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<document id="87008CB821227C1B3ACDE1159DF6FBF3" ID-CLB-Dataset="169754" ID-DOI="10.11646/zootaxa.5195.5.2" ID-GBIF-Dataset="79723b4d-3852-4d07-9abd-babc54b5728c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7223689" ID-ZooBank="7661F79E-F63F-4A0D-9BF1-7818E228B262" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666163692883" checkinUser="plazi" docAuthor="Mousavi-Sabet, Hamed, Eagderi, Soheil, Saemi-Komsari, Maryam, Kaya, Cüneyt &amp; Freyhof, Jörg" docDate="2022" docId="D409E1065102FFC0AFDD39D1FD683280" docLanguage="en" docName="zootaxa.5195.5.2.pdf" docOrigin="Zootaxa 5195 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Garra rezai Mousavi-Sabet &amp; Eagderi &amp; Saemi-Komsari &amp; Kaya &amp; Freyhof 2022, new species" docType="treatment" docVersion="8" lastPageNumber="433" masterDocId="2830997E5106FFCEAF4A3F02FFBB3005" masterDocTitle="Garra rezai, a new species from two widely disjunct areas in the Tigris drainage (Teleostei: Cyprinidae)" masterLastPageNumber="436" masterPageNumber="419" pageNumber="423" updateTime="1702000765933" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CA4CC3C47230803D4C0A0A8FB9169D52">Garra rezai, a new species from two widely disjunct areas in the Tigris drainage (Teleostei: Cyprinidae)</mods:title>
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<mods:namePart id="2ED1DD9C137BD44E747B61103FEDF901">Mousavi-Sabet, Hamed</mods:namePart>
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<mods:affiliation id="FE443ECCC86003C578D39E997D4B9003">Department of Fisheries Sciences, Faculty of Natural Resources, University of Guilan, Sowmeh Sara, Guilan, Iran. maryam _ saemi @ yahoo. com; https: // orcid. org / 0000 - 0002 - 2558 - 1644 &amp; The Caspian Sea Basin Research Center, University of Guilan, Rasht, Guilan, Iran.</mods:affiliation>
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<mods:namePart id="1C7DF3F15F45745801DEFEAFF2490C1B">Eagderi, Soheil</mods:namePart>
<mods:affiliation id="581AF2D1563EF6BCC5BDB36CB45066BC">Department of Fisheries, Faculty of Natural Resources, University of Tehran, Karaj, Iran.</mods:affiliation>
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<mods:namePart id="B45C105159B569D41C3FA6D8808E4508">Saemi-Komsari, Maryam</mods:namePart>
<mods:nameIdentifier id="DDBB2798577946C8E296F36F5C4A56D8" type="ORCID">0000-0002-2558-1644</mods:nameIdentifier>
<mods:affiliation id="62340EBC164259C733877C2339859DB8">Department of Fisheries Sciences, Faculty of Natural Resources, University of Guilan, Sowmeh Sara, Guilan, Iran. maryam _ saemi @ yahoo. com; https: // orcid. org / 0000 - 0002 - 2558 - 1644</mods:affiliation>
<mods:nameIdentifier id="93F4D3209C49CAA8386520065796D260" type="email">maryam_saemi@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="8A5C97E06BEF4679EFD5D8A2F2113143">Kaya, Cüneyt</mods:namePart>
<mods:affiliation id="AF10254B8B5A6D9B687200A0ADD7501A">Faculty of Fisheries, Recep Tayyip Erdogan University, 53100 Rize, Turkey.</mods:affiliation>
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<mods:namePart id="6CF872BA9A5613EB9764B6E15084C224">Freyhof, Jörg</mods:namePart>
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<mods:affiliation id="3A2E51BF1185C59416067BCB2A8C5288">Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, 10115 Berlin, Germany. joerg. freyhof @ mfn. berlin; https: // orcid. org / 0000 - 0002 - 7042 - 3127</mods:affiliation>
<mods:nameIdentifier id="5935F1A40CFDF4F533E16CACEA091394" type="email">joerg.freyhof@mfn.berlin</mods:nameIdentifier>
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<mods:title id="4D1BE92B46A0207201D7FE040F273F9D">Zootaxa</mods:title>
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<mods:date id="EF2B1F30487730D4C2F858189AFDA202">2022</mods:date>
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<taxonomicName id="9BA02B935102FFCAAFDD39D1FE9836E8" authority="Mousavi-Sabet &amp; Eagderi &amp; Saemi-Komsari &amp; Kaya &amp; Freyhof, 2022" authorityName="Mousavi-Sabet &amp; Eagderi &amp; Saemi-Komsari &amp; Kaya &amp; Freyhof" authorityYear="2022" box="[151,291,1747,1773]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="423" phylum="Chordata" rank="species" species="rezai" status="sp. nov.">
<emphasis id="6ED48C025102FFCAAFDD39D1FE9836E8" bold="true" box="[151,291,1747,1773]" italics="true" pageId="4" pageNumber="423">Garra rezai</emphasis>
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,
<taxonomicNameLabel id="75E731795102FFCAAE7B39D1FE7A36E8" box="[305,449,1747,1773]" pageId="4" pageNumber="423" rank="species">new species</taxonomicNameLabel>
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<paragraph id="5C1F50105102FFCAAFDD39F5FEA93714" blockId="4.[151,449,1747,1809]" box="[151,274,1783,1809]" pageId="4" pageNumber="423">
(
<figureCitation id="C49B4C955102FFCAAFD539F5FEB03714" box="[159,267,1783,1809]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStartId-0="5.[151,250,1944,1968]" captionStartId-1="7.[151,250,1969,1993]" captionStartId-2="8.[151,250,899,923]" captionStartId-3="8.[151,250,1833,1857]" captionStartId-4="9.[151,250,1914,1938]" captionStartId-5="10.[151,250,800,824]" captionStartId-6="10.[151,250,1782,1806]" captionStartId-7="11.[151,250,1785,1809]" captionTargetBox-0="[151,1436,813,1921]" captionTargetBox-1="[209,1377,1239,1945]" captionTargetBox-2="[152,1436,181,876]" captionTargetBox-3="[151,1436,984,1810]" captionTargetBox-4="[154,1433,741,1891]" captionTargetBox-5="[151,1437,181,776]" captionTargetBox-6="[151,1436,848,1758]" captionTargetBox-7="[151,1437,417,1762]" captionTargetId-0="figure-314@5.[151,1436,813,1921]" captionTargetId-1="figure-219@7.[209,1377,1236,1945]" captionTargetId-2="figure-17@8.[152,1436,181,876]" captionTargetId-3="figure-45@8.[151,1436,984,1810]" captionTargetId-4="figure-309@9.[154,1433,741,1891]" captionTargetId-5="figure-17@10.[151,1437,181,776]" captionTargetId-6="figure-42@10.[151,1436,848,1758]" captionTargetId-7="figure-138@11.[151,1437,417,1762]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="10" captionTargetPageId-7="11" captionText-0="FIGURE 2. Garra rezai, GUIC 7979, holotype, 80 mm SL; Iran: stream Bouein-Sofla." captionText-1="FIGURE 3. Garra rezai, paratypes; from above: FSJF 4111, 59 mm SL; VMFC GRE-P5216, 34 mm SL; 28 mm SL; Iran: stream Bouein-Sofla." captionText-2="FIGURE 4. Garra rezai, paratypes; from above: FSJF 4111, 59 mm SL; VMFC GRE-P5216, 34 mm SL; 28 mm SL; Iran: stream Bouein-Sofla." captionText-3="FIGURE 5. Garra rezai, paratypes; from above: FSJF 4111, 59 mm SL; VMFC GRE-P5216, 34 mm SL; 28 mm SL; Iran: stream Bouein-Sofla." captionText-4="FIGURE 6. Garra rezai; FFR 4038, 109 mm SL; Turkey: stream Büyük." captionText-5="FIGURE 7. Garra rezai; left, FFR 1273, 107 mm SL; stream Büyük; right, FFR 1302, 113 mm SL; stream Çıratan." captionText-6="FIGURE 8. Garra rezai, from above: IMNRF-UT 1245, paratype, 58 mm SL; Iran: stream Bouein-Sofla; FSJF 3824, 104 mm SL; Turkey: stream Çıratan." captionText-7="FIGURE 9. Garra rezai, FSJF 3824, 104 mm SL; Turkey: stream Çıratan." figureDoi-0="http://doi.org/10.5281/zenodo.7223693" figureDoi-1="http://doi.org/10.5281/zenodo.7223695" figureDoi-2="http://doi.org/10.5281/zenodo.7223697" figureDoi-3="http://doi.org/10.5281/zenodo.7223699" figureDoi-4="http://doi.org/10.5281/zenodo.7223703" figureDoi-5="http://doi.org/10.5281/zenodo.7223705" figureDoi-6="http://doi.org/10.5281/zenodo.7223707" figureDoi-7="http://doi.org/10.5281/zenodo.7223709" httpUri-0="https://zenodo.org/record/7223693/files/figure.png" httpUri-1="https://zenodo.org/record/7223695/files/figure.png" httpUri-2="https://zenodo.org/record/7223697/files/figure.png" httpUri-3="https://zenodo.org/record/7223699/files/figure.png" httpUri-4="https://zenodo.org/record/7223703/files/figure.png" httpUri-5="https://zenodo.org/record/7223705/files/figure.png" httpUri-6="https://zenodo.org/record/7223707/files/figure.png" httpUri-7="https://zenodo.org/record/7223709/files/figure.png" pageId="4" pageNumber="423">Figs. 29</figureCitation>
)
</paragraph>
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<paragraph id="5C1F50105102FFCAAFDD383DFAC4375F" blockId="4.[151,1437,1855,2025]" box="[151,1407,1855,1882]" pageId="4" pageNumber="423">
<materialsCitation id="ECC85A4D5102FFCAAFDD383DFAC4375F" ID-GBIF-Occurrence="3938572303" box="[151,1407,1855,1882]" collectionCode="GUIC" country="Iran" latitude="35.9378" location="Bane" longLatPrecision="6" longitude="45.9363" municipality="Bouein-Sofla" pageId="4" pageNumber="423" specimenCode="GUIC 7979" specimenCount="1" stateProvince="Kordestan" typeStatus="holotype">
<emphasis id="6ED48C025102FFCAAFDD383DFEB0375C" bold="true" box="[151,267,1855,1881]" pageId="4" pageNumber="423">
<typeStatus id="831BEEB25102FFCAAFDD383DFEBD375C" box="[151,262,1855,1881]" pageId="4" pageNumber="423" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="0C06F86B5102FFCAAE58383DFE23375C" box="[274,408,1855,1881]" collectionCode="GUIC" pageId="4" pageNumber="423">GUIC 7979</specimenCode>
,
<quantity id="9B58FDF55102FFCAAEE9383DFE49375C" box="[419,498,1855,1881]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="423" unit="mm" value="80.0">80 mm</quantity>
SL;
<collectingCountry id="24B710805102FFCAAD63383DFDE1375C" box="[553,602,1855,1881]" name="Iran" pageId="4" pageNumber="423">Iran</collectingCountry>
:
<collectingRegion id="9E649EF25102FFCAAD2E383DFCAC375C" box="[612,791,1855,1881]" country="Iran" name="Kordestan" pageId="4" pageNumber="423">Kurdistan prov.</collectingRegion>
: stream
<collectingMunicipality id="BC7BCA6A5102FFCAAC39383DFBB0375C" box="[883,1035,1855,1881]" pageId="4" pageNumber="423">Bouein-Sofla</collectingMunicipality>
, near
<location id="597F06CB5102FFCAAB07383DFB32375C" LSID="urn:lsid:plazi:treatment:D409E1065102FFC0AFDD39D1FD683280:597F06CB5102FFCAAB07383DFB32375C" box="[1101,1161,1855,1881]" country="Iran" latitude="35.9378" longLatPrecision="6" longitude="45.9363" municipality="Bouein-Sofla" name="Bane" pageId="4" pageNumber="423" stateProvince="Kordestan">Bane</location>
,
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<geoCoordinate id="399436D75102FFCAAA41383DFAC1375F" box="[1291,1402,1855,1882]" degrees="45.9363" direction="east" orientation="longitude" pageId="4" pageNumber="423" precision="5" value="45.9363">45.9363E</geoCoordinate>
.
</materialsCitation>
</paragraph>
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<materialsCitation id="ECC85A4D5102FFCAAF8D3861FC913778" ID-GBIF-Occurrence="3938572301" box="[199,810,1891,1918]" collectionCode="FSJF" pageId="4" pageNumber="423" specimenCode="IMNRF-UT 1245, 3" specimenCount="1" typeStatus="paratype">
<emphasis id="6ED48C025102FFCAAF8D3861FE843778" bold="true" box="[199,319,1891,1917]" pageId="4" pageNumber="423">
<typeStatus id="831BEEB25102FFCAAF8D3861FE843778" box="[199,319,1891,1917]" pageId="4" pageNumber="423" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<specimenCode id="0C06F86B5102FFCAAE073861FDAC377B" box="[333,535,1891,1918]" collectionCode="IMNRF-UT" pageId="4" pageNumber="423">IMNRF-UT 1245</specimenCode>
,
<specimenCode id="0C06F86B5102FFCAAD683861FD8A3778" box="[546,561,1891,1917]" collectionCode="IMNRF-UT" pageId="4" pageNumber="423">3</specimenCode>
,
<quantity id="9B58FDF55102FFCAAD753861FD023778" box="[575,697,1891,1918]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="6.7" metricValueMin="5.3" pageId="4" pageNumber="423" unit="mm" value="60.0" valueMax="67.0" valueMin="53.0">5367 mm</quantity>
SL;
<collectionCode id="3AB1C8D55102FFCAADBA3861FC913778" box="[752,810,1891,1917]" pageId="4" pageNumber="423">FSJF</collectionCode>
</materialsCitation>
<materialsCitation id="ECC85A4D5102FFCAAC7B3861FBDC3778" ID-GBIF-Occurrence="3938572306" box="[817,1127,1891,1918]" collectionCode="VMFC" pageId="4" pageNumber="423" specimenCode="1" specimenCount="1" typeStatus="paratype">
4111,
<specimenCode id="0C06F86B5102FFCAAC3F3861FC383778" box="[885,899,1891,1917]" collectionCode="FSJF" pageId="4" pageNumber="423">1</specimenCode>
,
<quantity id="9B58FDF55102FFCAACD83861FC5A3778" box="[914,993,1891,1918]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="4" pageNumber="423" unit="mm" value="59.0">59 mm</quantity>
SL;
<collectionCode id="3AB1C8D55102FFCAAB523861FBDC3778" box="[1048,1127,1891,1917]" pageId="4" pageNumber="423">VMFC</collectionCode>
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<materialsCitation id="ECC85A4D5102FFCAAB253861FE7A37A4" ID-GBIF-Occurrence="3938572302" collectionCode="GRE-P" country="Iran" latitude="35.9378" location="Bane" longLatPrecision="6" longitude="45.9363" municipality="Bouein-Sofla" pageId="4" pageNumber="423" specimenCode="GRE-P5216, 2" specimenCount="1" stateProvince="Kordestan" typeStatus="paratype">
<specimenCode id="0C06F86B5102FFCAAB253861FB40377B" box="[1135,1275,1891,1918]" collectionCode="GRE-P" pageId="4" pageNumber="423">GRE-P5216</specimenCode>
,
<specimenCode id="0C06F86B5102FFCAAA4D3861FAAC3778" box="[1287,1303,1891,1917]" collectionCode="GRE-P" pageId="4" pageNumber="423">2</specimenCode>
,
<quantity id="9B58FDF55102FFCAAA693861FA263778" box="[1315,1437,1891,1917]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.4" metricValueMin="2.8" pageId="4" pageNumber="423" unit="mm" value="31.0" valueMax="34.0" valueMin="28.0">2834 mm</quantity>
SL; same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="5C1F50105102FFCBAF8D38A9FE5330D0" blockId="4.[151,1437,1855,2025]" lastBlockId="5.[151,1436,150,789]" lastPageId="5" lastPageNumber="424" pageId="4" pageNumber="423">
<emphasis id="6ED48C025102FFCAAF8D38A9FE7F37C0" bold="true" box="[199,452,1963,1989]" pageId="4" pageNumber="423">Additional materials.</emphasis>
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FFR 1273, 6,
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SL;
<collectingCountry id="24B710805102FFCAAC5638A9FCCB37C0" box="[796,880,1963,1989]" name="Turkey" pageId="4" pageNumber="423">Turkey</collectingCountry>
:
<collectingRegion id="9E649EF25102FFCAAC3338A9FC4237C0" box="[889,1017,1963,1989]" country="Turkey" name="Bitlis" pageId="4" pageNumber="423">Bitlis prov.</collectingRegion>
: stream
<collectingMunicipality id="BC7BCA6A5102FFCAAB1938A9FB5A37C0" box="[1107,1249,1963,1989]" pageId="4" pageNumber="423">Gümüşkanat</collectingMunicipality>
about
<quantity id="9B58FDF55102FFCAAA6138A9FAD837C0" box="[1323,1379,1963,1989]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="423" unit="km" value="3.0">3 km</quantity>
west of
<location id="597F06CB5102FFCAAFF238CDFE9F37EC" LSID="urn:lsid:plazi:treatment:D409E1065102FFC0AFDD39D1FD683280:597F06CB5102FFCAAFF238CDFE9F37EC" box="[184,292,1999,2025]" country="Turkey" latitude="38.4028" longLatPrecision="6" longitude="41.7397" municipality="Gumuskanat" name="Tasbogaz" pageId="4" pageNumber="423" stateProvince="Bitlis">Taşboğaz</location>
,
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<geoCoordinate id="399436D75102FFCAAEE038CDFDA337EC" box="[426,536,1999,2025]" degrees="41.7397" direction="east" orientation="longitude" pageId="4" pageNumber="423" precision="5" value="41.7397">41.7397E</geoCoordinate>
.—FFR 1302, 22,
<quantity id="9B58FDF55102FFCAADA038CDFCC937EC" box="[746,882,1999,2025]" metricMagnitude="-2" metricUnit="m" metricValue="9.55" metricValueMax="11.7" metricValueMin="7.4" pageId="4" pageNumber="423" unit="mm" value="95.5" valueMax="117.0" valueMin="74.0">74117 mm</quantity>
SL; FFR 1344, 26,
<quantity id="9B58FDF55102FFCAAB1438CDFB5C37EC" box="[1118,1255,1999,2025]" metricMagnitude="-2" metricUnit="m" metricValue="9.15" metricValueMax="11.4" metricValueMin="6.9" pageId="4" pageNumber="423" unit="mm" value="91.5" valueMax="114.0" valueMin="69.0">69114 mm</quantity>
SL; FFR 4019
</materialsCitation>
, 8,
<quantity id="9B58FDF55103FFCBAFFF3F95FE8630B4" box="[181,317,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.035" metricValueMax="1.19" metricValueMin="0.88" pageId="5" pageNumber="424" unit="mm" value="103.5" valueMax="119.0" valueMin="88.0">88119 mm</quantity>
SL;
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<specimenCode id="0C06F86B5103FFCBAE323F95FE5530B4" box="[376,494,151,177]" collectionCode="FFR" pageId="5" pageNumber="424">FFR 4023</specimenCode>
,
<specimenCode id="0C06F86B5103FFCBAEB13F95FDB030B4" box="[507,523,151,177]" collectionCode="FFR" pageId="5" pageNumber="424">9</specimenCode>
,
<quantity id="9B58FDF55103FFCBAD533F95FD1930B4" box="[537,674,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.085" metricValueMax="1.27" metricValueMin="0.9" pageId="5" pageNumber="424" unit="mm" value="108.5" valueMax="127.0" valueMin="90.0">90127 mm</quantity>
SL
</materialsCitation>
;
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<specimenCode id="0C06F86B5103FFCBAD973F95FCE130B4" box="[733,858,151,177]" collectionCode="FSJF" pageId="5" pageNumber="424">FSJF 3824</specimenCode>
,
<specimenCode id="0C06F86B5103FFCBAC223F95FCC330B4" box="[872,888,151,177]" collectionCode="FSJF" pageId="5" pageNumber="424">7</specimenCode>
,
<quantity id="9B58FDF55103FFCBACCC3F94FBB430B4" box="[902,1039,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" metricValueMax="10.5" metricValueMin="7.7" pageId="5" pageNumber="424" unit="mm" value="91.0" valueMax="105.0" valueMin="77.0">77105 mm</quantity>
SL;
<collectingRegion id="9E649EF25103FFCBAB033F95FB7730B4" box="[1097,1228,151,177]" country="Turkey" name="Bitlis" pageId="5" pageNumber="424">Bitlis prov.</collectingRegion>
: stream
<collectingMunicipality id="BC7BCA6A5103FFCBAA663F95FAC530B4" box="[1324,1406,151,177]" pageId="5" pageNumber="424">Çıratan</collectingMunicipality>
at
<location id="597F06CB5103FFCBAFDD3FBEFF4830D2" LSID="urn:lsid:plazi:treatment:D409E1065102FFC0AFDD39D1FD683280:597F06CB5103FFCBAFDD3FBEFF4830D2" box="[151,243,188,215]" latitude="38.3547" longLatPrecision="6" longitude="41.7814" municipality="Ciratan" name="Ucadim" pageId="5" pageNumber="424" stateProvince="Bitlis">Üçadım</location>
,
<geoCoordinate id="399436D75103FFCBAFB43FB8FED630D0" box="[254,365,186,213]" degrees="38.3547" direction="north" orientation="latitude" pageId="5" pageNumber="424" precision="5" value="38.3547">38.3547N</geoCoordinate>
<geoCoordinate id="399436D75103FFCBAE3E3FB9FE5F30D0" box="[372,484,187,213]" degrees="41.7814" direction="east" orientation="longitude" pageId="5" pageNumber="424" precision="5" value="41.7814">41.7814E</geoCoordinate>
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.
