137 lines
18 KiB
XML
137 lines
18 KiB
XML
<document id="1A04DC856CFCA621906369018205DC29" ID-CLB-Dataset="5618" ID-DOI="10.11646/zootaxa.275.1.1" ID-GBIF-Dataset="9322f2b2-a384-40bf-9d57-d789c76ce5c4" ID-ISSN="11755334" ID-Zenodo-Dep="5019235" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624466010041" checkinUser="felipe" docAuthor="PRATHER, AYSHA L." docDate="2003" docId="546087A8FFD52A71FEA9FA40F894E477" docLanguage="en" docName="zt00275.pdf" docOrigin="Zootaxa 275 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.275.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Phylloicus llaviuco Prather 2003, new species" docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="A859FFD0FF952A30FFA1FF8CFC30E154" masterDocTitle="Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)" masterLastPageNumber="214" masterPageNumber="1" pageNumber="65" updateTime="1699020181560" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="769FFC69B8070D7186E97A6526EF66A5">Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)</mods:title>
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<mods:namePart id="D2718B3467E4EF711EF0CE2A4F68A6DC">PRATHER, AYSHA L.</mods:namePart>
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<mods:title id="AEBA91F681E5B94861C8C30691198751">Zootaxa</mods:title>
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<mods:date id="2E88CE1BD9099E53B04F7B276B2E8CE7">2003</mods:date>
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<mods:number id="15E149383555EA92155D7A8C18A3BD1E">2003-08-29</mods:number>
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<mods:number id="C930CF867182C14A0E6B6F09E72502F9">275</mods:number>
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<mods:url id="15904D08AC124318B54C526A82872D8B">https://biotaxa.org/Zootaxa/article/view/zootaxa.275.1.1</mods:url>
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<mods:identifier id="BA60F50204669AEA6A6BE5945228C65A" type="DOI">10.11646/zootaxa.275.1.1</mods:identifier>
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<treatment id="546087A8FFD52A71FEA9FA40F894E477" ID-DOI="http://doi.org/10.5281/zenodo.5081049" ID-GBIF-Taxon="183402338" ID-Zenodo-Dep="5081049" LSID="urn:lsid:plazi:treatment:546087A8FFD52A71FEA9FA40F894E477" httpUri="http://treatment.plazi.org/id/546087A8FFD52A71FEA9FA40F894E477" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
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<subSubSection id="94D36535FFD52A70FEA9FA40FEA0E4B2" box="[264,656,1484,1510]" pageId="64" pageNumber="65" type="nomenclature">
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<paragraph id="DC7636BEFFD52A70FEA9FA40FEA0E4B2" blockId="64.[264,656,1484,1545]" box="[264,656,1484,1510]" pageId="64" pageNumber="65">
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<heading id="873E81D2FFD52A70FEA9FA40FEA0E4B2" bold="true" box="[264,656,1484,1510]" fontSize="11" level="1" pageId="64" pageNumber="65" reason="1">
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<emphasis id="EEBDEAACFFD52A70FEA9FA40FEA0E4B2" bold="true" box="[264,656,1484,1510]" pageId="64" pageNumber="65">
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<taxonomicName id="1BC94D3DFFD52A70FEA9FA40FDC4E4B2" authority="Prather, 2003" authorityName="Prather" authorityYear="2003" box="[264,500,1484,1510]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="llaviuco" status="sp. nov.">Phylloicus llaviuco</taxonomicName>
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,
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<taxonomicNameLabel id="F58E57D7FFD52A70FDA1FA40FEA0E4B2" box="[512,656,1484,1510]" pageId="64" pageNumber="65" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="94D36535FFD52A71FEA9FA63FE28E5D7" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="description">
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<paragraph id="DC7636BEFFD52A70FEA9FA63FD6BE75D" blockId="64.[264,656,1484,1545]" box="[264,347,1519,1545]" pageId="64" pageNumber="65">
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<figureCitation id="44F22A3BFFD52A70FEA9FA63FD6BE75D" box="[264,347,1519,1545]" captionStart="FIGURE 63" captionStartId="166.[264,368,1512,1536]" captionTargetBox="[264,1319,333,1488]" captionTargetId="figure-35@166.[263,1323,326,1488]" captionTargetPageId="166" captionText="FIGURE 63. Phylloicus llaviuco. Male (holotype): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5019364" httpUri="https://zenodo.org/record/5019364/files/figure.png" pageId="64" pageNumber="65">Fig. 63</figureCitation>
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</paragraph>
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<paragraph id="DC7636BEFFD52A70FEA9F9B3FFEBE675" blockId="64.[264,1323,1599,1905]" pageId="64" pageNumber="65">
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This species is known only from a single male, with broken antennae and wings badly rubbed. Thus, wing ornamentation and antennal length are incompletely known. The phallic endotheca is not fully everted, so I am unable to describe the membranous lobes. This species is similar in size and wing coloration to
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<taxonomicName id="1BC94D3DFFD52A70FC96F93BFF9FE784" box="[823,943,1719,1744]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="magnus">
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<emphasis id="EEBDEAACFFD52A70FC96F93BFF9FE784" box="[823,943,1719,1744]" italics="true" pageId="64" pageNumber="65">P. magnus</emphasis>
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</taxonomicName>
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; however, the male genitalia are distinctly different.
