treatments-xml/data/45/11/E4/4511E41DD837FFD2FDF8FA7AF196FD75.xml

<document ID-DOI="http://dx.doi.org/10.5852/ejt.2013.37" ID-GBIF-Dataset="9682b45b-5e22-4965-a293-e203660e0ed3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3806770" ID-ZooBank="ADF6394E-77B5-4309-87FE-4097FDE0A3FD" checkinTime="1588796067459" checkinUser="carolina" docAuthor="Rowson, Ben &amp; Tattersfield, Peter" docDate="2013" docId="4511E41DD837FFD2FDF8FA7AF196FD75" docLanguage="en" docName="ejt-2013-37.pdf.imf" docOrigin="European Journal of Taxonomy 37" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article" docStyleVersion="14" docTitle="Dadagulella pembensis Rowson &amp; Tattersfield 2013, sp. nov." docType="treatment" docUuid="4442210F-A097-400D-B34F-96A4B7677CE7" docUuidSource="ZooBank" docVersion="8" lastPageNumber="22" masterDocId="B9289C65D824FFC7FFA4FF80F253FFE7" masterDocTitle="Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies" masterLastPageNumber="46" masterPageNumber="1" pageNumber="20" updateTime="1683580915233" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title>Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rowson, Ben</mods:namePart>
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<mods:affiliation>Biodiversity &amp; Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk &amp; urn: lsid: zoobank. org: author: 05 D 46259 - DA 45 - 45 A 4 - 9127 - 5 AF 361 D 566 A 7</mods:affiliation>
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<mods:namePart>Tattersfield, Peter</mods:namePart>
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<mods:affiliation>Biodiversity &amp; Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk &amp; urn: lsid: zoobank. org: author: 8 C 55642 E-B 18 A- 461 B- 9 BF 9 - 31 CB 6 DB 7 A 070</mods:affiliation>
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<mods:title>European Journal of Taxonomy</mods:title>
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<mods:date>2013</mods:date>
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<mods:number>2013-02-04</mods:number>
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<taxonomicName ID-CoL="33XQN" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[604,885,1530,1557]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="pembensis" status="sp. nov.">
<emphasis bold="true" box="[604,885,1530,1557]" italics="true" pageId="19" pageNumber="20">Dadagulella pembensis</emphasis>
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<taxonomicNameLabel box="[893,983,1531,1557]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act: 
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4 
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7677CE7
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<paragraph blockId="19.[372,1216,1530,1628]" box="[506,1082,1601,1628]" pageId="19" pageNumber="20">
<figureCitation box="[506,596,1601,1628]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson &amp; Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="19" pageNumber="20">Figs 32</figureCitation>
, 
<figureCitation box="[608,640,1601,1627]" captionStart="Figs 43-50" captionStartId="34.[189,241,1895,1921]" captionTargetBox="[189,1399,682,1855]" captionTargetId="figure@34.[189,1399,682,1855]" captionTargetPageId="34" captionText="Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza." figureDoi="http://doi.org/10.5281/zenodo.3806792" httpUri="https://zenodo.org/record/3806792/files/figure.png" pageId="19" pageNumber="20">46</figureCitation>
, 
<figureCitation box="[652,684,1601,1627]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="19" pageNumber="20">60</figureCitation>
, 
<figureCitation box="[696,767,1601,1627]" captionStart="Figs 65-68" captionStartId="36.[189,241,1895,1921]" captionTargetBox="[189,1399,903,1855]" captionTargetId="figure@36.[189,1399,903,1855]" captionTargetPageId="36" captionText="Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.3806800" httpUri="https://zenodo.org/record/3806800/files/figure.png" pageId="19" pageNumber="20">66-68</figureCitation>
, 
<figureCitation box="[779,855,1601,1628]" captionStart="Figs 69-76" captionStartId="37.[189,241,1789,1815]" captionTargetBox="[202,1390,364,1721]" captionTargetId="figure@37.[189,1398,356,1749]" captionTargetPageId="37" captionText="Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806804" httpUri="https://zenodo.org/record/3806804/files/figure.png" pageId="19" pageNumber="20">70, 75</figureCitation>
, 
<figureCitation box="[867,938,1602,1628]" captionStart="Figs 77-83" captionStartId="38.[189,241,1789,1815]" captionTargetBox="[201,1381,776,1726]" captionTargetId="figure@38.[189,1398,759,1748]" captionTargetPageId="38" captionText="Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806808" httpUri="https://zenodo.org/record/3806808/files/figure.png" pageId="19" pageNumber="20">79-80</figureCitation>
, 
<figureCitation box="[950,983,1602,1628]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="19" pageNumber="20">84</figureCitation>
; 
<tableCitation box="[995,1082,1601,1628]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="19" pageNumber="20" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
