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<document id="CB431618D4A5567AFDFC66049502787F" ID-DOI="10.5281/zenodo.4509293" ID-ISSN="2345-7600" ID-Zenodo-Dep="4509293" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612548681564" checkinUser="carolina" docAuthor="Marzuki, Mohammad Effendi bin &amp; Clements, Gopalasamy Reuben" docDate="2013" docId="0C348780FFE4FFA8B1A3FE7FB349FD8F" docLanguage="en" docName="RafflesBZool.61.1.21-24.pdf" docOrigin="Raffles Bulletin of Zoology 61 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Pearsonia tembatensis Marzuki &amp; Clements 2013, new species" docType="treatment" docVersion="6" lastPageNumber="23" masterDocId="F00DFFF8FFE5FFAAB148FFE2B15BFFBA" masterDocTitle="A New Species Of Cyclophorid Snail (Mollusca: Prosobranchia) From Terengganu, Peninsular Malaysia" masterLastPageNumber="24" masterPageNumber="21" pageNumber="22" updateTime="1698868034160" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="2876D69866F60B90C569D8A9F31EA7F9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0C28E327FA98965A607E70C7767EB5ED">
<mods:title id="5294167091DF9924D6DF0CCB6FB2CDBA">A New Species Of Cyclophorid Snail (Mollusca: Prosobranchia) From Terengganu, Peninsular Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="B551B310A2824B7190C103AE9BB96C24" type="personal">
<mods:role id="CB50F2BC8DB1E89C9DA0A4990338344F">
<mods:roleTerm id="13F026FB1A5A4EC721904A4B042D186E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5F04BA60618449DD4D9AFF69D997FD05">Marzuki, Mohammad Effendi bin</mods:namePart>
</mods:name>
<mods:name id="92DB7FA2FDDCEEC4AAD5731429D929F8" type="personal">
<mods:role id="287C620398C11B090B19C76DC1BC50DA">
<mods:roleTerm id="B5BDB8C4207AA59B257E95E7A769AC2F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="83E635BB0C5F0D68C00D5F861DB1BF85">Clements, Gopalasamy Reuben</mods:namePart>
</mods:name>
<mods:typeOfResource id="80FCA537151F7AF14B538171057E8CAE">text</mods:typeOfResource>
<mods:relatedItem id="D0F17BCAB070BA19FFB9B079484DA678" type="host">
<mods:titleInfo id="E34D7CA570C869D5DFCAF0769BFF65A1">
<mods:title id="391899952302FF817DDDF694FD6FA8F0">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="F6058BE924F263F006BEE805B486F5B4">
<mods:date id="EF338BACF95CE086DE4E70A691D48203">2013</mods:date>
<mods:detail id="2914F9128ED9CAFD0E303F875ADAAE87" type="pubDate">
<mods:number id="9D29C2010883DE87E20A116C0781B9B2">2013-02-28</mods:number>
</mods:detail>
<mods:detail id="AFC0CC08188E79884FE793B8888502A0" type="volume">
<mods:number id="F4E1B140261A071836B794554BBA0604">61</mods:number>
</mods:detail>
<mods:detail id="C307ACD788E0B3EE1011258298785BE7" type="issue">
<mods:number id="9965AA4C1E972E8A6D9D96B02B97611E">1</mods:number>
</mods:detail>
<mods:extent id="8EC0089C5F1A9AA27DD765BC2FC4E3A4" unit="page">
<mods:start id="D536C8DEDDC7102FDF109640CC211D64">21</mods:start>
<mods:end id="DE688E18D790F6B13ADC2ECC086984FE">24</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D2DC0A4906AC4D637F50F62DB313809F">journal article</mods:classification>
<mods:identifier id="DEA732E7760679950EBDED2E4B37D303" type="DOI">10.5281/zenodo.4509293</mods:identifier>
<mods:identifier id="4C9CDD6177A9845DE9DB6B60AD1EAF21" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="888C325532EAC944AB7A37E365C6BB2F" type="Zenodo-Dep">4509293</mods:identifier>
</mods:mods>
<treatment id="0C348780FFE4FFA8B1A3FE7FB349FD8F" ID-DOI="http://doi.org/10.5281/zenodo.4509786" ID-Zenodo-Dep="4509786" LSID="urn:lsid:plazi:treatment:0C348780FFE4FFA8B1A3FE7FB349FD8F" httpUri="http://treatment.plazi.org/id/0C348780FFE4FFA8B1A3FE7FB349FD8F" lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22">
<subSubSection id="CC87651DFFE4FFABB1A3FE7FB32CFE0F" box="[235,631,412,437]" pageId="1" pageNumber="22" type="nomenclature">
<paragraph id="84223696FFE4FFABB1A3FE7FB32CFE0F" blockId="1.[235,631,412,469]" box="[235,631,412,437]" pageId="1" pageNumber="22">
<heading id="DF6A81FAFFE4FFABB1A3FE7FB32CFE0F" bold="true" box="[235,631,412,437]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="22" reason="2">
<emphasis id="B6E9EA84FFE4FFABB1A3FE7FB32CFE0F" bold="true" box="[235,631,412,437]" pageId="1" pageNumber="22">
<taxonomicName id="439D4D15FFE4FFABB1A3FE7FB0B8FE0F" authority="Marzuki &amp; Clements, 2013" authorityName="Marzuki &amp; Clements" authorityYear="2013" box="[235,483,412,437]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="species" species="tembatensis" status="sp. nov.">
<emphasis id="B6E9EA84FFE4FFABB1A3FE7FB0B8FE0F" bold="true" box="[235,483,412,437]" italics="true" pageId="1" pageNumber="22">Pearsonia tembatensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADDA57FFFFE4FFABB0BAFE7FB32CFE0F" box="[498,631,413,437]" pageId="1" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CC87651DFFE4FFABB03DFE5FB0B7FE6F" box="[373,492,445,469]" pageId="1" pageNumber="22" type="description">
<paragraph id="84223696FFE4FFABB03DFE5FB0B7FE6F" blockId="1.[235,631,412,469]" box="[373,492,445,469]" pageId="1" pageNumber="22">
(
