxonomicName>
sp. nov., Judbarra National Park, Northern Territory. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="CFAF141F5EB8EAFB98786C9679FF72D6" doi="10.3897/zookeys.1150.93760.figure47" httpUri="https://binary.pensoft.net/fig/816435" pageId="0" pageNumber="1" start="Figure 47" startId="F55">
<paragraph id="DC01B300F610E6BE5DB9181CDEB84299" pageId="0" pageNumber="1">
<emphasis id="1E38B376AE924615FDC45542E7856161" bold="true" pageId="0" pageNumber="1">Figure 47.</emphasis>
<taxonomicName id="F35388D191BDD386AF4D6559868EB139" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi">
<emphasis id="611AAA43808CBCFC2D31D33760B17C35" italics="true" pageId="0" pageNumber="1">Karaops markharveyi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="C27D1C8AF3022F50725E0E7374FD0A64" bold="true" pageId="0" pageNumber="1">A</emphasis>
holotype female, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel_1249, WAM T155625)
<emphasis id="CC5717B989CC7BA2E75CC60F97DB5280" bold="true" pageId="0" pageNumber="1">B</emphasis>
habitat, Judbarra National Park, Northern Territory
<emphasis id="FFABE8586B50DAFB1C9EC6CF9D2BE815" bold="true" pageId="0" pageNumber="1">C</emphasis>
holotype female, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel_1249, WAM T155625)
<emphasis id="942F7F85E2AE063A5EFD2D475C034718" bold="true" pageId="0" pageNumber="1">D</emphasis>
epigyne, Judbarra National Park, Northern Territory (sel_1332, MAGNT A004888)
<emphasis id="951122C0B825A56495649DF83828F6ED" bold="true" pageId="0" pageNumber="1">E</emphasis>
same, endogyne. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="AA913E83115B3D2ABC9CA3CCB4A78B65" pageId="0" pageNumber="1">
<emphasis id="33FBA6A72E18CD0B6C16F2483A4749E1" bold="true" pageId="0" pageNumber="1">Male</emphasis>
(paratype). Total length 3.10. Carapace length 1.71, width 2.07. Chelicerae: promargin with three teeth, retromargin with two teeth (1-0-1). Eyes: AER recurved, PER strongly recurved; diameters, AME 0.12, ALE 0.06, PME 0.16, PLE 0.22; interdistances AME-PME 0.02, PME-ALE 0.11, ALE-PLE 0.15, PME-PME 0.63, ALE-ALE 0.88, AME-AME 0.33, PLE-PLE 1.09. Sternum: length 0.89, width 1.16. Abdomen length 1.39, width 1.29. Color (in life Figs
<figureCitation id="64EA7AC334319FE622CBAB587D8006F7" captionStart="Figure 43" captionStartId="F51" captionText="Figure 43. Karaops markharveyi sp. nov. A paratype female, Gibb River Road, Western Australia (sel _ 1253, WAM T 155629) B paratype male, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1250, WAM T 155626)." figureDoi="10.3897/zookeys.1150.93760.figure43" httpUri="https://binary.pensoft.net/fig/816431" pageId="0" pageNumber="1">43B</figureCitation>
,
<figureCitation id="D2F6B1FF2180800C396A29C1D82C345A" captionStart="Figure 44" captionStartId="F52" captionText="Figure 44. Karaops markharveyi sp. nov. A paratype male, 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1250, WAM T 155626) B adult male, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) C holotype female, epigyne, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1249, WAM T 155625) D same, endogyne E paratype female, epigyne, Gibb River Road, Western Australia (sel _ 1253, WAM T 155629) F same, endogyne G adult male, Judbarra National Park, Northern Territory (sel _ 1330, MAGNT A 004886) H adult female, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888). Scale bars: 0.5 mm (C-F); 1 mm (B, G, H)." figureDoi="10.3897/zookeys.1150.93760.figure44" httpUri="https://binary.pensoft.net/fig/816432" pageId="0" pageNumber="1">44A</figureCitation>
,
<figureCitation id="A3258720273BD70BFD5C1A9F9B2FA1D5" captionStart="Figure 45" captionStartId="F53" captionText="Figure 45. Karaops markharveyi sp. nov. from Judbarra National Park, Northern Territory A adult male (sel _ 1330, MAGNT A 004886) B adult male (sel _ 1329, MAGNT A 004885)." figureDoi="10.3897/zookeys.1150.93760.figure45" httpUri="https://binary.pensoft.net/fig/816433" pageId="0" pageNumber="1">45A, B</figureCitation>