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<taxonomicName id="1BC94D3DFFD52A70FE59F953FEEFE7AC" authorityName="Prather Table Of Contents" authorityYear="2003" box="[504,735,1759,1784]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="llaviuco">
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<emphasis id="EEBDEAACFFD52A70FE59F953FEEFE7AC" box="[504,735,1759,1784]" italics="true" pageId="64" pageNumber="65">Phylloicus llaviuco</emphasis>
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</taxonomicName>
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is distinguished by the long digitate basodorsal process of tergum X, and the absence of abdominal coremata.
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</paragraph>
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<paragraph id="DC7636BEFFD52A70FE99F8A3FF3BE61D" blockId="64.[264,1323,1599,1905]" box="[312,779,1839,1865]" pageId="64" pageNumber="65">Adult. Forewing length 16.2 mm, n = 1.</paragraph>
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<paragraph id="DC7636BEFFD52A71FE99F8DBFED3E37F" blockId="64.[264,1323,1599,1905]" lastBlockId="65.[264,1323,284,1315]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
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Head chestnut brown, with dorsomesal crest of dark brown setae. Maxillary palps golden brown, covered with dark brown setae. Antenna dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax chestnut brown; ventrolateral thorax golden brown. Legs golden brown. Metathoracic leg of male with posterior fringe of long setae, setae pale. Tibial spur formula 2,4,4. Forewing flat; dark brown; with color patches; with golden setae at chord, in patch distal to crossvein Cu, and marking vein A
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<subScript id="404D34FBFFD42A71FE57FE63FE32E350" attach="right" box="[502,514,495,516]" fontSize="9" pageId="65" pageNumber="66">3</subScript>
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; with small golden spots marking nygma and thyridium. Hind wing basal brush present in male, dark brown.
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</paragraph>
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<paragraph id="DC7636BEFFD42A71FE99FDB5FFB3E50F" blockId="65.[264,1323,284,1315]" pageId="65" pageNumber="66">
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Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures absent, terga IIIV unmodified, without membranous lobes or sclerotized processes. Sternum VII without anteromesal process. Sternum VIII enclosing base of elongate sternum IX, or similar to anterior sterna, sternum IX not elongate; posteromesal process absent and posterior of segment semimembranous (
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<figureCitation id="44F22A3BFFD42A71FD7FFD55FF5DE3A7" box="[734,877,729,755]" captionStart="FIGURE 63" captionStartId="166.[264,368,1512,1536]" captionTargetBox="[264,1319,333,1488]" captionTargetId="figure-35@166.[263,1323,326,1488]" captionTargetPageId="166" captionText="FIGURE 63. Phylloicus llaviuco. Male (holotype): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5019364" httpUri="https://zenodo.org/record/5019364/files/figure.png" pageId="65" pageNumber="66">Fig. 63A, C</figureCitation>
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). Tergum IX with mesal ridge extending full length of segment; posterior margin with round narrow mesal projection (
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<figureCitation id="44F22A3BFFD42A71FB5AFC8DFD0DE217" captionStart="FIGURE 63" captionStartId="166.[264,368,1512,1536]" captionTargetBox="[264,1319,333,1488]" captionTargetId="figure-35@166.[263,1323,326,1488]" captionTargetPageId="166" captionText="FIGURE 63. Phylloicus llaviuco. Male (holotype): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5019364" httpUri="https://zenodo.org/record/5019364/files/figure.png" pageId="65" pageNumber="66">Fig. 63B</figureCitation>
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); lateral ridge present; dorsal pleural setae absent, ventral pleural setae approximately 15 (
