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<mods:title id="6808B8B2FC33F5647DF9CC0CD68E4F58">Taxonomic revision of the Afrotropical Agabus raffrayi species group with the description of four new species (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="E689B761B9357248BA06FD8EE721D2C9">Englund, William F.</mods:namePart>
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<mods:namePart id="3D2E38651503A63D13557F6174FB5CC5">Njoroge, Laban</mods:namePart>
<mods:affiliation id="35F20515E50009130652540D034B8A34">National Museums of Kenya, Section of Invertebrate Zoology, Museum Hill, P. O. BOX 40658 - 00100, Nairobi, Kenya</mods:affiliation>
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<mods:namePart id="C63048F63435C2161B370E3E9859BDC7">Bistroem, Olof</mods:namePart>
<mods:affiliation id="EF51DF9098EDA3F672A4046D70E28A32">Finnish Museum of Natural History, Zoology Unit, P. O. Box 17, FI- 00014 University of Helsinki, Finland</mods:affiliation>
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<mods:namePart id="78BF9D80E0D79729E9395EA5952E790B">Miller, Kelly B.</mods:namePart>
<mods:affiliation id="EDC0588C9E9E491ACE9B57CE336D07EA">Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
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<mods:namePart id="9CDBB3CBF6EB59F6135B272C5182FD70">Bilton, David T.</mods:namePart>
<mods:affiliation id="2F341E45AE35FF7E5407A5870B826754">Marine Biology and Ecology Research Centre, School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth PL 4 8 AA, UK &amp; Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, Johannesburg 2006, South Africa</mods:affiliation>
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<mods:namePart id="5EDC407393A3504EFD4588DA834A751C">Bergsten, Johannes</mods:namePart>
<mods:affiliation id="D9E1F5C180C0E5DE621DCC5F6D64C95B">Swedish Museum of Natural History, Department of Zoology, Box 50007, SE- 10405 Stockholm, Sweden</mods:affiliation>
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<treatment id="2750ED0DFA485C54870787F934361C04" ID-GBIF-Taxon="167293858" LSID="urn:lsid:plazi:treatment:2750ED0DFA485C54870787F934361C04" httpUri="http://treatment.plazi.org/id/2750ED0DFA485C54870787F934361C04" lastPageNumber="45" pageId="0" pageNumber="45" scope_family="Dytiscidae" scope_order="Coleoptera">
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<paragraph id="271972191A07168135C3F7C6A57BB3CE" pageId="0" pageNumber="45">
<taxonomicName id="4B95EF454470F66223324CA9C6C6CE23" ID-CoL="7QDPQ" LSID="2750ED0D-FA48-5C54-8707-87F934361C04" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">Agabus raffrayi Sharp, 1882</taxonomicName>
<figureCitation id="F6930E0BACBF7C7FE32AF9D0A88B524D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the Agabus raffrayi group. Relief is expressed in meters above sea level." figureDoi="10.3897/zookeys.963.53470.figure1" httpUri="https://binary.pensoft.net/fig/444398" pageId="0" pageNumber="45">Figures 1</figureCitation>
<figureCitation id="8266D945A2AE6B34EA94D40DDEBD39F9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Agabus spp., metasternal wing and metacoxal plate in ventral view A A. pallidus B A. raffrayi. Dashed lines indicate measurements used; red: metasternal wing (WS), blue: metacoxal plate (WC)." figureDoi="10.3897/zookeys.963.53470.figure2" httpUri="https://binary.pensoft.net/fig/444399" pageId="0" pageNumber="45">, 2B</figureCitation>
<figureCitation id="5430073C5EB39E80A3DABDF82B593CCF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Agabus spp., protarsomeres in anterior view A A. dytiscoides B A. raffrayi. PT 4 L = measured distance for length of protarsomere 4; CL = measured distance for length of protarsal claw." figureDoi="10.3897/zookeys.963.53470.figure3" httpUri="https://binary.pensoft.net/fig/444400" pageId="0" pageNumber="45">, 3B</figureCitation>
<figureCitation id="384B6102B771C2343F9CD07ADE8B7265" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agabus spp., lateral view of tip of aedeagus A-C A. raffrayi, different specimens showing the variation in shape of the subapical tooth D A. austellus sp. nov. Note the prolonged section between subapical broadening and base of apical and subapical teeth in A. raffrayi (indicated with orange in the smaller outline illustrations)." figureDoi="10.3897/zookeys.963.53470.figure5" httpUri="https://binary.pensoft.net/fig/444402" pageId="0" pageNumber="45">, 5A-C</figureCitation>
<figureCitation id="6383B26847D6CD17FE829D85792E8AA5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">, 8D</figureCitation>
<figureCitation id="02E26AE02FE6173CA9C7B8526F108EC0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Agabus spp., hypomeron in lateral view A A. sjostedti B A. dytiscoides C A. anguluverpus sp. nov. D A. raffrayi. White arrows indicate hypomeron." figureDoi="10.3897/zookeys.963.53470.figure10" httpUri="https://binary.pensoft.net/fig/444407" pageId="0" pageNumber="45">, 10D</figureCitation>
<figureCitation id="2AF629B76EF0B3958F5EFDA64AE98F04" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">, 11D</figureCitation>
<figureCitation id="F76F64500AD52DE1DC9CDA53CEBB0767" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">, 11I</figureCitation>
<figureCitation id="4205CE2A5820096AEC909B6D8F8F73BE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">, 12</figureCitation>
<figureCitation id="75F18BD99F0CC03AE15F08FEB7B5E407" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">, 13</figureCitation>
<figureCitation id="AB7B834309C68DADAA4D29130F0D6031" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">, 14</figureCitation>
</paragraph>
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<subSubSection id="65564CC44083028B6A560E8324F99244" pageId="0" pageNumber="45" type="reference_group">
<paragraph id="FFDA6201582C3C1E0D7203CA7BF77CE9" pageId="0" pageNumber="45">
<taxonomicName id="205128A3C0CB9ADABCF137C3E304FB98" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">Agabus raffrayi</taxonomicName>
Sharp, 1882: 501-502
</paragraph>
<paragraph id="AF33A8FADF476DFD86FA03537963F619" pageId="0" pageNumber="45">
<taxonomicName id="D02328DB217AFFA199CB4203173C3637" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus limbicollis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="limbicollis">Agabus limbicollis</taxonomicName>
<normalizedToken id="FCC7D28DB33A4B450C5EB77232754E75" originalValue="Régimbart">Regimbart</normalizedToken>
, 1905: 224-225 (Syn. 
<bibRefCitation id="A9E9E7972B919B9F9A1149F346B5762D" DOI="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" author="Nilsson, AN" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="45" pagination="155 - 179" refId="B15" refString="Nilsson, AN, 1992a. A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists. Systematic Entomology 17: 155 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" title="A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists." url="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" volume="17" year="1992 a">Nilsson 1992a</bibRefCitation>
)
</paragraph>
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<subSubSection id="E66E47A1036CABD0E69174C883F8B930" pageId="0" pageNumber="45" type="type locality">
<paragraph id="772A934468178721A46B78A44409E7A9" pageId="0" pageNumber="45">Type locality.</paragraph>
<paragraph id="24D4C06D5C5DDA2C768278624F47F6AF" pageId="0" pageNumber="45">
<taxonomicName id="499F20F5EA48BF547BE0B5BE109D20E1" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="FDC8CFE61D0EC811511A567F80E92E3A" italics="true" pageId="0" pageNumber="45">raffrayi</emphasis>
</taxonomicName>
<normalizedToken id="15BDFF988DDFAC5322D2BC1F2026F345" originalValue="“Abyssinia”">&quot;Abyssinia&quot;</normalizedToken>
[Ethiopia]; 
<taxonomicName id="21C9FAA321B5AC8307512777FE41A9D8" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="limbicollis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="limbicollis">
<emphasis id="691964DDEE1B187A33533D550A491F79" italics="true" pageId="0" pageNumber="45">limbicollis</emphasis>
</taxonomicName>
&quot;Abyssinie: Auato, au bord du Nil Bleu, dans le Gindeberat&quot; [Ethiopia: Auato, on the banks of the Blue Nile, in the Gindeberat].
</paragraph>
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<subSubSection id="0BEC76D0B0ADE69D1553BADFB076CC94" pageId="0" pageNumber="45" type="type material">
<paragraph id="B86FA199FEC9100DF677E89EBF3DD6D1" pageId="0" pageNumber="45">Type material.</paragraph>
<paragraph id="745CF549928784EF4932D0B6AA807C67" pageId="0" pageNumber="45">
<emphasis id="39829AAA56C9A7DD57404CFE3BE76D6E" bold="true" italics="true" pageId="0" pageNumber="45">Lectotype</emphasis>
♂ of 
<taxonomicName id="02A26531019D79C78DD7763564115B5C" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="57CFE204096A95A98251BC99A7CA3C2B" italics="true" pageId="0" pageNumber="45">raffrayi</emphasis>
</taxonomicName>
(BMNH) labelled: &quot;♂ Abyssinia, Raffray 782&quot;, 
<normalizedToken id="3B7C725C590BAE08AC79660FE2B74314" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
, &quot;Sharp Coll 1905-313.&quot;, &quot;Type 782 
<taxonomicName id="D85A980194EBB3E1C834DCFED030F9A2" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">Agabus</taxonomicName>
Raffrayi n.sp. Abyssinia&quot;, &quot;LECTOTYPUS ♂ 
<taxonomicName id="18B6BFADF93C269849592BC05A43258B" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">Agabus raffrayi</taxonomicName>
Sharp, 1882 Des. A.Nilsson, 1989&quot;. 
