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<document id="AD9F4D650FE03F96EA50DB0426AB9385" ID-CLB-Dataset="5618" ID-DOI="10.11646/zootaxa.275.1.1" ID-GBIF-Dataset="9322f2b2-a384-40bf-9d57-d789c76ce5c4" ID-ISSN="1175­5334" ID-Zenodo-Dep="5019235" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624466010041" checkinUser="felipe" docAuthor="PRATHER, AYSHA L." docDate="2003" docId="546087A8FFCF2A6BFEA9F925FE13E44A" docLanguage="en" docName="zt00275.pdf" docOrigin="Zootaxa 275 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.275.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Phylloicus quadridigitatus Prather 2003, new species" docType="treatment" docVersion="9" lastPageNumber="92" masterDocId="A859FFD0FF952A30FFA1FF8CFC30E154" masterDocTitle="Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)" masterLastPageNumber="214" masterPageNumber="1" pageNumber="91" updateTime="1699020181560" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E5D1973433F6372E00442FA7833AAE28">Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)</mods:title>
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<mods:namePart id="51720FCEAB609F4D3D513FCF048E4C0B">PRATHER, AYSHA L.</mods:namePart>
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<treatment id="546087A8FFCF2A6BFEA9F925FE13E44A" ID-DOI="http://doi.org/10.5281/zenodo.5081033" ID-GBIF-Taxon="183402340" ID-Zenodo-Dep="5081033" LSID="urn:lsid:plazi:treatment:546087A8FFCF2A6BFEA9F925FE13E44A" httpUri="http://treatment.plazi.org/id/546087A8FFCF2A6BFEA9F925FE13E44A" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">
<subSubSection id="94D36535FFCF2A6AFEA9F925FEC0E797" box="[264,752,1705,1731]" pageId="90" pageNumber="91" type="nomenclature">
<paragraph id="DC7636BEFFCF2A6AFEA9F925FEC0E797" blockId="90.[264,752,1705,1766]" box="[264,752,1705,1731]" pageId="90" pageNumber="91">
<heading id="873E81D2FFCF2A6AFEA9F925FEC0E797" bold="true" box="[264,752,1705,1731]" fontSize="11" level="1" pageId="90" pageNumber="91" reason="1">
<emphasis id="EEBDEAACFFCF2A6AFEA9F925FEC0E797" bold="true" box="[264,752,1705,1731]" pageId="90" pageNumber="91">
<taxonomicName id="1BC94D3DFFCF2A6AFEA9F925FE64E797" authority="Prather, 2003" authorityName="Prather " authorityYear="2003" box="[264,596,1705,1731]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="quadridigitatus" status="sp. nov.">Phylloicus quadridigitatus</taxonomicName>
,
<taxonomicNameLabel id="F58E57D7FFCF2A6AFDC1F925FEC0E797" box="[608,752,1705,1731]" pageId="90" pageNumber="91" rank="species">new species</taxonomicNameLabel>
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</heading>
</paragraph>
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<subSubSection id="94D36535FFCF2A6BFEA9F940FE28E59A" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91" type="description">
<paragraph id="DC7636BEFFCF2A6AFEA9F940FD6BE7B2" blockId="90.[264,752,1705,1766]" box="[264,347,1740,1766]" pageId="90" pageNumber="91">
<figureCitation id="44F22A3BFFCF2A6AFEA9F940FD6BE7B2" box="[264,347,1740,1766]" captionStart="FIGURE 96" captionStartId="199.[264,367,1365,1389]" captionTargetBox="[265,1322,329,1341]" captionTargetId="figure-41@199.[265,1322,326,1341]" captionTargetPageId="199" captionText="FIGURE 96. Phylloicus quadridigitatus. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019466" httpUri="https://zenodo.org/record/5019466/files/figure.png" pageId="90" pageNumber="91">Fig. 96</figureCitation>
</paragraph>
<paragraph id="DC7636BEFFCF2A6BFEA9F88BFD42E00A" blockId="90.[264,1322,1799,1905]" lastBlockId="91.[264,1323,284,1310]" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">
This species is known only from a single male, with antennae and wings badly rubbed. Thus, any ornamentation of wings or antennae is unknown. The phallic endotheca is not fully everted, so I am unable to describe the membranous lobes.
<emphasis id="EEBDEAACFFCF2A6BFBABF8DBFD1AE061" italics="true" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">
<taxonomicName id="1BC94D3DFFCF2A6AFBABF8DBF8B5E624" box="[1034,1157,1879,1904]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">Phylloicus</taxonomicName>
quadridigitatus
</emphasis>
is distinguished by the two pairs of digitate processes of tergum X, from which it takes its name.