,
<figureCitation id="50CB9AC225F440CEA657E183D4AF279A" captionStart="Figure 48" captionStartId="F56" captionText="Figure 48. Karaops markharveyi sp. nov. and members of the Pilbara-Gascoyne species group A Karaops markharveyi sp. nov., adult male, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) B Karaops sp., Timber Creek, Nackeroo Lookout, Northern Territory. Arrow indicates setal tufts C Karaops burbidgei, paratype female, Barrow Island (WAM T 76698) D Karaops burbidgei, holotype male, Barrow Island (WAM T 55000). Scale bars: 1 mm." figureDoi="10.3897/zookeys.1150.93760.figure48" httpUri="https://binary.pensoft.net/fig/816436" pageId="0" pageNumber="1">48A</figureCitation>
/preserved Fig.
<figureCitation id="D9B7A6EBBE1AC0CD8E8B5C0847AC4B3E" captionStart="Figure 44" captionStartId="F52" captionText="Figure 44. Karaops markharveyi sp. nov. A paratype male, 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1250, WAM T 155626) B adult male, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) C holotype female, epigyne, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1249, WAM T 155625) D same, endogyne E paratype female, epigyne, Gibb River Road, Western Australia (sel _ 1253, WAM T 155629) F same, endogyne G adult male, Judbarra National Park, Northern Territory (sel _ 1330, MAGNT A 004886) H adult female, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888). Scale bars: 0.5 mm (C-F); 1 mm (B, G, H)." figureDoi="10.3897/zookeys.1150.93760.figure44" httpUri="https://binary.pensoft.net/fig/816432" pageId="0" pageNumber="1">44B, G</figureCitation>
): Carapace: reddish brown with two large dark patches originating behind PLEs, extending to middle of carapace next to furrow, some dark patches laterally, posterior to furrow, white setae behind eyes, in patches on carapace/yellow-brown, dark patches visible but dispersed, white setae not as conspicuous. Chelicerae: yellowish, anterior black except basolaterally, setae pale, darkened towards middle. Maxillae: yellowish with black marks anterolaterally. Labium tan, paler distally. Sternum: yellowish white. Abdomen dorsally brownish orange with dark areas medially, laterally, with four dots anteriorly, small patches of white setae at edges of dark medial mark; ventrally whitish gray. Spinnerets: anterior yellowish brown, posterior black except ventrally yellowish brown. Legs: yellowish with black marks anteriorly and posteriorly on Cx, anteriorly on Tr, black bands and spots on Fm, Pt dusky at Fm-Pt joint ventrally, black annulation at Pt-Ti joint, two additional dusky areas, dark annulation on Mt at Ti-Mt joint and at Mt-Ta joint, Ta dark at tip; white setae enlarged distally ventrally on Fm, Pt, Ti; spination leg I Fm d 1-1-1, pr 0-0-1, rl 0-0-1, Ti v 2-2-2-2-2-2, Mt v 2-2-2-2; leg II Fm d 1-1-1, pr 0-0-1, Ti v 2-2-2-2-2-2, Mt v 2-2-2-2; leg III Fm d 1-1-1, Ti v 2-2; leg IV Fm d 1-1-1, Ti v 2-2; leg formula 3241; measurements leg I 7.23 (2.19, 0.73, 1.85, 1.48, 0.98); leg II 8.71 (2.73, 0.79, 2.18, 1.89, 1.12); leg III 9.29 (2.99, 0.82, 2.33, 2.04, 1.11); leg IV 7.97 (2.60, 0.62, 1.84, 1.95, 0.96). Palp: spination Fm d 0-0-1; 1.73 (0.63, 0.22, 0.27, 0.61); dark spot retrolaterally on Fm, retrolaterally and dorsally on Ti, Cy with dark marking basodorsally, apically; dRTA very large, much larger than vRTA, wide, nearly as wide as Ti, pointed at tip, in ventral view, convex medially, plicate distally, vRTA narrow, spoon shaped in ventral view; rbcp smallish; Cy oval in ventral view, laterally, can see indentation along top with patch of chemosensory setae (Fig.