<figureCitation id="1CA62A13FFE4FFABB035FE5FB09CFE6F" box="[381,455,445,469]" captionStart="Fig" captionStartId="1.[114,149,1325,1347]" captionTargetBox="[121,744,933,1293]" captionTargetId="figure-126@1.[111,756,914,1310]" captionTargetPageId="1" captionText="Fig. 1. Shell of Pearsonia tembatensis, new species, Terengganu, Peninsular Malaysia. Holotype (ZRC.MOL.3074): a, ventral view; b, dorsal view; c, aperture view; d, side view; e, interior view of operculum; f, side view of operculum; and g, exterior view of operculum. Arrow indicates position of sutural tube." figureDoi="http://doi.org/10.5281/zenodo.4509295" httpUri="https://zenodo.org/record/4509295/files/figure.png" pageId="1" pageNumber="22">Figs. 1</figureCitation>
,
<figureCitation id="1CA62A13FFE4FFABB09EFE5FB0BFFE6F" box="[470,484,445,469]" captionStart="Fig" captionStartId="1.[114,149,2032,2054]" captionTargetBox="[114,754,1531,2001]" captionTargetId="figure-176@1.[113,755,1531,2004]" captionTargetPageId="1" captionText="Fig. 2.Animal of Pearsonia tembatensis, new species, Terengganu, Peninsular Malaysia. The body is grey and spotted and the foot is lightish brown. (Photograph by: Tan Siong Kiat)." figureDoi="http://doi.org/10.5281/zenodo.4509297" httpUri="https://zenodo.org/record/4509297/files/figure.png" pageId="1" pageNumber="22">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CC87651DFFE4FFABB139FE19B018FD60" pageId="1" pageNumber="22" type="materials_examined">
<paragraph id="84223696FFE4FFABB139FE19B3FCFDDD" blockId="1.[113,752,507,731]" pageId="1" pageNumber="22">
<emphasis id="B6E9EA84FFE4FFABB139FE19B06FFDAB" bold="true" box="[113,308,507,529]" pageId="1" pageNumber="22">
<emphasis id="B6E9EA84FFE4FFABB139FE19B075FDAB" bold="true" box="[113,302,507,529]" italics="true" pageId="1" pageNumber="22">Material examined</emphasis>
.
</emphasis>
—
<materialsCitation id="34F53CCBFFE4FFABB012FE19B3FCFDDD" collectingDate="2011-05" collectionCode="SH, ZRC, MOL, M, E, R" country="Malaysia" county="Clements" latitude="5.065528" location="Tembat Forest Reserve" longLatPrecision="1" longitude="102.52553" municipality="Marzuki" pageId="1" pageNumber="22" specimenCount="1" stateProvince="Terengganu" typeStatus="holotype">
<typeStatus id="5B268834FFE4FFABB012FE19B0E8FDAB" box="[346,435,507,529]" pageId="1" pageNumber="22" type="holotype">Holotype</typeStatus>
–
<specimenCount id="929BFD1FFFE4FFABB086FE1EB0A4FDAB" box="[462,511,508,529]" pageId="1" pageNumber="22" type="generic">1 ex.</specimenCount>
<collectionCode id="E28CAE53FFE4FFABB34DFE1EB37CFDAB" box="[517,551,508,529]" country="China" lsid="urn:lsid:biocol.org:col:12540" name="Academia Sinica" pageId="1" pageNumber="22" type="Herbarium">SH</collectionCode>
.
<quantity id="43659B73FFE4FFABB379FE19B320FDAB" box="[561,635,507,529]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="1" pageNumber="22" unit="mm" value="8.6">8.6 mm</quantity>
× SW.
<quantity id="43659B73FFE4FFABB38EFE1EB1CDFD94" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="1" pageNumber="22" unit="mm" value="23.8">23.8 mm</quantity>
(
<collectionCode id="E28CAE53FFE4FFABB1ECFDFAB18DFD94" box="[164,214,536,558]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="22">ZRC</collectionCode>
.
<collectionCode id="E28CAE53FFE4FFABB191FDFAB048FD94" box="[217,275,536,558]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="1" pageNumber="22" type="Herbarium">MOL</collectionCode>
.3074), on leaf litter near waterfall,
<location id="8142604DFFE4FFABB32EFDFAB19BFDF1" LSID="urn:lsid:plazi:treatment:0C348780FFE4FFA8B1A3FE7FB349FD8F:8142604DFFE4FFABB32EFDFAB19BFDF1" country="Malaysia" county="Clements" latitude="5.065528" longLatPrecision="1" longitude="102.52553" municipality="Marzuki" name="Tembat Forest Reserve" pageId="1" pageNumber="22" stateProvince="Terengganu">Tembat Forest Reserve</location>
(
<geoCoordinate id="E1A95051FFE4FFABB199FDD7B014FDF1" box="[209,335,565,587]" degrees="5" direction="north" minutes="03" orientation="latitude" pageId="1" pageNumber="22" precision="1" seconds="55.9" value="5.065528">5°03'55.9&quot;N</geoCoordinate>
,
<geoCoordinate id="E1A95051FFE4FFABB014FDD7B0ABFDF1" box="[348,496,565,587]" degrees="102" direction="east" minutes="31" orientation="longitude" pageId="1" pageNumber="22" precision="1" seconds="31.9" value="102.52553">102°31'31.9&quot;E</geoCoordinate>
),
<collectingRegion id="4659F874FFE4FFABB34CFDD4B320FDF1" box="[516,635,566,587]" country="Malaysia" name="Terengganu" pageId="1" pageNumber="22">Terengganu</collectingRegion>
,
<location id="8142604DFFE4FFABB3C0FDD7B3ABFDF1" LSID="urn:lsid:plazi:treatment:0C348780FFE4FFA8B1A3FE7FB349FD8F:8142604DFFE4FFABB3C0FDD7B3ABFDF1" box="[648,752,565,587]" country="Malaysia" county="Clements" latitude="5.065528" longLatPrecision="1" longitude="102.52553" municipality="Marzuki" name="Peninsular" pageId="1" pageNumber="22" stateProvince="Terengganu">Peninsular</location>
<collectingCountry id="FC8A7606FFE4FFABB139FDB0B190FDD2" box="[113,203,594,616]" name="Malaysia" pageId="1" pageNumber="22">Malaysia</collectingCountry>
, coll.