</paragraph>
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<paragraph blockId="19.[189,1168,1667,1694]" box="[189,1168,1667,1694]" pageId="19" pageNumber="20">
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““ 
<emphasis box="[215,319,1668,1694]" italics="true" pageId="19" pageNumber="20">
<taxonomicName ID-CoL="7P4LY" authorityName="L. Pfeiffer" authorityYear="1856" box="[215,308,1668,1694]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">Gulella</taxonomicName>
”
</emphasis>
<taxonomicName authority="(Preston, 1910)" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[327,593,1668,1694]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="radius">
<emphasis box="[327,402,1668,1694]" italics="true" pageId="19" pageNumber="20">radius</emphasis>
(
<bibRefCitation author="Preston H. B." box="[417,583,1668,1694]" pageId="19" pageNumber="20" pagination="526 - 536" refId="ref27068" refString="Preston H. B. 1910. Additions to the non-Marine Molluscan Fauna of British and German East Africa and Lake Albert Edward. Annals and Magazine of Natural History, Series 8 6: 526 - 536. http: // dx. doi. org / 10.1080 / 00222931008692885" type="journal article" year="1910">
Preston, 
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)
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” – 
<treatmentCitation author="Rowson B. &amp; Warren B. H. &amp; Ngereza C. F." box="[632,946,1667,1694]" page="9" pageId="19" pageNumber="20" year="2010">
<bibRefCitation author="Rowson B. &amp; Warren B. H. &amp; Ngereza C. F." box="[632,946,1667,1694]" pageId="19" pageNumber="20" pagination="1 - 39" refId="ref27399" refString="Rowson B., Warren B. H. &amp; Ngereza C. F. 2010 b. Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island. ZooKeys 70: 1 - 39. http: // dx. doi. org / 10.3897 / zookeys. 70.762" type="journal article" year="2010">
Rowson 
<emphasis box="[736,794,1668,1694]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
<date box="[802,865,1667,1693]" pageId="19" pageNumber="20" value="2010">2010</date>
b: 9-10
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, 28-29, figs 60-61.
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</paragraph>
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<paragraph blockId="19.[189,324,1734,1760]" box="[189,324,1734,1760]" pageId="19" pageNumber="20">
<heading bold="true" box="[189,324,1734,1760]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis bold="true" box="[189,324,1734,1760]" pageId="19" pageNumber="20">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[189,861,1780,1807]" box="[189,861,1780,1807]" pageId="19" pageNumber="20">
From 
<collectingRegion box="[260,422,1780,1806]" country="Tanzania" name="Kusini Pemba" pageId="19" pageNumber="20">
Pemba 
<collectingCountry box="[347,422,1780,1806]" name="Iceland" pageId="19" pageNumber="20">Island</collectingCountry>
</collectingRegion>
, with the final ‘a’ elided for euphony.
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<heading bold="true" box="[189,492,1846,1873]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis bold="true" box="[189,492,1846,1873]" pageId="19" pageNumber="20">
<typeStatus box="[189,250,1847,1873]" pageId="19" pageNumber="20">Type</typeStatus>
material examined
</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[189,1399,1893,2027]" lastBlockId="20.[189,1399,267,370]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<materialsCitation ID-GBIF-Occurrence="2610647301" collectingDate="2009-02-07" collectingDateMax="2010" collectingDateMin="2009-02-07" collectionCode="I" country="Tanzania" latitude="-4.94" location="Tondooni" longLatPrecision="783" longitude="39.69" municipality="Tondooni" pageId="19" pageNumber="20" specimenCode="NMW.Z.2009.013.00227" specimenCount="1" specimenCount-adult="1" stateProvince="Pemba" typeStatus="holotype">
<collectingCountry box="[189,343,1894,1920]" name="Tanzania" pageId="19" pageNumber="20">TANZANIA</collectingCountry>
: 
<typeStatus box="[352,455,1893,1919]" pageId="19" pageNumber="20">holotype</typeStatus>
<specimenCode box="[461,762,1894,1920]" pageId="19" pageNumber="20">NMW.Z.2009.013.00227</specimenCode>
: 
<specimenCount box="[771,829,1893,1920]" pageId="19" pageNumber="20" type="adult">1 ad.</specimenCount>
, Ngezi Forest Reserve (
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, 
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), 
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<collectionCode box="[189,201,1930,1956]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="19" pageNumber="20" type="Herbarium">I</collectionCode>
., 
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, dry forest on dark, sandy soil on coral rag on 
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peninsula within reserve, leg. BR, B. H. Warren &amp; CFN, 
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<collectingDate box="[378,517,1965,1992]" pageId="19" pageNumber="20" value="2009-02-07">7 Feb. 2009</collectingDate>
</date>
; fig. 