</paragraph>
<paragraph id="5C1F50105103FFCBAF8D3FDCFAB23144" blockId="5.[151,1436,150,789]" pageId="5" pageNumber="424">
<emphasis id="6ED48C025103FFCBAF8D3FDCFD0A30FD" bold="true" box="[199,689,222,249]" pageId="5" pageNumber="424">Material for molecular genetic analysis.</emphasis>
<materialsCitation id="ECC85A4D5103FFCBADF73FDDFC3C30FC" ID-GBIF-Occurrence="3938572309" box="[701,903,222,249]" collectionCode="IMNRF-UT" pageId="5" pageNumber="424" specimenCode="IMNRF-UT 1245" specimenCount="1">
<specimenCode id="0C06F86B5103FFCBADF73FDDFC3C30FC" box="[701,903,222,249]" collectionCode="IMNRF-UT" pageId="5" pageNumber="424">IMNRF-UT 1245</specimenCode>
</materialsCitation>
<materialsCitation id="ECC85A4D5103FFCBACDB3FDDFC7730FC" ID-GBIF-Occurrence="3938572311" box="[913,972,223,249]" collectionCode="AC" pageId="5" pageNumber="424" specimenCode="AC6" specimenCount="1">
<specimenCode id="0C06F86B5103FFCBACDB3FDDFC7230FC" box="[913,969,223,249]" collectionCode="AC" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="5" pageNumber="424" type="University or college">AC6</specimenCode>
,
</materialsCitation>
<materialsCitation id="ECC85A4D5103FFCBAC9D3FDDFD3B3118" ID-GBIF-Occurrence="3938572305" accessionNumber="OP184766, OP184767" collectionCode="AC" country="Iran" latitude="35.9378" location="Genbank" longLatPrecision="6" longitude="45.9363" municipality="Bouein-Sofla" pageId="5" pageNumber="424" specimenCode="AC7" specimenCount="1" stateProvince="Kordestan">
<specimenCode id="0C06F86B5103FFCBAC9D3FDDFBB430FC" box="[983,1039,223,249]" collectionCode="AC" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="5" pageNumber="424" type="University or college">AC7</specimenCode>
; same data as holotype (
<location id="597F06CB5103FFCBAA7F3FDDFA2730FC" LSID="urn:lsid:plazi:treatment:D409E1065102FFC0AFDD39D1FD683280:597F06CB5103FFCBAA7F3FDDFA2730FC" box="[1333,1436,223,249]" name="Genbank" pageId="5" pageNumber="424">Genbank</location>
accession numbers:
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,
<accessionNumber id="43F3CDF35103FFCBAD493E01FDC23118" box="[515,633,259,285]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OP184767" pageId="5" pageNumber="424">OP184767</accessionNumber>
)
</materialsCitation>
<materialsCitation id="ECC85A4D5103FFCBADD73E01FABE3144" ID-GBIF-Occurrence="3938572308" accessionNumber="OP184768 and OP184769" collectionCode="FSJF-DNA" collectorName="Genbank" country="Turkey" latitude="38.4028" location="Tasbogaz" longLatPrecision="6" longitude="41.7397" municipality="Gumuskanat" pageId="5" pageNumber="424" specimenCode="FSJF-DNA 2674" specimenCount="1" stateProvince="Bitlis">
<specimenCode id="0C06F86B5103FFCBADD73E01FCD83118" box="[669,867,259,285]" collectionCode="FSJF-DNA" pageId="5" pageNumber="424">FSJF-DNA 2674</specimenCode>
;
<collectingCountry id="24B710805103FFCBAC253E01FC783118" box="[879,963,259,285]" name="Turkey" pageId="5" pageNumber="424">Turkey</collectingCountry>
:
<collectingRegion id="9E649EF25103FFCBAC9A3E01FBE93118" box="[976,1106,259,285]" country="Turkey" name="Bitlis" pageId="5" pageNumber="424">Bitlis prov.</collectingRegion>
: stream
<collectingMunicipality id="BC7BCA6A5103FFCBABFB3E01FA843118" box="[1201,1343,259,285]" pageId="5" pageNumber="424">Gümüşkanat</collectingMunicipality>
about
<quantity id="9B58FDF55103FFCBAAC43E01FF003144" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="424" unit="km" value="3.0">3 km</quantity>
west of
<location id="597F06CB5103FFCBAE503E25FE3D3144" LSID="urn:lsid:plazi:treatment:D409E1065102FFC0AFDD39D1FD683280:597F06CB5103FFCBAE503E25FE3D3144" box="[282,390,295,321]" country="Turkey" latitude="38.4028" longLatPrecision="6" longitude="41.7397" municipality="Gumuskanat" name="Tasbogaz" pageId="5" pageNumber="424" stateProvince="Bitlis">Taşboğaz</location>
,
<geoCoordinate id="399436D75103FFCBAEDB3E25FDBA3144" box="[401,513,295,321]" degrees="38.4028" direction="north" orientation="latitude" pageId="5" pageNumber="424" precision="5" value="38.4028">38.4028N</geoCoordinate>
<geoCoordinate id="399436D75103FFCBAD423E25FDCF3144" box="[520,628,295,321]" degrees="41.7397" direction="east" orientation="longitude" pageId="5" pageNumber="424" precision="5" value="41.7397">41.7397E</geoCoordinate>
(
<collectorName id="F15535C65103FFCBADC83E25FD523144" box="[642,745,295,321]" pageId="5" pageNumber="424">Genbank</collectorName>
accession numbers:
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and
<accessionNumber id="43F3CDF35103FFCBABCA3E25FB463144" box="[1152,1277,295,321]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OP184769" pageId="5" pageNumber="424">OP184769</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="14BA039B5103FFCBAF8D3E48FE653310" pageId="5" pageNumber="424" type="diagnosis">
<paragraph id="5C1F50105103FFCBAF8D3E48FD4431D4" blockId="5.[151,1436,150,789]" pageId="5" pageNumber="424">
<emphasis id="6ED48C025103FFCBAF8D3E48FEF93161" bold="true" box="[199,322,330,356]" pageId="5" pageNumber="424">Diagnosis.</emphasis>
<taxonomicName id="9BA02B935103FFCBAE063E49FE6F3161" authorityName="Mousavi-Sabet &amp; Eagderi &amp; Saemi-Komsari &amp; Kaya &amp; Freyhof" authorityYear="2022" box="[332,468,331,357]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025103FFCBAE063E49FE6F3161" box="[332,468,331,357]" italics="true" pageId="5" pageNumber="424">Garra rezai</emphasis>
</taxonomicName>
is distinguished from the other species of the
<taxonomicName id="9BA02B935103FFCBACB13E49FB0C3160" box="[1019,1207,331,357]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="6ED48C025103FFCBACB13E49FB0C3160" box="[1019,1207,331,357]" italics="true" pageId="5" pageNumber="424">Garra variabilis</emphasis>
</taxonomicName>
species-group by a combination of characters, none unique to the species. It is distinguished from
<taxonomicName id="9BA02B935103FFCBAB413E6DFB23318C" box="[1035,1176,367,393]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="6ED48C025103FFCBAB413E6DFB23318C" box="[1035,1176,367,393]" italics="true" pageId="5" pageNumber="424">G. variabilis</emphasis>
</taxonomicName>
by having two pairs of barbels (vs. one), 1519 scales on the predorsal midline between the dorsal-fin origin and the nape (vs. 1115), and 1116 total gill rakers on the first gill arch (vs. 2026).
</paragraph>
<paragraph id="5C1F50105103FFCBAF8D3ED9FE653310" blockId="5.[151,1436,150,789]" pageId="5" pageNumber="424">
The new species is distinguished from
<taxonomicName id="9BA02B935103FFCBADC83ED9FD4B31F0" box="[642,752,475,501]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="kemali">
<emphasis id="6ED48C025103FFCBADC83ED9FD4B31F0" box="[642,752,475,501]" italics="true" pageId="5" pageNumber="424">G. kemali</emphasis>
</taxonomicName>
and
<taxonomicName id="9BA02B935103FFCBAC623ED9FC3D31F0" box="[808,902,475,501]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="klatti">
<emphasis id="6ED48C025103FFCBAC623ED9FC3D31F0" box="[808,902,475,501]" italics="true" pageId="5" pageNumber="424">G. klatti</emphasis>
</taxonomicName>
from Central
<collectingCountry id="24B710805103FFCBAB6F3ED9FB3E31F0" box="[1061,1157,475,501]" name="Turkey" pageId="5" pageNumber="424">Anatolia</collectingCountry>
by having a mental disc (vs. absent), and two pairs of barbels (vs. none). It is distinguished from
<taxonomicName id="9BA02B935103FFCBAC843EFDFBD4321C" box="[974,1135,511,537]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="nudiventris">
<emphasis id="6ED48C025103FFCBAC843EFDFBD4321C" box="[974,1135,511,537]" italics="true" pageId="5" pageNumber="424">G. nudiventris</emphasis>
</taxonomicName>
,
<taxonomicName id="9BA02B935103FFCBAB343EFDFB50321D" box="[1150,1259,511,537]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="roseae">
<emphasis id="6ED48C025103FFCBAB343EFDFB50321D" box="[1150,1259,511,537]" italics="true" pageId="5" pageNumber="424">G. roseae</emphasis>
</taxonomicName>
, and
<taxonomicName id="9BA02B935103FFCBAA633EFDFA27321D" box="[1321,1436,511,537]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="rossica">
<emphasis id="6ED48C025103FFCBAA633EFDFA27321D" box="[1321,1436,511,537]" italics="true" pageId="5" pageNumber="424">G. rossica</emphasis>
</taxonomicName>
by having two pairs of barbels (vs. no barbels in
<taxonomicName id="9BA02B935103FFCBAD893D21FC8B3239" box="[707,816,547,573]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="roseae">
<emphasis id="6ED48C025103FFCBAD893D21FC8B3239" box="[707,816,547,573]" italics="true" pageId="5" pageNumber="424">G. roseae</emphasis>
</taxonomicName>
, one pair in
<taxonomicName id="9BA02B935103FFCBAC8A3D21FBE23238" box="[960,1113,547,573]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="nudiventris">
<emphasis id="6ED48C025103FFCBAC8A3D21FBE23238" box="[960,1113,547,573]" italics="true" pageId="5" pageNumber="424">G.nudiventris</emphasis>
</taxonomicName>
, and most populations of
<taxonomicName id="9BA02B935103FFCBAACB3D21FF5C3265" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="rossica">
<emphasis id="6ED48C025103FFCBAACB3D21FF5C3265" italics="true" pageId="5" pageNumber="424">G. rossica</emphasis>
</taxonomicName>
; two pairs in some populations of
<taxonomicName id="9BA02B935103FFCBADCA3D45FD4E3265" box="[640,757,583,609]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="rossica">
<emphasis id="6ED48C025103FFCBADCA3D45FD4E3265" box="[640,757,583,609]" italics="true" pageId="5" pageNumber="424">G. rossica</emphasis>
</taxonomicName>
), a well-developed mental disc (vs. reduced in
<taxonomicName id="9BA02B935103FFCBAA553D45FA2E3265" box="[1311,1429,583,609]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="rossica">
<emphasis id="6ED48C025103FFCBAA553D45FA2E3265" box="[1311,1429,583,609]" italics="true" pageId="5" pageNumber="424">G. rossica</emphasis>
</taxonomicName>
;
<figureCitation id="C49B4C955103FFCBAFDD3D69FF513280" box="[151,234,619,645]" captionStart="FIGURE 10" captionStartId="12.[151,250,729,753]" captionTargetBox="[151,1436,181,705]" captionTargetId="figure-17@12.[151,1436,181,705]" captionTargetPageId="12" captionText="FIGURE 10. From left: Garra rezai, GUIC 7979, holotype, 80 mm SL; Iran: stream Bouein-Sofla; Garra rossica, VMFC GROS, 86 mm SL; Iran: spring Golbahar." figureDoi="http://doi.org/10.5281/zenodo.7223711" httpUri="https://zenodo.org/record/7223711/files/figure.png" pageId="5" pageNumber="424">Fig. 10</figureCitation>
), the predorsal mid-line fully covered by scales (vs. naked in
<taxonomicName id="9BA02B935103FFCBACE43D69FBEB3280" box="[942,1104,619,645]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="nudiventris">
<emphasis id="6ED48C025103FFCBACE43D69FBEB3280" box="[942,1104,619,645]" italics="true" pageId="5" pageNumber="424">G. nudiventris</emphasis>
</taxonomicName>
), the belly covered by scales (vs. naked in
<taxonomicName id="9BA02B935103FFCBAE793D8DFE6E32AC" box="[307,469,655,681]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="nudiventris">
<emphasis id="6ED48C025103FFCBAE793D8DFE6E32AC" box="[307,469,655,681]" italics="true" pageId="5" pageNumber="424">G. nudiventris</emphasis>
</taxonomicName>
).
<taxonomicName id="9BA02B935103FFCBAEA43D8DFDCF32AD" authorityName="Mousavi-Sabet &amp; Eagderi &amp; Saemi-Komsari &amp; Kaya &amp; Freyhof" authorityYear="2022" box="[494,628,655,681]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025103FFCBAEA43D8DFDCF32AD" box="[494,628,655,681]" italics="true" pageId="5" pageNumber="424">Garra rezai</emphasis>
</taxonomicName>
is further distinguished from
<taxonomicName id="9BA02B935103FFCBAC813D8DFB8232AD" box="[971,1081,655,681]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="roseae">
<emphasis id="6ED48C025103FFCBAC813D8DFB8232AD" box="[971,1081,655,681]" italics="true" pageId="5" pageNumber="424">G. roseae</emphasis>
</taxonomicName>
by having 3540 scales on the lateral line (vs. 4258), 5½6½ transverse scale rows between the lateral line and the dorsal-fin origin (vs. 7½8½), 4½5½ transverse scale rows between the lateral line and the pelvic-fin origin (vs. 6½), axillary scale present at the pelvic-fin origin (vs. absent).