<collectionCode id="E28CAE53FFE4FFABB040FDB0B07BFDDD" box="[264,288,594,615]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="22" type="Herbarium">M</collectionCode>
.
<collectionCode id="E28CAE53FFE4FFABB062FDB0B060FDDD" box="[298,315,594,615]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="1" pageNumber="22" type="Herbarium">E</collectionCode>
.
<collectingMunicipality id="6446ACECFFE4FFABB00EFDB0B0CCFDD2" box="[326,407,594,616]" pageId="1" pageNumber="22">Marzuki</collectingMunicipality>
&amp;
<collectionCode id="E28CAE53FFE4FFABB0F1FDB0B097FDDD" box="[441,460,594,615]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="22">R</collectionCode>
.
<collectingCounty id="6D434E1AFFE4FFABB09EFDB0B368FDD2" box="[470,563,594,616]" pageId="1" pageNumber="22">Clements</collectingCounty>
,
<date id="F0231056FFE4FFABB376FDB0B3F8FDDD" box="[574,675,594,615]" pageId="1" pageNumber="22" value="2011-05">
<collectingDate id="E067E9BEFFE4FFABB376FDB0B3F8FDDD" box="[574,675,594,615]" pageId="1" pageNumber="22" value="2011-05">May 2011</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="84223696FFE4FFABB139FD8DB018FD60" blockId="1.[113,752,507,731]" pageId="1" pageNumber="22">
<materialsCitation id="34F53CCBFFE4FFABB139FD8DB065FD60" collectingDate="2011-05" collectionCode="SH, M, E, R" country="Malaysia" county="Clements" latitude="5.065528" location="Tembat Forest Reserve" longLatPrecision="1" longitude="102.52553" municipality="Marzuki" pageId="1" pageNumber="22" specimenCode="ME2011" specimenCount="1" stateProvince="Terengganu" typeStatus="paratype">
<typeStatus id="5B268834FFE4FFABB139FD8DB19EFD3E" box="[113,197,623,644]" pageId="1" pageNumber="22" type="paratype">Paratype</typeStatus>
—
<specimenCount id="929BFD1FFFE4FFABB1AFFD8DB049FD3E" box="[231,274,623,644]" pageId="1" pageNumber="22" type="generic">1 ex.</specimenCount>
<collectionCode id="E28CAE53FFE4FFABB05EFD8DB063FD3E" box="[278,312,623,644]" country="China" lsid="urn:lsid:biocol.org:col:12540" name="Academia Sinica" pageId="1" pageNumber="22" type="Herbarium">SH</collectionCode>
.
<quantity id="43659B73FFE4FFABB009FD8DB0CFFD3E" box="[321,404,623,644]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="1" pageNumber="22" unit="mm" value="11.8">11.8 mm</quantity>
x SW. 27.0 mm (
<specimenCode id="D43B9EEDFFE4FFABB371FD8DB3D4FD3F" box="[569,655,623,645]" collectionCode="ME" pageId="1" pageNumber="22">ME2011</specimenCode>
/0186), on leaf litter,
<location id="8142604DFFE4FFABB192FD69B0E5FD1B" LSID="urn:lsid:plazi:treatment:0C348780FFE4FFA8B1A3FE7FB349FD8F:8142604DFFE4FFABB192FD69B0E5FD1B" box="[218,446,651,673]" country="Malaysia" county="Clements" latitude="5.065528" longLatPrecision="1" longitude="102.52553" municipality="Marzuki" name="Tembat Forest Reserve" pageId="1" pageNumber="22" stateProvince="Terengganu">Tembat Forest Reserve</location>
(
<geoCoordinate id="E1A95051FFE4FFABB086FD69B313FD1B" box="[462,584,651,673]" degrees="5" direction="north" minutes="03" orientation="latitude" pageId="1" pageNumber="22" precision="1" seconds="55.9" value="5.065528">5°03'55.9&quot;N</geoCoordinate>
,
<geoCoordinate id="E1A95051FFE4FFABB31DFD6EB3BEFD18" box="[597,741,652,674]" degrees="102" direction="east" minutes="31" orientation="longitude" pageId="1" pageNumber="22" precision="1" seconds="31.9" value="102.52553">102°31'31.9&quot;E</geoCoordinate>
),
<collectingRegion id="4659F874FFE4FFABB139FD4BB1B7FD04" box="[113,236,681,702]" country="Malaysia" name="Terengganu" pageId="1" pageNumber="22">Terengganu</collectingRegion>
,
<location id="8142604DFFE4FFABB1B2FD4AB03DFD04" LSID="urn:lsid:plazi:treatment:0C348780FFE4FFA8B1A3FE7FB349FD8F:8142604DFFE4FFABB1B2FD4AB03DFD04" box="[250,358,680,702]" country="Malaysia" county="Clements" latitude="5.065528" longLatPrecision="1" longitude="102.52553" municipality="Marzuki" name="Peninsular" pageId="1" pageNumber="22" stateProvince="Terengganu">Peninsular</location>
<collectingCountry id="FC8A7606FFE4FFABB038FD4AB08BFD04" box="[368,464,680,702]" name="Malaysia" pageId="1" pageNumber="22">Malaysia</collectingCountry>
, coll.