<emphasis id="72DC9420126EB129BC283CA72C68FB5F" bold="true" italics="true" pageId="0" pageNumber="45">Lectotype</emphasis>
♂ of 
<taxonomicName id="6107C0ABCF2DAE63380E838E1338BA4F" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="limbicollis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="limbicollis">
<emphasis id="34722A7815FA89DC2018F7AE817A075D" italics="true" pageId="0" pageNumber="45">limbicollis</emphasis>
</taxonomicName>
(MNB) labelled: &quot;N.O. - Africa, Schoa, Falle O. Neumann S.&quot;, &quot;610 Falle&quot;, &quot; 
<taxonomicName id="280A7D13157148FCC3B746AB0A4A6C7A" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus limbicollis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="limbicollis">Agabus limbicollis</taxonomicName>
<normalizedToken id="142138F06CE456F7331B292E16CFEE98" originalValue="Rég">Reg</normalizedToken>
. Type.&quot;, &quot;LECTOTYPUS ♂ 
<taxonomicName id="9C4ED200ED36CE513C9E5E771E063CF3" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus limbicollis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="limbicollis">Agabus limbicollis</taxonomicName>
<normalizedToken id="F0CF474606AF27891DE8B64A61D783B6" originalValue="Régimbart">Regimbart</normalizedToken>
, 1905. Des. Nilsson -90&quot;.
</paragraph>
</subSubSection>
<subSubSection id="F6269A82CC3D357F1F1EC7D4723C4CEE" pageId="0" pageNumber="45" type="diagnosis">
<paragraph id="91B92158C3CDE3ECF9BD0BDBCF93D6EF" pageId="0" pageNumber="45">Diagnosis.</paragraph>
<paragraph id="CF3D1627C860E740BEC50D0C40E8EB30" pageId="0" pageNumber="45">
With a prolonged preapical section of male aedeagus and a pronotal hypomeron which is not visible in lateral view, this species is most similar to 
<taxonomicName id="98F5AE7B6896DAD5DB6EFC53ED6DF850" lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="44C3B691B1F07921C8C570D33C90CE9E" italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="1BC7925CED88FDB023270F0994D9ABEA" lsidName="A. ruwenzoricus" pageId="0" pageNumber="45" rank="species" species="ruwenzoricus">
<emphasis id="A1723D5BDC1005187CD63048417A0EE2" italics="true" pageId="0" pageNumber="45">A. ruwenzoricus</emphasis>
</taxonomicName>
. From the former it is separated by its broader metasternal wing (Figs 
<figureCitation id="1F456693ED317E31D276EF83822721BC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Agabus spp., metasternal wing and metacoxal plate in ventral view A A. pallidus B A. raffrayi. Dashed lines indicate measurements used; red: metasternal wing (WS), blue: metacoxal plate (WC)." figureDoi="10.3897/zookeys.963.53470.figure2" httpUri="https://binary.pensoft.net/fig/444399" pageId="0" pageNumber="45">2</figureCitation>
, 
<figureCitation id="8CD324046ADB7A2BFC8FB64E058F13F1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
) and from the latter by the lack of interocular spots (compare Fig. 
<figureCitation id="F13ED99320109E91D1DB7725FB6D8386" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agabus spp., head and pronotum in dorsal view A A. dytiscoides B A. ruwenzoricus C A. sjostedti D A. pallidus. Red dashed lines indicate measurements used for interocular distance and pronotal width." figureDoi="10.3897/zookeys.963.53470.figure4" httpUri="https://binary.pensoft.net/fig/444401" pageId="0" pageNumber="45">4D, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FFC603474E762CFA76DD6CAA2E6E07AD" pageId="0" pageNumber="45" type="description">
<paragraph id="B39D9EEFA08C5DF7B7C209A52328CED8" pageId="0" pageNumber="45">Description.</paragraph>
<paragraph id="3216FB6F0B6705D9C1AD219D37D59703" pageId="0" pageNumber="45">
Habitus as in Fig. 
<figureCitation id="D5A7708D69F950573B07462EF4FAB732" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">11D, I</figureCitation>
.
</paragraph>
<paragraph id="58B0F70D35B1BF7DDFF8B4931DE4831C" pageId="0" pageNumber="45">
<emphasis id="355FFA3DA881FD978A6764283C31A96B" bold="true" italics="true" pageId="0" pageNumber="45">Colour</emphasis>
: Head black, most specimens with a small rufous anterior area, interocular spots not present. Pronotum black with rufous margins. Elytra rufotestaceous to brown. Ventral surface black, hypomeron and epipleuron testaceous. Legs rufous to rufopiceous. Antennae and palpi testaceous to rufotestaceous.
</paragraph>
<paragraph id="60381FF136CC93B16C43B9A4396238DB" pageId="0" pageNumber="45">
<taxonomicName id="5DA172943758CC7C7C378273D362DC93" genus="Microreticulation" lsidName="Microreticulation" pageId="0" pageNumber="45" rank="genus">
<emphasis id="193FC76C47231D11BE4164EAD0447A84" bold="true" italics="true" pageId="0" pageNumber="45">Microreticulation</emphasis>
</taxonomicName>
: Medium impressed on head, pronotum and elytra, similar in both sexes. Composed of a mixture of small and somewhat larger, uneven meshes.
</paragraph>
<paragraph id="3D5F22F267A2BB0156EC4CBB3658274D" pageId="0" pageNumber="45">
<taxonomicName id="C51201EA311F4906410F33A3E52A33BB" genus="Structural" lsidName="Structural features" pageId="0" pageNumber="45" rank="species" species="features">
<emphasis id="5A32576C567485BF0719E4E1BA97DFAF" bold="true" italics="true" pageId="0" pageNumber="45">Structural features</emphasis>
</taxonomicName>
: Body length: 6.96-8.24 mm (see Table 
<tableCitation id="AF938185BE44C501448D06248161E2CA" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
). Hypomeron not visible in strict lateral view (Fig. 
<figureCitation id="76D5B828EEFDCF99A125AF399DC381AB" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Agabus spp., hypomeron in lateral view A A. sjostedti B A. dytiscoides C A. anguluverpus sp. nov. D A. raffrayi. White arrows indicate hypomeron." figureDoi="10.3897/zookeys.963.53470.figure10" httpUri="https://binary.pensoft.net/fig/444407" pageId="0" pageNumber="45">10D</figureCitation>
), lateral bead of pronotum well defined (see Fig. 
<figureCitation id="EF46992E03ECFFF9A8B18CDB660B8BB7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Agabus spp., hypomeron in lateral view A A. sjostedti B A. dytiscoides C A. anguluverpus sp. nov. D A. raffrayi. White arrows indicate hypomeron." figureDoi="10.3897/zookeys.963.53470.figure10" httpUri="https://binary.pensoft.net/fig/444407" pageId="0" pageNumber="45">10D</figureCitation>
). Metasternal wing broad, WC/WS less than 2.9 (see Table 
<tableCitation id="411598937E5B34716464E4BA05C4A05D" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
, Figs 
<figureCitation id="C8DBED16FF1D2D89D7B9C23F6CB0BA0E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Agabus spp., metasternal wing and metacoxal plate in ventral view A A. pallidus B A. raffrayi. Dashed lines indicate measurements used; red: metasternal wing (WS), blue: metacoxal plate (WC)." figureDoi="10.3897/zookeys.963.53470.figure2" httpUri="https://binary.pensoft.net/fig/444399" pageId="0" pageNumber="45">2B</figureCitation>
, 
<figureCitation id="A2B9E8D9A7254CC333998B4BE08D6C1E" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
). Pronotum broad, more than twice as broad as interocular distance (see Table 
<tableCitation id="01F0A5D0F66349655CCF4947D6229213" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
, as in Fig. 
<figureCitation id="5726482EA9E8FC106A2AB02051BEB1CA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agabus spp., head and pronotum in dorsal view A A. dytiscoides B A. ruwenzoricus C A. sjostedti D A. pallidus. Red dashed lines indicate measurements used for interocular distance and pronotal width." figureDoi="10.3897/zookeys.963.53470.figure4" httpUri="https://binary.pensoft.net/fig/444401" pageId="0" pageNumber="45">4B</figureCitation>
).
</paragraph>
<paragraph id="8DF4821C6ABB0CF228FAAA3E239E70EE" pageId="0" pageNumber="45">
<emphasis id="7863555A36DF23D028BB55ACB0EA458E" bold="true" italics="true" pageId="0" pageNumber="45">Legs</emphasis>
: Protarsal claws long,&gt; 1.6 
<normalizedToken id="CACA575CB07DEC26D5DA1C58DC215DE7" originalValue="×">x</normalizedToken>
as long as protarsomere 4 in all males and most females (see Table 
<tableCitation id="3DD56800168CA40D47BD02EC9CEAAA55" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
, Fig. 
<figureCitation id="A87E7020D57153BF5550F40BD067C103" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">14</figureCitation>
). Metatarsomeres short and broad; metatarsomere 2 &lt;1.6 
<normalizedToken id="9CB1E4BC177C1EE295BD621305C71408" originalValue="×">x</normalizedToken>
as long as broad (see Table 
<tableCitation id="26A276C5608D07004BE6F8C2BDA79F5D" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
), metatarsomere 5 &lt;3.0 times as long as broad (see Table 
<tableCitation id="ED4D3AAAD92E42BEBAB3BD107A3AFE1A" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
).