<figureCitation id="F5CD0C916CD1176653AA1B0A5A0EF415" captionStart="Figure 46" captionStartId="F54" captionText="Figure 46. Karaops markharveyi sp. nov. and Karaops sp. A paratype male, palp, ventral, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1250, WAM T 155626) B same, retrolateral C same, dorsal D habitat of Karaops sp., Timber Creek, Nackeroo Lookout, Northern Territory E Karaops markharveyi sp. nov., palp, dorsal, tilted forward to see chemosensory brush, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) F habitat of Karaops markharveyi sp. nov., Judbarra National Park, Northern Territory. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure46" httpUri="https://binary.pensoft.net/fig/816434" pageId="0" pageNumber="1">46A, B</figureCitation>
); C large, heavily sclerotized, arced along top, similar to a wave, obscuring ~ 1/2 of E in CS, C covered by TS; E arises from large TL with anterior extension, hook shaped, originating at 7
<normalizedToken id="BF1CFF13C67BBB0B767D26DB7BA4C831" originalValue="oclock">o'clock</normalizedToken>
, ending at 1
<normalizedToken id="2CDBE92CE8DDBFA9B538741564F942F4" originalValue="oclock">o'clock</normalizedToken>
; MA large, roundish base, with small lobe retrolaterally, tapers only slightly to tip shaped like a
<normalizedToken id="0CDFD4A01F157EA8EBC5C43C8B81967E" originalValue="birds">bird's</normalizedToken>
head, heavily sclerotized at base, base covered with Sp.
</paragraph>
<caption id="34961BDD0BBD24B21343844CB75B53BE" doi="10.3897/zookeys.1150.93760.figure48" httpUri="https://binary.pensoft.net/fig/816436" pageId="0" pageNumber="1" start="Figure 48" startId="F56">
<paragraph id="E27C58AB7964AC964D4310BA8E736ED9" pageId="0" pageNumber="1">
<emphasis id="158C14F5A091FAD6A58BD50071755D63" bold="true" pageId="0" pageNumber="1">Figure 48.</emphasis>
<taxonomicName id="6AE3AE59E9669609EC1623026FD06DC4" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi">
<emphasis id="4B1FBBA834386C2223F037A69B30E24B" italics="true" pageId="0" pageNumber="1">Karaops markharveyi</emphasis>
</taxonomicName>
sp. nov. and members of the Pilbara-Gascoyne species group
<emphasis id="21B9EE19DF6E45CC961FF26063A09EDB" bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName id="9A2B0B69E67BC5D269B19D5B212ECEB1" authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops markharveyi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="markharveyi">
<emphasis id="CC31D37A31DCBE1A0575E6B20989A859" italics="true" pageId="0" pageNumber="1">Karaops markharveyi</emphasis>
</taxonomicName>
sp. nov., adult male, Judbarra National Park, Northern Territory (sel_1329, MAGNT A004885)
<emphasis id="5465A4732CDB0C7E09A3FFEE0A63FFFA" bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName id="21871E386A4A3EFC7490712E57B899DD" authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="1EBDBA68D53A2A852306B2638F874328" italics="true" pageId="0" pageNumber="1">Karaops</emphasis>
</taxonomicName>
sp., Timber Creek, Nackeroo Lookout, Northern Territory. Arrow indicates setal tufts
<emphasis id="3949D4B13B6A9D758F448D63322B53CC" bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName id="5EE9D2553B74D48568B8A941975BEEF7" authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops burbidgei" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="burbidgei">
<emphasis id="4E9D68FDABDFDEDE35B2CD339D2ED732" italics="true" pageId="0" pageNumber="1">Karaops burbidgei</emphasis>
</taxonomicName>