</paragraph>
<paragraph id="DC7636BEFFCE2A6BFE99FEE0FECDE0D2" blockId="91.[264,1323,284,1310]" box="[312,765,364,390]" pageId="91" pageNumber="92">Adult. Forewing length 7.3 mm, n = 1.</paragraph>
<paragraph id="DC7636BEFFCE2A6BFE99FE18F894E31A" blockId="91.[264,1323,284,1310]" pageId="91" pageNumber="92">Head black, including setal warts. Maxillary palps black. Antenna twice forewing length; dark brown. Prothorax black; dorsal pterothorax black; ventrolateral thorax chestnut brown. Femora chestnut brown; tibiae golden brown; tarsi golden brown. Metathoracic leg of male with posterior fringe of long setae. Tibial spur formula 2,2,2. Forewing flat; dark brown; without colored markings. Hind wing basal brush present in male.</paragraph>
<paragraph id="DC7636BEFFCE2A6BFE99FDD0F81DE5F2" blockId="91.[264,1323,284,1310]" pageId="91" pageNumber="92">
Male. Preterminalic abdominal terga without anteromesal notches. Corematic structures absent, terga III­V unmodified, without membranous lobes or sclerotized processes. Sternum VII without anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX with paired mesal ridges extending length of segment; posterior margin obtuse (
<figureCitation id="44F22A3BFFCE2A6BFE52FD70FE50E242" box="[499,608,764,790]" captionStart="FIGURE 96" captionStartId="199.[264,367,1365,1389]" captionTargetBox="[265,1322,329,1341]" captionTargetId="figure-41@199.[265,1322,326,1341]" captionTargetPageId="199" captionText="FIGURE 96. Phylloicus quadridigitatus. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019466" httpUri="https://zenodo.org/record/5019466/files/figure.png" pageId="91" pageNumber="92">Fig. 96B</figureCitation>
); lateral ridge absent; dorsal pleural setae approximately 10, ventral pleural setae absent (
<figureCitation id="44F22A3BFFCE2A6BFDF5FCA8FEF1E26A" box="[596,705,804,830]" captionStart="FIGURE 96" captionStartId="199.[264,367,1365,1389]" captionTargetBox="[265,1322,329,1341]" captionTargetId="figure-41@199.[265,1322,326,1341]" captionTargetPageId="199" captionText="FIGURE 96. Phylloicus quadridigitatus. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019466" httpUri="https://zenodo.org/record/5019466/files/figure.png" pageId="91" pageNumber="92">Fig. 96A</figureCitation>
); sternum IX without mesolateral ridges; sternum IX (
<figureCitation id="44F22A3BFFCE2A6BFEB0FCC0FD4FE232" box="[273,383,844,870]" captionStart="FIGURE 96" captionStartId="199.[264,367,1365,1389]" captionTargetBox="[265,1322,329,1341]" captionTargetId="figure-41@199.[265,1322,326,1341]" captionTargetPageId="199" captionText="FIGURE 96. Phylloicus quadridigitatus. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019466" httpUri="https://zenodo.org/record/5019466/files/figure.png" pageId="91" pageNumber="92">Fig. 96C</figureCitation>
). Preanal appendage less than 2/3 length of tergum X, widest near base, setae long, but not filamentous or longer than appendage (
<figureCitation id="44F22A3BFFCE2A6BFCCCFCF8FFCBE2DA" box="[877,1019,884,910]" captionStart="FIGURE 96" captionStartId="199.[264,367,1365,1389]" captionTargetBox="[265,1322,329,1341]" captionTargetId="figure-41@199.[265,1322,326,1341]" captionTargetPageId="199" captionText="FIGURE 96. Phylloicus quadridigitatus. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019466" httpUri="https://zenodo.org/record/5019466/files/figure.png" pageId="91" pageNumber="92">Fig. 96A, B</figureCitation>
). Tergum X without basal lobes; basodorsal process absent; basolateral processes long, length at least twice diameter; apex, in lateral view, with long digitate posterodorsal projection and short ventral projection, in dorsal view, cleft; with paired spine­like mesal setae at mid­length; posteriorly with two pairs of digitate processes, dorsomesal processes twice length of ventrolateral processes (