</paragraph>
<paragraph id="C0B8074359BA7C979D80BA6CDD758909" pageId="0" pageNumber="45">
<emphasis id="5BECC1950735613D408BAC86CA10B89F" bold="true" italics="true" pageId="0" pageNumber="45">Male genitalia</emphasis>
: Subapically broadened, and prolonged between the subapical broadening and the apical and subapical teeth. Subapical tooth with quite variable shape (see Figs 
<figureCitation id="D81AAE6F55523C8F69D398BCE1F065DC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">8D</figureCitation>
, 
<figureCitation id="44F49B191E9A18D2A4741A1B743E60CB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agabus spp., lateral view of tip of aedeagus A-C A. raffrayi, different specimens showing the variation in shape of the subapical tooth D A. austellus sp. nov. Note the prolonged section between subapical broadening and base of apical and subapical teeth in A. raffrayi (indicated with orange in the smaller outline illustrations)." figureDoi="10.3897/zookeys.963.53470.figure5" httpUri="https://binary.pensoft.net/fig/444402" pageId="0" pageNumber="45">5A-C</figureCitation>
).
</paragraph>
<paragraph id="3A4A9E72733058AE942E49BFF314F898" pageId="0" pageNumber="45">
<emphasis id="B907719DFED5B7050AE5A56CCD29E170" bold="true" pageId="0" pageNumber="45">Female</emphasis>
: Externally similar to males but colour of the elytra tends to be slightly lighter.
</paragraph>
<caption id="58C2E4E516604DBBF3E07D97904B1BE0" doi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45" start="Figure 8" startId="F8">
<paragraph id="98C86D3732BA22FCF53B82F1786ABE75" pageId="0" pageNumber="45">
<emphasis id="61FFE3CC20525373D750E233591B9F11" bold="true" pageId="0" pageNumber="45">Figure 8.</emphasis>
<taxonomicName id="D8E58911C3BF4152B69221A8FC1891CA" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="F5BC9AA3C42992D1E2EE99FB5D164863" italics="true" pageId="0" pageNumber="45">Agabus</emphasis>
</taxonomicName>
spp., aedeagus in lateral view 
<emphasis id="C48C907242E5F0331F7C890595A95F46" bold="true" pageId="0" pageNumber="45">A</emphasis>
<taxonomicName id="63D823F05F3FDA9D6FC39B7C3800039E" lsidName="A. sjostedti" pageId="0" pageNumber="45" rank="species" species="sjostedti">
<emphasis id="CD29EE7AE78BB083FEAF4B2A6C966E2C" italics="true" pageId="0" pageNumber="45">A. sjostedti</emphasis>
</taxonomicName>
<emphasis id="9C19C7BB5F3EB44FA6CEE36954B20352" bold="true" pageId="0" pageNumber="45">B</emphasis>
<taxonomicName id="296CBCED085F439275F4E2B2B2D9DCFF" lsidName="A. dytiscoides" pageId="0" pageNumber="45" rank="species" species="dytiscoides">
<emphasis id="6B29CBC6F3A7F45151645B6275ED5CDA" italics="true" pageId="0" pageNumber="45">A. dytiscoides</emphasis>
</taxonomicName>
<emphasis id="13C0EB9D8B0EBDF2DA173969F1A803D6" bold="true" pageId="0" pageNumber="45">C</emphasis>
<taxonomicName id="E4E6249513447E93853DFD6E8E2DD686" lsidName="A. ruwenzoricus" pageId="0" pageNumber="45" rank="species" species="ruwenzoricus">
<emphasis id="D5CB779EC3B7844FAA6F399EA5C2B3C6" italics="true" pageId="0" pageNumber="45">A. ruwenzoricus</emphasis>
</taxonomicName>
<emphasis id="959A233303C0EA90BCDE7BC6BBE0E9EE" bold="true" pageId="0" pageNumber="45">D</emphasis>
<taxonomicName id="204611F0404C6447385EB753A6EAC5D9" lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis id="C49C75DB58522A3281370DE18193BC68" italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
<emphasis id="17DC39462563610E4BCF13F87B5A8729" bold="true" pageId="0" pageNumber="45">E</emphasis>
<taxonomicName id="1B56D13D98333982F290C04FEA21CE74" lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="37BBBEBBF2E12E59C36F8218029F696D" italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
<emphasis id="4F2373C90C04D3499289DD337D40AB14" bold="true" pageId="0" pageNumber="45">F</emphasis>
<taxonomicName id="902EA3B04753B2048D815564D2F27A36" lsidName="A. anguluverpus" pageId="0" pageNumber="45" rank="species" species="anguluverpus">
<emphasis id="05B254128FC86B982515B684922AC2D6" italics="true" pageId="0" pageNumber="45">A. anguluverpus</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="D2928F284FEBCC72EADC96D7D6597E6C" bold="true" pageId="0" pageNumber="45">G</emphasis>
<taxonomicName id="B26D242161431952A3F1C124C02E7005" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="03665133B92339DD355C4B0A717D275D" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="FDCE2078977742A1992FF26F56B77E7B" bold="true" pageId="0" pageNumber="45">H</emphasis>
<taxonomicName id="6FB8DC30462334670F0CC694037B6795" lsidName="A. riberae" pageId="0" pageNumber="45" rank="species" species="riberae">
<emphasis id="3B8348EC57F510336966F3E968236BDB" italics="true" pageId="0" pageNumber="45">A. riberae</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="BB3B42ECE16D453CFBFFD4BB162DAA5C" bold="true" pageId="0" pageNumber="45">I</emphasis>
<taxonomicName id="665FA6B91AD562763F0D4E6C8803E089" lsidName="A. agulhas" pageId="0" pageNumber="45" rank="species" species="agulhas">
<emphasis id="90F74E2B19F18A235810ED75472B20B5" italics="true" pageId="0" pageNumber="45">A. agulhas</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="67159E0D7AF50D15AEBFDE7055555332" pageId="0" pageNumber="45" type="distribution">
<paragraph id="B6591511DA530F7D944E87E5EC82CC8C" pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph id="1D3B75AE65F13E1951E0FBE252F65279" pageId="0" pageNumber="45">
Ethiopia (see Fig. 
<figureCitation id="BFA57321EDC7995DF34062A70644652D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the Agabus raffrayi group. Relief is expressed in meters above sea level." figureDoi="10.3897/zookeys.963.53470.figure1" httpUri="https://binary.pensoft.net/fig/444398" pageId="0" pageNumber="45">1</figureCitation>
). 
<bibRefCitation id="0CB62CAA44F4B6F90970A5FA43E90C60" DOI="https://doi.org/10.1080/03749444.1975.10736808" author="Rocchi, S" journalOrPublisher="): Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906. Uppsala: Almquist &amp; Wiksells Boktryckeri-A. B." pageId="0" pageNumber="45" refId="B23" refString="Rocchi, S, 1975. Coleotteri ditiscidi dell'Africa orientale. Monitore Zoologico Italiano (N.S. Suppl. 6) 3: 45-56. https://doi.org/10.1080/03749444.1975.10736808" title="Coleotteri ditiscidi dell'Africa orientale. Monitore Zoologico Italiano (N. S. Suppl. 6) 3: 45 - 56." url="https://doi.org/10.1080/03749444.1975.10736808" year="1975">Rocchi (1975)</bibRefCitation>
listed the distribution of 
<taxonomicName id="9A5233923C70B8FEEDCCF149F5A5BAC0" lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis id="AA3683978CA8E6675286D6A5AB31278C" italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
to also include the Democratic Republic of the Congo, Uganda, Rwanda, Tanzania, Zimbabwe, and South Africa but these specimens are likely to belong to other species.
</paragraph>
</subSubSection>
<subSubSection id="550CFC1E36A88549D81498A1B676E4CD" pageId="0" pageNumber="45" type="habitat">
<paragraph id="81B01CFD0A55680729551E181BAE8178" pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph id="476CA9F14ED82F6761CB592A3D3C1373" pageId="0" pageNumber="45">
Found in small, often temporary, streams and pools in streambeds at elevations between 2100 to 3200 m (
<bibRefCitation id="3AA77A5C50114BDE67EBFB6BF3696AFF" DOI="https://doi.org/10.1080/01650429009361398" author="Nilsson, AN" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="45" pagination="135 - 144" refId="B17" refString="Nilsson, AN, Persson, S, 1990. Dimorphism of the metasternal wing in Agabus raffrayi and A. labiatus (Coleoptera: Dytiscidae) questioned. Aquatic Insects 12 (3): 135 - 144, DOI: https://doi.org/10.1080/01650429009361398" title="Dimorphism of the metasternal wing in Agabus raffrayi and A. labiatus (Coleoptera: Dytiscidae) questioned." url="https://doi.org/10.1080/01650429009361398" volume="12" year="1990">Nilsson and Persson 1990</bibRefCitation>
, 
<bibRefCitation id="9A6A18CB97A8664367FB479FA709EE78" DOI="https://doi.org/10.33338/ef.83753" author="Nilsson, AN" journalOrPublisher="Entomologica Fennica" pageId="0" pageNumber="45" pagination="57 - 94" refId="B18" refString="Nilsson, AN, Persson, S, 1993. Taxonomy, distribution and habitats of the Dytiscidae (Coleoptera) of Ethiopia. Entomologica Fennica 4: 57 - 94, DOI: https://doi.org/10.33338/ef.83753" title="Taxonomy, distribution and habitats of the Dytiscidae (Coleoptera) of Ethiopia." url="https://doi.org/10.33338/ef.83753" volume="4" year="1993">1993</bibRefCitation>
; 
<bibRefCitation id="DFB10717BD021A3D2E0BDD09DB0C5867" DOI="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" author="Nilsson, AN" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="45" pagination="155 - 179" refId="B15" refString="Nilsson, AN, 1992a. A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists. Systematic Entomology 17: 155 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" title="A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists." url="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" volume="17" year="1992 a">Nilsson 1992a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A68961F899722E7E52D6E7C9BF9AB60C" pageId="0" pageNumber="45" type="etymology">
<paragraph id="BBD1849C9E492DECB07F868793655220" pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph id="F09B062D6A8AC0268E4342EBBC2526EB" pageId="0" pageNumber="45">
The name refers to the collector of the type specimens, Achille Raffray. The name of the synonym 
<taxonomicName id="4DCE76CA1A70F24EA4DC1B9859670263" lsidName="A. limbicollis" pageId="0" pageNumber="45" rank="species" species="limbicollis">
<emphasis id="B2680C47E656C310F486EF98EB7109E1" italics="true" pageId="0" pageNumber="45">A. limbicollis</emphasis>
</taxonomicName>
refers to the well-defined lateral bead of the pronotum (Latin: 
<emphasis id="5B1AAC825EFE433E0C0309F7DBBAA650" italics="true" pageId="0" pageNumber="45">limbus</emphasis>
= border, 
<emphasis id="0E89C9774A3E08643B547A5FC1319EFC" italics="true" pageId="0" pageNumber="45">collum</emphasis>
= neck).