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<bibRefCitation author="Rowson B. &amp; Warren B. H. &amp; Ngereza C. F." box="[634,894,1965,1992]" pageId="19" pageNumber="20" pagination="1 - 39" refId="ref27399" refString="Rowson B., Warren B. H. &amp; Ngereza C. F. 2010 b. Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island. ZooKeys 70: 1 - 39. http: // dx. doi. org / 10.3897 / zookeys. 70.762" type="journal article" year="2010">
Rowson 
<emphasis box="[736,792,1965,1991]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
(
<date box="[806,870,1965,1992]" pageId="19" pageNumber="20" value="2010">
<collectingDate box="[806,870,1965,1992]" pageId="19" pageNumber="20" value="2010">2010</collectingDate>
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b)
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</materialsCitation>
. 
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<collectionCode box="[1023,1101,1965,1991]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="19" pageNumber="20">NMW</collectionCode>
. 
<collectionCode box="[1104,1122,1965,1991]" country="Switzerland" name="Universität Zürich" pageId="19" pageNumber="20" type="Herbarium">Z</collectionCode>
.2009.013.00228: 2 ads, data as 
<typeStatus box="[271,378,2001,2027]" pageId="19" pageNumber="20">holotype</typeStatus>
. 
<typeStatus box="[385,499,2001,2027]" pageId="19" pageNumber="20">Paratypes</typeStatus>
<collectionCode box="[503,595,2001,2027]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="19" pageNumber="20" type="Museum">
<collectionCode45 box="[503,595,2001,2027]" pageId="19" pageNumber="20">MRAC</collectionCode45>
</collectionCode>
. 
<collectionCode box="[599,645,2001,2027]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="19" pageNumber="20">MT</collectionCode>
.803796-7: 2 ads, data as 
<typeStatus box="[922,1029,2001,2027]" pageId="19" pageNumber="20">holotype</typeStatus>
. 
<typeStatus box="[1037,1151,2001,2027]" pageId="19" pageNumber="20">Paratypes</typeStatus>
<collectionCode box="[1155,1271,2001,2027]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="20" type="Museum">
<collectionCode45 box="[1155,1271,2001,2027]" pageId="19" pageNumber="20">NHMUK</collectionCode45>
</collectionCode>
.20120261: 2 ads, data as 
<typeStatus box="[346,453,267,293]" pageId="20" pageNumber="21">holotype</typeStatus>
. 
<typeStatus box="[461,575,268,294]" pageId="20" pageNumber="21">Paratypes</typeStatus>
<collectionCode box="[581,654,268,294]" country="Kenya" httpUri="http://grbio.org/cool/h6fg-emd8" name="National Museums of Kenya" pageId="20" pageNumber="21" type="Museum">NMK</collectionCode>
: 2 ads, data as 
<typeStatus box="[819,926,267,293]" pageId="20" pageNumber="21">holotype</typeStatus>
. 
<typeStatus box="[935,1049,268,294]" pageId="20" pageNumber="21">Paratypes</typeStatus>
<collectionCode box="[1054,1122,268,294]" pageId="20" pageNumber="21">NMT</collectionCode>
: 2 ads, data as 
<typeStatus box="[1288,1395,267,293]" pageId="20" pageNumber="21">holotype</typeStatus>
. 
<typeStatus box="[189,303,306,332]" pageId="20" pageNumber="21">Paratypes</typeStatus>
<collectionCode box="[312,401,305,331]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="20" pageNumber="21" type="Museum">
<collectionCode45 box="[312,401,305,331]" pageId="20" pageNumber="21">NMSA</collectionCode45>
</collectionCode>
. 
<collectionCode box="[405,423,305,331]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="20" pageNumber="21" type="Herbarium">L</collectionCode>
8694/ 
<collectionCode box="[493,511,305,331]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="21">T</collectionCode>
3063: 2 ads, data as 
<typeStatus box="[754,861,305,331]" pageId="20" pageNumber="21">holotype</typeStatus>
. 
<typeStatus box="[873,987,306,332]" pageId="20" pageNumber="21">Paratypes</typeStatus>
<collectionCode box="[997,1090,306,332]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21" type="Museum">
<collectionCode45 box="[997,1090,306,332]" pageId="20" pageNumber="21">RMNH</collectionCode45>
</collectionCode>
. 
<collectionCode box="[1093,1162,306,332]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="20" pageNumber="21" type="Herbarium">MOL</collectionCode>
.288090: 2 ads, data as 
<typeStatus box="[221,328,344,370]" pageId="20" pageNumber="21">holotype</typeStatus>
.