</paragraph>
</subSubSection>
<caption id="08DF00985103FFCBAFDD389AFBB637B4" ID-DOI="http://doi.org/10.5281/zenodo.7223693" ID-Zenodo-Dep="7223693" box="[151,1037,1944,1969]" httpUri="https://zenodo.org/record/7223693/files/figure.png" pageId="5" pageNumber="424" startId="5.[151,250,1944,1968]" targetBox="[151,1436,813,1921]" targetPageId="5">
<paragraph id="5C1F50105103FFCBAFDD389AFBB637B4" blockId="5.[151,1037,1944,1969]" box="[151,1037,1944,1969]" pageId="5" pageNumber="424">
<emphasis id="6ED48C025103FFCBAFDD389AFEA837B5" bold="true" box="[151,275,1944,1968]" pageId="5" pageNumber="424">FIGURE 2.</emphasis>
<taxonomicName id="9BA02B935103FFCBAE53389BFE6037B4" authority=", GUIC" authorityName="GUIC" box="[281,475,1945,1969]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="424" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025103FFCBAE53389BFE2A37B5" box="[281,401,1945,1968]" italics="true" pageId="5" pageNumber="424">Garra rezai</emphasis>
, GUIC
</taxonomicName>
7979, holotype, 80 mm SL; Iran: stream Bouein-Sofla.
</paragraph>
</caption>
<subSubSection id="14BA039B5100FFC5AF8D3F94FB78318C" lastPageId="11" lastPageNumber="430" pageId="6" pageNumber="425" type="description">
<paragraph id="5C1F50105100FFC8AF8D3F94FAB13384" blockId="6.[151,1437,150,897]" pageId="6" pageNumber="425">
<emphasis id="6ED48C025100FFC8AF8D3F94FEE230B5" bold="true" box="[199,345,150,176]" pageId="6" pageNumber="425">Description.</emphasis>
For general appearance, see
<figureCitation id="C49B4C955100FFC8ADFF3F95FC9D30B4" box="[693,806,151,177]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStartId-0="5.[151,250,1944,1968]" captionStartId-1="7.[151,250,1969,1993]" captionStartId-2="8.[151,250,899,923]" captionStartId-3="8.[151,250,1833,1857]" captionStartId-4="9.[151,250,1914,1938]" captionStartId-5="10.[151,250,800,824]" captionStartId-6="10.[151,250,1782,1806]" captionStartId-7="11.[151,250,1785,1809]" captionTargetBox-0="[151,1436,813,1921]" captionTargetBox-1="[209,1377,1239,1945]" captionTargetBox-2="[152,1436,181,876]" captionTargetBox-3="[151,1436,984,1810]" captionTargetBox-4="[154,1433,741,1891]" captionTargetBox-5="[151,1437,181,776]" captionTargetBox-6="[151,1436,848,1758]" captionTargetBox-7="[151,1437,417,1762]" captionTargetId-0="figure-314@5.[151,1436,813,1921]" captionTargetId-1="figure-219@7.[209,1377,1236,1945]" captionTargetId-2="figure-17@8.[152,1436,181,876]" captionTargetId-3="figure-45@8.[151,1436,984,1810]" captionTargetId-4="figure-309@9.[154,1433,741,1891]" captionTargetId-5="figure-17@10.[151,1437,181,776]" captionTargetId-6="figure-42@10.[151,1436,848,1758]" captionTargetId-7="figure-138@11.[151,1437,417,1762]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="10" captionTargetPageId-7="11" captionText-0="FIGURE 2. Garra rezai, GUIC 7979, holotype, 80 mm SL; Iran: stream Bouein-Sofla." captionText-1="FIGURE 3. Garra rezai, paratypes; from above: FSJF 4111, 59 mm SL; VMFC GRE-P5216, 34 mm SL; 28 mm SL; Iran: stream Bouein-Sofla." captionText-2="FIGURE 4. Garra rezai, paratypes; from above: FSJF 4111, 59 mm SL; VMFC GRE-P5216, 34 mm SL; 28 mm SL; Iran: stream Bouein-Sofla." captionText-3="FIGURE 5. Garra rezai, paratypes; from above: FSJF 4111, 59 mm SL; VMFC GRE-P5216, 34 mm SL; 28 mm SL; Iran: stream Bouein-Sofla." captionText-4="FIGURE 6. Garra rezai; FFR 4038, 109 mm SL; Turkey: stream Büyük." captionText-5="FIGURE 7. Garra rezai; left, FFR 1273, 107 mm SL; stream Büyük; right, FFR 1302, 113 mm SL; stream Çıratan." captionText-6="FIGURE 8. Garra rezai, from above: IMNRF-UT 1245, paratype, 58 mm SL; Iran: stream Bouein-Sofla; FSJF 3824, 104 mm SL; Turkey: stream Çıratan." captionText-7="FIGURE 9. Garra rezai, FSJF 3824, 104 mm SL; Turkey: stream Çıratan." figureDoi-0="http://doi.org/10.5281/zenodo.7223693" figureDoi-1="http://doi.org/10.5281/zenodo.7223695" figureDoi-2="http://doi.org/10.5281/zenodo.7223697" figureDoi-3="http://doi.org/10.5281/zenodo.7223699" figureDoi-4="http://doi.org/10.5281/zenodo.7223703" figureDoi-5="http://doi.org/10.5281/zenodo.7223705" figureDoi-6="http://doi.org/10.5281/zenodo.7223707" figureDoi-7="http://doi.org/10.5281/zenodo.7223709" httpUri-0="https://zenodo.org/record/7223693/files/figure.png" httpUri-1="https://zenodo.org/record/7223695/files/figure.png" httpUri-2="https://zenodo.org/record/7223697/files/figure.png" httpUri-3="https://zenodo.org/record/7223699/files/figure.png" httpUri-4="https://zenodo.org/record/7223703/files/figure.png" httpUri-5="https://zenodo.org/record/7223705/files/figure.png" httpUri-6="https://zenodo.org/record/7223707/files/figure.png" httpUri-7="https://zenodo.org/record/7223709/files/figure.png" pageId="6" pageNumber="425">Figs. 29</figureCitation>
; morphometric data are provided in
<tableCitation id="112265AB5100FFC8AB933F95FAEB30B4" box="[1241,1360,151,177]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="6.[152,245,944,968]" captionStartId-1="7.[152,245,151,175]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="TABLE 3. Morphometric data of Garra rezai. Holotype, GUIC 7979, paratypes, IMNRF-UT 1245 (n=3), VMFC GR-P (n=2), FSJF 4111 (n=1). The holotype is included in the calculations." captionText-1="TABLE 4. Morphometric data of Garra rezai. Additional materials, FFR 4019 (n=11); FFR 1344 (n=6); FFR 1307 (n=3); FFR 1273 (n=3)." httpUri-0="http://table.plazi.org/id/08DF00985100FFC8AFD23CB2FCC233E9" pageId="6" pageNumber="425" tableUuid-0="08DF00985100FFC8AFD23CB2FCC233E9">Table 34</tableCitation>
. Body cylindrical, compressed laterally at caudal peduncle, body height almost equal to its width. Dorsal head profile rising gently, slightly convex. Predorsal contour slightly convex between nape and dorsal-fin origin. Ventral profile almost straight between pelvic and anal-fin origins. Body deepest at about dorsal-fin origin or about middle between nape and dorsal-fin origin, depth decreasing towards caudal-fin base. Greatest body width at about middle between pectoral- and pelvic-fin bases, decreasing towards caudal-fin base. Caudal peduncle 1.11.4 times longer than deep. Head moderately small, section roundish, flattened on ventral surface, slightly depressed, almost conical. Head slightly convex or flat in interorbital space. Height at nape shorter than head length. Width at nape greater than depth at nape. Head length 0.91.1 times in body depth. Snout length 1.01.3 times in postorbital length. Small tubercles scattered on transverse lobe, transverse lobe demarcated posteriorly by a shallow transverse groove. No tubercle on proboscis and lateral surface of snout. Depressed rostral surface with scattered, small tubercles, slightly or not separating transverse lobe from lateral surface. No groove between transverse lobe and lateral surface. Eyes large, its diameter 2.34.1 times in head depth at eye, 2.13.3 times in interorbital width. Two pairs of short barbels, rostral barbel antero-laterally located, shorter than eye diameter. Maxillary barbel at corner of mouth, same size as rostral barbel. Rostral cap well-developed, fimbriate, papillate on ventral surface. Upper lip present. Upper jaw usually covered by rostral cap. Gular disc elliptical, shorter than wide and narrower than head width. Papillae on anterior fold of gular disc of same size, regularly arranged. A deep and narrow groove between antero-median fold and central callous-pad. Latero-posterior flap present. Surface of central callous pad without or with sparsely arranged small papillae. Posterior margin of central callous pad extending vertically to anterior edge of eye. Nostril located immediately anterior to eye. Anterior nostril opening developed as a low, pointed and flap-like tube. Posterior nostril narrow, nostrils adjacent, posterior tip of anterior nostril reaching posterior nostril when folded down.
</paragraph>
<caption id="08DF00985100FFC8AFD23CB2FCC233E9" ID-Table-UUID="08DF00985100FFC8AFD23CB2FCC233E9" httpUri="http://table.plazi.org/id/08DF00985100FFC8AFD23CB2FCC233E9" pageId="6" pageNumber="425" startId="6.[152,245,944,968]" targetBox="[166,1386,1021,1970]" targetIsTable="true" targetPageId="6">
<paragraph id="5C1F50105100FFC8AFD23CB2FCC233E9" blockId="6.[151,1436,944,1005]" pageId="6" pageNumber="425">
<emphasis id="6ED48C025100FFC8AFD23CB2FEB433CD" bold="true" box="[152,271,944,968]" pageId="6" pageNumber="425">TABLE 3.</emphasis>
Morphometric data of
<taxonomicName id="9BA02B935100FFC8AD4D3CB3FD3E33CD" box="[519,645,945,968]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="425" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025100FFC8AD4D3CB3FD3E33CD" box="[519,645,945,968]" italics="true" pageId="6" pageNumber="425">Garra rezai</emphasis>
</taxonomicName>
. Holotype, GUIC 7979, paratypes, IMNRF-UT 1245 (n=3), VMFC GR-P (n=2), FSJF 4111 (n=1). The holotype is included in the calculations.
</paragraph>
</caption>
<paragraph id="5C1F50105100FFC8AC203CFFFAD137B7" pageId="6" pageNumber="425">
<table id="2EA0A2B051000031AFEC3CFFFAD137B7" box="[166,1386,1021,1970]" gridcols="6" gridrows="24" pageId="6" pageNumber="425" rowsContinueIn="7.[166,1376,228,1177]">
<tr id="E290525251000031AFEC3CFFFAD13410" box="[166,1386,1021,1045]" gridrow="0" pageId="6" pageNumber="425" rowspan-0="1" rowspan-5="1">
<th id="A1413B2E51000031AC203CFFFC783410" box="[874,963,1021,1045]" gridcol="1" gridrow="0" pageId="6" pageNumber="425">holotype</th>
<th id="A1413B2E51000031AB4C3CFFFA8B3410" box="[1030,1328,1021,1045]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="6" pageNumber="425">holotype and paratypes (n=7)</th>
</tr>
<tr id="E290525251000031AFEC3B27FAD13438" box="[166,1386,1061,1085]" gridrow="1" pageId="6" pageNumber="425" rowspan-0="1" rowspan-1="1">
<td id="A1413B2E51000031AB4C3B27FB893438" box="[1030,1074,1061,1085]" gridcol="2" gridrow="1" pageId="6" pageNumber="425">min</td>
<td id="A1413B2E51000031AB2C3B27FB293438" box="[1126,1170,1061,1085]" gridcol="3" gridrow="1" pageId="6" pageNumber="425">max</td>
<td id="A1413B2E51000031AB913B27FA8B3438" box="[1243,1328,1061,1085]" gridcol="4" gridrow="1" pageId="6" pageNumber="425">mean</td>
<td id="A1413B2E51000031AA003B27FAD13438" box="[1354,1386,1061,1085]" gridcol="5" gridrow="1" pageId="6" pageNumber="425">SD</td>
</tr>
<tr id="E290525251000031AFEC3B4FFAD13460" box="[166,1386,1101,1125]" gridrow="2" pageId="6" pageNumber="425" rowspan-4="1" rowspan-5="1">
<th id="A1413B2E51000031AFEC3B4FFD2C3460" box="[166,663,1101,1125]" gridcol="0" gridrow="2" pageId="6" pageNumber="425">Standard length (mm)</th>
<td id="A1413B2E51000031AC203B4FFC783460" box="[874,963,1101,1125]" gridcol="1" gridrow="2" pageId="6" pageNumber="425">80.0</td>
<td id="A1413B2E51000031AB4C3B4FFB893460" box="[1030,1074,1101,1125]" gridcol="2" gridrow="2" pageId="6" pageNumber="425">28.0</td>
<td id="A1413B2E51000031AB2C3B4FFB293460" box="[1126,1170,1101,1125]" gridcol="3" gridrow="2" pageId="6" pageNumber="425">67.0</td>
</tr>
<tr id="E290525251000031AFEC3B77FAD13488" box="[166,1386,1141,1165]" gridrow="3" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3B77FAD13488" box="[166,1386,1141,1165]" colspan="6" colspanRight="5" gridcol="0" gridrow="3" pageId="6" pageNumber="425">
<emphasis id="6ED48C025100FFC8AFEC3B77FE593488" bold="true" box="[166,482,1141,1165]" pageId="6" pageNumber="425">In percent of standard length</emphasis>
</th>
</tr>
<tr id="E290525251000031AFEC3B9CFAD134B3" box="[166,1386,1182,1206]" gridrow="4" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3B9CFD2C34B3" box="[166,663,1182,1206]" gridcol="0" gridrow="4" pageId="6" pageNumber="425">Head length</th>
<td id="A1413B2E51000031AC203B9CFC7834B3" box="[874,963,1182,1206]" gridcol="1" gridrow="4" pageId="6" pageNumber="425">19.4</td>
<td id="A1413B2E51000031AB4C3B9CFB8934B3" box="[1030,1074,1182,1206]" gridcol="2" gridrow="4" pageId="6" pageNumber="425">18.7</td>
<td id="A1413B2E51000031AB2C3B9CFB2934B3" box="[1126,1170,1182,1206]" gridcol="3" gridrow="4" pageId="6" pageNumber="425">23.2</td>
<td id="A1413B2E51000031AB913B9CFA8B34B3" box="[1243,1328,1182,1206]" gridcol="4" gridrow="4" pageId="6" pageNumber="425">20.8</td>
<td id="A1413B2E51000031AA003B9CFAD134B3" box="[1354,1386,1182,1206]" gridcol="5" gridrow="4" pageId="6" pageNumber="425">2.1</td>
</tr>
<tr id="E290525251000031AFEC3BC4FAD134DB" box="[166,1386,1222,1246]" gridrow="5" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3BC4FD2C34DB" box="[166,663,1222,1246]" gridcol="0" gridrow="5" pageId="6" pageNumber="425">Body depth at dorsal-fin origin</th>
<td id="A1413B2E51000031AC203BC4FC7834DB" box="[874,963,1222,1246]" gridcol="1" gridrow="5" pageId="6" pageNumber="425">21.9</td>
<td id="A1413B2E51000031AB4C3BC4FB8934DB" box="[1030,1074,1222,1246]" gridcol="2" gridrow="5" pageId="6" pageNumber="425">17.4</td>
<td id="A1413B2E51000031AB2C3BC4FB2934DB" box="[1126,1170,1222,1246]" gridcol="3" gridrow="5" pageId="6" pageNumber="425">22.4</td>
<td id="A1413B2E51000031AB913BC4FA8B34DB" box="[1243,1328,1222,1246]" gridcol="4" gridrow="5" pageId="6" pageNumber="425">20.0</td>
<td id="A1413B2E51000031AA003BC4FAD134DB" box="[1354,1386,1222,1246]" gridcol="5" gridrow="5" pageId="6" pageNumber="425">2.5</td>
</tr>
<tr id="E290525251000031AFEC3BECFAD13503" box="[166,1386,1262,1286]" gridrow="6" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3BECFD2C3503" box="[166,663,1262,1286]" gridcol="0" gridrow="6" pageId="6" pageNumber="425">Predorsal length</th>
<td id="A1413B2E51000031AC203BECFC783503" box="[874,963,1262,1286]" gridcol="1" gridrow="6" pageId="6" pageNumber="425">47.2</td>
<td id="A1413B2E51000031AB4C3BECFB893503" box="[1030,1074,1262,1286]" gridcol="2" gridrow="6" pageId="6" pageNumber="425">47.1</td>
<td id="A1413B2E51000031AB2C3BECFB293503" box="[1126,1170,1262,1286]" gridcol="3" gridrow="6" pageId="6" pageNumber="425">55.6</td>
<td id="A1413B2E51000031AB913BECFA8B3503" box="[1243,1328,1262,1286]" gridcol="4" gridrow="6" pageId="6" pageNumber="425">49.9</td>
<td id="A1413B2E51000031AA003BECFAD13503" box="[1354,1386,1262,1286]" gridcol="5" gridrow="6" pageId="6" pageNumber="425">4.0</td>
</tr>
<tr id="E290525251000031AFEC3A14FAD1352B" box="[166,1386,1302,1326]" gridrow="7" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3A14FD2C352B" box="[166,663,1302,1326]" gridcol="0" gridrow="7" pageId="6" pageNumber="425">Postdorsal length</th>
<td id="A1413B2E51000031AC203A14FC78352B" box="[874,963,1302,1326]" gridcol="1" gridrow="7" pageId="6" pageNumber="425">36.7</td>
<td id="A1413B2E51000031AB4C3A14FB89352B" box="[1030,1074,1302,1326]" gridcol="2" gridrow="7" pageId="6" pageNumber="425">31.5</td>
<td id="A1413B2E51000031AB2C3A14FB29352B" box="[1126,1170,1302,1326]" gridcol="3" gridrow="7" pageId="6" pageNumber="425">38.1</td>
<td id="A1413B2E51000031AB913A14FA8B352B" box="[1243,1328,1302,1326]" gridcol="4" gridrow="7" pageId="6" pageNumber="425">35.8</td>
<td id="A1413B2E51000031AA003A14FAD1352B" box="[1354,1386,1302,1326]" gridcol="5" gridrow="7" pageId="6" pageNumber="425">2.9</td>
</tr>