</paragraph>
</subSubSection>
<subSubSection id="0627CAE3857D7D7CA87314262D77B962" pageId="0" pageNumber="45" type="comments">
<paragraph id="92CD0102D8B6786C047714AC19A60956" pageId="0" pageNumber="45">Comments.</paragraph>
<paragraph id="F8C522AF79C9EEC1DB09E40CE85F3EBF" pageId="0" pageNumber="45">
The fact that 
<taxonomicName id="9C2E054B232779A9F14B71F48C5BE37C" lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis id="89C8469ED5E5B38B2B0FB99DF0E882A6" italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
and 
<taxonomicName id="DF44F01CA5BCD1BBA4CBF932CDA52DE1" lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="221A21FF5B9A61F1C61BD9C5D98E6345" italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
are distinguishable only on the width of the metasternal wing led some previous authors to suggest the occurrence of a single species which was dimorphic with regard to this character (
<bibRefCitation id="F654E71FD856805840A46678D7EB519F" DOI="https://doi.org/10.1111/j.1365-3032.1956.tb00191.x" author="Jackson, DJ" journalOrPublisher="Proceedings of the Royal entomological Society of London (A)" pageId="0" pageNumber="45" pagination="1 - 11" refId="B9" refString="Jackson, DJ, 1956. Dimorphism of the metasternal wings in Agabus raffrayi Sharp and A. labiatus Brahm (Col., Dytiscidae) and its relation to capacity for flight. Proceedings of the Royal entomological Society of London (A) 31: 1 - 11, DOI: https://doi.org/10.1111/j.1365-3032.1956.tb00191.x" title="Dimorphism of the metasternal wings in Agabus raffrayi Sharp and A. labiatus Brahm (Col., Dytiscidae) and its relation to capacity for flight." url="https://doi.org/10.1111/j.1365-3032.1956.tb00191.x" volume="31" year="1956">Jackson 1956</bibRefCitation>
). 
<bibRefCitation id="20BC71C904716AE95B86351AF5976799" DOI="https://doi.org/10.1080/01650429009361398" author="Nilsson, AN" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="45" pagination="135 - 144" refId="B17" refString="Nilsson, AN, Persson, S, 1990. Dimorphism of the metasternal wing in Agabus raffrayi and A. labiatus (Coleoptera: Dytiscidae) questioned. Aquatic Insects 12 (3): 135 - 144, DOI: https://doi.org/10.1080/01650429009361398" title="Dimorphism of the metasternal wing in Agabus raffrayi and A. labiatus (Coleoptera: Dytiscidae) questioned." url="https://doi.org/10.1080/01650429009361398" volume="12" year="1990">Nilsson and Persson (1990)</bibRefCitation>
provided a detailed account of this argument, analysed a large series of specimens and concluded that the variation should rather be interpreted as two separate species. We agree with this assessment and concur that male genitalia are not diagnostic for these two species, only the width of the metasternal wing being reliable. In our measurements, the pronotum is marginally broader in 
<taxonomicName id="CBF196220C94A591BD3B478C74B45D3C" lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="6592476044410CA4A400926C6185DBD4" italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
but the small sample size forbids any strong conclusions at present (Fig. 
<figureCitation id="661F7CFA61442219CA871447032BF810" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">13</figureCitation>
).
</paragraph>
<paragraph id="9D848EE832A2ED9E8B9A039E694EF58C" pageId="0" pageNumber="45">
<bibRefCitation id="9AAD4A00ECEB36F31BBA303EA6E842A6" DOI="https://doi.org/10.1080/03946975.1992.10539193" author="Nilsson, AN" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="45" pagination="207 - 217" refId="B16" refString="Nilsson, AN, 1992b. Larval morphology of six species of Afrotropical Agabus Leach 1817 (coleoptera Dytiscidae). Tropical Zoology 5 (2): 207 - 217, DOI: https://doi.org/10.1080/03946975.1992.10539193" title="Larval morphology of six species of Afrotropical Agabus Leach 1817 (coleoptera Dytiscidae)." url="https://doi.org/10.1080/03946975.1992.10539193" volume="5" year="1992 b">Nilsson (1992b)</bibRefCitation>
described the larval morphology of 
<taxonomicName id="5F9DA9739BF655974A5DA37F9CCC6F6C" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="D9EE427B0CF62F675A0757C6EDA715A9" italics="true" pageId="0" pageNumber="45">Agabus raffrayi</emphasis>
</taxonomicName>
along with some representatives of two other Afrotropical 
<taxonomicName id="E775B05AAA3EF372CD7B81CB20CA2C41" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="7C295DF59F933550E461005C9B4B94AE" italics="true" pageId="0" pageNumber="45">Agabus</emphasis>
</taxonomicName>
groups.
</paragraph>
<caption id="32CD6FE83289B95C4463DF364384C979" doi="10.3897/zookeys.963.53470.figure9" httpUri="https://binary.pensoft.net/fig/444406" pageId="0" pageNumber="45" start="Figure 9" startId="F9">
<paragraph id="9289883735135BDE2897F71A80E7284C" pageId="0" pageNumber="45">
<emphasis id="CF1CCB94E43383ABF34FAEF756CE45B4" bold="true" pageId="0" pageNumber="45">Figure 9.</emphasis>
South African 
<taxonomicName id="1D046A185FA75ABB5D069DF9FE148288" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="3206C2E244A947D93681E1F2BA92A689" italics="true" pageId="0" pageNumber="45">Agabus</emphasis>
</taxonomicName>
spp., microreticulation of male elytral disc 
<emphasis id="4059DF766F314C99AF24249865F5C302" bold="true" pageId="0" pageNumber="45">A</emphasis>
<taxonomicName id="565E253A4AC72F1E95E0A48D19B7FD99" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="44B6CE3C141977ACC03DAC2D134D7DE2" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. holotype, Keurboom, Western Cape 
<emphasis id="19519B5BD95E4A8A93A4EFC6339664DD" bold="true" pageId="0" pageNumber="45">B</emphasis>
<taxonomicName id="1B9DD422B9186230A8A363C21ED398B7" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="E47F60389AA6D85072DB8F0C3A71F478" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. Groote Swartberg, Western Cape 
<emphasis id="43A1923FA2C7017A20DDA2AAFC374F4B" bold="true" pageId="0" pageNumber="45">C</emphasis>
<taxonomicName id="E94508F55766C6244B418D593B9ADCBE" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="9764B56224EB5727F54E5257F125AB6D" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. Sentinel Peak, KZN Drakensberg 
<emphasis id="E1EDA427351615669F59C1F64DDF628C" bold="true" pageId="0" pageNumber="45">D</emphasis>
<taxonomicName id="0686D6D2896BD5B093CC1BE923720CB8" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="DDB552F940DB4567CC4EAB16C8BB7EA7" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. Harkerville Forest, Western Cape 
<emphasis id="2C384559068F4F72CAE95CA1B5773B03" bold="true" pageId="0" pageNumber="45">E</emphasis>
<taxonomicName id="262F434B168AC508C2336FD495FA4A69" lsidName="A. riberae" pageId="0" pageNumber="45" rank="species" species="riberae">
<emphasis id="CA393DDF24CBE115169C3139EEF3074C" italics="true" pageId="0" pageNumber="45">A. riberae</emphasis>
</taxonomicName>
sp. nov. Kamiesberg, Northern Cape 
<emphasis id="2D5ECFB9500AFDF387F3380E3FE41862" bold="true" pageId="0" pageNumber="45">F</emphasis>
<taxonomicName id="68DEED2BFCE5B1EDEFB8B1C2304A04D4" lsidName="A. agulhas" pageId="0" pageNumber="45" rank="species" species="agulhas">
<emphasis id="7AB619754015F9FAD7BD47A4627CAAEF" italics="true" pageId="0" pageNumber="45">A. agulhas</emphasis>
</taxonomicName>
sp. nov. holotype, Rattelrivier, Western Cape.