</paragraph>
<paragraph blockId="20.[189,505,412,438]" box="[189,505,412,438]" pageId="20" pageNumber="21">
<heading bold="true" box="[189,505,412,438]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis bold="true" box="[189,505,412,438]" pageId="20" pageNumber="21">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[189,1399,461,794]" pageId="20" pageNumber="21">
<collectingCountry box="[189,343,462,488]" name="Tanzania" pageId="20" pageNumber="21">TANZANIA</collectingCountry>
: 
<collectionCode box="[357,435,462,488]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="21">NMW</collectionCode>
. 
<collectionCode box="[438,456,462,488]" country="Switzerland" name="Universität Zürich" pageId="20" pageNumber="21" type="Herbarium">Z</collectionCode>
.2009.013.00229: 15 juvs, data as 
<typeStatus box="[874,980,461,487]" pageId="20" pageNumber="21">holotype</typeStatus>
; one figured as fig. 
<quantity box="[1228,1292,461,487]" metricMagnitude="0" metricUnit="m" metricValue="1.5493999999999999" pageId="20" pageNumber="21" unit="in" value="61.0">61 in</quantity>
<bibRefCitation author="Rowson B. &amp; Warren B. H. &amp; Ngereza C. F." pageId="20" pageNumber="21" pagination="1 - 39" refId="ref27399" refString="Rowson B., Warren B. H. &amp; Ngereza C. F. 2010 b. Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island. ZooKeys 70: 1 - 39. http: // dx. doi. org / 10.3897 / zookeys. 70.762" type="journal article" year="2010">
Rowson 
<emphasis box="[189,251,500,526]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
(
<date box="[269,333,500,527]" pageId="20" pageNumber="21" value="2010">2010</date>
b)
</bibRefCitation>
. 
<collectionCode box="[371,449,500,526]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="21">NMW</collectionCode>
. 
<collectionCode box="[452,470,500,526]" country="Switzerland" name="Universität Zürich" pageId="20" pageNumber="21" type="Herbarium">Z</collectionCode>
.2009.013.00230: 3 ads, near Chwaka (
<geoCoordinate box="[942,1025,499,526]" degrees="5.39" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="555" value="-5.39">
5.39° 
<collectionCode box="[1010,1025,499,526]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="20" pageNumber="21" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate box="[1040,1140,500,526]" degrees="39.77" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="555" value="39.77">
39.77° 
<collectionCode box="[1121,1140,500,526]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="20" pageNumber="21" type="Herbarium">E</collectionCode>
</geoCoordinate>
), 
<collectingRegion box="[1163,1243,500,526]" country="Tanzania" name="Kusini Pemba" pageId="20" pageNumber="21">Pemba</collectingRegion>
<collectionCode box="[1253,1266,500,526]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="20" pageNumber="21" type="Herbarium">I</collectionCode>
., 
<collectingRegion box="[1288,1394,500,526]" country="Tanzania" name="Kaskazini Unguja" pageId="20" pageNumber="21">
<collectingCountry box="[1288,1394,500,526]" name="Tanzania" pageId="20" pageNumber="21">Zanzibar</collectingCountry>
</collectingRegion>
, clove and fruit tree woodland on dark, sandy soil, leg. 
<collectionCode box="[886,928,538,564]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="20" pageNumber="21">BR</collectionCode>
, 
<collectionCode box="[946,968,538,564]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="20" pageNumber="21" type="Herbarium">B</collectionCode>
. 
<collectionCode box="[971,997,538,564]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="20" pageNumber="21" type="Herbarium">H</collectionCode>
. Warren &amp; 
<collectionCode box="[1148,1208,538,564]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12862" lsid="urn:lsid:biocol.org:col:12862" name="Clifton College" pageId="20" pageNumber="21" type="Herbarium">CFN</collectionCode>
, 
<date box="[1226,1394,538,565]" pageId="20" pageNumber="21" value="2009-02-15">15 Feb. 2009</date>
. 
<collectionCode box="[189,267,576,602]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="21">NMW</collectionCode>
. 
<collectionCode box="[271,289,576,602]" country="Switzerland" name="Universität Zürich" pageId="20" pageNumber="21" type="Herbarium">Z</collectionCode>
.2009.013.00231: 3 ads, Msitu Mkuu Forest Reserve (
<geoCoordinate box="[942,1024,576,603]" degrees="5.00" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="555" value="-5.0">
5.00° 
<collectionCode box="[1009,1024,576,603]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="20" pageNumber="21" type="Herbarium">S</collectionCode>
</geoCoordinate>
, 
<geoCoordinate box="[1040,1139,576,602]" degrees="39.83" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="555" value="39.83">
39.83° 
<collectionCode box="[1120,1139,576,602]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="20" pageNumber="21" type="Herbarium">E</collectionCode>
</geoCoordinate>
), 
<collectingRegion box="[1163,1243,576,602]" country="Tanzania" name="Kusini Pemba" pageId="20" pageNumber="21">Pemba</collectingRegion>
<collectionCode box="[1253,1266,576,602]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="20" pageNumber="21" type="Herbarium">I</collectionCode>
., 
<collectingRegion box="[1288,1394,576,602]" country="Tanzania" name="Kaskazini Unguja" pageId="20" pageNumber="21">
<collectingCountry box="[1288,1394,576,602]" name="Tanzania" pageId="20" pageNumber="21">Zanzibar</collectingCountry>
</collectingRegion>
, moist forest on dark (not sandy) soil on coral rag in high forest in north of reserve, leg. 