<collectionCode id="E28CAE53FFE4FFABB35EFD4BB374FD04" box="[534,559,681,702]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="22" type="Herbarium">M</collectionCode>
.
<collectionCode id="E28CAE53FFE4FFABB374FD4BB315FD04" box="[572,590,681,702]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="1" pageNumber="22" type="Herbarium">E</collectionCode>
.
<collectingMunicipality id="6446ACECFFE4FFABB314FD4AB3E9FD04" box="[604,690,680,702]" pageId="1" pageNumber="22">Marzuki</collectingMunicipality>
&amp;
<collectionCode id="E28CAE53FFE4FFABB391FD4BB3B7FD04" box="[729,748,681,702]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="22">R</collectionCode>
.
<collectingCounty id="6D434E1AFFE4FFABB139FD27B195FD61" box="[113,206,709,731]" pageId="1" pageNumber="22">Clements</collectingCounty>
,
<date id="F0231056FFE4FFABB191FD24B065FD60" box="[217,318,709,731]" pageId="1" pageNumber="22" value="2011-05">
<collectingDate id="E067E9BEFFE4FFABB191FD24B065FD60" box="[217,318,709,731]" pageId="1" pageNumber="22" value="2011-05">May 2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CC87651DFFE4FFABB139FCE1B223FD0F" pageId="1" pageNumber="22" type="diagnosis">
<paragraph id="84223696FFE4FFABB139FCE1B223FD0F" blockId="1.[113,752,771,859]" lastBlockId="1.[812,1453,189,693]" pageId="1" pageNumber="22">
<emphasis id="B6E9EA84FFE4FFABB139FCE1B1B0FCA1" bold="true" box="[113,235,771,795]" pageId="1" pageNumber="22">
<emphasis id="B6E9EA84FFE4FFABB139FCE1B1B8FCA1" bold="true" box="[113,227,771,795]" italics="true" pageId="1" pageNumber="22">Diagnosis</emphasis>
.
</emphasis>
— Shell medium-sized, widely umbilicated, dextral, rather thick. Colour brown, translucent, shiny, crossed by narrow peripheral band. Whorls five, slowly increasing; periphery rounded. Radial sculpture ornamented with fine, irregularly growth lines below suture and around umbilicus; spiral sculpture absent. Apex smooth, more or less rounded with inconspicuous growth lines. Wrinkled, irregular diffuse blotches appear after 3½ whorl, becoming inconspicuous after 4½ whorls. Spire flatly discoid, very low, slightly raised above body whorl, ornamented with faint, pale brown stripe at 4½ whorl. Suture deeply impressed; sutural tube short, opening slightly backwards behind peristome, about
<quantity id="43659B73FFE4FFABB57EFE3FB52FFE4F" box="[1078,1140,477,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="22" unit="mm" value="2.0">2 mm</quantity>
in length. Aperture circular, oblique, white with double peristome; inner thickened while outer peristome reflected and expanded at suture forming an open descending wing identical to
<taxonomicName id="439D4D15FFE4FFABB5E0FDDDB47EFDEC" baseAuthorityName="Benson" baseAuthorityYear="1832" box="[1192,1317,575,598]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="22" phylum="Mollusca" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB5E0FDDDB47EFDEC" box="[1192,1317,575,598]" italics="true" pageId="1" pageNumber="22">Pterocyclos</emphasis>
</taxonomicName>
. Operculum corneous, roundly convex, multispiral, ciliated on raised edge, smooth at centre. Animal grey, spotted, foot light brown.
</paragraph>
</subSubSection>
<subSubSection id="CC87651DFFE4FFABB264FD3EB22EFCCF" pageId="1" pageNumber="22" type="etymology">
<paragraph id="84223696FFE4FFABB264FD3EB22EFCCF" blockId="1.[812,1453,732,885]" pageId="1" pageNumber="22">
<emphasis id="B6E9EA84FFE4FFABB264FD3EB2F7FD4F" bold="true" box="[812,940,732,757]" pageId="1" pageNumber="22">
<emphasis id="B6E9EA84FFE4FFABB264FD3EB2FEFD4F" bold="true" box="[812,933,732,757]" italics="true" pageId="1" pageNumber="22">Etymology</emphasis>
.
</emphasis>
— This new species is named after its
<typeStatus id="5B268834FFE4FFABB435FD3CB4F7FD4C" box="[1405,1452,734,758]" pageId="1" pageNumber="22">type</typeStatus>
locality, Tembat Forest Reserve,
<collectingRegion id="4659F874FFE4FFABB5C5FD1CB455FCAC" box="[1165,1294,766,790]" country="Malaysia" name="Terengganu" pageId="1" pageNumber="22">Terengganu</collectingRegion>
. We chose this name to highlight the biological importance of this forest reserve, portions of which are currently being cleared for a dam.
</paragraph>
</subSubSection>
<subSubSection id="CC87651DFFE4FFA8B264FC7EB349FD8F" lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22" type="discussion">
<paragraph id="84223696FFE4FFABB264FC7EB5B5FB6F" blockId="1.[812,1453,924,1237]" pageId="1" pageNumber="22">
<emphasis id="B6E9EA84FFE4FFABB264FC7EB2C7FC0F" bold="true" box="[812,924,924,949]" pageId="1" pageNumber="22">
<emphasis id="B6E9EA84FFE4FFABB264FC7EB2CFFC0F" bold="true" box="[812,916,924,949]" italics="true" pageId="1" pageNumber="22">Remarks</emphasis>
.