, paratype female, Barrow Island (WAM T76698)
<emphasis id="EA66D4DDBE854DA2BEB2DADD80496F75" bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName id="FF4703D7BF07090C6EF79F0C1B59E624" authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops burbidgei" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="burbidgei">
<emphasis id="FAA2F7F8B4B2778337BD6C58083680AB" italics="true" pageId="0" pageNumber="1">Karaops burbidgei</emphasis>
</taxonomicName>
, holotype male, Barrow Island (WAM T55000). Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E92F36282080C6DF99F9B6FF144EB8D7" pageId="0" pageNumber="1" type="variation">
<paragraph id="5DEEC7877E224AF5BB6ADDFEB76A6F75" pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph id="693F86B1A6EE48D4CBDE6F46151C1C01" pageId="0" pageNumber="1">
Females (
<emphasis id="4D63566F0BB0378291D83EEB74BDC112" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 3): In sel_1253, the AME and PME are equal in size, but in sel_1249 and sel_1332, the PME are larger than the AME. Leg III is longest in sel_1253, leg II is longest in sel_1249; however, leg II and leg III are very similar in length. Body length 3.07-4.09. Colors slightly vary, sel_1253 is paler than sel_1249. There is a lot of variability in genitalia - in fact, of the three, none are identical (Figs
<figureCitation id="B96FBF41763676CF532ECE456C934911" captionStart="Figure 44" captionStartId="F52" captionText="Figure 44. Karaops markharveyi sp. nov. A paratype male, 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1250, WAM T 155626) B adult male, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) C holotype female, epigyne, ~ 4.6 km from Drysdale River Road, Gibb River Road, Western Australia (sel _ 1249, WAM T 155625) D same, endogyne E paratype female, epigyne, Gibb River Road, Western Australia (sel _ 1253, WAM T 155629) F same, endogyne G adult male, Judbarra National Park, Northern Territory (sel _ 1330, MAGNT A 004886) H adult female, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888). Scale bars: 0.5 mm (C-F); 1 mm (B, G, H)." figureDoi="10.3897/zookeys.1150.93760.figure44" httpUri="https://binary.pensoft.net/fig/816432" pageId="0" pageNumber="1">44C-F</figureCitation>
,
<figureCitation id="ECFB2C8BAD8F555C5143456A2B5BFCEC" captionStart="Figure 47" captionStartId="F55" captionText="Figure 47. Karaops markharveyi sp. nov. A holotype female, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1249, WAM T 155625) B habitat, Judbarra National Park, Northern Territory C holotype female, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1249, WAM T 155625) D epigyne, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888) E same, endogyne. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure47" httpUri="https://binary.pensoft.net/fig/816435" pageId="0" pageNumber="1">47D, E</figureCitation>
). In sel_1249 the accessory bulbs are very close together, the accessory bulbs are slanted anteriorly from medial to lateral, in sel_1253 the accessory bulbs are much further apart - nearly 3/4 of one accessory bulb diameter. The spermathecae are horizontal. In sel_1249, the copulatory openings are located in a small depression, in sel_1253, there is not much of a depression, and in sel_1332, the depression more conspicuous, uterus externus and pdf quite different, the accessory bulbs are very close together, the spermathecae are horizontal, and the fertilization ducts are of a completely different shape.