<collectionCode box="[1173,1215,615,641]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="20" pageNumber="21">BR</collectionCode>
, 
<collectionCode box="[1225,1247,615,641]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="20" pageNumber="21" type="Herbarium">B</collectionCode>
. 
<collectionCode box="[1251,1277,615,641]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="20" pageNumber="21" type="Herbarium">H</collectionCode>
. Warren &amp; 
<collectionCode box="[189,249,652,678]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12862" lsid="urn:lsid:biocol.org:col:12862" name="Clifton College" pageId="20" pageNumber="21" type="Herbarium">CFN</collectionCode>
, 
<date box="[264,427,652,678]" pageId="20" pageNumber="21" value="2009-02-10">10 Feb. 2009</date>
. 
<collectionCode box="[441,519,652,678]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="21">NMW</collectionCode>
. 
<collectionCode box="[522,540,652,678]" country="Switzerland" name="Universität Zürich" pageId="20" pageNumber="21" type="Herbarium">Z</collectionCode>
.2009.013.00232: 3 ads, Ngezi Forest Reserve, 
<collectingRegion box="[1114,1194,652,678]" country="Tanzania" name="Kusini Pemba" pageId="20" pageNumber="21">Pemba</collectingRegion>
<collectionCode box="[1204,1217,653,679]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="20" pageNumber="21" type="Herbarium">I</collectionCode>
., 
<collectingRegion box="[1239,1345,652,678]" country="Tanzania" name="Kaskazini Unguja" pageId="20" pageNumber="21">
<collectingCountry box="[1239,1345,652,678]" name="Tanzania" pageId="20" pageNumber="21">Zanzibar</collectingCountry>
</collectingRegion>
, dry forest and thicket on dark, sandy soil on coral rag on coast of Tondooni peninsula within reserve, leg. 
<collectionCode box="[189,231,729,755]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="20" pageNumber="21">BR</collectionCode>
, 
<collectionCode box="[243,265,729,755]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="20" pageNumber="21" type="Herbarium">B</collectionCode>
. 
<collectionCode box="[269,295,729,755]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="20" pageNumber="21" type="Herbarium">H</collectionCode>
. Warren &amp; 
<collectionCode box="[429,489,729,755]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12862" lsid="urn:lsid:biocol.org:col:12862" name="Clifton College" pageId="20" pageNumber="21" type="Herbarium">CFN</collectionCode>
, 
<date box="[501,644,728,755]" pageId="20" pageNumber="21" value="2009-02-08">8 Feb. 2009</date>
. 
<collectionCode box="[656,735,729,755]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="20" pageNumber="21">NMW</collectionCode>
. 
<collectionCode box="[738,756,729,755]" country="Switzerland" name="Universität Zürich" pageId="20" pageNumber="21" type="Herbarium">Z</collectionCode>
.2009.013: Many additional ads and juvs from the 
<typeStatus box="[1348,1399,729,755]" pageId="20" pageNumber="21">type</typeStatus>
locality and other sites on 
<collectingRegion box="[498,578,767,793]" country="Tanzania" name="Kusini Pemba" pageId="20" pageNumber="21">Pemba</collectingRegion>
<date box="[585,603,767,793]" pageId="20" pageNumber="21" value="2009-01">
<collectionCode box="[585,598,767,793]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="20" pageNumber="21" type="Herbarium">I</collectionCode>
.