<tr id="E290525251000031AFEC3A3DFAD13552" box="[166,1386,1343,1367]" gridrow="8" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3A3DFD2C3552" box="[166,663,1343,1367]" gridcol="0" gridrow="8" pageId="6" pageNumber="425">Preanal length</th>
<td id="A1413B2E51000031AC203A3DFC783552" box="[874,963,1343,1367]" gridcol="1" gridrow="8" pageId="6" pageNumber="425">74.9</td>
<td id="A1413B2E51000031AB4C3A3DFB893552" box="[1030,1074,1343,1367]" gridcol="2" gridrow="8" pageId="6" pageNumber="425">74.0</td>
<td id="A1413B2E51000031AB2C3A3DFB293552" box="[1126,1170,1343,1367]" gridcol="3" gridrow="8" pageId="6" pageNumber="425">82.1</td>
<td id="A1413B2E51000031AB913A3DFA8B3552" box="[1243,1328,1343,1367]" gridcol="4" gridrow="8" pageId="6" pageNumber="425">76.9</td>
<td id="A1413B2E51000031AA003A3DFAD13552" box="[1354,1386,1343,1367]" gridcol="5" gridrow="8" pageId="6" pageNumber="425">3.7</td>
</tr>
<tr id="E290525251000031AFEC3A65FAD1357A" box="[166,1386,1383,1407]" gridrow="9" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3A65FD2C357A" box="[166,663,1383,1407]" gridcol="0" gridrow="9" pageId="6" pageNumber="425">Prepelvic length</th>
<td id="A1413B2E51000031AC203A65FC78357A" box="[874,963,1383,1407]" gridcol="1" gridrow="9" pageId="6" pageNumber="425">52.9</td>
<td id="A1413B2E51000031AB4C3A65FB89357A" box="[1030,1074,1383,1407]" gridcol="2" gridrow="9" pageId="6" pageNumber="425">50.6</td>
<td id="A1413B2E51000031AB2C3A65FB29357A" box="[1126,1170,1383,1407]" gridcol="3" gridrow="9" pageId="6" pageNumber="425">61.2</td>
<td id="A1413B2E51000031AB913A65FA8B357A" box="[1243,1328,1383,1407]" gridcol="4" gridrow="9" pageId="6" pageNumber="425">55.0</td>
<td id="A1413B2E51000031AA003A65FAD1357A" box="[1354,1386,1383,1407]" gridcol="5" gridrow="9" pageId="6" pageNumber="425">4.6</td>
</tr>
<tr id="E290525251000031AFEC3A8DFAD135A2" box="[166,1386,1423,1447]" gridrow="10" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3A8DFD2C35A2" box="[166,663,1423,1447]" gridcol="0" gridrow="10" pageId="6" pageNumber="425">Distance between pectoral and pelvic-fin origins</th>
<td id="A1413B2E51000031AC203A8DFC7835A2" box="[874,963,1423,1447]" gridcol="1" gridrow="10" pageId="6" pageNumber="425">36.2</td>
<td id="A1413B2E51000031AB4C3A8DFB8935A2" box="[1030,1074,1423,1447]" gridcol="2" gridrow="10" pageId="6" pageNumber="425">32.0</td>
<td id="A1413B2E51000031AB2C3A8DFB2935A2" box="[1126,1170,1423,1447]" gridcol="3" gridrow="10" pageId="6" pageNumber="425">36.2</td>
<td id="A1413B2E51000031AB913A8DFA8B35A2" box="[1243,1328,1423,1447]" gridcol="4" gridrow="10" pageId="6" pageNumber="425">34.5</td>
<td id="A1413B2E51000031AA003A8DFAD135A2" box="[1354,1386,1423,1447]" gridcol="5" gridrow="10" pageId="6" pageNumber="425">2.0</td>
</tr>
<tr id="E290525251000031AFEC3AB5FAD135CA" box="[166,1386,1463,1487]" gridrow="11" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3AB5FD2C35CA" box="[166,663,1463,1487]" gridcol="0" gridrow="11" pageId="6" pageNumber="425">Distance between pelvic and anal-fin origins</th>
<td id="A1413B2E51000031AC203AB5FC7835CA" box="[874,963,1463,1487]" gridcol="1" gridrow="11" pageId="6" pageNumber="425">21.7</td>
<td id="A1413B2E51000031AB4C3AB5FB8935CA" box="[1030,1074,1463,1487]" gridcol="2" gridrow="11" pageId="6" pageNumber="425">21.3</td>
<td id="A1413B2E51000031AB2C3AB5FB2935CA" box="[1126,1170,1463,1487]" gridcol="3" gridrow="11" pageId="6" pageNumber="425">23.4</td>
<td id="A1413B2E51000031AB913AB5FA8B35CA" box="[1243,1328,1463,1487]" gridcol="4" gridrow="11" pageId="6" pageNumber="425">22.4</td>
<td id="A1413B2E51000031AA003AB5FAD135CA" box="[1354,1386,1463,1487]" gridcol="5" gridrow="11" pageId="6" pageNumber="425">1.1</td>
</tr>
<tr id="E290525251000031AFEC3AE2FAD135FD" box="[166,1386,1504,1528]" gridrow="12" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3AE2FD2C35FD" box="[166,663,1504,1528]" gridcol="0" gridrow="12" pageId="6" pageNumber="425">Depth of caudal peduncle</th>
<td id="A1413B2E51000031AC203AE2FC7835FD" box="[874,963,1504,1528]" gridcol="1" gridrow="12" pageId="6" pageNumber="425">12.4</td>
<td id="A1413B2E51000031AB4C3AE2FB8935FD" box="[1030,1074,1504,1528]" gridcol="2" gridrow="12" pageId="6" pageNumber="425">11.0</td>
<td id="A1413B2E51000031AB2C3AE2FB2935FD" box="[1126,1170,1504,1528]" gridcol="3" gridrow="12" pageId="6" pageNumber="425">12.8</td>
<td id="A1413B2E51000031AB913AE2FA8B35FD" box="[1243,1328,1504,1528]" gridcol="4" gridrow="12" pageId="6" pageNumber="425">11.9</td>
<td id="A1413B2E51000031AA003AE2FAD135FD" box="[1354,1386,1504,1528]" gridcol="5" gridrow="12" pageId="6" pageNumber="425">0.8</td>
</tr>
<tr id="E290525251000031AFEC390AFAD13625" box="[166,1386,1544,1568]" gridrow="13" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC390AFD2C3625" box="[166,663,1544,1568]" gridcol="0" gridrow="13" pageId="6" pageNumber="425">Length of caudal peduncle</th>
<td id="A1413B2E51000031AC20390AFC783625" box="[874,963,1544,1568]" gridcol="1" gridrow="13" pageId="6" pageNumber="425">15.8</td>
<td id="A1413B2E51000031AB4C390AFB893625" box="[1030,1074,1544,1568]" gridcol="2" gridrow="13" pageId="6" pageNumber="425">13.5</td>
<td id="A1413B2E51000031AB2C390AFB293625" box="[1126,1170,1544,1568]" gridcol="3" gridrow="13" pageId="6" pageNumber="425">15.8</td>
<td id="A1413B2E51000031AB91390AFA8B3625" box="[1243,1328,1544,1568]" gridcol="4" gridrow="13" pageId="6" pageNumber="425">14.8</td>
<td id="A1413B2E51000031AA00390AFAD13625" box="[1354,1386,1544,1568]" gridcol="5" gridrow="13" pageId="6" pageNumber="425">1.1</td>
</tr>
<tr id="E290525251000031AFEC3932FAD1364D" box="[166,1386,1584,1608]" gridrow="14" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3932FD2C364D" box="[166,663,1584,1608]" gridcol="0" gridrow="14" pageId="6" pageNumber="425">Anal-fin base length</th>
<td id="A1413B2E51000031AC203932FC78364D" box="[874,963,1584,1608]" gridcol="1" gridrow="14" pageId="6" pageNumber="425">7.8</td>
<td id="A1413B2E51000031AB4C3932FB89364D" box="[1030,1074,1584,1608]" gridcol="2" gridrow="14" pageId="6" pageNumber="425">7.0</td>
<td id="A1413B2E51000031AB2C3932FB29364D" box="[1126,1170,1584,1608]" gridcol="3" gridrow="14" pageId="6" pageNumber="425">9.4</td>
<td id="A1413B2E51000031AB913932FA8B364D" box="[1243,1328,1584,1608]" gridcol="4" gridrow="14" pageId="6" pageNumber="425">8.2</td>
<td id="A1413B2E51000031AA003932FAD1364D" box="[1354,1386,1584,1608]" gridcol="5" gridrow="14" pageId="6" pageNumber="425">1.0</td>
</tr>
<tr id="E290525251000031AFEC395AFAD13675" box="[166,1386,1624,1648]" gridrow="15" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC395AFD2C3675" box="[166,663,1624,1648]" gridcol="0" gridrow="15" pageId="6" pageNumber="425">Pectoral-fin length</th>
<td id="A1413B2E51000031AC20395AFC783675" box="[874,963,1624,1648]" gridcol="1" gridrow="15" pageId="6" pageNumber="425">17.2</td>
<td id="A1413B2E51000031AB4C395AFB893675" box="[1030,1074,1624,1648]" gridcol="2" gridrow="15" pageId="6" pageNumber="425">17.2</td>
<td id="A1413B2E51000031AB2C395AFB293675" box="[1126,1170,1624,1648]" gridcol="3" gridrow="15" pageId="6" pageNumber="425">19.9</td>
<td id="A1413B2E51000031AB91395AFA8B3675" box="[1243,1328,1624,1648]" gridcol="4" gridrow="15" pageId="6" pageNumber="425">19.0</td>
<td id="A1413B2E51000031AA00395AFAD13675" box="[1354,1386,1624,1648]" gridcol="5" gridrow="15" pageId="6" pageNumber="425">1.2</td>
</tr>
<tr id="E290525251000031AFEC3982FAD1369C" box="[166,1386,1664,1689]" gridrow="16" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3982FD2C369C" box="[166,663,1664,1689]" gridcol="0" gridrow="16" pageId="6" pageNumber="425">Pelvic-fin length</th>
<td id="A1413B2E51000031AC203982FC78369C" box="[874,963,1664,1689]" gridcol="1" gridrow="16" pageId="6" pageNumber="425">15.8</td>
<td id="A1413B2E51000031AB4C3982FB89369C" box="[1030,1074,1664,1689]" gridcol="2" gridrow="16" pageId="6" pageNumber="425">15.5</td>
<td id="A1413B2E51000031AB2C3982FB29369C" box="[1126,1170,1664,1689]" gridcol="3" gridrow="16" pageId="6" pageNumber="425">17.1</td>
<td id="A1413B2E51000031AB913982FA8B369C" box="[1243,1328,1664,1689]" gridcol="4" gridrow="16" pageId="6" pageNumber="425">16.2</td>
<td id="A1413B2E51000031AA003982FAD1369C" box="[1354,1386,1664,1689]" gridcol="5" gridrow="16" pageId="6" pageNumber="425">0.7</td>
</tr>
<tr id="E290525251000031AFEC39AAFAD136C4" box="[166,1386,1704,1729]" gridrow="17" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC39AAFAD136C4" box="[166,1386,1704,1729]" colspan="6" colspanRight="5" gridcol="0" gridrow="17" pageId="6" pageNumber="425">
<emphasis id="6ED48C025100FFC8AFEC39ABFE0E36C4" bold="true" box="[166,437,1704,1729]" pageId="6" pageNumber="425">In percent of head length</emphasis>
</th>
</tr>
<tr id="E290525251000031AFEC39D3FAD136EF" box="[166,1386,1745,1770]" gridrow="18" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC39D3FD2C36EF" box="[166,663,1745,1770]" gridcol="0" gridrow="18" pageId="6" pageNumber="425">Head depth at eye</th>
<td id="A1413B2E51000031AC2039D3FC7836EF" box="[874,963,1745,1770]" gridcol="1" gridrow="18" pageId="6" pageNumber="425">61</td>
<td id="A1413B2E51000031AB4C39D3FB8936EF" box="[1030,1074,1745,1770]" gridcol="2" gridrow="18" pageId="6" pageNumber="425">60</td>
<td id="A1413B2E51000031AB2C39D3FB2936EF" box="[1126,1170,1745,1770]" gridcol="3" gridrow="18" pageId="6" pageNumber="425">61</td>
<td id="A1413B2E51000031AB9139D3FA8B36EF" box="[1243,1328,1745,1770]" gridcol="4" gridrow="18" pageId="6" pageNumber="425">60.5</td>
<td id="A1413B2E51000031AA0039D3FAD136EF" box="[1354,1386,1745,1770]" gridcol="5" gridrow="18" pageId="6" pageNumber="425">0.2</td>
</tr>
<tr id="E290525251000031AFEC39FBFAD13714" box="[166,1386,1785,1809]" gridrow="19" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC39FBFD2C3714" box="[166,663,1785,1809]" gridcol="0" gridrow="19" pageId="6" pageNumber="425">Snout length</th>
<td id="A1413B2E51000031AC2039FBFC783714" box="[874,963,1785,1809]" gridcol="1" gridrow="19" pageId="6" pageNumber="425">35</td>
<td id="A1413B2E51000031AB4C39FBFB893714" box="[1030,1074,1785,1809]" gridcol="2" gridrow="19" pageId="6" pageNumber="425">35</td>
<td id="A1413B2E51000031AB2C39FBFB293714" box="[1126,1170,1785,1809]" gridcol="3" gridrow="19" pageId="6" pageNumber="425">37</td>
<td id="A1413B2E51000031AB9139FBFA8B3714" box="[1243,1328,1785,1809]" gridcol="4" gridrow="19" pageId="6" pageNumber="425">36.0</td>
<td id="A1413B2E51000031AA0039FBFAD13714" box="[1354,1386,1785,1809]" gridcol="5" gridrow="19" pageId="6" pageNumber="425">0.9</td>
</tr>
<tr id="E290525251000031AFEC3823FAD1373F" box="[166,1386,1825,1850]" gridrow="20" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3823FD2C373F" box="[166,663,1825,1850]" gridcol="0" gridrow="20" pageId="6" pageNumber="425">Eye diameter</th>
<td id="A1413B2E51000031AC203823FC78373F" box="[874,963,1825,1850]" gridcol="1" gridrow="20" pageId="6" pageNumber="425">20</td>
<td id="A1413B2E51000031AB4C3823FB89373F" box="[1030,1074,1825,1850]" gridcol="2" gridrow="20" pageId="6" pageNumber="425">20</td>
<td id="A1413B2E51000031AB2C3823FB29373F" box="[1126,1170,1825,1850]" gridcol="3" gridrow="20" pageId="6" pageNumber="425">25</td>
<td id="A1413B2E51000031AB913823FA8B373F" box="[1243,1328,1825,1850]" gridcol="4" gridrow="20" pageId="6" pageNumber="425">23.0</td>
<td id="A1413B2E51000031AA003823FAD1373F" box="[1354,1386,1825,1850]" gridcol="5" gridrow="20" pageId="6" pageNumber="425">2.2</td>
</tr>
<tr id="E290525251000031AFEC3848FAD13767" box="[166,1386,1866,1890]" gridrow="21" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3848FD2C3767" box="[166,663,1866,1890]" gridcol="0" gridrow="21" pageId="6" pageNumber="425">Postorbital distance</th>
<td id="A1413B2E51000031AC203848FC783767" box="[874,963,1866,1890]" gridcol="1" gridrow="21" pageId="6" pageNumber="425">43</td>
<td id="A1413B2E51000031AB4C3848FB893767" box="[1030,1074,1866,1890]" gridcol="2" gridrow="21" pageId="6" pageNumber="425">36</td>
<td id="A1413B2E51000031AB2C3848FB293767" box="[1126,1170,1866,1890]" gridcol="3" gridrow="21" pageId="6" pageNumber="425">43</td>
<td id="A1413B2E51000031AB913848FA8B3767" box="[1243,1328,1866,1890]" gridcol="4" gridrow="21" pageId="6" pageNumber="425">40.1</td>
<td id="A1413B2E51000031AA003848FAD13767" box="[1354,1386,1866,1890]" gridcol="5" gridrow="21" pageId="6" pageNumber="425">3.0</td>
</tr>
<tr id="E290525251000031AFEC3870FAD1378F" box="[166,1386,1906,1930]" gridrow="22" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3870FD2C378F" box="[166,663,1906,1930]" gridcol="0" gridrow="22" pageId="6" pageNumber="425">Maximum head width</th>
<td id="A1413B2E51000031AC203870FC78378F" box="[874,963,1906,1930]" gridcol="1" gridrow="22" pageId="6" pageNumber="425">89</td>
<td id="A1413B2E51000031AB4C3870FB89378F" box="[1030,1074,1906,1930]" gridcol="2" gridrow="22" pageId="6" pageNumber="425">77</td>
<td id="A1413B2E51000031AB2C3870FB29378F" box="[1126,1170,1906,1930]" gridcol="3" gridrow="22" pageId="6" pageNumber="425">89</td>
<td id="A1413B2E51000031AB913870FA8B378F" box="[1243,1328,1906,1930]" gridcol="4" gridrow="22" pageId="6" pageNumber="425">84.6</td>
<td id="A1413B2E51000031AA003870FAD1378F" box="[1354,1386,1906,1930]" gridcol="5" gridrow="22" pageId="6" pageNumber="425">5.5</td>
</tr>
<tr id="E290525251000031AFEC3898FAD137B7" box="[166,1386,1946,1970]" gridrow="23" pageId="6" pageNumber="425">
<th id="A1413B2E51000031AFEC3898FD2C37B7" box="[166,663,1946,1970]" gridcol="0" gridrow="23" pageId="6" pageNumber="425">Interorbital width</th>
<td id="A1413B2E51000031AC203898FC7837B7" box="[874,963,1946,1970]" gridcol="1" gridrow="23" pageId="6" pageNumber="425">56</td>
<td id="A1413B2E51000031AB4C3898FB8937B7" box="[1030,1074,1946,1970]" gridcol="2" gridrow="23" pageId="6" pageNumber="425">43</td>
<td id="A1413B2E51000031AB2C3898FB2937B7" box="[1126,1170,1946,1970]" gridcol="3" gridrow="23" pageId="6" pageNumber="425">56</td>
<td id="A1413B2E51000031AB913898FA8B37B7" box="[1243,1328,1946,1970]" gridcol="4" gridrow="23" pageId="6" pageNumber="425">49.3</td>
<td id="A1413B2E51000031AA003898FAD137B7" box="[1354,1386,1946,1970]" gridcol="5" gridrow="23" pageId="6" pageNumber="425">5.6</td>
</tr>
</table>
</paragraph>
<caption id="08DF00985101FFC9AFD23F95FEF730D1" pageId="7" pageNumber="426" startId="7.[152,245,151,175]" targetBox="[166,1376,228,1177]" targetIsTable="true" targetPageId="7">
<paragraph id="5C1F50105101FFC9AFD23F95FEF730D1" blockId="7.[151,1437,151,212]" pageId="7" pageNumber="426">
<emphasis id="6ED48C025101FFC9AFD23F95FEB630AA" bold="true" box="[152,269,151,175]" pageId="7" pageNumber="426">TABLE 4.</emphasis>
Morphometric data of
<taxonomicName id="9BA02B935101FFC9AD4A3F9AFDC630AA" box="[512,637,152,175]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025101FFC9AD4A3F9AFDC630AA" box="[512,637,152,175]" italics="true" pageId="7" pageNumber="426">Garra rezai</emphasis>
</taxonomicName>
. Additional materials, FFR 4019 (n=11); FFR 1344 (n=6); FFR 1307 (n=3); FFR 1273 (n=3).