<figureCitation id="44F22A3BFFCE2A6BFE29FBB0FE25E502" box="[392,533,1084,1110]" captionStart="FIGURE 96" captionStartId="199.[264,367,1365,1389]" captionTargetBox="[265,1322,329,1341]" captionTargetId="figure-41@199.[265,1322,326,1341]" captionTargetPageId="199" captionText="FIGURE 96. Phylloicus quadridigitatus. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019466" httpUri="https://zenodo.org/record/5019466/files/figure.png" pageId="91" pageNumber="92">Fig. 96A, B</figureCitation>
). Harpago large, rounded; peglike setae many, apical (
<figureCitation id="44F22A3BFFCE2A6BFB2EFBB0F92DE502" box="[1167,1309,1084,1110]" captionStart="FIGURE 96" captionStartId="199.[264,367,1365,1389]" captionTargetBox="[265,1322,329,1341]" captionTargetId="figure-41@199.[265,1322,326,1341]" captionTargetPageId="199" captionText="FIGURE 96. Phylloicus quadridigitatus. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019466" httpUri="https://zenodo.org/record/5019466/files/figure.png" pageId="91" pageNumber="92">Fig. 96A, C</figureCitation>
). Phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped (
<figureCitation id="44F22A3BFFCE2A6BFC33FB00F810E5F2" box="[914,1056,1164,1190]" captionStart="FIGURE 96" captionStartId="199.[264,367,1365,1389]" captionTargetBox="[265,1322,329,1341]" captionTargetId="figure-41@199.[265,1322,326,1341]" captionTargetPageId="199" captionText="FIGURE 96. Phylloicus quadridigitatus. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019466" httpUri="https://zenodo.org/record/5019466/files/figure.png" pageId="91" pageNumber="92">Fig. 96D, E</figureCitation>
).
</paragraph>
<paragraph id="DC7636BEFFCE2A6BFE99FB38FE28E59A" blockId="91.[264,1323,284,1310]" box="[312,536,1204,1230]" pageId="91" pageNumber="92">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="94D36535FFCE2A6BFE99FB50FE13E44A" pageId="91" pageNumber="92" type="materials_examined">
<paragraph id="DC7636BEFFCE2A6BFE99FB50F926E5A2" blockId="91.[264,1323,284,1310]" box="[312,1302,1244,1270]" pageId="91" pageNumber="92">
<materialsCitation id="6CA13CE3FFCE2A6BFE99FB50F926E5A2" ID-GBIF-Occurrence="3319300378" box="[312,1302,1244,1270]" collectingDate="1927-10-29" collectingDateMax="1927-10-30" collectingDateMin="1927-10-29" collectionCode="NMW" collectorName="da Serra" country="Brazil" location="Alto da Serra" pageId="91" pageNumber="92" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="EEBDEAACFFCE2A6BFE99FB50FF36E5A2" bold="true" box="[312,774,1244,1270]" pageId="91" pageNumber="92">
<typeStatus id="0372881CFFCE2A6BFE99FB50FD9AE5A2" box="[312,426,1244,1270]" pageId="91" pageNumber="92" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="A4DE762EFFCE2A6BFE5FFB50FE43E5A2" box="[510,627,1244,1270]" name="Brazil" pageId="91" pageNumber="92">BRAZIL</collectingCountry>
:
<collectingRegion id="1E0DF85CFFCE2A6BFD21FB50FF31E5A2" box="[640,769,1244,1270]" country="Brazil" name="Sao Paulo" pageId="91" pageNumber="92">São Paulo</collectingRegion>
:
</emphasis>
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Alto
<collectorName id="713C5368FFCE2A6BFCE8FB50FF9DE5A2" box="[841,941,1244,1270]" pageId="91" pageNumber="92">da Serra</collectorName>
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,
<date id="A877107EFFCE2A6BFC19FB50F867E5A2" box="[952,1111,1244,1270]" pageId="91" pageNumber="92" value="1927-10-29" valueMax="1927-10-30" valueMin="1927-10-29">
<collectingDate id="B833E996FFCE2A6BFC19FB50F867E5A2" box="[952,1111,1244,1270]" pageId="91" pageNumber="92" value="1927-10-29" valueMax="1927-10-30" valueMin="1927-10-29">29­30.x.1927</collectingDate>
</date>
, Zerny (
<collectionCode id="BAD8AE7BFFCE2A6BFB18FB50F93AE5A2" box="[1209,1290,1244,1270]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="91" pageNumber="92">NMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="DC7636BEFFCE2A6BFE99FA88FE13E44A" blockId="91.[264,1323,284,1310]" box="[312,547,1284,1310]" pageId="91" pageNumber="92">
Distribution.
<collectingCountry id="A4DE762EFFCE2A6BFE75FA88FE2FE44A" box="[468,543,1284,1310]" name="Brazil" pageId="91" pageNumber="92">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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