</date>
, 
<date box="[617,737,767,793]" pageId="20" pageNumber="21" value="2009-02">Feb. 2009</date>
; see table 2 of 
<bibRefCitation author="Rowson B. &amp; Warren B. H. &amp; Ngereza C. F." box="[912,1178,767,794]" pageId="20" pageNumber="21" pagination="1 - 39" refId="ref27399" refString="Rowson B., Warren B. H. &amp; Ngereza C. F. 2010 b. Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island. ZooKeys 70: 1 - 39. http: // dx. doi. org / 10.3897 / zookeys. 70.762" type="journal article" year="2010">
Rowson 
<emphasis box="[1016,1075,767,793]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
(
<date box="[1090,1154,767,794]" pageId="20" pageNumber="21" value="2010">2010</date>
b)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="description">
<paragraph blockId="20.[189,334,835,861]" box="[189,334,835,861]" pageId="20" pageNumber="21">
<heading bold="true" box="[189,334,835,861]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis bold="true" box="[189,334,835,861]" pageId="20" pageNumber="21">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[189,1399,884,1293]" pageId="20" pageNumber="21">
<collectionCode box="[189,257,885,911]" pageId="20" pageNumber="21">
<collectionCode45 box="[189,257,885,911]" pageId="20" pageNumber="21">SHELL</collectionCode45>
</collectionCode>
(
<figureCitation box="[277,369,884,911]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson &amp; Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="20" pageNumber="21">Figs 32</figureCitation>
, 
<figureCitation box="[383,415,884,910]" captionStart="Figs 43-50" captionStartId="34.[189,241,1895,1921]" captionTargetBox="[189,1399,682,1855]" captionTargetId="figure@34.[189,1399,682,1855]" captionTargetPageId="34" captionText="Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza." figureDoi="http://doi.org/10.5281/zenodo.3806792" httpUri="https://zenodo.org/record/3806792/files/figure.png" pageId="20" pageNumber="21">46</figureCitation>
, 
<figureCitation box="[429,462,884,911]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="20" pageNumber="21">60</figureCitation>
). Large (
<quantity box="[570,740,884,911]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.5" metricValueMin="4.8" pageId="20" pageNumber="21" unit="mm" value="5.15" valueMax="5.5" valueMin="4.8">
4.80 
<emphasis box="[624,634,885,910]" italics="true" pageId="20" pageNumber="21">-</emphasis>
5.50 mm
</quantity>
high x 2.00 
<emphasis box="[886,896,885,910]" italics="true" pageId="20" pageNumber="21">-</emphasis>
<quantity box="[896,1003,884,911]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="mm" value="2.5">2.50 mm</quantity>
wide), of 6.5 
<emphasis box="[1166,1176,885,910]" italics="true" pageId="20" pageNumber="21">-</emphasis>
7.5 whorls. Ovateacuminate, spire narrowly to broadly acuminate (spire angle 51 
<emphasis box="[928,938,924,949]" italics="true" pageId="20" pageNumber="21">-</emphasis>
65°). Apex sharply pointed. Embryonic whorls smoothly granulate. Later whorls with relatively coarse ribs (12 
<emphasis box="[999,1009,962,987]" italics="true" pageId="20" pageNumber="21">-</emphasis>
15 per mm on penultimate whorl). Sutures shallow, shell appearing relatively straight-sided. Umbilicus closed or nearly so. Peristome complete, or incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth and often another corresponding to the basal tooth. Dentition 6-fold, consisting of: one lamella-like parietal tooth; one slab-like palatal tooth, forming a parieto-palatal sinus; one basal denticle; a deep-set columellar baffle, sometimes folded, sub-bifid or sub-trifid; and two shallower, broad columellar denticles. Juvenile shells (
<figureCitation box="[707,794,1190,1217]" captionStart="Figs 43-50" captionStartId="34.[189,241,1895,1921]" captionTargetBox="[189,1399,682,1855]" captionTargetId="figure@34.[189,1399,682,1855]" captionTargetPageId="34" captionText="Figs 43-50. Juvenile shells of Dadagulella gen. nov. species, with dentition emphasized in basal view including earlier dentition visible through the shell. 43-45. D. radius radius, Mbudya I. 46. D. pembensis sp. nov., Ngezi. 47-48. D. browni browni, Mwanihana. 49.? D. ecclesiola sp. nov., Kimboza. 50. D. minareta sp. nov., Kimboza." figureDoi="http://doi.org/10.5281/zenodo.3806792" httpUri="https://zenodo.org/record/3806792/files/figure.png" pageId="20" pageNumber="21">Fig. 46</figureCitation>
) with 3-fold dentition, similar to some forms of 
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis italics="true" pageId="20" pageNumber="21">D. r. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[294,428,1228,1254]" pageId="20" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
: one parietal lamella; one basal tooth; and one columellar thickening. Earlier basal teeth are retained in juveniles.
</paragraph>
<paragraph blockId="20.[189,1018,1334,1361]" box="[189,1018,1334,1361]" pageId="20" pageNumber="21">CEPHALOPODIUM. Pale yellow, with apricot to orange tentacle retractors.</paragraph>
<paragraph blockId="20.[189,1398,1403,1467]" pageId="20" pageNumber="21">
SALIVARY GLANDS (
<figureCitation box="[402,489,1403,1429]" captionStart="Figs 69-76" captionStartId="37.[189,241,1789,1815]" captionTargetBox="[202,1390,364,1721]" captionTargetId="figure@37.[189,1398,356,1749]" captionTargetPageId="37" captionText="Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806804" httpUri="https://zenodo.org/record/3806804/files/figure.png" pageId="20" pageNumber="21">Fig. 70</figureCitation>
). United, soft, not tumid, elongate, Y-shaped; each duct leaving at the apex of the lobe and evenly thick throughout.