</paragraph>
</caption>
<caption id="0C7CFABE0614D9459C4EEE6945B64D99" doi="10.3897/zookeys.963.53470.figure10" httpUri="https://binary.pensoft.net/fig/444407" pageId="0" pageNumber="45" start="Figure 10" startId="F10">
<paragraph id="32D4EE0102201D989D5E4F57CE231096" pageId="0" pageNumber="45">
<emphasis id="4D522519FC288535592FCD8F0DF669EE" bold="true" pageId="0" pageNumber="45">Figure 10.</emphasis>
<taxonomicName id="42A7D81CA90C94BDF59BF6D4277ADA33" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="46879067D1D82E223EC487D967165BF7" italics="true" pageId="0" pageNumber="45">Agabus</emphasis>
</taxonomicName>
spp., hypomeron in lateral view 
<emphasis id="79DB24831CEC49EB9852EF2165BE04F1" bold="true" pageId="0" pageNumber="45">A</emphasis>
<taxonomicName id="43EC17B94387BA4E065454937740FA53" lsidName="A. sjostedti" pageId="0" pageNumber="45" rank="species" species="sjostedti">
<emphasis id="FCEDB53F04B545FE39ADC1E347200016" italics="true" pageId="0" pageNumber="45">A. sjostedti</emphasis>
</taxonomicName>
<emphasis id="E38CDAA2B9AF5B419A41208C700F5A0A" bold="true" pageId="0" pageNumber="45">B</emphasis>
<taxonomicName id="B1F9F4CB5CCBBE07C7B3689240078B01" lsidName="A. dytiscoides" pageId="0" pageNumber="45" rank="species" species="dytiscoides">
<emphasis id="00C4762E2022825E6DA5B5D8051A6300" italics="true" pageId="0" pageNumber="45">A. dytiscoides</emphasis>
</taxonomicName>
<emphasis id="8779FBC97BDEFA5A8FCDD3F21D401E8F" bold="true" pageId="0" pageNumber="45">C</emphasis>
<taxonomicName id="F9332E82F68057E732105A233BE4163B" lsidName="A. anguluverpus" pageId="0" pageNumber="45" rank="species" species="anguluverpus">
<emphasis id="B1DC38A7B82C1C4980B30370FB6D3F1E" italics="true" pageId="0" pageNumber="45">A. anguluverpus</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="0C721A348A70CA1DE39EDBF4CBAD9955" bold="true" pageId="0" pageNumber="45">D</emphasis>
<taxonomicName id="3E7CE1729EA1A92059A17992EF0DA796" lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis id="A5677D0D598C3CF05A8C2AC28F4E0DCF" italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
. White arrows indicate hypomeron.
</paragraph>
</caption>
<caption id="EC20EA9B30576BE179C66F15DBC0C3C6" ID-Table-UUID="EC20EA9B30576BE179C66F15DBC0C3C6" httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" start="Table 1" startId="T1">
<paragraph id="D8B05D470CB7B525AD7801C0A111D596" pageId="0" pageNumber="45">
<emphasis id="FEAEACD0929B33DBFD26966D36380AB9" bold="true" pageId="0" pageNumber="45">Table 1.</emphasis>
Morphological characters in the 
<taxonomicName id="1061E43794C175A8ADC5C2DE9FC098CC" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="75D84460AFF59E81ABF811B534B4D93C" italics="true" pageId="0" pageNumber="45">Agabus raffrayi</emphasis>
</taxonomicName>
group. WC/WS = width of metacoxal plate /width of metasternal wing, PW/ID = width of pronotum/interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable.
</paragraph>
</caption>
<paragraph id="E48A54E27C36FF6D8729042F7FEEA221" pageId="0" pageNumber="45">
<table id="AE3E3AF3FCCF84956060AF4644878F64" pageId="0" pageNumber="45">
<tr id="AE8C59DF61E9C472E5ACE6FCFD52CEB1" pageId="0" pageNumber="45">
<th id="F2B1CD8C67FD4073491010BC41C95C9D" colspan="1" pageId="0" pageNumber="45" rowspan="2">Species</th>
<th id="EDBA57B8027B28A8138E564C6E5AFE52" colspan="1" pageId="0" pageNumber="45" rowspan="2">Sex</th>
<th id="F956BE01CE185208A2E7F0E92BECB51C" colspan="5" pageId="0" pageNumber="45" rowspan="1">WC/WS</th>
<th id="9A124D460A7DA25DEA37E589DEA523FC" colspan="5" pageId="0" pageNumber="45" rowspan="1">PW/ID</th>
<th id="9114E407E3536D72DA8A9A77C3B7A317" colspan="5" pageId="0" pageNumber="45" rowspan="1">TL</th>
</tr>
<tr id="A23E4C09B2E4F5EA7F546B213E2B24D2" pageId="0" pageNumber="45">
<th id="0A232A50E04F3F8DFC1998BFC3FA0AE0" colspan="1" pageId="0" pageNumber="45" rowspan="1">N</th>
<th id="0538D5D86BAEABDA22F653DA10C3C8CA" colspan="1" pageId="0" pageNumber="45" rowspan="1">Min</th>
<th id="B805569B72BD4D1CEDE25FEEBFEF98BE" colspan="1" pageId="0" pageNumber="45" rowspan="1">Max</th>
<th id="3E63A7C4F8C67758BED4FF93341996B9" colspan="1" pageId="0" pageNumber="45" rowspan="1">Mean</th>
<th id="A60419280501D9D4ED2A6CF8BE0C4D38" colspan="1" pageId="0" pageNumber="45" rowspan="1">SD</th>
<th id="1EBD555CBA9B5710D0F21FA149A41E23" colspan="1" pageId="0" pageNumber="45" rowspan="1">N</th>
<th id="B9652305660DE45E2EF5859E7365F998" colspan="1" pageId="0" pageNumber="45" rowspan="1">Min</th>
<th id="A43E713E9B87F58CA2B2A3C2CDA383B5" colspan="1" pageId="0" pageNumber="45" rowspan="1">Max</th>
<th id="F230A0276A885C41659B4586487F21A4" colspan="1" pageId="0" pageNumber="45" rowspan="1">Mean</th>
<th id="A0573FFD5AF50DFA9D42029E24196574" colspan="1" pageId="0" pageNumber="45" rowspan="1">SD</th>
<th id="BBA5B2596038F6CE2C52656C90BAF786" colspan="1" pageId="0" pageNumber="45" rowspan="1">N</th>
<th id="3E99325F2F4ADA6DDC74EADE24D76BA1" colspan="1" pageId="0" pageNumber="45" rowspan="1">Min</th>
<th id="7D51CA096CA947F59991695F9D3466EA" colspan="1" pageId="0" pageNumber="45" rowspan="1">Max</th>
<th id="BC78A3E5F796D09260F5A9DCEF7E1B55" colspan="1" pageId="0" pageNumber="45" rowspan="1">Mean</th>
<th id="DDC9F9D630250412795DA31D942F6452" colspan="1" pageId="0" pageNumber="45" rowspan="1">SD</th>
</tr>
<tr id="E33B9AB7760331D382E9D4DF108CBFC8" pageId="0" pageNumber="45">
<td id="6806A20545E38FB4F077B5773606F59B" colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName id="846810BF7A82424A897005D3F896A8B8" lsidName="A. sjostedti" pageId="0" pageNumber="45" rank="species" species="sjostedti">
<emphasis id="150B85D44C0F33F985EE01EE406C7F56" italics="true" pageId="0" pageNumber="45">A. sjostedti</emphasis>
</taxonomicName>
</td>
<td id="3B3D92BD1F3D3EA480C92826C269C8CB" colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td id="4FB2BB3BB492157CC3FE9688CD430870" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="85E1BB4B669A604DF4BC6EB3F8C26BED" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.60</td>
<td id="0A6A42A2E03A69278BE6B510FCF390C3" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.69</td>
<td id="3836A660EED8A183E3BB80CCCA9F8890" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.64</td>
<td id="3A6C6A2BF00183FB51B583822D02B339" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
<td id="38AA67EB110F94CC6D7F7BE98281FF59" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="07A9703A99E5F9DCA153B53E049FAA38" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.05</td>
<td id="9EE3A6E2CC08CE2D8A76725F2B89B7D8" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td id="61F83003F7B727D4229F48D2160222DF" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.07</td>
<td id="A2C9FBC148C9C2F9257FFD3E829E336F" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.04</td>
<td id="9F9A9F8F75686777A7BC67FDD36F832A" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="A6B53568E8F46A085F0B45CAC9A5F3F6" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.08</td>
<td id="F7532AA63DA81D36AD469DCF22CF464C" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.24</td>
<td id="6DC8C13DD7D7A1651AA1937A05AD0B0D" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.16</td>
<td id="073AC77D3DBE46AE8D660FDF38C46440" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.11</td>
</tr>
<tr id="CF62B3B257569E7537CAAB9F712B6726" pageId="0" pageNumber="45">
<td id="D91AE24119A96DDEE6D2A78CA43FF989" colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td id="78BA2E144183ED8C175A8F882B33B33F" colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td id="15857C76A988265CF8A75856B3A5A42C" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.29</td>
<td id="8DA71EB3780DC5F94E2466516CBADE56" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.65</td>
<td id="56881D23D93E797DF42FF0539C664AD3" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.55</td>
<td id="397A985E3A64B785A6119F33CCD69F8A" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.17</td>
<td id="CAF96EDB0362E66785891FA70E2E4332" colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td id="810C3E2E9BAEF668CE9299715D0B8BF5" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.07</td>
<td id="9B7CA896206EDB271430A4211A860C99" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.24</td>
<td id="E6EC80AD9173CAAB5D845EDCA6929336" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.13</td>