</paragraph>
</caption>
<paragraph id="5C1F50105101FFC9AC843FE6FADB349C" pageId="7" pageNumber="426">
<table id="2EA0A2B051010031AFEC3FE6FADB349C" box="[166,1376,228,1177]" gridcols="5" gridrows="24" pageId="7" pageNumber="426" rowsContinueFrom="6.[166,1386,1021,1970]">
<tr id="E290525251010031AFEC3FE6FADB30F9" box="[166,1376,228,252]" gridrow="0" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3FE6FADB30F9" box="[166,1376,228,252]" colspan="5" colspanRight="4" gridcol="0" gridrow="0" pageId="7" pageNumber="426">Additional materials (n=23)</th>
</tr>
<tr id="E290525251010031AFEC3E0EFADB3121" box="[166,1376,268,292]" gridrow="1" pageId="7" pageNumber="426" rowspan-0="1">
<td id="A1413B2E51010031AC843E0EFC413121" box="[974,1018,268,292]" gridcol="1" gridrow="1" pageId="7" pageNumber="426">min</td>
<td id="A1413B2E51010031AB713E0EFBC83121" box="[1083,1139,268,292]" gridcol="2" gridrow="1" pageId="7" pageNumber="426">max</td>
<td id="A1413B2E51010031AB883E0EFB423121" box="[1218,1273,268,292]" gridcol="3" gridrow="1" pageId="7" pageNumber="426">mean</td>
<td id="A1413B2E51010031AA0A3E0EFADB3121" box="[1344,1376,268,292]" gridcol="4" gridrow="1" pageId="7" pageNumber="426">SD</td>
</tr>
<tr id="E290525251010031AFEC3E36FADB3149" box="[166,1376,308,332]" gridrow="2" pageId="7" pageNumber="426" rowspan-3="1" rowspan-4="1">
<th id="A1413B2E51010031AFEC3E36FD2C3149" box="[166,663,308,332]" gridcol="0" gridrow="2" pageId="7" pageNumber="426">Standard length (mm)</th>
<td id="A1413B2E51010031AC843E36FC413149" box="[974,1018,308,332]" gridcol="1" gridrow="2" pageId="7" pageNumber="426">82.0</td>
<td id="A1413B2E51010031AB713E36FBC83149" box="[1083,1139,308,332]" gridcol="2" gridrow="2" pageId="7" pageNumber="426">114.0</td>
</tr>
<tr id="E290525251010031AFEC3E5EFADB3171" box="[166,1376,348,372]" gridrow="3" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3E5EFADB3171" box="[166,1376,348,372]" colspan="5" colspanRight="4" gridcol="0" gridrow="3" pageId="7" pageNumber="426">
<emphasis id="6ED48C025101FFC9AFEC3E5EFE593171" bold="true" box="[166,482,348,372]" pageId="7" pageNumber="426">In percent of standard length</emphasis>
</th>
</tr>
<tr id="E290525251010031AFEC3E87FADB3198" box="[166,1376,389,413]" gridrow="4" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3E87FD2C3198" box="[166,663,389,413]" gridcol="0" gridrow="4" pageId="7" pageNumber="426">Head length</th>
<td id="A1413B2E51010031AC843E87FC413198" box="[974,1018,389,413]" gridcol="1" gridrow="4" pageId="7" pageNumber="426">21.8</td>
<td id="A1413B2E51010031AB713E87FBC83198" box="[1083,1139,389,413]" gridcol="2" gridrow="4" pageId="7" pageNumber="426">24.0</td>
<td id="A1413B2E51010031AB883E87FB423198" box="[1218,1273,389,413]" gridcol="3" gridrow="4" pageId="7" pageNumber="426">23.0</td>
<td id="A1413B2E51010031AA0A3E87FADB3198" box="[1344,1376,389,413]" gridcol="4" gridrow="4" pageId="7" pageNumber="426">0.6</td>
</tr>
<tr id="E290525251010031AFEC3EAFFADB31C0" box="[166,1376,429,453]" gridrow="5" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3EAFFD2C31C0" box="[166,663,429,453]" gridcol="0" gridrow="5" pageId="7" pageNumber="426">Body depth at dorsal-fin origin</th>
<td id="A1413B2E51010031AC843EAFFC4131C0" box="[974,1018,429,453]" gridcol="1" gridrow="5" pageId="7" pageNumber="426">19.3</td>
<td id="A1413B2E51010031AB713EAFFBC831C0" box="[1083,1139,429,453]" gridcol="2" gridrow="5" pageId="7" pageNumber="426">22.4</td>
<td id="A1413B2E51010031AB883EAFFB4231C0" box="[1218,1273,429,453]" gridcol="3" gridrow="5" pageId="7" pageNumber="426">20.5</td>
<td id="A1413B2E51010031AA0A3EAFFADB31C0" box="[1344,1376,429,453]" gridcol="4" gridrow="5" pageId="7" pageNumber="426">0.7</td>
</tr>
<tr id="E290525251010031AFEC3ED7FADB31E8" box="[166,1376,469,493]" gridrow="6" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3ED7FD2C31E8" box="[166,663,469,493]" gridcol="0" gridrow="6" pageId="7" pageNumber="426">Predorsal length</th>
<td id="A1413B2E51010031AC843ED7FC4131E8" box="[974,1018,469,493]" gridcol="1" gridrow="6" pageId="7" pageNumber="426">47.3</td>
<td id="A1413B2E51010031AB713ED7FBC831E8" box="[1083,1139,469,493]" gridcol="2" gridrow="6" pageId="7" pageNumber="426">52.2</td>
<td id="A1413B2E51010031AB883ED7FB4231E8" box="[1218,1273,469,493]" gridcol="3" gridrow="6" pageId="7" pageNumber="426">49.6</td>
<td id="A1413B2E51010031AA0A3ED7FADB31E8" box="[1344,1376,469,493]" gridcol="4" gridrow="6" pageId="7" pageNumber="426">1.0</td>
</tr>
<tr id="E290525251010031AFEC3EFFFADB3213" box="[166,1376,509,534]" gridrow="7" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3EFFFD2C3213" box="[166,663,509,534]" gridcol="0" gridrow="7" pageId="7" pageNumber="426">Postdorsal length</th>
<td id="A1413B2E51010031AC843EFFFC413213" box="[974,1018,509,534]" gridcol="1" gridrow="7" pageId="7" pageNumber="426">35.0</td>
<td id="A1413B2E51010031AB713EFFFBC83213" box="[1083,1139,509,534]" gridcol="2" gridrow="7" pageId="7" pageNumber="426">39.6</td>
<td id="A1413B2E51010031AB883EFFFB423213" box="[1218,1273,509,534]" gridcol="3" gridrow="7" pageId="7" pageNumber="426">37.5</td>
<td id="A1413B2E51010031AA0A3EFFFADB3213" box="[1344,1376,509,534]" gridcol="4" gridrow="7" pageId="7" pageNumber="426">1.3</td>
</tr>
<tr id="E290525251010031AFEC3D24FADB323B" box="[166,1376,550,574]" gridrow="8" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3D24FD2C323B" box="[166,663,550,574]" gridcol="0" gridrow="8" pageId="7" pageNumber="426">Preanal length</th>
<td id="A1413B2E51010031AC843D24FC41323B" box="[974,1018,550,574]" gridcol="1" gridrow="8" pageId="7" pageNumber="426">75.0</td>
<td id="A1413B2E51010031AB713D24FBC8323B" box="[1083,1139,550,574]" gridcol="2" gridrow="8" pageId="7" pageNumber="426">79.9</td>
<td id="A1413B2E51010031AB883D24FB42323B" box="[1218,1273,550,574]" gridcol="3" gridrow="8" pageId="7" pageNumber="426">77.6</td>
<td id="A1413B2E51010031AA0A3D24FADB323B" box="[1344,1376,550,574]" gridcol="4" gridrow="8" pageId="7" pageNumber="426">1.3</td>
</tr>
<tr id="E290525251010031AFEC3D4CFADB3263" box="[166,1376,590,614]" gridrow="9" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3D4CFD2C3263" box="[166,663,590,614]" gridcol="0" gridrow="9" pageId="7" pageNumber="426">Prepelvic length</th>
<td id="A1413B2E51010031AC843D4CFC413263" box="[974,1018,590,614]" gridcol="1" gridrow="9" pageId="7" pageNumber="426">48.2</td>
<td id="A1413B2E51010031AB713D4CFBC83263" box="[1083,1139,590,614]" gridcol="2" gridrow="9" pageId="7" pageNumber="426">56.2</td>
<td id="A1413B2E51010031AB883D4CFB423263" box="[1218,1273,590,614]" gridcol="3" gridrow="9" pageId="7" pageNumber="426">53.3</td>
<td id="A1413B2E51010031AA0A3D4CFADB3263" box="[1344,1376,590,614]" gridcol="4" gridrow="9" pageId="7" pageNumber="426">2.0</td>
</tr>
<tr id="E290525251010031AFEC3D74FADB328B" box="[166,1376,630,654]" gridrow="10" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3D74FD2C328B" box="[166,663,630,654]" gridcol="0" gridrow="10" pageId="7" pageNumber="426">Distance between pectoral and pelvic-fin origins</th>
<td id="A1413B2E51010031AC843D74FC41328B" box="[974,1018,630,654]" gridcol="1" gridrow="10" pageId="7" pageNumber="426">32.5</td>
<td id="A1413B2E51010031AB713D74FBC8328B" box="[1083,1139,630,654]" gridcol="2" gridrow="10" pageId="7" pageNumber="426">37.8</td>
<td id="A1413B2E51010031AB883D74FB42328B" box="[1218,1273,630,654]" gridcol="3" gridrow="10" pageId="7" pageNumber="426">35.0</td>
<td id="A1413B2E51010031AA0A3D74FADB328B" box="[1344,1376,630,654]" gridcol="4" gridrow="10" pageId="7" pageNumber="426">1.5</td>
</tr>
<tr id="E290525251010031AFEC3D9CFADB32B2" box="[166,1376,670,695]" gridrow="11" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3D9CFD2C32B2" box="[166,663,670,695]" gridcol="0" gridrow="11" pageId="7" pageNumber="426">Distance between pelvic and anal-fin origins</th>
<td id="A1413B2E51010031AC843D9CFC4132B2" box="[974,1018,670,695]" gridcol="1" gridrow="11" pageId="7" pageNumber="426">21.7</td>
<td id="A1413B2E51010031AB713D9CFBC832B2" box="[1083,1139,670,695]" gridcol="2" gridrow="11" pageId="7" pageNumber="426">28.8</td>
<td id="A1413B2E51010031AB883D9CFB4232B2" box="[1218,1273,670,695]" gridcol="3" gridrow="11" pageId="7" pageNumber="426">24.3</td>
<td id="A1413B2E51010031AA0A3D9CFADB32B2" box="[1344,1376,670,695]" gridcol="4" gridrow="11" pageId="7" pageNumber="426">1.6</td>
</tr>
<tr id="E290525251010031AFEC3DC5FADB32DA" box="[166,1376,711,735]" gridrow="12" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3DC5FD2C32DA" box="[166,663,711,735]" gridcol="0" gridrow="12" pageId="7" pageNumber="426">Depth of caudal peduncle</th>
<td id="A1413B2E51010031AC843DC5FC4132DA" box="[974,1018,711,735]" gridcol="1" gridrow="12" pageId="7" pageNumber="426">11.9</td>
<td id="A1413B2E51010031AB713DC5FBC832DA" box="[1083,1139,711,735]" gridcol="2" gridrow="12" pageId="7" pageNumber="426">13.4</td>
<td id="A1413B2E51010031AB883DC5FB4232DA" box="[1218,1273,711,735]" gridcol="3" gridrow="12" pageId="7" pageNumber="426">12.6</td>
<td id="A1413B2E51010031AA0A3DC5FADB32DA" box="[1344,1376,711,735]" gridcol="4" gridrow="12" pageId="7" pageNumber="426">0.3</td>
</tr>
<tr id="E290525251010031AFEC3DEDFADB3302" box="[166,1376,751,775]" gridrow="13" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3DEDFD2C3302" box="[166,663,751,775]" gridcol="0" gridrow="13" pageId="7" pageNumber="426">Length of caudal peduncle</th>
<td id="A1413B2E51010031AC843DEDFC413302" box="[974,1018,751,775]" gridcol="1" gridrow="13" pageId="7" pageNumber="426">14.4</td>
<td id="A1413B2E51010031AB713DEDFBC83302" box="[1083,1139,751,775]" gridcol="2" gridrow="13" pageId="7" pageNumber="426">19.4</td>
<td id="A1413B2E51010031AB883DEDFB423302" box="[1218,1273,751,775]" gridcol="3" gridrow="13" pageId="7" pageNumber="426">16.3</td>
<td id="A1413B2E51010031AA0A3DEDFADB3302" box="[1344,1376,751,775]" gridcol="4" gridrow="13" pageId="7" pageNumber="426">1.1</td>
</tr>
<tr id="E290525251010031AFEC3C15FADB332A" box="[166,1376,791,815]" gridrow="14" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3C15FD2C332A" box="[166,663,791,815]" gridcol="0" gridrow="14" pageId="7" pageNumber="426">Anal-fin base length</th>
<td id="A1413B2E51010031AC843C15FC41332A" box="[974,1018,791,815]" gridcol="1" gridrow="14" pageId="7" pageNumber="426">6.4</td>
<td id="A1413B2E51010031AB713C15FBC8332A" box="[1083,1139,791,815]" gridcol="2" gridrow="14" pageId="7" pageNumber="426">8.1</td>
<td id="A1413B2E51010031AB883C15FB42332A" box="[1218,1273,791,815]" gridcol="3" gridrow="14" pageId="7" pageNumber="426">7.2</td>
<td id="A1413B2E51010031AA0A3C15FADB332A" box="[1344,1376,791,815]" gridcol="4" gridrow="14" pageId="7" pageNumber="426">0.5</td>
</tr>
<tr id="E290525251010031AFEC3C3DFADB335D" box="[166,1376,831,856]" gridrow="15" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3C3DFD2C335D" box="[166,663,831,856]" gridcol="0" gridrow="15" pageId="7" pageNumber="426">Pectoral-fin length</th>
<td id="A1413B2E51010031AC843C3DFC41335D" box="[974,1018,831,856]" gridcol="1" gridrow="15" pageId="7" pageNumber="426">19.8</td>
<td id="A1413B2E51010031AB713C3DFBC8335D" box="[1083,1139,831,856]" gridcol="2" gridrow="15" pageId="7" pageNumber="426">26.5</td>
<td id="A1413B2E51010031AB883C3DFB42335D" box="[1218,1273,831,856]" gridcol="3" gridrow="15" pageId="7" pageNumber="426">22.7</td>
<td id="A1413B2E51010031AA0A3C3DFADB335D" box="[1344,1376,831,856]" gridcol="4" gridrow="15" pageId="7" pageNumber="426">1.5</td>
</tr>
<tr id="E290525251010031AFEC3C6AFADB3385" box="[166,1376,872,896]" gridrow="16" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3C6AFD2C3385" box="[166,663,872,896]" gridcol="0" gridrow="16" pageId="7" pageNumber="426">Pelvic-fin length</th>
<td id="A1413B2E51010031AC843C6AFC413385" box="[974,1018,872,896]" gridcol="1" gridrow="16" pageId="7" pageNumber="426">17.1</td>
<td id="A1413B2E51010031AB713C6AFBC83385" box="[1083,1139,872,896]" gridcol="2" gridrow="16" pageId="7" pageNumber="426">22.6</td>
<td id="A1413B2E51010031AB883C6AFB423385" box="[1218,1273,872,896]" gridcol="3" gridrow="16" pageId="7" pageNumber="426">19.4</td>
<td id="A1413B2E51010031AA0A3C6AFADB3385" box="[1344,1376,872,896]" gridcol="4" gridrow="16" pageId="7" pageNumber="426">0.7</td>
</tr>
<tr id="E290525251010031AFEC3C8DFADB33AD" box="[166,1376,911,936]" gridrow="17" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3C8DFADB33AD" box="[166,1376,911,936]" colspan="5" colspanRight="4" gridcol="0" gridrow="17" pageId="7" pageNumber="426">
<emphasis id="6ED48C025101FFC9AFEC3C92FE0E33AD" bold="true" box="[166,437,911,936]" pageId="7" pageNumber="426">In percent of head length</emphasis>
</th>
</tr>
<tr id="E290525251010031AFEC3CBAFADB33D4" box="[166,1376,952,977]" gridrow="18" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3CBAFD2C33D4" box="[166,663,952,977]" gridcol="0" gridrow="18" pageId="7" pageNumber="426">Head depth at eye</th>
<td id="A1413B2E51010031AC843CBAFC4133D4" box="[974,1018,952,977]" gridcol="1" gridrow="18" pageId="7" pageNumber="426">53</td>
<td id="A1413B2E51010031AB713CBAFBC833D4" box="[1083,1139,952,977]" gridcol="2" gridrow="18" pageId="7" pageNumber="426">60</td>
<td id="A1413B2E51010031AB883CBAFB4233D4" box="[1218,1273,952,977]" gridcol="3" gridrow="18" pageId="7" pageNumber="426">56.1</td>
<td id="A1413B2E51010031AA0A3CBAFADB33D4" box="[1344,1376,952,977]" gridcol="4" gridrow="18" pageId="7" pageNumber="426">1.5</td>
</tr>
<tr id="E290525251010031AFEC3CE2FADB33FD" box="[166,1376,992,1016]" gridrow="19" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3CE2FD2C33FD" box="[166,663,992,1016]" gridcol="0" gridrow="19" pageId="7" pageNumber="426">Snout length</th>
<td id="A1413B2E51010031AC843CE2FC4133FD" box="[974,1018,992,1016]" gridcol="1" gridrow="19" pageId="7" pageNumber="426">47</td>
<td id="A1413B2E51010031AB713CE2FBC833FD" box="[1083,1139,992,1016]" gridcol="2" gridrow="19" pageId="7" pageNumber="426">54</td>
<td id="A1413B2E51010031AB883CE2FB4233FD" box="[1218,1273,992,1016]" gridcol="3" gridrow="19" pageId="7" pageNumber="426">49.9</td>
<td id="A1413B2E51010031AA0A3CE2FADB33FD" box="[1344,1376,992,1016]" gridcol="4" gridrow="19" pageId="7" pageNumber="426">2.0</td>
</tr>
<tr id="E290525251010031AFEC3B0AFADB3424" box="[166,1376,1032,1057]" gridrow="20" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3B0AFD2C3424" box="[166,663,1032,1057]" gridcol="0" gridrow="20" pageId="7" pageNumber="426">Eye diameter</th>