</paragraph>
<paragraph blockId="20.[189,1399,1509,1612]" pageId="20" pageNumber="21">
RADULA (
<figureCitation box="[298,429,1509,1536]" captionStart="Figs 65-68" captionStartId="36.[189,241,1895,1921]" captionTargetBox="[189,1399,903,1855]" captionTargetId="figure@36.[189,1399,903,1855]" captionTargetPageId="36" captionText="Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.3806800" httpUri="https://zenodo.org/record/3806800/files/figure.png" pageId="20" pageNumber="21">Figs 66-68</figureCitation>
). With a unicuspid central tooth and around 15 laterals in each half-row, gradually diminishing in size laterally. All laterals bicuspid, tricuspid, or quadricuspid, with outer cusps much smaller than inner cusps. Teeth delicate, short and flake-like at the ventral end of the radular ribbon.
</paragraph>
<paragraph blockId="20.[189,1399,1654,1909]" pageId="20" pageNumber="21">
GENITALIA (
<figureCitation box="[323,414,1654,1681]" captionStart="Figs 69-76" captionStartId="37.[189,241,1789,1815]" captionTargetBox="[202,1390,364,1721]" captionTargetId="figure@37.[189,1398,356,1749]" captionTargetPageId="37" captionText="Figs 69-76. Salivary glands and genitalia of Dadagulella gen. nov. species. 69. D. radius radius, Amboni. 70. D. pembensis sp. nov., Ngezi. 71. D. browni browni, Kosi Bay. 72. D. delta sp. nov., Mwanihana. 73. D. radius radius, Amboni. 74. D. browni browni, Kosi Bay. 75. D. pembensis sp. nov., Ngezi. 76. D. delta sp. nov., Mwanihana. Abbreviations: ad = anterior duct of salivary gland; ag = albumen gland; at = atrium; bc = bursa copulatrix; bd = bursa duct; dv = diverticulum of vas deferens; e = egg; fo = free oviduct; hdd/t = hermaphroditic duct diverticulum / talon; ma = muscular apex; p = penis; pd = posterior duct of salivary gland; pr = penial retractor muscle; v = vagina; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806804" httpUri="https://zenodo.org/record/3806804/files/figure.png" pageId="20" pageNumber="21">Figs 75</figureCitation>
, 
<figureCitation box="[428,506,1654,1680]" captionStart="Figs 77-83" captionStartId="38.[189,241,1789,1815]" captionTargetBox="[201,1381,776,1726]" captionTargetId="figure@38.[189,1398,759,1748]" captionTargetPageId="38" captionText="Figs 77-83. Penes and spermatophores of Dadagulella gen. nov. species. 77. D. radius radius, Amboni, penis with spermatophore in situ. 78. Same, hook and scoop in situ when hook lifted back to expose scoop. 79. D. pembensis sp. nov., Ngezi, penis. 80, same, spermatophore from penis of another individual. 81. D. delta, Mwanihana, penis. 82. D. browni browni, Kosi Bay, penis (diagrammatic). 83. D. radius crassa, Amboni, scoop magnified to show serrated edge. Abbreviations: cl = chitinized lobe; lh = large apical hook; ma = muscular apex; pr = penial retractor muscle; sc = apical scoop; st = spermatophore tail; sr = spermatophore reservoir; vd = vas deferens." figureDoi="http://doi.org/10.5281/zenodo.3806808" httpUri="https://zenodo.org/record/3806808/files/figure.png" pageId="20" pageNumber="21">79, 80</figureCitation>
). Vas deferens appearing thickened prior to insertion on penis, but actually with a short, broad parallel diverticulum. Penial sheath absent but with a thin sheath-like layer contiguous with wall of lower penis. Interior of penis with weak radial pilasters and small rhombic pads. Apical, muscular part of penis with a single large hook, associated with a spatulate “scoop” with microscopically serrated tip. Elsewhere in penis, one or two longitudinal rows of short, simple hooks mounted on rhombic pads. “Spermatophore” (see Discussion) present in penis of two Ngezi specimens; partially digested remains in bursa of another.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="multiple">
<paragraph blockId="20.[189,421,1951,1977]" box="[189,421,1951,1977]" pageId="20" pageNumber="21">
<heading bold="true" box="[189,421,1951,1977]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis bold="true" box="[189,421,1951,1977]" pageId="20" pageNumber="21">Range and habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[189,1142,2001,2027]" box="[189,1142,2001,2027]" pageId="20" pageNumber="21">
<collectingRegion box="[189,351,2001,2027]" country="Tanzania" name="Kusini Pemba" pageId="20" pageNumber="21">
Pemba 
<collectingCountry box="[276,351,2001,2027]" name="Iceland" pageId="20" pageNumber="21">Island</collectingCountry>
</collectingRegion>
, 
<collectingCountry box="[362,471,2001,2027]" name="Tanzania" pageId="20" pageNumber="21">Tanzania</collectingCountry>
, where widespread in forest and other vegetated habitats.