<td id="C2161E596334CBAB7C6DC5E39A0513B5" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.08</td>
<td id="B2A6A94B5417E80C016E3DE1D1A7F7B5" colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td id="457996BC1B964BBB42AB922078023490" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.40</td>
<td id="EC43F1290A17B345062850387FE41795" colspan="1" pageId="0" pageNumber="45" rowspan="1">9.12</td>
<td id="08E2423EF884D0E281AA80E98DC8EFBC" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.70</td>
<td id="5CD62037F701B9D48CDF3B1722929B5D" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.31</td>
</tr>
<tr id="A1A87169023B014B14A0731DF65287B7" pageId="0" pageNumber="45">
<td id="254171A9A73DE1321889B608D606DF52" colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName id="BC0DBABAD18F127A4FB3A54D0B03AC8D" lsidName="A. dytiscoides" pageId="0" pageNumber="45" rank="species" species="dytiscoides">
<emphasis id="E36A66D6E5C9B492BE32355803847B81" italics="true" pageId="0" pageNumber="45">A. dytiscoides</emphasis>
</taxonomicName>
</td>
<td id="F7453FC52CB943F30130548A1822F4F7" colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td id="68906BD18CE61EAE5544A0C68512938D" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="26924E2BEFB52D2FAC910B673E56F34C" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.00</td>
<td id="BC8C021471BDD96C328818A4FE5AA0FC" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.27</td>
<td id="60162F612BA162DB5AB13F94ABDC9059" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.13</td>
<td id="B9345A4A43B9EC3F625D69FF666E8F12" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.19</td>
<td id="C3F0EA03A4B7B269758B04D7EEDBE490" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="21785102B423C365EB2F9FBA61544288" colspan="1" pageId="0" pageNumber="45" rowspan="1">1.93</td>
<td id="2D0C830B2BB350A9AF3DAF3843AEE1AE" colspan="1" pageId="0" pageNumber="45" rowspan="1">1.93</td>
<td id="AF288E5EC8DEA2499B1702EB3E0C050C" colspan="1" pageId="0" pageNumber="45" rowspan="1">1.93</td>
<td id="17CEBE98DE33C4432F3EE58370B825C1" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.00</td>
<td id="BECE292CF248425ADB6EAA4B3E12B842" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="C765E48A805AAA2CC480A0E68BC8D358" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.36</td>
<td id="EAB174499EE1165AC1BD0533F2D8006E" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.60</td>
<td id="9AB525F5C8BCB90A1D74869B11F423F2" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.48</td>
<td id="A5F6335B45B737157F0F6F3C3A50BBDC" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.17</td>
</tr>
<tr id="C1F38064B788A88F46EB48B0FB02A99B" pageId="0" pageNumber="45">
<td id="28EDB3E420923E9F8EAE1967A36C5BF8" colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td id="C5DD3BF251FAE1C2F2EAFAD4B15374BC" colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td id="3F7687A20EEB96F1453FD5DC43AFF519" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.88</td>
<td id="919861395B0F1D4548528849174B9327" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.40</td>
<td id="CA98DD435A9C010DA141933F46BB1A02" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.14</td>
<td id="F034C40EE27B54863C76B7DC034A9D4A" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.22</td>
<td id="062FB478576159877DF59498C0E4D817" colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td id="AECF801CC2EBAAECB7916AFCDB1C46A5" colspan="1" pageId="0" pageNumber="45" rowspan="1">1.93</td>
<td id="E3CA6A4A1612DF8C89A3B2CF8DCA6945" colspan="1" pageId="0" pageNumber="45" rowspan="1">1.98</td>
<td id="D4600448688233F5052404D937F42664" colspan="1" pageId="0" pageNumber="45" rowspan="1">1.96</td>
<td id="0BA25AD5A4AA8357A04E34BD43EF709D" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.02</td>
<td id="A02709EF789CBB41513026A6BBEA2D3D" colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td id="54BC95499407011F6BBD0ACA865DD6A6" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.36</td>
<td id="3FBD14F933DBF39F335C2DE081FE7D76" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.76</td>
<td id="C52FDF49BC7BCD7369F67B2C3D459A61" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.55</td>
<td id="6DCB59BAAB71B20288FF21B3D19A9A24" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.18</td>
</tr>
<tr id="6B8984977A18180FBB138BB07EAB2C00" pageId="0" pageNumber="45">
<td id="6A6325AF22B0196D1F04FC6FFE9F229E" colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName id="FE00280835DFDCD9ECBF7E139EEF8F22" lsidName="A. anguluverpus" pageId="0" pageNumber="45" rank="species" species="anguluverpus">
<emphasis id="A25CB018D9C26A265783B618300C2654" italics="true" pageId="0" pageNumber="45">A. anguluverpus</emphasis>
</taxonomicName>
sp. nov.
</td>
<td id="EC11A052D2B666A4F5DB6FBEE7436CEA" colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td id="DAA86F3DB3CFA04C7FBFA93AFB008E20" colspan="1" pageId="0" pageNumber="45" rowspan="1">1</td>
<td id="72123E34EE45178C13DB65C906C0DA8C" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.47</td>
<td id="A296C987EF1BFE47D96ED90FAFCAE44F" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.47</td>
<td id="6CDED7EEAFEA29AC8C95061E2BFBCD67" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.47</td>
<td id="DFC684F629F4619E8F8071139BA8F7AD" colspan="1" pageId="0" pageNumber="45" rowspan="1">NA</td>
<td id="513E1FB9BD894A848B55E02C04764358" colspan="1" pageId="0" pageNumber="45" rowspan="1">1</td>
<td id="8C080176522CC79AF49BEF7928E632DC" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td id="623DD8E5345C8F39A15CA7B87C889A28" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td id="52FCE85DBE1AADC281571E02C0F8768E" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td id="B7AB9E2BA762DF0772DC5F506C272BE4" colspan="1" pageId="0" pageNumber="45" rowspan="1">NA</td>
<td id="3B8FBFB9744CF96E214F9011D4A1AA9B" colspan="1" pageId="0" pageNumber="45" rowspan="1">1</td>
<td id="36D0194CD68E2129D450354444B0F1EA" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td id="E413458700791D3F190C11D681A8F844" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td id="4E39A907D7C4F3343940E03F08B843AB" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td id="42062FABAEBAE19C85DC758D3F0262AF" colspan="1" pageId="0" pageNumber="45" rowspan="1">NA</td>
</tr>
<tr id="85AAB269AA8FECE69567886B16DF061A" pageId="0" pageNumber="45">
<td id="16762DCC46077519B8836BCA388CB295" colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td id="33391A686FA1B66B11388A3190A9E0DA" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="880C49F79AF8A9D3EBDD53E84344EA26" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.19</td>
<td id="E689A4D7BD15761232D19F8466594E17" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.53</td>
<td id="93B9052193CA84C31857317A9304B61C" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.36</td>
<td id="5B2B35A9C8755A795C453CC950CE9A19" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.25</td>
<td id="4509CC7476A58400E55733DF9AC89106" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="031F31945F89462FC0592E4B426B2785" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.05</td>
<td id="E719A4759B2C40FFF97F7CE0C06E8DCA" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.15</td>
<td id="7A807AF8E8DDE1C32E95C34C17FC8D56" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td id="9F2F26FCD84953B82ADC1367CD05FE52" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.07</td>
<td id="754DC826E51C994D57C7139E92A0638E" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="21FE310547479BCF4FF02F4A42986B78" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.36</td>
<td id="F85D4FF4AA244A6B6BE22CDDE3BCD38F" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.44</td>
<td id="747EDD49352754A3B2AB4B28007E931B" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.40</td>
<td id="EE6E09AD50DBA7AE3243343353EE4135" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
</tr>
<tr id="E97EA8BC5B3E995C663DA95BDF33616F" pageId="0" pageNumber="45">
<td id="6A5FFD480ED8967FFD59C9B7CE01A19B" colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName id="278F610174BB0C751AB53A5756EC5498" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="CCC3DDCDB8153DE600D638FB2CAE1A82" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov.