<td id="A1413B2E51010031AC843B0AFC413424" box="[974,1018,1032,1057]" gridcol="1" gridrow="20" pageId="7" pageNumber="426">14</td>
<td id="A1413B2E51010031AB713B0AFBC83424" box="[1083,1139,1032,1057]" gridcol="2" gridrow="20" pageId="7" pageNumber="426">17</td>
<td id="A1413B2E51010031AB883B0AFB423424" box="[1218,1273,1032,1057]" gridcol="3" gridrow="20" pageId="7" pageNumber="426">15.8</td>
<td id="A1413B2E51010031AA0A3B0AFADB3424" box="[1344,1376,1032,1057]" gridcol="4" gridrow="20" pageId="7" pageNumber="426">0.9</td>
</tr>
<tr id="E290525251010031AFEC3B33FADB344C" box="[166,1376,1073,1097]" gridrow="21" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3B33FD2C344C" box="[166,663,1073,1097]" gridcol="0" gridrow="21" pageId="7" pageNumber="426">Postorbital distance</th>
<td id="A1413B2E51010031AC843B33FC41344C" box="[974,1018,1073,1097]" gridcol="1" gridrow="21" pageId="7" pageNumber="426">38</td>
<td id="A1413B2E51010031AB713B33FBC8344C" box="[1083,1139,1073,1097]" gridcol="2" gridrow="21" pageId="7" pageNumber="426">44</td>
<td id="A1413B2E51010031AB883B33FB42344C" box="[1218,1273,1073,1097]" gridcol="3" gridrow="21" pageId="7" pageNumber="426">40.0</td>
<td id="A1413B2E51010031AA0A3B33FADB344C" box="[1344,1376,1073,1097]" gridcol="4" gridrow="21" pageId="7" pageNumber="426">1.3</td>
</tr>
<tr id="E290525251010031AFEC3B5BFADB3474" box="[166,1376,1113,1137]" gridrow="22" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3B5BFD2C3474" box="[166,663,1113,1137]" gridcol="0" gridrow="22" pageId="7" pageNumber="426">Maximum head width</th>
<td id="A1413B2E51010031AC843B5BFC413474" box="[974,1018,1113,1137]" gridcol="1" gridrow="22" pageId="7" pageNumber="426">72</td>
<td id="A1413B2E51010031AB713B5BFBC83474" box="[1083,1139,1113,1137]" gridcol="2" gridrow="22" pageId="7" pageNumber="426">81</td>
<td id="A1413B2E51010031AB883B5BFB423474" box="[1218,1273,1113,1137]" gridcol="3" gridrow="22" pageId="7" pageNumber="426">77.7</td>
<td id="A1413B2E51010031AA0A3B5BFADB3474" box="[1344,1376,1113,1137]" gridcol="4" gridrow="22" pageId="7" pageNumber="426">2.4</td>
</tr>
<tr id="E290525251010031AFEC3B83FADB349C" box="[166,1376,1153,1177]" gridrow="23" pageId="7" pageNumber="426">
<th id="A1413B2E51010031AFEC3B83FD2C349C" box="[166,663,1153,1177]" gridcol="0" gridrow="23" pageId="7" pageNumber="426">Interorbital width</th>
<td id="A1413B2E51010031AC843B83FC41349C" box="[974,1018,1153,1177]" gridcol="1" gridrow="23" pageId="7" pageNumber="426">42</td>
<td id="A1413B2E51010031AB713B83FBC8349C" box="[1083,1139,1153,1177]" gridcol="2" gridrow="23" pageId="7" pageNumber="426">51</td>
<td id="A1413B2E51010031AB883B83FB42349C" box="[1218,1273,1153,1177]" gridcol="3" gridrow="23" pageId="7" pageNumber="426">46.1</td>
<td id="A1413B2E51010031AA0A3B83FADB349C" box="[1344,1376,1153,1177]" gridcol="4" gridrow="23" pageId="7" pageNumber="426">2.6</td>
</tr>
</table>
</paragraph>
<caption id="08DF00985101FFC9AFDD38B3FECB37E8" ID-DOI="http://doi.org/10.5281/zenodo.7223695" ID-Zenodo-Dep="7223695" httpUri="https://zenodo.org/record/7223695/files/figure.png" pageId="7" pageNumber="426" startId="7.[151,250,1969,1993]" targetBox="[209,1377,1239,1945]" targetPageId="7">
<paragraph id="5C1F50105101FFC9AFDD38B3FECB37E8" blockId="7.[151,1437,1969,2029]" pageId="7" pageNumber="426">
<emphasis id="6ED48C025101FFC9AFDD38B3FEAE37CC" bold="true" box="[151,277,1969,1993]" pageId="7" pageNumber="426">FIGURE 3.</emphasis>
<taxonomicName id="9BA02B935101FFC9AE5738B0FE2C37CC" box="[285,407,1970,1993]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="426" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025101FFC9AE5738B0FE2C37CC" box="[285,407,1970,1993]" italics="true" pageId="7" pageNumber="426">Garra rezai</emphasis>
</taxonomicName>
, paratypes; from above: FSJF 4111, 59 mm SL; VMFC GRE-P5216, 34 mm SL; 28 mm SL; Iran: stream Bouein-Sofla.
</paragraph>
</caption>
<caption id="08DF0098510EFFC6AFDD3C81FECB33C5" ID-DOI="http://doi.org/10.5281/zenodo.7223697" ID-Zenodo-Dep="7223697" httpUri="https://zenodo.org/record/7223697/files/figure.png" pageId="8" pageNumber="427" startId="8.[151,250,899,923]" targetBox="[152,1436,181,876]" targetPageId="8">
<paragraph id="5C1F5010510EFFC6AFDD3C81FECB33C5" blockId="8.[151,1437,899,960]" pageId="8" pageNumber="427">
<emphasis id="6ED48C02510EFFC6AFDD3C81FEAE3399" bold="true" box="[151,277,899,924]" pageId="8" pageNumber="427">FIGURE 4.</emphasis>
<taxonomicName id="9BA02B93510EFFC6AE573C86FE2C339E" box="[285,407,900,923]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510EFFC6AE573C86FE2C339E" box="[285,407,900,923]" italics="true" pageId="8" pageNumber="427">Garra rezai</emphasis>
</taxonomicName>
, paratypes; from above: FSJF 4111, 59 mm SL; VMFC GRE-P5216, 34 mm SL; 28 mm SL; Iran: stream Bouein-Sofla.
</paragraph>
</caption>
<caption id="08DF0098510EFFC6AFDD382BFECB3760" ID-DOI="http://doi.org/10.5281/zenodo.7223699" ID-Zenodo-Dep="7223699" httpUri="https://zenodo.org/record/7223699/files/figure.png" pageId="8" pageNumber="427" startId="8.[151,250,1833,1857]" targetBox="[151,1436,984,1810]" targetPageId="8">
<paragraph id="5C1F5010510EFFC6AFDD382BFECB3760" blockId="8.[151,1437,1833,1893]" pageId="8" pageNumber="427">
<emphasis id="6ED48C02510EFFC6AFDD382BFEAE3744" bold="true" box="[151,277,1833,1857]" pageId="8" pageNumber="427">FIGURE 5.</emphasis>
<taxonomicName id="9BA02B93510EFFC6AE573828FE2C3744" box="[285,407,1834,1857]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="427" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510EFFC6AE573828FE2C3744" box="[285,407,1834,1857]" italics="true" pageId="8" pageNumber="427">Garra rezai</emphasis>
</taxonomicName>
, paratypes; from above: FSJF 4111, 59 mm SL; VMFC GRE-P5216, 34 mm SL; 28 mm SL; Iran: stream Bouein-Sofla.
</paragraph>
</caption>
<paragraph id="5C1F5010510EFFC7AF8D3891FAF032C8" blockId="8.[151,1437,1939,2038]" lastBlockId="9.[151,1437,151,717]" lastPageId="9" lastPageNumber="428" pageId="8" pageNumber="427">
Dorsal fin with 3 simple and 7½ (54), rarely 8½ (4), branched rays, last simple ray shorter than head length. Posterior dorsal-fin margin slightly concave. Dorsal-fin origin situated at about middle between caudal-fin base and snout tip. First branched dorsal-fin ray longest, tip of last branched dorsal-fin ray reaching vertical to, or a point slightly behind of anus when folded down. Pectoral fin with one simple and 11 (21) or 12 (11) branched rays. Pectoral fin reaching 5183% of distance between pectoral-fin origin and pelvic-fin origin, its length equal to or slightly shorter than head length. Pelvic fin with one simple and 7 (27) or 8 (5) branched rays. Pelvic fin exceed anus, not reaching anal-fin base. Pelvic-fin origin closer to anal-fin origin than pectoral-fin origin, inserted below second or third branched dorsal-fin ray. Anal fin short, with 3 simple and 5½ (30) or 6½ (2) branched rays, first branched ray longest. Posterior anal-fin margin straight or slightly concave. Anal-fin origin at about middle between pelvic-fin origin and caudal-fin base, closer to pelvic-fin origin than to caudal-fin base in some individuals. Anal fin reaching approximately
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<subScript id="C0245255510FFFC7AEF33EA7FE7A31B6" attach="left" box="[441,449,421,435]" fontSize="6" pageId="9" pageNumber="428">2</subScript>
to
<superScript id="ABD5FD58510FFFC7AEAD3E90FE5431A5" attach="right" box="[487,495,402,416]" fontSize="6" pageId="9" pageNumber="428">4</superScript>
/
<subScript id="C0245255510FFFC7AEBD3EA7FE4431B6" attach="left" box="[503,511,421,435]" fontSize="6" pageId="9" pageNumber="428">5</subScript>
of caudal peduncle when folded down. Caudal fin forked with rounded lobes and 9+8 branched rays. Length of middle caudal-fin ray 5266% of longest branched ray in upper lobe. Total gill rakers on first branchial arch 1116 [11(1), 12(2), 13(2), 14(1), 15(3), 16(3)]. Lateral line complete, with 3540 [35(4), 36(3), 37(15), 38(17), 39(9), 40(3)] scales, 13 of them were on caudal-fin base. Transverse scale rows above lateral line 46 [4(3), 5(43), 6(5)], between lateral line and pelvic-fin origin 34 [3(3), 4(22)], and 45 [4(33), 5(16)] between lateral line and anal-fin origin. Circumpeduncular scale rows 1518 [15(2), 16(17), 17(3), 18(4)], 1519 [15(2), 16(3), 17(10), 18(8), 19(2)] scales on predorsal midline between dorsal-fin origin and nape, 34 scales between anus and anal-fin origin. Scales on flank regularly arranged. Chest with embedded scales and belly covered by scales. A well-developed axillary scale present at pelvic-fin base. Largest known individual
<quantity id="9B58FDF5510FFFC7ABF53DB1FAA732C8" box="[1215,1308,691,717]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="9" pageNumber="428" unit="mm" value="127.0">127 mm</quantity>
SL.
</paragraph>
<caption id="08DF0098510FFFC7AFDD3878FC3B3796" ID-DOI="http://doi.org/10.5281/zenodo.7223703" ID-Zenodo-Dep="7223703" box="[151,896,1914,1939]" httpUri="https://zenodo.org/record/7223703/files/figure.png" pageId="9" pageNumber="428" startId="9.[151,250,1914,1938]" targetBox="[154,1433,741,1891]" targetPageId="9">
<paragraph id="5C1F5010510FFFC7AFDD3878FC3B3796" blockId="9.[151,896,1914,1939]" box="[151,896,1914,1939]" pageId="9" pageNumber="428">
<emphasis id="6ED48C02510FFFC7AFDD3878FEA83797" bold="true" box="[151,275,1914,1938]" pageId="9" pageNumber="428">FIGURE 6.</emphasis>
<taxonomicName id="9BA02B93510FFFC7AE533879FE2A3797" box="[281,401,1915,1938]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="428" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510FFFC7AE533879FE2A3797" box="[281,401,1915,1938]" italics="true" pageId="9" pageNumber="428">Garra rezai</emphasis>
</taxonomicName>
; FFR 4038, 109 mm SL; Turkey: stream Büyük.
</paragraph>
</caption>
<caption id="08DF0098510CFFC4AFDD3C22FA95333D" ID-DOI="http://doi.org/10.5281/zenodo.7223705" ID-Zenodo-Dep="7223705" box="[151,1326,800,824]" httpUri="https://zenodo.org/record/7223705/files/figure.png" pageId="10" pageNumber="429" startId="10.[151,250,800,824]" targetBox="[151,1437,181,776]" targetPageId="10">
<paragraph id="5C1F5010510CFFC4AFDD3C22FA95333D" blockId="10.[151,1326,800,824]" box="[151,1326,800,824]" pageId="10" pageNumber="429">
<emphasis id="6ED48C02510CFFC4AFDD3C22FEA8333D" bold="true" box="[151,275,800,824]" pageId="10" pageNumber="429">FIGURE 7.</emphasis>
<taxonomicName id="9BA02B93510CFFC4AE533C22FE2A333D" box="[281,401,800,824]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="429" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510CFFC4AE533C22FE2A333D" box="[281,401,800,824]" italics="true" pageId="10" pageNumber="429">Garra rezai</emphasis>
</taxonomicName>
; left, FFR 1273, 107 mm SL; stream Büyük; right, FFR 1302, 113 mm SL; stream Çıratan.
</paragraph>
</caption>
<caption id="08DF0098510CFFC4AFDD39F4FE083737" ID-DOI="http://doi.org/10.5281/zenodo.7223707" ID-Zenodo-Dep="7223707" httpUri="https://zenodo.org/record/7223707/files/figure.png" pageId="10" pageNumber="429" startId="10.[151,250,1782,1806]" targetBox="[151,1436,848,1758]" targetPageId="10">
<paragraph id="5C1F5010510CFFC4AFDD39F4FE083737" blockId="10.[151,1436,1782,1842]" pageId="10" pageNumber="429">
<emphasis id="6ED48C02510CFFC4AFDD39F4FEA8370B" bold="true" box="[151,275,1782,1806]" pageId="10" pageNumber="429">FIGURE 8.</emphasis>
<taxonomicName id="9BA02B93510CFFC4AE5339F4FE2A370B" box="[281,401,1782,1806]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="429" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510CFFC4AE5339F4FE2A370B" box="[281,401,1782,1806]" italics="true" pageId="10" pageNumber="429">Garra rezai</emphasis>
</taxonomicName>
, from above: IMNRF-UT 1245, paratype, 58 mm SL; Iran: stream Bouein-Sofla; FSJF 3824, 104 mm SL; Turkey: stream Çıratan.
</paragraph>
</caption>
<paragraph id="5C1F5010510CFFC5AF8D3862FB9930D0" blockId="10.[151,1437,1888,2023]" lastBlockId="11.[151,1437,151,393]" lastPageId="11" lastPageNumber="430" pageId="10" pageNumber="429">
<emphasis id="6ED48C02510CFFC4AF8D3862FEF4377F" bold="true" box="[199,335,1888,1914]" pageId="10" pageNumber="429">Coloration.</emphasis>
In preserved individuals: background colour grey, pale- or dark-brown. Scales with a very narrow beige margin. Isolated individual or patches of dark-brown scales, darker than background, often arranged in horizontal rows. Flank above lateral line darker than below. Elongated, dark grey dots at base of dorsal-fin rays in some individuals. A blackish spot at lateral-line origin and a vertically elongated blotch at caudal-fin base in juveniles. Cheek pale plain yellowish or whitish. Mouth, chest and abdomen yellowish. All fins hyaline, grey or pale-brown, rays darker than membranes with irregularly set black spots on rays.
</paragraph>
<paragraph id="5C1F5010510DFFC5AF8D3FDDFB78318C" blockId="11.[151,1437,151,393]" pageId="11" pageNumber="430">In life: background colour pale- or dark-brown, fins beige or pale-brown, with a reddish hue in many individuals, rays darker than membranes. Isolated or patches of dark-brown scales, scale margins of some scales much darker than center of scales creating an irregular, mottled pattern. Elongated, dark grey dots at base of dorsal-fin rays. Head plain beige or greyish brown. A blackish spot at lateral-line origin and a vertically elongated blotch at caudal-fin base in juveniles. Iris silvery orange to golden with dark grey spots, internal ring without spots.</paragraph>
</subSubSection>
<caption id="08DF0098510DFFC5AFDD39FBFC363717" ID-DOI="http://doi.org/10.5281/zenodo.7223709" ID-Zenodo-Dep="7223709" box="[151,909,1785,1810]" httpUri="https://zenodo.org/record/7223709/files/figure.png" pageId="11" pageNumber="430" startId="11.[151,250,1785,1809]" targetBox="[151,1437,417,1762]" targetPageId="11">
<paragraph id="5C1F5010510DFFC5AFDD39FBFC363717" blockId="11.[151,909,1785,1810]" box="[151,909,1785,1810]" pageId="11" pageNumber="430">
<emphasis id="6ED48C02510DFFC5AFDD39FBFEA83714" bold="true" box="[151,275,1785,1809]" pageId="11" pageNumber="430">FIGURE 9.</emphasis>
<taxonomicName id="9BA02B93510DFFC5AE5339F8FE6A3717" authority=", FSJF" authorityName="FSJF" box="[281,465,1786,1810]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="430" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510DFFC5AE5339F8FE2A3714" box="[281,401,1786,1809]" italics="true" pageId="11" pageNumber="430">Garra rezai</emphasis>
, FSJF
</taxonomicName>
3824, 104 mm SL; Turkey: stream Çıratan.