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="discussion">
<paragraph blockId="21.[189,303,267,293]" box="[189,303,267,293]" pageId="21" pageNumber="22">
<heading bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis bold="true" box="[189,303,267,293]" pageId="21" pageNumber="22">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="21.[189,1399,314,659]" pageId="21" pageNumber="22">
This species was discussed by 
<bibRefCitation author="Rowson B. &amp; Warren B. H. &amp; Ngereza C. F." box="[554,821,314,341]" pageId="21" pageNumber="22" pagination="1 - 39" refId="ref27399" refString="Rowson B., Warren B. H. &amp; Ngereza C. F. 2010 b. Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island. ZooKeys 70: 1 - 39. http: // dx. doi. org / 10.3897 / zookeys. 70.762" type="journal article" year="2010">
Rowson 
<emphasis box="[658,717,314,340]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(
<date box="[734,798,314,341]" pageId="21" pageNumber="22" value="2010">2010</date>
b)
</bibRefCitation>
, who suggested a thorough revision of the group to establish whether the 
<collectingRegion box="[472,552,349,375]" country="Tanzania" name="Kusini Pemba" pageId="21" pageNumber="22">Pemba</collectingRegion>
populations were part of a variable 
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[970,1080,349,375]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis box="[970,1080,349,375]" italics="true" pageId="21" pageNumber="22">D. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1087,1216,349,376]" pageId="21" pageNumber="22" rank="species">comb. nov.</taxonomicNameLabel>
or a subspecies or species endemic to 
<collectingRegion box="[459,543,384,410]" country="Tanzania" name="Kusini Pemba" pageId="21" pageNumber="22">Pemba</collectingRegion>
. This uncertainty was incorporated into their conclusions on endemism on the island (
<bibRefCitation author="Rowson B. &amp; Warren B. H. &amp; Ngereza C. F." box="[359,698,420,446]" pageId="21" pageNumber="22" pagination="1 - 39" refId="ref27399" refString="Rowson B., Warren B. H. &amp; Ngereza C. F. 2010 b. Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island. ZooKeys 70: 1 - 39. http: // dx. doi. org / 10.3897 / zookeys. 70.762" type="journal article" year="2010">
Rowson 
<emphasis box="[466,527,420,446]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
<date box="[536,599,420,446]" pageId="21" pageNumber="22" value="2010">2010</date>
b: 28-29
</bibRefCitation>
, 32). We find here that there is little or no overlap with 
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis italics="true" pageId="21" pageNumber="22">D. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[272,402,455,482]" pageId="21" pageNumber="22" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[409,444,455,481]" italics="true" pageId="21" pageNumber="22">s.l.</emphasis>
or other species, so conclude that this is an island endemic, which we here name 
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[189,348,491,517]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="pembensis" status="sp. nov.">
<emphasis box="[189,348,491,517]" italics="true" pageId="21" pageNumber="22">D. pembensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[357,449,491,517]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
Like specimens of 
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[683,819,491,517]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis box="[683,819,491,517]" italics="true" pageId="21" pageNumber="22">
D. r. 
<taxonomicName box="[744,819,491,517]" pageId="21" pageNumber="22" rank="species" species="radius" status="comb. nov.">radius</taxonomicName>
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[827,958,491,517]" pageId="21" pageNumber="22" rank="species">comb. nov.</taxonomicNameLabel>
on the opposite mainland at Amboni (e.g. 
<figureCitation box="[248,336,526,553]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="21" pageNumber="22">Figs 15</figureCitation>
, 
<figureCitation box="[348,380,526,553]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="21" pageNumber="22">53</figureCitation>
), 
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[401,559,526,552]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="pembensis" status="sp. nov.">
<emphasis box="[401,559,526,552]" italics="true" pageId="21" pageNumber="22">D. pembensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[566,656,527,553]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
is large and has a complex pattern of 6-fold dentition. However it is distinguishable from them by its even larger size, shallower sutures, and having on average an extra 0.5 whorls. It differs from 
<taxonomicName box="[501,786,597,623]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="subSpecies" species="minuscula" subSpecies="mahorana">
<emphasis box="[501,786,597,623]" italics="true" pageId="21" pageNumber="22">D. minuscula mahorana</emphasis>
</taxonomicName>
subsp. nov. which has similar dentition, in its larger size, shallower sutures, and stronger and more widely spaced ribs.
</paragraph>
</subSubSection>
</treatment>
</document>