</emphasis>
— Based on our comparisons with a fairly complete collation of cyclophorid literature (i.e.,
<bibRefCitation id="E00C4B67FFE4FFABB414FC5FB23DFC4F" author="Morlet, L" pageId="1" pageNumber="22" pagination="82 - 86" refId="ref2631" refString="Morlet, L., 1892. Diagnoses Molluscorum novorum, in Indo-China collectorum. Journal de Conchyliologie, 40: 82 - 86." type="journal article" year="1892">Morlet, 1892</bibRefCitation>
;
<bibRefCitation id="E00C4B67FFE4FFABB23EFC3FB555FC4F" author="Kobelt, W" box="[886,1038,989,1013]" pageId="1" pageNumber="22" refId="ref2412" refString="Kobelt, W., 1902. Das Tierreich-Cyclophoridae. Koniglich Preussischen Akademie der Wissenschaften zu Berlin, R. Friedlander &amp; Sohn, Berlin. 662 pp." type="book" year="1902">Kobelt, 1902</bibRefCitation>
,
<bibRefCitation id="E00C4B67FFE4FFABB555FC3FB50CFC4F" author="Kobelt, W" box="[1053,1111,989,1013]" pageId="1" pageNumber="22" pagination="712 - 980" refId="ref2441" refString="Kobelt, W., 1908. Die gedeckelten Lungenschnecken (Cyclostomacea). Systematisches Conchylien-Cabinet von Martini und Chemnitz, 1: 712 - 980, pls. 104 - 144." type="journal article" year="1908">1908</bibRefCitation>
;
<bibRefCitation id="E00C4B67FFE4FFABB52FFC3FB5B5FC4F" author="Gude, G" box="[1127,1262,989,1013]" pageId="1" pageNumber="22" refId="ref2370" refString="Gude, G. K., 1921. Mollusca III Land Operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicidae). The Fauna of British India including Ceylon and Burma. Taylor &amp; Francis, London. 386 pp." type="book" year="1921">Gude, 1921</bibRefCitation>
;
<bibRefCitation id="E00C4B67FFE4FFABB5B5FC3CB42EFC4F" author="Yen" box="[1277,1397,989,1013]" pageId="1" pageNumber="22" pagination="1 - 233" refId="ref2794" refString="Yen, T- Ch., 1939. Die Chinesischen Land-und Susswasser- Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 444: 1 - 233 + 16 pls." type="book chapter" year="1939">Yen, 1939</bibRefCitation>
), we assigned our new species to the genus
<taxonomicName id="439D4D15FFE4FFABB59EFC1DB41FFBAC" authorityName="Kobelt" authorityYear="1902" box="[1238,1348,1023,1046]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB59EFC1DB41FFBAC" box="[1238,1348,1023,1046]" italics="true" pageId="1" pageNumber="22">Pearsonia</emphasis>
</taxonomicName>
based on a general agreement with the original shell description in German (translated by F. Köhler) by
<bibRefCitation id="E00C4B67FFE4FFABB58AFBDFB405FBEF" author="Kobelt, W" box="[1218,1374,1085,1109]" pageId="1" pageNumber="22" refId="ref2412" refString="Kobelt, W., 1902. Das Tierreich-Cyclophoridae. Koniglich Preussischen Akademie der Wissenschaften zu Berlin, R. Friedlander &amp; Sohn, Berlin. 662 pp." type="book" year="1902">Kobelt (1902)</bibRefCitation>
: “Shell discoid, with thick, sometimes hairy periostracum. Aperture circular with thin small tubular pore behind the lip opening backwards. Operculum, non-calcareous, multispiral, outside convex, inside flat. Outer whorls flaring.”
</paragraph>
<paragraph id="84223696FFE4FFABB264FB1DB29DF9CF" blockId="1.[812,1453,1277,1653]" pageId="1" pageNumber="22">
<taxonomicName id="439D4D15FFE4FFABB264FB1DB2C1FAAC" authorityName="Kobelt" authorityYear="1902" box="[812,922,1279,1302]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB264FB1DB2C1FAAC" box="[812,922,1279,1302]" italics="true" pageId="1" pageNumber="22">Pearsonia</emphasis>
</taxonomicName>
(and our new species) can be differentiated from
<taxonomicName id="439D4D15FFE4FFABB264FAFDB530FA8F" authority="(Benson, 1832)" baseAuthorityName="Benson" baseAuthorityYear="1832" box="[812,1131,1309,1334]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="22" phylum="Mollusca" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB264FAFDB2F4FA8C" box="[812,943,1311,1334]" italics="true" pageId="1" pageNumber="22">Pterocyclos</emphasis>
(Benson, 1832)
</taxonomicName>
and
<taxonomicName id="439D4D15FFE4FFABB5E1FAFDB236FAEF" authority="(Martens, 1891)" baseAuthorityName="Martens" baseAuthorityYear="1891" class="Gastropoda" family="Cyclophoridae" genus="Crossopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="22" phylum="Mollusca" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB5E1FAFDB46EFA8C" box="[1193,1333,1311,1334]" italics="true" pageId="1" pageNumber="22">Crossopoma</emphasis>
(
<bibRefCitation id="E00C4B67FFE4FFABB400FAFFB23FFAEF" author="Martens, E" pageId="1" pageNumber="22" pagination="209 - 264" refId="ref2558" refString="Martens, E. von, 1891. Landschnecken des Indischen Archipels. Zoologische Ergebnisse einer reise in Niederlandisch Ost- Indien, 2: 209 - 264." type="journal article" year="1891">Martens, 1891</bibRefCitation>
)
</taxonomicName>
by the presence of a sutural tube just behind the aperture (
<bibRefCitation id="E00C4B67FFE4FFABB2DDFABFB572FACF" author="Kobelt, W" box="[917,1065,1373,1397]" pageId="1" pageNumber="22" refId="ref2412" refString="Kobelt, W., 1902. Das Tierreich-Cyclophoridae. Koniglich Preussischen Akademie der Wissenschaften zu Berlin, R. Friedlander &amp; Sohn, Berlin. 662 pp." type="book" year="1902">Kobelt, 1902</bibRefCitation>
;
<bibRefCitation id="E00C4B67FFE4FFABB57FFABFB5E1FACF" author="Gude, G" box="[1079,1210,1373,1397]" pageId="1" pageNumber="22" refId="ref2370" refString="Gude, G. K., 1921. Mollusca III Land Operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicidae). The Fauna of British India including Ceylon and Burma. Taylor &amp; Francis, London. 386 pp." type="book" year="1921">Gude, 1921</bibRefCitation>
).