</paragraph>
</caption>
<subSubSection id="14BA039B510DFFC5AF8D383DFD8637EF" pageId="11" pageNumber="430" type="distribution">
<paragraph id="5C1F5010510DFFC5AF8D383DFD8637EF" blockId="11.[151,1436,1855,2026]" pageId="11" pageNumber="430">
<emphasis id="6ED48C02510DFFC5AF8D383DFEDA375C" bold="true" box="[199,353,1855,1881]" pageId="11" pageNumber="430">Distribution.</emphasis>
The species is currently only known from two areas in the Tigris drainage, a small stream in the Chooman drainage in
<collectingCountry id="24B71080510DFFC5AEC53866FE07377B" box="[399,444,1892,1918]" name="Iran" pageId="11" pageNumber="430">Iran</collectingCountry>
and the upper Yanarsu in
<collectingCountry id="24B71080510DFFC5AD943866FC8B377B" box="[734,816,1892,1918]" name="Turkey" pageId="11" pageNumber="430">Turkey</collectingCountry>
. In the Chooman, it was found in a spring in the village Boein-e-Olia and the outflowing Boein-e-Sofla stream. The Chooman flows from
<collectingCountry id="24B71080510DFFC5AB63388AFBED37A7" box="[1065,1110,1928,1954]" name="Iran" pageId="11" pageNumber="430">Iran</collectingCountry>
to
<collectingCountry id="24B71080510DFFC5AB33388AFB1D37A7" box="[1145,1190,1928,1954]" name="Iraq" pageId="11" pageNumber="430">Iraq</collectingCountry>
and the species might also occur in this country. The Yanarsu is one of the tributaries of the Tigris in
<collectingCountry id="24B71080510DFFC5AB4D38AEFBE237C3" box="[1031,1113,1964,1990]" name="Turkey" pageId="11" pageNumber="430">Turkey</collectingCountry>
. Here, the species was found in three streams in the upper Yanarsu.
</paragraph>
</subSubSection>
<caption id="08DF0098510AFFC2AFDD3DDBFDFB3310" ID-DOI="http://doi.org/10.5281/zenodo.7223711" ID-Zenodo-Dep="7223711" httpUri="https://zenodo.org/record/7223711/files/figure.png" pageId="12" pageNumber="431" startId="12.[151,250,729,753]" targetBox="[151,1436,181,705]" targetPageId="12">
<paragraph id="5C1F5010510AFFC2AFDD3DDBFDFB3310" blockId="12.[151,1436,729,789]" pageId="12" pageNumber="431">
<emphasis id="6ED48C02510AFFC2AFDD3DDBFE9932F4" bold="true" box="[151,290,729,753]" pageId="12" pageNumber="431">FIGURE 10.</emphasis>
From left:
<taxonomicName id="9BA02B93510AFFC2AED73DDBFDDE32F4" authority=", GUIC" authorityName="GUIC" box="[413,613,729,753]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2AED73DDBFDA332F4" box="[413,536,729,753]" italics="true" pageId="12" pageNumber="431">Garra rezai</emphasis>
, GUIC
</taxonomicName>
7979, holotype, 80 mm SL; Iran: stream Bouein-Sofla;
<taxonomicName id="9BA02B93510AFFC2ABFF3DDBFF663310" authority=", VMFC GROS" authorityName="VMFC GROS" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rossica">
<emphasis id="6ED48C02510AFFC2ABFF3DDBFAFE32F4" box="[1205,1349,729,753]" italics="true" pageId="12" pageNumber="431">Garra rossica</emphasis>
, VMFC GROS
</taxonomicName>
, 86 mm SL; Iran: spring Golbahar.
</paragraph>
</caption>
<subSubSection id="14BA039B510AFFC2AF8D3C3CFF6E3555" pageId="12" pageNumber="431" type="etymology">
<paragraph id="5C1F5010510AFFC2AF8D3C3CFBCA33A5" blockId="12.[151,1437,830,2009]" pageId="12" pageNumber="431">
<emphasis id="6ED48C02510AFFC2AF8D3C3CFEF5335D" bold="true" box="[199,334,830,856]" pageId="12" pageNumber="431">Etymology.</emphasis>
The species is named after Reza Mousavi-Sabet, father of the first author, who was injured by chemical weapons in the region of the
<typeStatus id="831BEEB2510AFFC2AD043C60FDC53379" box="[590,638,866,892]" pageId="12" pageNumber="431">type</typeStatus>
locality of this species. We named this taxon to respect and remember all Iranian and Iraqi victims of chemical weapons during the Iran-Iraq war (1980-1988).
</paragraph>
<paragraph id="5C1F5010510AFFC2AF8D3CA8FF6E3555" blockId="12.[151,1437,830,2009]" pageId="12" pageNumber="431">
<emphasis id="6ED48C02510AFFC2AF8D3CA8FE2C33C6" bold="true" box="[199,407,937,964]" pageId="12" pageNumber="431">Notes on habitat.</emphasis>
In
<collectingCountry id="24B71080510AFFC2AE8B3CA8FE4A33C1" box="[449,497,938,964]" name="Iran" pageId="12" pageNumber="431">Iran</collectingCountry>
,
<taxonomicName id="9BA02B93510AFFC2AEB53CA8FDE133C1" box="[511,602,938,964]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2AEB53CA8FDE133C1" box="[511,602,938,964]" italics="true" pageId="12" pageNumber="431">G. rezai</emphasis>
</taxonomicName>
was found in the spring in Boein-e-Olia in
<date id="281E76D0510AFFC2AB1C3CA8FB6D33C1" box="[1110,1238,938,964]" pageId="12" pageNumber="431" value="2016-04">April 2016</date>
. The spring (
<figureCitation id="C49B4C95510AFFC2AA253CA8FF0933ED" captionStart="FIGURE 11" captionStartId="13.[151,250,1941,1965]" captionTargetBox="[151,1436,273,1918]" captionTargetId="figure-75@13.[151,1436,273,1918]" captionTargetPageId="13" captionText="FIGURE 11. Type locality of G. rezai; from above: spring in Boein-e-Olia; Boein-e-Sofla stream." figureDoi="http://doi.org/10.5281/zenodo.7223713" httpUri="https://zenodo.org/record/7223713/files/figure.png" pageId="12" pageNumber="431">Fig. 11</figureCitation>
) was about 23 meters wide, and the outflowing stream had a low velocity and the bed was covered by gravel. Revisiting the place in
<date id="281E76D0510AFFC2AEEA3CF0FDAD3409" box="[416,534,1010,1036]" pageId="12" pageNumber="431" value="2021-06">June 2021</date>
, the species could not be found anymore in the spring. We were able to collect
<taxonomicName id="9BA02B93510AFFC2AFDD3B14FF493435" box="[151,242,1046,1072]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2AFDD3B14FF493435" box="[151,242,1046,1072]" italics="true" pageId="12" pageNumber="431">G. rezai</emphasis>
</taxonomicName>
from the stream Boein-e-Sofla (which is fed by the spring) in
<date id="281E76D0510AFFC2AC8D3B14FB843435" box="[967,1087,1046,1072]" pageId="12" pageNumber="431" value="2021-06">June 2021</date>
. At the
<typeStatus id="831BEEB2510AFFC2ABD03B14FB713435" box="[1178,1226,1046,1072]" pageId="12" pageNumber="431">type</typeStatus>
locality (Boein-eOlia spring and Boein-e-Sofla stream) no
<taxonomicName id="9BA02B93510AFFC2AD243B38FD043451" box="[622,703,1082,1108]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rufa">
<emphasis id="6ED48C02510AFFC2AD243B38FD043451" box="[622,703,1082,1108]" italics="true" pageId="12" pageNumber="431">G. rufa</emphasis>
</taxonomicName>
was found, but
<taxonomicName id="9BA02B93510AFFC2AC393B38FC7F3451" box="[883,964,1082,1108]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rufa">
<emphasis id="6ED48C02510AFFC2AC393B38FC7F3451" box="[883,964,1082,1108]" italics="true" pageId="12" pageNumber="431">G. rufa</emphasis>
</taxonomicName>
is widespread in the Lesser Zab drainage, including the Chooman River drainage. In
<collectingCountry id="24B71080510AFFC2ADCD3B5CFD63347D" box="[647,728,1118,1144]" name="Turkey" pageId="12" pageNumber="431">Turkey</collectingCountry>
, all three sampling sites were shallow, with clear and swiftly flowing water and cobbles and pebbles substrate. Among the three sampling sites, Çıratan (
<figureCitation id="C49B4C95510AFFC2ABFD3B80FAB03499" box="[1207,1291,1154,1180]" captionStart="FIGURE 12" captionStartId="14.[151,250,1550,1574]" captionTargetBox="[151,1436,669,1527]" captionTargetId="figure-269@14.[151,1436,669,1527]" captionTargetPageId="14" captionText="FIGURE 12. Stream Çıratan at Üçadım, Turkey; habitat of G. rezai." figureDoi="http://doi.org/10.5281/zenodo.7223715" httpUri="https://zenodo.org/record/7223715/files/figure.png" pageId="12" pageNumber="431">Fig. 12</figureCitation>
) was visited several times during different seasons and
<taxonomicName id="9BA02B93510AFFC2ADCB3BA4FD6734C5" box="[641,732,1190,1216]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2ADCB3BA4FD6734C5" box="[641,732,1190,1216]" italics="true" pageId="12" pageNumber="431">G. rezai</emphasis>
</taxonomicName>
is abundant in its Turkish distribution area. The stream hosts several regionally endemic species, such as
<emphasis id="6ED48C02510AFFC2ADCB3BC8FB4234E1" box="[641,1273,1226,1252]" italics="true" pageId="12" pageNumber="431">
<taxonomicName id="9BA02B93510AFFC2ADCB3BC8FBF634E1" authority=", Paracobitis" authorityName="Paracobitis" box="[641,1101,1226,1252]" class="Actinopterygii" family="Nemacheilidae" genus="Turcinoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="kosswigi">Turcinoemacheilus kosswigi, Paracobitis</taxonomicName>
zabgawraensis
</emphasis>
,
<taxonomicName id="9BA02B93510AFFC2AA4D3BC8FF4B350D" authorityName="Freyhof, Kaya, Abdullah &amp; Geiger" authorityYear="2021" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="daemon">
<emphasis id="6ED48C02510AFFC2AA4D3BC8FF4B350D" italics="true" pageId="12" pageNumber="431">Glyptothorax daemon</emphasis>
</taxonomicName>
, and
<taxonomicName id="9BA02B93510AFFC2AE653BECFE53350D" baseAuthorityName="Berg" baseAuthorityYear="1931" box="[303,488,1262,1288]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis id="6ED48C02510AFFC2AE653BECFE53350D" box="[303,488,1262,1288]" italics="true" pageId="12" pageNumber="431">G. kurdistanicus</emphasis>
</taxonomicName>
(
<bibRefCitation id="38312DE1510AFFC2AEBD3BECFD03350D" author="Kaya, C. &amp; Turan, D. &amp; Unlu, E." box="[503,696,1262,1288]" pageId="12" pageNumber="431" pagination="545 - 562" refId="ref8707" refString="Kaya, C., Turan, D. &amp; Unlu, E. (2016) The latest status and distribution of fishes in upper Tigris River and two new records for Turkish freshwaters. Turkish Journal of Fisheries and Aquatic Sciences, 16, 545 - 562. https: // doi. org / 10.4194 / 1303 - 2712 - v 16 _ 3 _ 07" type="journal article" year="2016">
Kaya
<emphasis id="6ED48C02510AFFC2AD763BEDFDCE350D" box="[572,629,1262,1288]" italics="true" pageId="12" pageNumber="431">et al.</emphasis>
2016
</bibRefCitation>
,
<bibRefCitation id="38312DE1510AFFC2AD893BECFC1A350D" author="Freyhof, J. &amp; Kaya, C. &amp; Abdullah, Y. S. &amp; Geiger, M. F." box="[707,929,1262,1288]" pageId="12" pageNumber="431" pagination="453 - 491" refId="ref8010" refString="Freyhof, J., Kaya, C., Abdullah, Y. S. &amp; Geiger, M. F. (2021) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453 - 491. https: // doi. org / 10.11646 / zootaxa. 4969.3.2" type="journal article" year="2021">
Freyhof
<emphasis id="6ED48C02510AFFC2AC6E3BEDFCE6350D" box="[804,861,1262,1288]" italics="true" pageId="12" pageNumber="431">et al.</emphasis>
2021
</bibRefCitation>
). In Çıratan,
<specimenCount id="4AA69B99510AFFC2AB7F3BECFB24350D" box="[1077,1183,1262,1288]" pageId="12" pageNumber="431" type="adult">two adult</specimenCount>
individuals of
<taxonomicName id="9BA02B93510AFFC2AA003BECFA26350D" box="[1354,1437,1262,1288]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rufa">
<emphasis id="6ED48C02510AFFC2AA003BECFA26350D" box="[1354,1437,1262,1288]" italics="true" pageId="12" pageNumber="431">G. rufa</emphasis>
</taxonomicName>
were found, and one half-grown individual of
<taxonomicName id="9BA02B93510AFFC2ADFA3A10FCBF3529" box="[688,772,1298,1324]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rufa">
<emphasis id="6ED48C02510AFFC2ADFA3A10FCBF3529" box="[688,772,1298,1324]" italics="true" pageId="12" pageNumber="431">G. rufa</emphasis>
</taxonomicName>
was found in the stream Büyük, both in syntopy with
<taxonomicName id="9BA02B93510AFFC2AACB3A10FF753555" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2AACB3A10FF753555" italics="true" pageId="12" pageNumber="431">G. rezai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="14BA039B510AFFC0AF8D3A58FD683280" lastPageId="14" lastPageNumber="433" pageId="12" pageNumber="431" type="discussion">
<paragraph id="5C1F5010510AFFC2AF8D3A58FA9E35B9" blockId="12.[151,1437,830,2009]" pageId="12" pageNumber="431">
<emphasis id="6ED48C02510AFFC2AF8D3A58FE803571" bold="true" box="[199,315,1370,1396]" pageId="12" pageNumber="431">Remarks.</emphasis>
The large direct distance of
<quantity id="9B58FDF5510AFFC2AD313A58FD6B3571" box="[635,720,1370,1397]" metricMagnitude="5" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="431" unit="km" value="450.0">450 km</quantity>
between both areas of occurrence of
<taxonomicName id="9BA02B93510AFFC2AB383A58FB773571" box="[1138,1228,1370,1396]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2AB383A58FB773571" box="[1138,1228,1370,1396]" italics="true" pageId="12" pageNumber="431">G. rezai</emphasis>
</taxonomicName>
is very surprising. We suspect that
<taxonomicName id="9BA02B93510AFFC2AE053A7CFE13359D" box="[335,424,1406,1432]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2AE053A7CFE13359D" box="[335,424,1406,1432]" italics="true" pageId="12" pageNumber="431">G. rezai</emphasis>
</taxonomicName>
might be much more widespread, at least in the Tigris drainage. Indeed, it is superficially not very different from the ubiquitous
<taxonomicName id="9BA02B93510AFFC2AD0F3AA0FD2C35B9" box="[581,663,1442,1468]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rufa">
<emphasis id="6ED48C02510AFFC2AD0F3AA0FD2C35B9" box="[581,663,1442,1468]" italics="true" pageId="12" pageNumber="431">G. rufa</emphasis>
</taxonomicName>
and might be misidentified as this species at many places.
</paragraph>
<paragraph id="5C1F5010510AFFC2AF8D3AC4FC793721" blockId="12.[151,1437,830,2009]" pageId="12" pageNumber="431">
<taxonomicName id="9BA02B93510AFFC2AF8D3AC4FEF835E5" box="[199,323,1478,1504]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rufa">
<emphasis id="6ED48C02510AFFC2AF8D3AC4FEF835E5" box="[199,323,1478,1504]" italics="true" pageId="12" pageNumber="431">Garra rufa</emphasis>
</taxonomicName>
and
<taxonomicName id="9BA02B93510AFFC2AE333AC4FE4A35E5" box="[377,497,1478,1504]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="elegans">
<emphasis id="6ED48C02510AFFC2AE333AC4FE4A35E5" box="[377,497,1478,1504]" italics="true" pageId="12" pageNumber="431">G. elegans</emphasis>
</taxonomicName>
are two other species of
<taxonomicName id="9BA02B93510AFFC2AC423AC4FCF535E5" box="[776,846,1478,1504]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="6ED48C02510AFFC2AC423AC4FCF535E5" box="[776,846,1478,1504]" italics="true" pageId="12" pageNumber="431">Garra</emphasis>
</taxonomicName>
known from the Lesser Zab drainage.
<taxonomicName id="9BA02B93510AFFC2ABB53AC4FA3835E5" box="[1279,1411,1478,1504]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2ABB53AC4FA3835E5" box="[1279,1411,1478,1504]" italics="true" pageId="12" pageNumber="431">Garra rezai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="9BA02B93510AFFC2AE3A3AE8FE533601" box="[368,488,1514,1540]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="elegans">
<emphasis id="6ED48C02510AFFC2AE3A3AE8FE533601" box="[368,488,1514,1540]" italics="true" pageId="12" pageNumber="431">G. elegans</emphasis>
</taxonomicName>
by having the mental disc fully developed with free edges (vs. a shallow mental disc completely fused with the chin).