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<figureCitation id="44F22A3BFFD42A71FE9BFCDDFD9EE23F" box="[314,430,849,875]" captionStart="FIGURE 63" captionStartId="166.[264,368,1512,1536]" captionTargetBox="[264,1319,333,1488]" captionTargetId="figure-35@166.[263,1323,326,1488]" captionTargetPageId="166" captionText="FIGURE 63. Phylloicus llaviuco. Male (holotype): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5019364" httpUri="https://zenodo.org/record/5019364/files/figure.png" pageId="65" pageNumber="66">Fig. 63A</figureCitation>
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); sternum IX (
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<figureCitation id="44F22A3BFFD42A71FDC6FCDDFEE8E23F" box="[615,728,849,875]" captionStart="FIGURE 63" captionStartId="166.[264,368,1512,1536]" captionTargetBox="[264,1319,333,1488]" captionTargetId="figure-35@166.[263,1323,326,1488]" captionTargetPageId="166" captionText="FIGURE 63. Phylloicus llaviuco. Male (holotype): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5019364" httpUri="https://zenodo.org/record/5019364/files/figure.png" pageId="65" pageNumber="66">Fig. 63C</figureCitation>
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). Preanal appendage shorter than tergum X, but greater than 2/3 length, widest near base, setae long, but not filamentous or longer than appendage (
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<figureCitation id="44F22A3BFFD42A71FE34FC2DFE14E2EF" box="[405,548,929,955]" captionStart="FIGURE 63" captionStartId="166.[264,368,1512,1536]" captionTargetBox="[264,1319,333,1488]" captionTargetId="figure-35@166.[263,1323,326,1488]" captionTargetPageId="166" captionText="FIGURE 63. Phylloicus llaviuco. Male (holotype): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5019364" httpUri="https://zenodo.org/record/5019364/files/figure.png" pageId="65" pageNumber="66">Fig. 63A, B</figureCitation>
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). Tergum X without basal lobes; basodorsal process long and digitate; basolateral processes absent; apex, in lateral view, truncate, in dorsal view, entire (
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<figureCitation id="44F22A3BFFD42A71FEB0FC7DFD97E55F" box="[273,423,1009,1035]" captionStart="FIGURE 63" captionStartId="166.[264,368,1512,1536]" captionTargetBox="[264,1319,333,1488]" captionTargetId="figure-35@166.[263,1323,326,1488]" captionTargetPageId="166" captionText="FIGURE 63. Phylloicus llaviuco. Male (holotype): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5019364" httpUri="https://zenodo.org/record/5019364/files/figure.png" pageId="65" pageNumber="66">Fig. 63A, B</figureCitation>
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). Harpago rounded; peglike setae few, apical (
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<figureCitation id="44F22A3BFFD42A71FC45FC7DF84BE55F" box="[996,1147,1009,1035]" captionStart="FIGURE 63" captionStartId="166.[264,368,1512,1536]" captionTargetBox="[264,1319,333,1488]" captionTargetId="figure-35@166.[263,1323,326,1488]" captionTargetPageId="166" captionText="FIGURE 63. Phylloicus llaviuco. Male (holotype): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5019364" httpUri="https://zenodo.org/record/5019364/files/figure.png" pageId="65" pageNumber="66">Fig. 63A, C</figureCitation>
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). Phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoeshaped (
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<figureCitation id="44F22A3BFFD42A71FD46FBCDFF46E50F" box="[743,886,1089,1115]" captionStart="FIGURE 63" captionStartId="166.[264,368,1512,1536]" captionTargetBox="[264,1319,333,1488]" captionTargetId="figure-35@166.[263,1323,326,1488]" captionTargetPageId="166" captionText="FIGURE 63. Phylloicus llaviuco. Male (holotype): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5019364" httpUri="https://zenodo.org/record/5019364/files/figure.png" pageId="65" pageNumber="66">Fig. 63D, E</figureCitation>
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).