<taxonomicName id="439D4D15FFE4FFABB598FABDB465FACC" authorityName="Kobelt" authorityYear="1902" box="[1232,1342,1375,1398]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB598FABDB465FACC" box="[1232,1342,1375,1398]" italics="true" pageId="1" pageNumber="22">Pearsonia</emphasis>
</taxonomicName>
is clearly distinct from
<taxonomicName id="439D4D15FFE4FFABB29AFA9DB44EFA2F" authority="(Swainson, 1840)" baseAuthorityName="Swainson" baseAuthorityYear="1840" box="[978,1301,1405,1430]" class="Insecta" family="Psocidae" genus="Cyclotus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB29AFA9DB56CFA2C" box="[978,1079,1407,1430]" italics="true" pageId="1" pageNumber="22">Cyclotus</emphasis>
(
<bibRefCitation id="E00C4B67FFE4FFABB503FA9FB456FA2F" author="Swainson, W" box="[1099,1293,1405,1429]" pageId="1" pageNumber="22" refId="ref2730" refString="Swainson, W., 1840. A Treatise of Malacology: The Natural Classification of Shell and Shell-Fish. New-Street-Square, London. 419 pp." type="book" year="1840">Swainson, 1840</bibRefCitation>
)
</taxonomicName>
, which does not have a sutural tube. Although both
<taxonomicName id="439D4D15FFE4FFABB44CFA7DB422FA0C" authorityName="Kobelt" authorityYear="1902" box="[1284,1401,1439,1462]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB44CFA7DB422FA0C" box="[1284,1401,1439,1462]" italics="true" pageId="1" pageNumber="22">Pearsonia</emphasis>
</taxonomicName>
and
<taxonomicName id="439D4D15FFE4FFABB264FA5DB524FA6F" authority="(Benson, 1851)" baseAuthorityName="Benson" baseAuthorityYear="1851" box="[812,1151,1469,1494]" class="Gastropoda" family="Cyclophoridae" genus="Opisthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="22" phylum="Mollusca" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB264FA5DB290FA6C" box="[812,971,1471,1494]" italics="true" pageId="1" pageNumber="22">Opisthophorus</emphasis>
(Benson, 1851)
</taxonomicName>
possess a sutural tube, the tube of the latter genus is significantly longer, and turns either upwards or downwards as it decreases in length. Unlike
<taxonomicName id="439D4D15FFE4FFABB234F9FDB2B1F98C" authorityName="Kobelt" authorityYear="1902" box="[892,1002,1567,1590]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB234F9FDB2B1F98C" box="[892,1002,1567,1590]" italics="true" pageId="1" pageNumber="22">Pearsonia</emphasis>
</taxonomicName>
(
<bibRefCitation id="E00C4B67FFE4FFABB2B0F9FFB5B1F98F" author="Godwin-Austen, H" box="[1016,1258,1565,1589]" pageId="1" pageNumber="22" pagination="35 - 39" refId="ref2273" refString="Godwin-Austen, H. H., 1889. On a collection of land-shells made in Borneo by Mr. A. Everett, with descriptions of supposed new species. Part 1. Cyclostomacae. Proceedings of the Zoological Society of London: 332 - 355, pls. 35 - 39." type="book chapter" year="1889">Godwin-Austen, 1889</bibRefCitation>
;
<bibRefCitation id="E00C4B67FFE4FFABB5BEF9FFB4FAF98F" author="Stoliczka, F" box="[1270,1441,1565,1589]" pageId="1" pageNumber="22" pagination="261 - 271" refId="ref2684" refString="Stoliczka, F., 1872. On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part first, Cyclostomacea. Journal of Asiatic Society of Bengal, 41: 261 - 271, pl. 10." type="journal article" year="1872">Stoliczka, 1872</bibRefCitation>
), the edges of the operculum in
<taxonomicName id="439D4D15FFE4FFABB531F9DDB58DF9EC" baseAuthorityName="Swainson" baseAuthorityYear="1840" box="[1145,1238,1599,1622]" class="Insecta" family="Psocidae" genus="Cyclotus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB531F9DDB58DF9EC" box="[1145,1238,1599,1622]" italics="true" pageId="1" pageNumber="22">Cyclotus</emphasis>
</taxonomicName>
and
<taxonomicName id="439D4D15FFE4FFABB446F9DDB4F6F9EC" baseAuthorityName="Benson" baseAuthorityYear="1851" box="[1294,1453,1599,1622]" class="Gastropoda" family="Cyclophoridae" genus="Opisthophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="22" phylum="Mollusca" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB446F9DDB4F6F9EC" box="[1294,1453,1599,1622]" italics="true" pageId="1" pageNumber="22">Opisthophorus</emphasis>
</taxonomicName>
are not raised.