</paragraph>
<paragraph id="1FE3AC94B51C495E0ED7E0D87BE17B59" pageId="0" pageNumber="1">
Males (
<emphasis id="68B6B65906E9D88362384BE53826ABC2" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 4): In sel_1329, the AME=PME, whereas in sel_1250 and sel_1330, the PME are larger than the AME, and there is some variation in leg spination. Body length is 3.10-3.36. sel_1250 is a little paler, more orangey pink, but the pattern the same.
</paragraph>
</subSubSection>
<subSubSection id="4EC81D7205FCF2E02CC819D141F6A354" pageId="0" pageNumber="1" type="etymology">
<paragraph id="2EE76981099573EDEF32309A3EF415CC" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="9C423BD9CE7D13C357B881E64676B6FF" pageId="0" pageNumber="1">This species is named after Mark S. Harvey, friend, mentor, and person responsible for getting me into the mess of Strayan selenopids. Name in genitive case.</paragraph>
</subSubSection>
<subSubSection id="F0F54B6FCFEF524C5662DF9A67C3E52D" pageId="0" pageNumber="1" type="distribution">
<paragraph id="5442C43705BBCD04392113026F780FA8" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="96828477C9A1FF7ED00D9E9C9B3C905B" pageId="0" pageNumber="1">
Known from along the Gibb River Road, northeastern Western Australia (Fig.
<figureCitation id="6539C7C03205A5D8A057BBE1FAFF1A31" captionStart="Figure 42" captionStartId="F50" captionText="Figure 42. Karaops mareeba sp. nov. and Karaops markharveyi sp. nov. A Karaops mareeba sp. nov., paratype female, Desailly Range, Queensland (QMS 120908) B Karaops mareeba sp. nov., holotype female, Desailly Range, Queensland (QMS 110872) C same, epigyne D Karaops markharveyi sp. nov., 4.6 km from Drysdale River Road, Gibb River Road, Western Australia, holotype female (sel _ 1249, WAM T 155625) E Karaops mareeba sp. nov., holotype female, endogyne, Desailly Range, Queensland (QMS 110872) F sandstone ridge habitat of Karaops markharveyi sp. nov. G Karaops mareeba sp. nov., holotype female, epigyne, caudal, Desailly Range, Queensland (QMS 110872). Scale bars: 0.5 mm (C, E, G); 2 mm (A, B, D)." figureDoi="10.3897/zookeys.1150.93760.figure42" httpUri="https://binary.pensoft.net/fig/816430" pageId="0" pageNumber="1">42F</figureCitation>
), and Judbarra National Park, northwestern Northern Territory (Figs
<figureCitation id="83F04C900485D7B7BA9F37AA176B717E" captionStart="Figure 46" captionStartId="F54" captionText="Figure 46. Karaops markharveyi sp. nov. and Karaops sp. A paratype male, palp, ventral, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1250, WAM T 155626) B same, retrolateral C same, dorsal D habitat of Karaops sp., Timber Creek, Nackeroo Lookout, Northern Territory E Karaops markharveyi sp. nov., palp, dorsal, tilted forward to see chemosensory brush, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) F habitat of Karaops markharveyi sp. nov., Judbarra National Park, Northern Territory. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure46" httpUri="https://binary.pensoft.net/fig/816434" pageId="0" pageNumber="1">46F</figureCitation>
,
<figureCitation id="E8B3E6D07F107995DFC2C9051541DECB" captionStart="Figure 47" captionStartId="F55" captionText="Figure 47. Karaops markharveyi sp. nov. A holotype female, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1249, WAM T 155625) B habitat, Judbarra National Park, Northern Territory C holotype female, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1249, WAM T 155625) D epigyne, Judbarra National Park, Northern Territory (sel _ 1332, MAGNT A 004888) E same, endogyne. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure47" httpUri="https://binary.pensoft.net/fig/816435" pageId="0" pageNumber="1">47B</figureCitation>