</td>
<td id="749B4D8DE946628D3060AF5E2D0046C8" colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td id="1AB9AFCD01B1C94EB928BAA3FF04DB3F" colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td id="C516C9BF2C54109752AB66636194F5B9" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.11</td>
<td id="0F17B767F97C30B2CEEE95FC4D07B37E" colspan="1" pageId="0" pageNumber="45" rowspan="1">4.15</td>
<td id="417648F8A7244EF43562A9F234BB7B55" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.55</td>
<td id="052F90A308C06718372D3DB982A2FD64" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.23</td>
<td id="3D4E7F6426B8FF984DC5A4FA7129D6A0" colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td id="2D2355220BF8F5F93F86985B8CEE895D" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.12</td>
<td id="7E9919C8F01C6331D82AC6F323FB63C5" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.30</td>
<td id="B2B1AE1E6D0EB38EBD3CAEFE2C687C3E" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.22</td>
<td id="04674495B72550D068F1356CB8AF230A" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.05</td>
<td id="CC853B47DF1DA8527FFFF15CE40807AF" colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td id="031F61D4FD70EA81E96005B43D9CD7ED" colspan="1" pageId="0" pageNumber="45" rowspan="1">6.80</td>
<td id="91BDAEF6C6F29BF64917B0541D604D49" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.16</td>
<td id="C7764ADD72A9F3E5C1616B93F9F0757F" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.53</td>
<td id="66839CB609540ED227F49C14CAB78162" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.40</td>
</tr>
<tr id="70FBBAEF2F102FAE2F274A17BE318CEB" pageId="0" pageNumber="45">
<td id="10D3371C380BA4C25D2852D0102FA294" colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td id="B54DBF8EE14669210DB1D3779845B334" colspan="1" pageId="0" pageNumber="45" rowspan="1">27</td>
<td id="C66579F856B5467824BB4D0CBCEC5AF5" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.17</td>
<td id="1CFA4BC048610E7B8B3C2508D3F5132A" colspan="1" pageId="0" pageNumber="45" rowspan="1">4.0</td>
<td id="C4B67DD5B0507E4408EBB47B9D43E005" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.52</td>
<td id="6AF74B1A933F815B908A0C477105A08F" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.25</td>
<td id="4439F1D47698D5509E49F278EEE2E64B" colspan="1" pageId="0" pageNumber="45" rowspan="1">27</td>
<td id="1FC7D19E7E1B820EDA1D61F777BBA170" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.12</td>
<td id="D7ADF8D3241E3650CA450308800AFDAA" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.49</td>
<td id="493D4572476E5AE20447A853D4FC82B7" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.28</td>
<td id="8892E6EDDA279AA2F512D97642C5EB6D" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.08</td>
<td id="0FE99F68D510B05A0B29D952A6E12CE9" colspan="1" pageId="0" pageNumber="45" rowspan="1">27</td>
<td id="41C69C4E335186E9C7F3638B5D22D6FE" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.04</td>
<td id="B3261F5AA2971B66FF92920422D84E1E" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.40</td>
<td id="59487A5943F31C1858EDF845FF705E59" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.67</td>
<td id="E4CA115D5206F286F1C007FE7E434E2C" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.36</td>
</tr>
<tr id="4A193021C151C1EFE4DF3F398988E733" pageId="0" pageNumber="45">
<td id="B57D80C5F7B5E6E4530A75DB829EE205" colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName id="82B5F5FD7A855A5F4F68EBD2B62C734A" lsidName="A. ruwenzoricus" pageId="0" pageNumber="45" rank="species" species="ruwenzoricus">
<emphasis id="EF42DB3C8881AFEE78F6DD7CC813C1CB" italics="true" pageId="0" pageNumber="45">A. ruwenzoricus</emphasis>
</taxonomicName>
</td>
<td id="5530EB61FFDE0415CC6DC44B54EFB705" colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td id="6FD96BDAA6AB34B796ECA513FF48DED4" colspan="1" pageId="0" pageNumber="45" rowspan="1">9</td>
<td id="E380FBF6608706ED1873F0DE48541C65" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.90</td>
<td id="D36F37036BD39AB5678B4738525B36BD" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.41</td>
<td id="4F4E38D828348261F49B4DB16D013A9C" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.16</td>
<td id="324DB47BA3FB08D9E8815CE8C097F3E9" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.19</td>
<td id="174109BA10F48AD82E6726600562EAFD" colspan="1" pageId="0" pageNumber="45" rowspan="1">9</td>
<td id="FAF6E6D590CEC6B6D44557EF660D1A0C" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.05</td>
<td id="4C2DCC8BF95811B2B7B55EB4A0B80E3E" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.33</td>
<td id="D70E7C2F293394ECC6E7F0DBB65E4F3D" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.19</td>
<td id="301AF09145BE0F6877F3F297FC74B740" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.10</td>
<td id="00C803DFC35D1DDE7AD4A0203104DCED" colspan="1" pageId="0" pageNumber="45" rowspan="1">9</td>
<td id="DE6B4DDA695EB42014B9A3A045A6CF82" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.36</td>
<td id="220616E99B1B47610589E34EF2192D23" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.08</td>
<td id="8E2B75FE101F23B052C36135C3610A52" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.67</td>
<td id="6E13D04ACD6C8A5BD5177B5948A438D9" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.21</td>
</tr>
<tr id="3F7CFB35A22A8220F056383D33313BAC" pageId="0" pageNumber="45">
<td id="E709321CA258B55B6FA110F8BB6D9C9A" colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td id="E52BEAC41A4A55951A00D767608C67CB" colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td id="FFD354893E0DBF2135ACEC1825514854" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.73</td>
<td id="5B6AF7B8680382B5DAC9F509C0472079" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.63</td>
<td id="0BDB7B94123F8E1CA85CAB5ED21BFF54" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.05</td>
<td id="BCA912FDD67D687E28B6238D19170502" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.28</td>
<td id="2D5470B6A8CBD9D7F49428C5C4E41D80" colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td id="E5976DC13124E0926EC2E176820D79EB" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.09</td>
<td id="B330BD4EA0A48B68C3F7F84B90F256C1" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.33</td>
<td id="394923329F1628F9B530F1F84019ED54" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.24</td>
<td id="17B3A2E9F5FB41BF94D22DAF6E1D659F" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.07</td>
<td id="F67E068C6E4FFC4DC9454B214B943D11" colspan="1" pageId="0" pageNumber="45" rowspan="1">17</td>
<td id="10AEEE55EEB0DEF919E5564D71BDDE18" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td id="AF0E527B47A05D690564269D86CA1374" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.08</td>
<td id="62600294440C43A06310D402151399A0" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.86</td>
<td id="B27736D0845FDB7F9A8E0209EE8A4FD2" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.20</td>
</tr>
<tr id="65F1B89962B19CE0CFED062E723C7163" pageId="0" pageNumber="45">
<td id="7C9581909EE853AD7C4A9BD84DB9CFB0" colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName id="7976E051F6D874F84FED39D13CEA2DB4" lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis id="74A0F0DED9DEEAB10D40EDC4E0A73BAB" italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
</td>
<td id="A410FDFAAA0F9683B33BE6CD50F9A1A5" colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td id="1FD2C884437C59E1D6B08FF2CBB2B9AC" colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td id="50FE953A016A7D3B4CF4E5BDE1B05552" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.11</td>
<td id="F0FF2FE2209DFF9D3A87BCBD3961587B" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.84</td>
<td id="C8288383E74FCDECF0696AC286D3CCC2" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.58</td>
<td id="B0016422EEDA3BEA77FFB6FAF691BCE2" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.28</td>
<td id="D2D694A4E8D8C7526966DD74BA5B577E" colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td id="3D81C6B8918E87752F67278F74A8384E" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.16</td>
<td id="B6B873D629B4CEEE5E5E900F2C24CE09" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.33</td>
<td id="355E77C441E3D0734DC246C3A49C927F" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.25</td>
<td id="3D904619DA498ADCB623AD0F770731C0" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
<td id="E2866DC0D96ADBCBAFD291BE06A8CC59" colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td id="2971DE6BA5085BA2107D07E17D6A37E1" colspan="1" pageId="0" pageNumber="45" rowspan="1">6.96</td>
<td id="0FEA0D7C387B90B8D07F31AF4DFE8073" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.16</td>
<td id="86ACA8D16C131AE6D2A3BDAA5659F4D3" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.70</td>
<td id="479BFF744155B3E623A3A174A18AB64D" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.45</td>
</tr>
<tr id="8C60A931F01E4425889CD31FF4271C98" pageId="0" pageNumber="45">
<td id="510D3F732108D6F8FE3F5EE2C2AC3D44" colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td id="FCA82B06399AEC559C813C77ED8F8D7D" colspan="1" pageId="0" pageNumber="45" rowspan="1">7</td>
<td id="FF77A390DF1D81882D0AB49BA2FBE435" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.46</td>
<td id="3575FEE4DD212D7FA4A4FC67125719F2" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.71</td>
<td id="A15B577D6D08D89E9A6FE97B321322D5" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.61</td>
<td id="630662CF7427EB4DC99C473CB47AC69F" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.10</td>
<td id="A238092AAE544994A4E962966C5909BE" colspan="1" pageId="0" pageNumber="45" rowspan="1">7</td>
<td id="8504F26BE2EC158B0534D703CED9F1AC" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.25</td>
<td id="2F52C3B932A2FD7106C484CEF3D08E2F" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.29</td>
<td id="374B3BF944B7CD9E3B3BDDC872691839" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.27</td>
<td id="F0ACB5CD9A5C19BB703E62F8BBE98681" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.01</td>
<td id="6F46D731975413E86D4142E822EA2F0D" colspan="1" pageId="0" pageNumber="45" rowspan="1">7</td>
<td id="271E6025DF3713B83518859D915DEC4E" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td id="9F031AE47ED0E22D8A2C18730C21C379" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.24</td>
<td id="C090E993F9789B26AE7B59438707394E" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.82</td>
<td id="EEDBCE7FBA81D7DD9ABC75AB3893596D" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.31</td>
</tr>
<tr id="335C953E9AF5E1673251105690C5F3E8" pageId="0" pageNumber="45">
<td id="7EC47457244DDD5D4BE9239132CABBC3" colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName id="E04C09D3533A63431830C882BA1933BD" lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="31D09840F8E4421EC0F1059ED55AC076" italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
</td>
<td id="ACCCF75A79E19AB8AB2C5A901D03A713" colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td id="C68E42405C3D827287670B37C2748915" colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td id="EF59B46BB30A8322C4887BE3C352E0D2" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.06</td>
<td id="9CE87E2B6E970A241ACF085D2A3CED49" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.53</td>
<td id="299A2146DFCB327D4C58F7BA9FD2FF55" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.29</td>
<td id="F884E4C2828D812AE50ADC25EEBC0639" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.16</td>
<td id="7152E74D33DDE9ED3A7659E99FA42251" colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td id="6CAF26DB0F3EA5B5CE723C01CFD4EFB4" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.15</td>
<td id="40444D5C4F2A9F17F1A189944977E9C9" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.29</td>
<td id="B77F739C74C49C7E4543853210A0A02A" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.22</td>
<td id="ECB554ABAAF4DAF085EE1BB10C8C7442" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.05</td>
<td id="244D9D69D1A17FAFDB1A4DF730DE3E98" colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td id="820F1D0ADDC221F9C1998E218EB389A6" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.52</td>
<td id="AEC39CAF4367B1B98960DDCA01D207A8" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.08</td>
<td id="4AAB0D72EF092AF34945B85CC25887DC" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.79</td>
<td id="579D8D78EBABD9241D9379BBFFADF6FE" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.25</td>
</tr>
<tr id="9BBB7DE633A669078094FEAFCD8CE614" pageId="0" pageNumber="45">
<td id="5C0584D85237AA4F8E75B1D6187EA9C3" colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td id="B45ADFA96C0FAE62102815110B1363CB" colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td id="4092DB31FD88E4243B201CA449A22F5E" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.05</td>
<td id="C0C4A0CC1D4DEC442E0D5E8CAE7B3878" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.81</td>
<td id="E0F2C8465A8324C354FD2DB9BCBABD61" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.39</td>
<td id="52A748085DC15581097D3636D433D8FE" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.27</td>
<td id="F5326E214C7C7B1FF59F145F4D247BAD" colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td id="DFA585F78EA0687BFB0CAEC306A427BE" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.30</td>
<td id="BDAEAFD4FE79DA90A17FCF4A9EF9DDC8" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.45</td>
<td id="16F33893BB3BA58C5A4195AA81532470" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.34</td>
<td id="C69E710BCEB7D55738A4E269308E3E66" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
<td id="1F1B9E1A970DF6E61981C6D7BB0370A2" colspan="1" pageId="0" pageNumber="45" rowspan="1">5</td>
<td id="F4531B696D7B431796295BC90F7BB22A" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.92</td>
<td id="0CE5343553850EE6896F1E00FFDE3DF2" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.40</td>
<td id="490E86CA881FB8445FA414881A234694" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.14</td>
<td id="2464007EC69B41DCE4A9927A249F7AFA" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.18</td>
</tr>
<tr id="157CA149996CF385C57B5AE0F6C2222D" pageId="0" pageNumber="45">
<td id="B5253F579B26A04484E24AB79BC553F1" colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName id="A284B61683F85678E271B44875C05C01" lsidName="A. riberae" pageId="0" pageNumber="45" rank="species" species="riberae">
<emphasis id="84DA7426B82CA6053511724728FE3ADC" italics="true" pageId="0" pageNumber="45">A. riberae</emphasis>
</taxonomicName>
sp. nov.