</paragraph>
<caption id="D0E2661EFFE4FFABB13AFACFB328FA0C" ID-DOI="http://doi.org/10.5281/zenodo.4509295" ID-Zenodo-Dep="4509295" httpUri="https://zenodo.org/record/4509295/files/figure.png" pageId="1" pageNumber="22" startId="1.[114,149,1325,1347]" targetBox="[121,744,933,1293]" targetPageId="1">
<paragraph id="84223696FFE4FFABB13AFACFB328FA0C" blockId="1.[114,758,1325,1462]" pageId="1" pageNumber="22">
Fig. 1. Shell of
<taxonomicName id="439D4D15FFE4FFABB046FACCB0B2FAF8" authority="Marzuki &amp; Clements, 2013" authorityName="Marzuki &amp; Clements" authorityYear="2013" box="[270,489,1326,1346]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="species" species="tembatensis" status="sp. nov.">
<emphasis id="B6E9EA84FFE4FFABB046FACCB0B2FAF8" box="[270,489,1326,1346]" italics="true" pageId="1" pageNumber="22">Pearsonia tembatensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADDA57FFFFE4FFABB0BFFACCB334FAF9" box="[503,623,1325,1347]" pageId="1" pageNumber="22" rank="species">new species</taxonomicNameLabel>
, Terengganu, Peninsular Malaysia. Holotype (ZRC.MOL.3074): a, ventral view; b, dorsal view; c, aperture view; d, side view; e, interior view of operculum; f, side view of operculum; and g, exterior view of operculum. Arrow indicates position of sutural tube.
</paragraph>
</caption>
<paragraph id="84223696FFE4FFABB264F97FB43EF8EF" blockId="1.[812,1453,1693,1877]" pageId="1" pageNumber="22">
There is no other representative of
<taxonomicName id="439D4D15FFE4FFABB5A5F97DB43CF90C" authorityName="Kobelt" authorityYear="1902" box="[1261,1383,1695,1718]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB5A5F97DB43CF90C" box="[1261,1383,1695,1718]" italics="true" pageId="1" pageNumber="22">Pearsonia</emphasis>
</taxonomicName>
from
<collectingCountry id="FC8A7606FFE4FFABB264F95FB2D5F96F" box="[812,910,1725,1749]" name="Malaysia" pageId="1" pageNumber="22">Malaysia</collectingCountry>
for comparison. Among Malaysian cyclophorids,
<taxonomicName id="439D4D15FFE4FFABB264F93DB296F94C" authority="Marzuki &amp; Clements, 2013" authorityName="Marzuki &amp; Clements" authorityYear="2013" box="[812,973,1759,1782]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="species" species="tembatensis" status="sp. nov.">
<emphasis id="B6E9EA84FFE4FFABB264F93DB296F94C" box="[812,973,1759,1782]" italics="true" pageId="1" pageNumber="22">P. tembatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADDA57FFFFE4FFABB290F93CB539F94F" box="[984,1122,1757,1781]" pageId="1" pageNumber="22" rank="species">new species</taxonomicNameLabel>
, has the closest affinity with
<taxonomicName id="439D4D15FFE4FFABB264F91DB45EF8AF" authority="Benthem-Jutting, 1949" authorityName="Benthem-Jutting" authorityYear="1949" box="[812,1285,1789,1814]" class="Insecta" family="Psocidae" genus="Cyclotus" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="22" phylum="Arthropoda" rank="species" species="umbraticus">
<emphasis id="B6E9EA84FFE4FFABB264F91DB552F8AC" box="[812,1033,1791,1814]" italics="true" pageId="1" pageNumber="22">Cyclotus umbraticus</emphasis>
Benthem-Jutting, 1949
</taxonomicName>
from Larut hill [
<collectingRegion id="4659F874FFE4FFABB27BF8FFB2D2F88F" box="[819,905,1821,1845]" country="Malaysia" name="Pahang" pageId="1" pageNumber="22">Pahang</collectingRegion>
], but again, the latter species does not have a sutural tube and its shell sculpture has a zigzag-like pattern.
</paragraph>
<paragraph id="84223696FFE4FFA8B264F89FB059FEEF" blockId="1.[812,1456,1917,2102]" lastBlockId="2.[136,776,189,341]" lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22">
Within
<taxonomicName id="439D4D15FFE4FFABB2CEF89DB2A0F82C" authorityName="Kobelt" authorityYear="1902" box="[902,1019,1919,1942]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E9EA84FFE4FFABB2CEF89DB2A0F82C" box="[902,1019,1919,1942]" italics="true" pageId="1" pageNumber="22">Pearsonia</emphasis>
</taxonomicName>
,
<taxonomicName id="439D4D15FFE4FFABB546F89DB5EEF82C" authorityName="Marzuki &amp; Clements" authorityYear="2013" box="[1038,1205,1919,1942]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="species" species="tembatensis">
<emphasis id="B6E9EA84FFE4FFABB546F89DB5EEF82C" box="[1038,1205,1919,1942]" italics="true" pageId="1" pageNumber="22">P. tembatensis</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="439D4D15FFE4FFABB264F87DB5BFF80F" authority="(Godwin-Austen, 1915)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1915" box="[812,1252,1949,1974]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="species" species="putaoensis">
<emphasis id="B6E9EA84FFE4FFABB264F87DB29DF80C" box="[812,966,1951,1974]" italics="true" pageId="1" pageNumber="22">P. putaoensis</emphasis>
(
<bibRefCitation id="E00C4B67FFE4FFABB291F87FB586F80F" author="Godwin-Austen, H" box="[985,1245,1949,1973]" pageId="1" pageNumber="22" pagination="494 - 503" refId="ref2327" refString="Godwin-Austen, H. H., 1915. Zoological results of the Abor Expedition: Mollusca III, Cyclophoridae (in part). Records of the Indian Museum, 8: 494 - 503, pls. 38 - 40." type="journal article" year="1915">Godwin-Austen, 1915</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="439D4D15FFE4FFABB46CF87DB574F86F" authority="(Godwin-Austen, 1915)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1915" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="species" species="minimum">
<emphasis id="B6E9EA84FFE4FFABB46CF87DB4F7F80C" box="[1316,1452,1951,1974]" italics="true" pageId="1" pageNumber="22">P. minimum</emphasis>
(
<bibRefCitation id="E00C4B67FFE4FFABB27BF85FB573F86F" author="Godwin-Austen, H" box="[819,1064,1981,2005]" pageId="1" pageNumber="22" pagination="494 - 503" refId="ref2327" refString="Godwin-Austen, H. H., 1915. Zoological results of the Abor Expedition: Mollusca III, Cyclophoridae (in part). Records of the Indian Museum, 8: 494 - 503, pls. 38 - 40." type="journal article" year="1915">Godwin-Austen, 1915</bibRefCitation>
)
</taxonomicName>
from the Indian region. However, the shells of the latter two species are significantly smaller and their shell sculpture comprises fine transverse striae on the epidermis. Furthermore, the outer peristome of
<taxonomicName id="439D4D15FFE4FFA8B4D1F7FDB1B0FF6C" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1915" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" lastPageId="2" lastPageNumber="23" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="species" species="minimum">
<emphasis id="B6E9EA84FFE4FFA8B4D1F7FDB1B0FF6C" italics="true" lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22">P. minimum</emphasis>
</taxonomicName>
is very simple and only slightly reflected. We hope that future phylogenetic studies incorporating molecular, conchological and anatomical data on cyclophorids will include this new species in order to validate our hypothesis.