) (Map
<figureCitation id="D554BEA89A79D15E3D515F9AF5CBAFC3" captionStart="Map 8" captionStartId="F44" captionText="Map 8. Species that are not currently placed in a species group. Diamonds (juveniles): blue = Little Mertens, red = Ruby Plains, pink = Nackeroo Lookout; Circles: yellow = Karaops yumbubaarnji sp. nov., green = Karaops markharveyi sp. nov., purple = Karaops kennerleyorum sp. nov., blue = Karaops madhawundu sp. nov., red = Karaops mareeba sp. nov. Juveniles are treated as distinct species based on molecular data." figureDoi="10.3897/zookeys.1150.93760.map8" httpUri="https://binary.pensoft.net/fig/816424" pageId="0" pageNumber="1">8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="48DB306A15B6E2C1313FAFF8C4A9251B" pageId="0" pageNumber="1" type="natural history">
<paragraph id="6EC15F848D40B69B3280F5617B8B5F9F" pageId="0" pageNumber="1">Natural history.</paragraph>
<paragraph id="03D813BE1E3BD86532A479EF24A939DE" pageId="0" pageNumber="1">The collecting localities for this species occur in the Pentecost subregion of the Central Kimberley, the Mitchell subregion of the Northern Kimberley (but very close to a junction with subregion Pentecost from the Central Kimberley region and the Berkeley subregion of the North Kimberley), and the Victoria Bonaparte bioregion and the Keep subregion. The Pentecost subregion has a dry, hot tropical and sub-humid to semi-arid climate with summer rainfall of 750-1000 mm. The area is a savannah woodland of eucalypts and hummock grasses. These spiders were found under sandstone, and it is suggested that these sandstone communities may be areas of high diversity. There have been no systematic faunal surveys of the area, and ecological and life history data of organisms are missing.</paragraph>
<paragraph id="03016537996A239D8AA3F7364D1A7F08" pageId="0" pageNumber="1">
The climate of the Mitchell subregion of the Northern Kimberley bioregion is dry, hot tropical to sub-humid with 1100-1500 mm of mostly summer rain. The area is home to many endemic vertebrates and plants, though little is known about the terrestrial arthropods. Like the Pentecost subregion of the Central Kimberley, it is thought that the sandstone communities may be highly diverse. The Berkeley subregion of the North Kimberley bioregion has a dry, hot tropical sub-humid climate with high summer rainfall (
<bibRefCitation id="4E1810D5160E7281CB6A4AAF28419AFB" author="Graham, G" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="1" publicationUrl="https://www.dpaw.wa.gov.au/images/documents/about/science/projects/waaudit/north_kimberley02_p506-512.pdf" refId="B29" refString="Graham, G, 2001g. North Kimberley 2 (NK2 - Berkeley subregion). In: Department of Conservation and Land Management A Biodiversity Audit of Western Australia's 53 Biogeographic Subregions in 2002, 506-512. https://www.dpaw.wa.gov.au/images/documents/about/science/projects/waaudit/north_kimberley02_p506-512.pdf" title="North Kimberley 2 (NK 2 - Berkeley subregion). In: Department of Conservation and Land Management A Biodiversity Audit of Western Australia's 53 Biogeographic Subregions in 2002, 506 - 512." url="https://www.dpaw.wa.gov.au/images/documents/about/science/projects/waaudit/north_kimberley02_p506-512.pdf" year="2001 g">Graham 2001g</bibRefCitation>
).