</td>
<td id="D077BEB158B188C12A7D6F6DCF075186" colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td id="B286999DD39293451DD70D82A01AC5A0" colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td id="CC2A1409F334804F0E60205CFE8CCC37" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.00</td>
<td id="C7D693E5EE834B65E699184B772F6203" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.17</td>
<td id="D379C09E6DB35B1849BE637C0BE1D58A" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.05</td>
<td id="74BF4CB00F9F33E85A4CAAC63C0662B9" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.07</td>
<td id="79DBBD869F4AD145C4255ADE7355A6AE" colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td id="2B7E890F06F3FC9AB4D9ACD42F6C982B" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.24</td>
<td id="3299B19EA63D4D8A5F0C062408E0F895" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.33</td>
<td id="7B0BC1E71139F9C6A01895CCF0AD2071" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.27</td>
<td id="BA69AC3E66733E29613D892A5F50A1BA" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.03</td>
<td id="2869996C5A4CF60C5816108CE6E17992" colspan="1" pageId="0" pageNumber="45" rowspan="1">6</td>
<td id="64B324B6CE2BE8E4DC2AB0C495F97219" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.21</td>
<td id="7C18DBED4A6971B4A1B70C1FDD42E478" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.08</td>
<td id="7D731BA5F9F866E64C26629135EE4261" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.70</td>
<td id="14B188EED7269474F4BE85FC9B83AD9A" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.35</td>
</tr>
<tr id="30C87B209102ADA3F47FE4DF07B56B52" pageId="0" pageNumber="45">
<td id="CF9FA19F513C569F7A3F9AD3832D3055" colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td id="A91F12ADF28EC9ABB87C2BA60A8A1A59" colspan="1" pageId="0" pageNumber="45" rowspan="1">16</td>
<td id="9900B9A16807A6D9C06AAAF57AFAAA01" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.65</td>
<td id="5505CC1B0DC0B06872409798F6203854" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.53</td>
<td id="33AB87205BD5913244C49F46B14031B0" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.91</td>
<td id="859DF61FFCAF4DDF0893D43CEFE1D793" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.19</td>
<td id="4A54EF4797F3ACBA35BD9BCEFE5069AB" colspan="1" pageId="0" pageNumber="45" rowspan="1">16</td>
<td id="D6BDC1D0BBAEB359C48DC22C5CBF5E0F" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.21</td>
<td id="5ADB4BFA2D7F54870CB2ACD4EFE9BF9B" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.37</td>
<td id="11A84FF495E78C06A8219A5AF87CE0C7" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.28</td>
<td id="4CBCE7F4E7FF7132E14A3B98C6A4506C" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.05</td>
<td id="7B73EB2220313C91C695B76E9B806561" colspan="1" pageId="0" pageNumber="45" rowspan="1">16</td>
<td id="C3A2EE30F4BDD7C8973A5794EE66C4DD" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.21</td>
<td id="C0094BB4DB2A52AADFD7B2395829FEF7" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.24</td>
<td id="7F1E49A655E3832A4772215EBADBBBCE" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.83</td>
<td id="EAD70291D59E2985BFB8BE3AD8ACFB1A" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.29</td>
</tr>
<tr id="5889325F9A5F2CE585A9E41270F0973A" pageId="0" pageNumber="45">
<td id="1E1867CAE823E848220B733B266A9404" colspan="1" pageId="0" pageNumber="45" rowspan="2">
<taxonomicName id="F9C9AA0B630A3F731CA870F943DADD09" lsidName="A. agulhas" pageId="0" pageNumber="45" rank="species" species="agulhas">
<emphasis id="AFFA64DC93890A7D634BAA5AE654C0E4" italics="true" pageId="0" pageNumber="45">A. agulhas</emphasis>
</taxonomicName>
sp. nov.
</td>
<td id="5B264C376FA67A253E7DB13498989709" colspan="1" pageId="0" pageNumber="45" rowspan="1">♀</td>
<td id="9E5C1BCF652038C9618D6213B9DF1403" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="0DDFD7A4B8F0AD159D6D8814984B980E" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.67</td>
<td id="E978343EF466A8987A33AEEDD4DEC8A3" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.75</td>
<td id="7097C10EEF429F6522276A38460D2FB2" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.71</td>
<td id="0CD7503F87ECCF90D523856B49C2FAF2" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
<td id="C02691D09C9963B869784E7F9C5390A1" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="8ED7A21A6C805F3894AA6304FA9CC908" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.15</td>
<td id="A41E750A98558DF0E0BFE490A9A77B37" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.18</td>
<td id="7DB818BFD1B344D564E1DEDC1AAD92D8" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.16</td>
<td id="81C3C353E93F7BA6929AF6E9463F141C" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.02</td>
<td id="000680FB5394AB7DAAEB300005E54A03" colspan="1" pageId="0" pageNumber="45" rowspan="1">2</td>
<td id="C62D20C16D4FE917B8F7B082A79DDA98" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.94</td>
<td id="1CB4AF8EBEC3E5154219F5B132E1A3E2" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.94</td>
<td id="388F6A434037D77F3588D3075B361393" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.94</td>
<td id="EBC8910E27B4DF5307F0EEC3E87122C5" colspan="1" pageId="0" pageNumber="45" rowspan="1">0</td>
</tr>
<tr id="A598EF4E19B109F1519C2AA471163E4E" pageId="0" pageNumber="45">
<td id="A0BC3C9FAE4509A4B8178D1C9FA32B68" colspan="1" pageId="0" pageNumber="45" rowspan="1">♂</td>
<td id="7508B4E61C919D9B6660CC62DF8E67D1" colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td id="1F7A66828693D113D92192333B529B94" colspan="1" pageId="0" pageNumber="45" rowspan="1">3.67</td>
<td id="FA57BEEECF9D82854E7AEC1D68E0720D" colspan="1" pageId="0" pageNumber="45" rowspan="1">4.54</td>
<td id="0CFDCF74200F171ABB4721B57E3496FF" colspan="1" pageId="0" pageNumber="45" rowspan="1">4.01</td>
<td id="91909CD43925D9BAC212570C8BF8A3AB" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.38</td>
<td id="8C998DE383DE486EC9F8F73668717F55" colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td id="D945B69556657DF3FE3BDFDA491E1598" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.10</td>
<td id="1AE24DF84376F8BBB157ABF365BE4C7E" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.24</td>
<td id="9AB0170867A53BA958BE4959C3B6D340" colspan="1" pageId="0" pageNumber="45" rowspan="1">2.17</td>
<td id="373FE57190872856189D39B074FABA14" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.06</td>
<td id="30EAB08DCA217DA2A14D7CC709FDC281" colspan="1" pageId="0" pageNumber="45" rowspan="1">4</td>
<td id="CD17D856571F98E2F25627A2B24CFF0C" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.60</td>
<td id="8059590158B272F07118189F8BCCA84A" colspan="1" pageId="0" pageNumber="45" rowspan="1">8.00</td>
<td id="8DA9AB7B731BD5E9A38E0E76ECCE0494" colspan="1" pageId="0" pageNumber="45" rowspan="1">7.86</td>
<td id="CF33FA0F8C40F365C9C3A50E0963DEF9" colspan="1" pageId="0" pageNumber="45" rowspan="1">0.19</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>