</paragraph>
<caption id="D0E2661EFFE4FFABB13AF812B30EF785" ID-DOI="http://doi.org/10.5281/zenodo.4509297" ID-Zenodo-Dep="4509297" httpUri="https://zenodo.org/record/4509297/files/figure.png" pageId="1" pageNumber="22" startId="1.[114,149,2032,2054]" targetBox="[114,754,1531,2001]" targetPageId="1">
<paragraph id="84223696FFE4FFABB13AF812B30EF785" blockId="1.[114,753,2032,2111]" pageId="1" pageNumber="22">
Fig. 2. Animal of
<taxonomicName id="439D4D15FFE4FFABB050F813B0AAF7BF" authority="Marzuki &amp; Clements, 2013" authorityName="Marzuki &amp; Clements" authorityYear="2013" box="[280,497,2033,2053]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="species" species="tembatensis" status="sp. nov.">
<emphasis id="B6E9EA84FFE4FFABB050F813B0AAF7BF" box="[280,497,2033,2053]" italics="true" pageId="1" pageNumber="22">Pearsonia tembatensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ADDA57FFFFE4FFABB0B5F812B329F7BC" box="[509,626,2032,2054]" pageId="1" pageNumber="22" rank="species">new species</taxonomicNameLabel>
, Terengganu, Peninsular Malaysia. The body is grey and spotted and the foot is lightish brown. (Photograph by: Tan Siong Kiat).
</paragraph>
</caption>
<paragraph id="84223696FFE7FFA8B1C0FE9FB349FD8F" blockId="2.[136,777,381,565]" pageId="2" pageNumber="23">
Based on the geographical distribution of
<taxonomicName id="439D4D15FFE7FFA8B316FE9DB397FE2C" authorityName="Kobelt" authorityYear="1902" box="[606,716,383,406]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E9EA84FFE7FFA8B316FE9DB397FE2C" box="[606,716,383,406]" italics="true" pageId="2" pageNumber="23">Pearsonia</emphasis>
</taxonomicName>
, it is highly likely that our new species belongs to this genus. To date, there are 22 species in the genus
<taxonomicName id="439D4D15FFE7FFA8B3DAFE5DB25CFE6C" authorityName="Kobelt" authorityYear="1902" box="[658,775,447,470]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E9EA84FFE7FFA8B3DAFE5DB25CFE6C" box="[658,775,447,470]" italics="true" pageId="2" pageNumber="23">Pearsonia</emphasis>
</taxonomicName>
found mostly within the Indo-Malayan region. This new species represents the southernmost extent of
<taxonomicName id="439D4D15FFE7FFA8B3C0FE1DB3ADFDAC" authorityName="Kobelt" authorityYear="1902" box="[648,758,511,534]" class="Magnoliopsida" family="Fabaceae" genus="Pearsonia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E9EA84FFE7FFA8B3C0FE1DB3ADFDAC" box="[648,758,511,534]" italics="true" pageId="2" pageNumber="23">Pearsonia</emphasis>
</taxonomicName>
’s distribution in Indo-Malaya (
<figureCitation id="1CA62A13FFE7FFA8B08AFDFFB35FFD8F" box="[450,516,541,565]" captionStart="Fig" captionStartId="2.[136,171,1918,1940]" captionTargetBox="[146,1467,958,1891]" captionTargetId="figure-333@2.[136,1475,949,1895]" captionTargetPageId="2" captionText="Fig. 3. Distribution of worldwide Pearsonia spp. based on approximate location of type localities: 1, Pearsonia travancorica (Blanford, 1880); 2, P. fairbanki (Blanford, 1869); 3, P. beddomei Blanford, 1866; 4, P. hispida (Pearson, 1833); 5, P. assamensis Fulton, 1900; 6, P. nagaensis (Godwin-Austen &amp; Beddome, 1894); 7, P. mastersi (Hanley &amp; Theobald, 1870); 8, P. simplex (Nevill, 1878); 9, P. nevilli (Godwin-Austen, 1876); 10, P. plana (Godwin-Austen, 1915); 11, P. luyorensis Godwin-Austen, 1915; 12, P. oakesi (Godwin-Austen, 1915); 13, P. minima (Godwin-Austen, 1915); 14, P. kempi Godwin-Austen, 1915; 15, P. putaoensis (Godwin-Austen, 1915); 16, P. andersoni Blanford, 1869; 17, P. bhamoensis Theobald, 1876; 18, P. bitubifera Theobald, 1876; 19, P. avana; 20, P. massiei (Morlet, 1892); 21, P. gredleri Yen, 1939; and 22, P. tembatensis Marzuki &amp; Clements, 2013." figureDoi="http://doi.org/10.5281/zenodo.4509299" httpUri="https://zenodo.org/record/4509299/files/figure.png" pageId="2" pageNumber="23">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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