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</paragraph>
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<paragraph id="DC7636BEFFD42A71FE99FBE5FE28E5D7" blockId="65.[264,1323,284,1315]" box="[312,536,1129,1155]" pageId="65" pageNumber="66">Female. Unknown.</paragraph>
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</subSubSection>
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||
<subSubSection id="94D36535FFD42A71FE99FB1DF894E477" pageId="65" pageNumber="66" type="materials_examined">
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||
<paragraph id="DC7636BEFFD42A71FE99FB1DFE78E587" blockId="65.[264,1323,284,1315]" pageId="65" pageNumber="66">
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<materialsCitation id="6CA13CE3FFD42A71FE99FB1DFE78E587" ID-GBIF-Occurrence="3319300537" collectingDate="1990-09-18" collectionCode="NMNH" country="Ecuador" elevation="3010" location="Rio Llaviuco" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Azuay" typeStatus="holotype">
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<emphasis id="EEBDEAACFFD42A71FE99FB1DFF3FE5FF" bold="true" box="[312,783,1169,1195]" pageId="65" pageNumber="66">
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||
<typeStatus id="0372881CFFD42A71FE99FB1DFD9AE5FF" box="[312,426,1169,1195]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
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male:
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||
<collectingCountry id="A4DE762EFFD42A71FDABFB1DFE93E5FF" box="[522,675,1169,1195]" name="Ecuador" pageId="65" pageNumber="66">ECUADOR</collectingCountry>
|
||
:
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||
<collectingRegion id="1E0DF85CFFD42A71FD17FB1DFF3AE5FF" box="[694,778,1169,1195]" country="Ecuador" name="Azuay" pageId="65" pageNumber="66">Azuay</collectingRegion>
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||
:
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||
</emphasis>
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||
<location id="D9166065FFD42A71FCBDFB1DFF8FE5FF" LSID="urn:lsid:plazi:treatment:546087A8FFD52A71FEA9FA40F894E477:D9166065FFD42A71FCBDFB1DFF8FE5FF" box="[796,959,1169,1195]" country="Ecuador" name="Rio Llaviuco" pageId="65" pageNumber="66" stateProvince="Azuay">Río Llaviuco</location>
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||
,
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||
<locationDeviation id="CDC42F9CFFD42A71FC70FB1DF884E5FF" box="[977,1204,1169,1195]" location="Quenca" pageId="65" pageNumber="66">
|
||
<quantity id="1B319B5BFFD42A71FC70FB1DF810E5FF" box="[977,1056,1169,1195]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="65" pageNumber="66" unit="km" value="16.0">16 km</quantity>
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||
W Quenca
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||
</locationDeviation>
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||
,
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||
<quantity id="1B319B5BFFD42A71FB64FB1DF916E5FF" box="[1221,1318,1169,1195]" metricMagnitude="3" metricUnit="m" metricValue="3.01" pageId="65" pageNumber="66" unit="m" value="3010.0">
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||
<elevation id="57E4D18DFFD42A71FB64FB1DF916E5FF" box="[1221,1318,1169,1195]" metricMagnitude="3" metricUnit="m" metricValue="3.01" pageId="65" pageNumber="66" unit="m" value="3010.0">3010 m</elevation>
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||
</quantity>
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||
,
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||
<date id="A877107EFFD42A71FEA9FB35FDB8E587" box="[264,392,1209,1235]" pageId="65" pageNumber="66" value="1990-09-18">
|
||
<collectingDate id="B833E996FFD42A71FEA9FB35FDB8E587" box="[264,392,1209,1235]" pageId="65" pageNumber="66" value="1990-09-18">18.ix.1990</collectingDate>
|
||
</date>
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||
, Flint (
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||
<collectionCode id="BAD8AE7BFFD42A71FE7AFB35FE0CE587" box="[475,572,1209,1235]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="65" pageNumber="66" type="Museum">NMNH</collectionCode>
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||
).
|
||
</materialsCitation>
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||
</paragraph>
|
||
<paragraph id="DC7636BEFFD42A71FE99FB6DFE0CE5AF" blockId="65.[264,1323,284,1315]" box="[312,572,1249,1275]" pageId="65" pageNumber="66">
|
||
Distribution.
|
||
<collectingCountry id="A4DE762EFFD42A71FE75FB6DFE08E5AF" box="[468,568,1249,1275]" name="Ecuador" pageId="65" pageNumber="66">Ecuador</collectingCountry>
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||
.
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||
</paragraph>
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<paragraph id="DC7636BEFFD42A71FE99FA85F894E477" blockId="65.[264,1323,284,1315]" box="[312,1188,1289,1315]" pageId="65" pageNumber="66">Etymology. Named for the Río Llaviuco, where this species was collected.</paragraph>
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</subSubSection>
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</treatment>
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</document> |