</paragraph>
<paragraph id="71AE0C33C935DE9298E14A8EDDFBB2A3" pageId="0" pageNumber="1">The third locality is the Keep subregion of the Victoria Bonaparte bioregion. The part of this bioregion in Western Australia is not divided into subregions. In the Northern Territory, this is considered the Keep subregion. Judbarra National Park is in the Victoria Basin and has many sandstone ranges. It has a tropical semi-arid climate with summer monsoons.</paragraph>
<paragraph id="147B153C62160AC60C74C0D95D550F0C" pageId="0" pageNumber="1">All specimens were collected as immatures beneath sandstone rocks during the day or on/under the rocks at dusk in the drier, cooler time of the year. The were reared in captivity, with most maturing in the wetter, hotter time of the year. Although this is a small sampling, the females matured after almost all of the adult males were dead. This could be indicative of many things - in captivity, the molting/feeding/growing does not reflect what happens in nature, there are other males that were not collected that would overlap with the females, in nature the females would live until the following season when males matured, etc. Given what we know from other studies, it is likely that there are adults all year round, and overlaps do occur at different times which is not reflected in the small sample (Suppl. material 2: tables S1, S14).</paragraph>
</subSubSection>
<subSubSection id="1DAB86E5D07F5E54FA55B3E4B5D6E8E6" pageId="0" pageNumber="1" type="discussion">
<paragraph id="1F5FB1D6467D2E7BDB062C7A69C3ED54" pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph id="1D0C54E77AB094C708E99B5F63D3482E" pageId="0" pageNumber="1">
Given the dissimilarity of the female genitalia of specimens from the three different localities, it was assumed that they were three different species. Therefore, finding corresponding differences in the males was expected. However, there are no differences in the males, including the palps, which have been thoroughly examined and expanded. DNA data indicate that there is no mixing between the three populations, and that they may be related to a species from Timber Creek collected at Nackeroo Lookout (Figs
<figureCitation id="579696B6AD03169477D3D8FA3B3F9B34" captionStart="Figure 46" captionStartId="F54" captionText="Figure 46. Karaops markharveyi sp. nov. and Karaops sp. A paratype male, palp, ventral, Gibb River Road, ~ 4.6 km from Drysdale River Road, Western Australia (sel _ 1250, WAM T 155626) B same, retrolateral C same, dorsal D habitat of Karaops sp., Timber Creek, Nackeroo Lookout, Northern Territory E Karaops markharveyi sp. nov., palp, dorsal, tilted forward to see chemosensory brush, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) F habitat of Karaops markharveyi sp. nov., Judbarra National Park, Northern Territory. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure46" httpUri="https://binary.pensoft.net/fig/816434" pageId="0" pageNumber="1">46D</figureCitation>
,
<figureCitation id="63520ECEB3B151F00AE57B4439330CC5" captionStart="Figure 48" captionStartId="F56" captionText="Figure 48. Karaops markharveyi sp. nov. and members of the Pilbara-Gascoyne species group A Karaops markharveyi sp. nov., adult male, Judbarra National Park, Northern Territory (sel _ 1329, MAGNT A 004885) B Karaops sp., Timber Creek, Nackeroo Lookout, Northern Territory. Arrow indicates setal tufts C Karaops burbidgei, paratype female, Barrow Island (WAM T 76698) D Karaops burbidgei, holotype male, Barrow Island (WAM T 55000). Scale bars: 1 mm." figureDoi="10.3897/zookeys.1150.93760.figure48" httpUri="https://binary.pensoft.net/fig/816436" pageId="0" pageNumber="1">48B</figureCitation>
; Suppl. material 1). Some analyses even render the Judbarra population and the Gibb populations paraphyletic (Suppl. material 1). Because there are no corroborating morphological data, these three populations are currently considered to be the same species, but with more sampling, this is likely to change. These are among the smallest
<taxonomicName id="6E3D021362D49B053338E75822EF840E" authorityName="Simon" authorityYear="1897" class="Arachnida" family="Selenopidae" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Selenopidae</taxonomicName>
, with the male paratype being the smallest known (3.10 mm).
</paragraph>
</subSubSection>
</treatment>
</document>