<taxonomicName id="9BA02B93510AFFC2AD5B390CFD2C362D" box="[529,663,1550,1576]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2AD5B390CFD2C362D" box="[529,663,1550,1576]" italics="true" pageId="12" pageNumber="431">Garra rezai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="9BA02B93510AFFC2ACD3390CFC57362D" box="[921,1004,1550,1576]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rufa">
<emphasis id="6ED48C02510AFFC2ACD3390CFC57362D" box="[921,1004,1550,1576]" italics="true" pageId="12" pageNumber="431">G. rufa</emphasis>
</taxonomicName>
by having 1116 total gill rakers (vs. 2029), 7½ (rarely 8½) branched dorsal-fin rays (vs. usually 8½, rarely 7½ or 9½), 1518 circumpeduncular scales (vs. 1113), 5½ (rarely 4½ and 6½) transverse scale rows between the lateral line and the dorsal-fin origin (vs. 4½), and 1519 scales on predorsal midline between dorsal-fin origin and nape (vs. 1114). It is further distinguished from
<taxonomicName id="9BA02B93510AFFC2AF9F399CFE9C36BD" box="[213,295,1694,1720]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rufa">
<emphasis id="6ED48C02510AFFC2AF9F399CFE9C36BD" box="[213,295,1694,1720]" italics="true" pageId="12" pageNumber="431">G. rufa</emphasis>
</taxonomicName>
by a minimum K2P distance of 14.9% in the mtDNA COI barcode region. We examined
<specimenCount id="4AA69B99510AFFC2AA52399CFA2736BD" box="[1304,1436,1694,1720]" pageId="12" pageNumber="431" type="generic" typeStatus="syntypes">11 syntypes</specimenCount>
of
<taxonomicName id="9BA02B93510AFFC2AFFD39C0FE9D36D9" baseAuthorityName="Preston" baseAuthorityYear="1908" box="[183,294,1730,1756]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="obtusa">
<emphasis id="6ED48C02510AFFC2AFFD39C0FE9D36D9" box="[183,294,1730,1756]" italics="true" pageId="12" pageNumber="431">G. obtusa</emphasis>
</taxonomicName>
at NMW; all have 8½ branched dorsal-fin rays. We could count the gill rakers in two individuals of NMW 53257 (25, 25 total gill rakers) and one individual of NMW 53238 (26 total gill rakers). These
<typeStatus id="831BEEB2510AFFC2AA5B39E4FAC83705" box="[1297,1395,1766,1792]" pageId="12" pageNumber="431" type="syntype">syntypes</typeStatus>
are identified as
<taxonomicName id="9BA02B93510AFFC2AE623808FEC23721" box="[296,377,1802,1828]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rufa">
<emphasis id="6ED48C02510AFFC2AE623808FEC23721" box="[296,377,1802,1828]" italics="true" pageId="12" pageNumber="431">G. rufa</emphasis>
</taxonomicName>
, and
<taxonomicName id="9BA02B93510AFFC2AEFC3808FD983721" baseAuthorityName="Preston" baseAuthorityYear="1908" box="[438,547,1802,1828]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="obtusa">
<emphasis id="6ED48C02510AFFC2AEFC3808FD983721" box="[438,547,1802,1828]" italics="true" pageId="12" pageNumber="431">G. obtusa</emphasis>
</taxonomicName>
remains in the synonymy of
<taxonomicName id="9BA02B93510AFFC2AC203808FC003721" box="[874,955,1802,1828]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rufa">
<emphasis id="6ED48C02510AFFC2AC203808FC003721" box="[874,955,1802,1828]" italics="true" pageId="12" pageNumber="431">G. rufa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5C1F5010510AFFC3AF8D382CFAEB30FC" blockId="12.[151,1437,830,2009]" lastBlockId="13.[151,1436,150,249]" lastPageId="13" lastPageNumber="432" pageId="12" pageNumber="431">
<taxonomicName id="9BA02B93510AFFC2AF8D382CFEF4374D" box="[199,335,1838,1864]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2AF8D382CFEF4374D" box="[199,335,1838,1864]" italics="true" pageId="12" pageNumber="431">Garra rezai</emphasis>
</taxonomicName>
is distinguished from its subterranean congeners in the Tigris and Euphrates drainage (i.e.,
<taxonomicName id="9BA02B93510AFFC2AACB382CFE913769" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="lorestanensis">
<emphasis id="6ED48C02510AFFC2AACB382CFE913769" italics="true" pageId="12" pageNumber="431">G. lorestanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="9BA02B93510AFFC2AE7D3850FE673769" box="[311,476,1874,1900]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="tashanensis">
<emphasis id="6ED48C02510AFFC2AE7D3850FE673769" box="[311,476,1874,1900]" italics="true" pageId="12" pageNumber="431">G. tashanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="9BA02B93510AFFC2AEA33850FDD13769" box="[489,618,1874,1900]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="typhlops">
<emphasis id="6ED48C02510AFFC2AEA33850FDD13769" box="[489,618,1874,1900]" italics="true" pageId="12" pageNumber="431">G. typhlops</emphasis>
</taxonomicName>
and
<taxonomicName id="9BA02B93510AFFC2ADD53850FCFC3769" box="[671,839,1874,1900]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="widdowsoni">
<emphasis id="6ED48C02510AFFC2ADD53850FCFC3769" box="[671,839,1874,1900]" italics="true" pageId="12" pageNumber="431">G. widdowsoni</emphasis>
</taxonomicName>
) by having a brown or grey body (vs. whitish, orange or pink) and a fully developed eye (vs. eye externally invisible).
<taxonomicName id="9BA02B93510AFFC2AC213874FC543795" box="[875,1007,1910,1936]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2AC213874FC543795" box="[875,1007,1910,1936]" italics="true" pageId="12" pageNumber="431">Garra rezai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="9BA02B93510AFFC2ABA03874FA273795" box="[1258,1436,1910,1936]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="amirhosseini">
<emphasis id="6ED48C02510AFFC2ABA03874FA273795" box="[1258,1436,1910,1936]" italics="true" pageId="12" pageNumber="431">G. amirhosseini</emphasis>
</taxonomicName>
(data based on
<bibRefCitation id="38312DE1510AFFC2AE753898FD9A37B1" author="Esmaeili, H. R. &amp; Sayyadzadeh, G. &amp; Coad, B. W. &amp; Eagderi, S." box="[319,545,1946,1972]" pageId="12" pageNumber="431" pagination="82 - 121" refId="ref7944" refString="Esmaeili, H. R., Sayyadzadeh, G., Coad, B. W. &amp; Eagderi, S. (2016) Review of the genus Garra Hamilton, 1822 in Iran with description of a new species: a morpho-molecular approach (Teleostei: Cyprinidae). Iranian Journal of Ichthyology, 3 (2), 82 - 121." type="journal article" year="2016">
Esmaeili
<emphasis id="6ED48C02510AFFC2AEE23899FE5B37B1" box="[424,480,1946,1972]" italics="true" pageId="12" pageNumber="431">et al.</emphasis>
2016
</bibRefCitation>
) by having 3540 total lateral line scales (vs. 3336), 1116 gill rakers (vs. 16 20), and 1518 circumpeduncular scales (vs. 1314).
<taxonomicName id="9BA02B93510AFFC2ADBA38BCFCCE37DD" box="[752,885,1982,2008]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510AFFC2ADBA38BCFCCE37DD" box="[752,885,1982,2008]" italics="true" pageId="12" pageNumber="431">Garra rezai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="9BA02B93510AFFC2AB3A38BCFA9D37DD" box="[1136,1318,1982,2008]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="431" phylum="Chordata" rank="species" species="gymnothorax">
<emphasis id="6ED48C02510AFFC2AB3A38BCFA9D37DD" box="[1136,1318,1982,2008]" italics="true" pageId="12" pageNumber="431">G. gymnothorax</emphasis>
</taxonomicName>
by having 1518 circumpeduncular scales (vs. 1213), 1116 gill rakers (vs. 1621), and 1519 scales along the predorsal midline (vs 1012). It should be noted, that
<bibRefCitation id="38312DE1510BFFC3ADCE3FB9FCCC30D0" author="Esmaeili, H. R. &amp; Sayyadzadeh, G. &amp; Coad, B. W. &amp; Eagderi, S." box="[644,887,187,213]" pageId="13" pageNumber="432" pagination="82 - 121" refId="ref7944" refString="Esmaeili, H. R., Sayyadzadeh, G., Coad, B. W. &amp; Eagderi, S. (2016) Review of the genus Garra Hamilton, 1822 in Iran with description of a new species: a morpho-molecular approach (Teleostei: Cyprinidae). Iranian Journal of Ichthyology, 3 (2), 82 - 121." type="journal article" year="2016">
Esmaeili
<emphasis id="6ED48C02510BFFC3ADA73FB9FC9E30D0" box="[749,805,187,213]" italics="true" pageId="13" pageNumber="432">et al.</emphasis>
(2016)
</bibRefCitation>
diagnosed
<taxonomicName id="9BA02B93510BFFC3ACBC3FB9FB1630D0" box="[1014,1197,187,213]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="gymnothorax">
<emphasis id="6ED48C02510BFFC3ACBC3FB9FB1630D0" box="[1014,1197,187,213]" italics="true" pageId="13" pageNumber="432">G. gymnothorax</emphasis>
</taxonomicName>
lacking scales on the breast, a character state confirmed in only a part of our materials and therefore not diagnostic to the species.
</paragraph>
<caption id="08DF0098510BFFC3AFDD3897FBC037A8" ID-DOI="http://doi.org/10.5281/zenodo.7223713" ID-Zenodo-Dep="7223713" box="[151,1147,1941,1966]" httpUri="https://zenodo.org/record/7223713/files/figure.png" pageId="13" pageNumber="432" startId="13.[151,250,1941,1965]" targetBox="[151,1436,273,1918]" targetPageId="13">
<paragraph id="5C1F5010510BFFC3AFDD3897FBC037A8" blockId="13.[151,1147,1941,1966]" box="[151,1147,1941,1966]" pageId="13" pageNumber="432">
<emphasis id="6ED48C02510BFFC3AFDD3897FEA537A8" bold="true" box="[151,286,1941,1965]" pageId="13" pageNumber="432">FIGURE 11.</emphasis>
Type locality of
<taxonomicName id="9BA02B93510BFFC3AE813894FDA737A8" box="[459,540,1942,1965]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="432" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C02510BFFC3AE813894FDA737A8" box="[459,540,1942,1965]" italics="true" pageId="13" pageNumber="432">G. rezai</emphasis>
</taxonomicName>
; from above: spring in Boein-e-Olia; Boein-e-Sofla stream.
</paragraph>
</caption>
<paragraph id="5C1F50105108FFC0AF8D3F95FD683280" blockId="14.[151,1437,150,645]" pageId="14" pageNumber="433">
<taxonomicName id="9BA02B935108FFC0AF8D3F95FEF430B5" box="[199,335,151,177]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025108FFC0AF8D3F95FEF430B5" box="[199,335,151,177]" italics="true" pageId="14" pageNumber="433">Garra rezai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="9BA02B935108FFC0AD123F95FCB130B4" box="[600,778,151,177]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="hormuzensis">
<emphasis id="6ED48C025108FFC0AD123F95FCB130B4" box="[600,778,151,177]" italics="true" pageId="14" pageNumber="433">G. hormuzensis</emphasis>
</taxonomicName>
by having 1519 scales along the predorsal midline (vs 1014), 5½ (rarely 4½ and 6½) transverse scale rows between the lateral line and the dorsal-fin origin (vs. usually 4½), and 1116 gill rakers (vs. 1621).
<taxonomicName id="9BA02B935108FFC0AD1C3FDDFD6130FD" box="[598,730,223,249]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025108FFC0AD1C3FDDFD6130FD" box="[598,730,223,249]" italics="true" pageId="14" pageNumber="433">Garra rezai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="9BA02B935108FFC0AC9D3FDDFB3630FC" box="[983,1165,223,249]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="meymehensis">
<emphasis id="6ED48C025108FFC0AC9D3FDDFB3630FC" box="[983,1165,223,249]" italics="true" pageId="14" pageNumber="433">G. meymehensis</emphasis>
</taxonomicName>
by having 1519 scales along the predorsal midline (vs. 11), 5½ (rarely 4½ and 6½) transverse scale rows between the lateral line and the dorsal-fin origin (vs. usually 4½), and 1116 gill rakers (vs. 1621) (data on
<taxonomicName id="9BA02B935108FFC0ACA53E25FB1E3144" box="[1007,1189,295,321]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="meymehensis">
<emphasis id="6ED48C025108FFC0ACA53E25FB1E3144" box="[1007,1189,295,321]" italics="true" pageId="14" pageNumber="433">G. meymehensis</emphasis>
</taxonomicName>
are based on ZamaniFaradonbe
<emphasis id="6ED48C025108FFC0AE5E3E49FEF03160" box="[276,331,331,357]" italics="true" pageId="14" pageNumber="433">et al.</emphasis>
2021a).
<taxonomicName id="9BA02B935108FFC0AEE63E49FD943161" box="[428,559,331,357]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025108FFC0AEE63E49FD943161" box="[428,559,331,357]" italics="true" pageId="14" pageNumber="433">Garra rezai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="9BA02B935108FFC0AC6C3E49FC1C3160" box="[806,935,331,357]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="mondica">
<emphasis id="6ED48C025108FFC0AC6C3E49FC1C3160" box="[806,935,331,357]" italics="true" pageId="14" pageNumber="433">G. mondica</emphasis>
</taxonomicName>
, by having 3540 total lateral line scales (vs. 2832), and 1116 gill rakers (vs. 1823). It should be noted that
<bibRefCitation id="38312DE15108FFC0AC3C3E6DFB21318C" author="Sayyadzadeh, G. &amp; Esmaeili, H. R. &amp; Freyhof, J." box="[886,1178,366,393]" pageId="14" pageNumber="433" pagination="1" refId="ref9706" refString="Sayyadzadeh, G., Esmaeili, H. R. &amp; Freyhof, J. (2015) Garra mondica, a new species from the Mond River drainage with remarks on the genus Garra from the Persian Gulf basin in Iran (Teleostei: Cyprinidae). Zootaxa, 4048 (1), 075 - 089. https: // doi. org / 10.11646 / zootaxa. 4048.1.4" type="journal article" year="2015">
Sayyadzadeh
<emphasis id="6ED48C025108FFC0AB5B3E6DFBF2318C" box="[1041,1097,367,393]" italics="true" pageId="14" pageNumber="433">et al.</emphasis>
(2015)
</bibRefCitation>
diagnosed
<taxonomicName id="9BA02B935108FFC0AA503E6DFA27318C" box="[1306,1436,367,393]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="mondica">
<emphasis id="6ED48C025108FFC0AA503E6DFA27318C" box="[1306,1436,367,393]" italics="true" pageId="14" pageNumber="433">G. mondica</emphasis>
</taxonomicName>
as lacking scales on the breast, the belly and the back in front of the dorsal-fin origin. However, in our materials, as the one examined by
<bibRefCitation id="38312DE15108FFC0AEE43EB5FC9231D4" author="Zamani-Faradonbe, M. &amp; Zhang, E. &amp; Keivany, Y." box="[430,809,439,465]" pageId="14" pageNumber="433" pagination="380 - 394" refId="ref10080" refString="Zamani-Faradonbe, M., Zhang, E. &amp; Keivany, Y. (2021 b) Garra hormuzensis, a new species from the upper Kol River drainage in the Persian Gulf basin (Teleostei: Cyprinidae). Zootaxa, 5052 (3), 380 - 394. https: // doi. org / 10.11646 / zootaxa. 5052.3.4" type="journal article" year="2021">
Zamani-Faradonbe
<emphasis id="6ED48C025108FFC0ADC63EB5FD7D31D4" box="[652,710,439,465]" italics="true" pageId="14" pageNumber="433">et al.</emphasis>
(2021b)
</bibRefCitation>
, the breast is covered by embedded scales, the belly is cover by non-embedded scales and there are 1115 scales along the predorsal midline.
<taxonomicName id="9BA02B935108FFC0AB293ED9FB5C31F1" box="[1123,1255,475,501]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025108FFC0AB293ED9FB5C31F1" box="[1123,1255,475,501]" italics="true" pageId="14" pageNumber="433">Garra rezai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="9BA02B935108FFC0AF9E3EFDFEF3321D" box="[212,328,511,537]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="persica">
<emphasis id="6ED48C025108FFC0AF9E3EFDFEF3321D" box="[212,328,511,537]" italics="true" pageId="14" pageNumber="433">G. persica</emphasis>
</taxonomicName>
by having 9+8 branched caudal-fin rays (vs. 8+8), and 1116 gill rakers (vs. 1719).
<taxonomicName id="9BA02B935108FFC0AA4A3EFDFA38321D" box="[1280,1411,511,537]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="rezai">
<emphasis id="6ED48C025108FFC0AA4A3EFDFA38321D" box="[1280,1411,511,537]" italics="true" pageId="14" pageNumber="433">Garra rezai</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="9BA02B935108FFC0AE253D21FE793239" box="[367,450,547,573]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="tiam">
<emphasis id="6ED48C025108FFC0AE253D21FE793239" box="[367,450,547,573]" italics="true" pageId="14" pageNumber="433">G. tiam</emphasis>
</taxonomicName>
by having 3540 total lateral line scales (vs 3034), 1116 gill rakers (vs. 1822), 1519 scales along predorsal midline (vs 911, or embedded), and 1518 circumpeduncular scales (vs. 1213) (data on
<taxonomicName id="9BA02B935108FFC0AACB3D45FF723281" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="433" phylum="Chordata" rank="species" species="tiam">
<emphasis id="6ED48C025108FFC0AACB3D45FF723281" italics="true" pageId="14" pageNumber="433">G. tiam</emphasis>
</taxonomicName>
are based on
<bibRefCitation id="38312DE15108FFC0AE293D69FD7C3280" author="Zamani-Faradonbe, M. &amp; Keivany, Y. &amp; Dorafshan, S. &amp; Zhang, E." box="[355,711,619,645]" pageId="14" pageNumber="433" pagination="249 - 270" refId="ref10019" refString="Zamani-Faradonbe, M., Keivany, Y., Dorafshan, S. &amp; Zhang, E. (2021 a) Two new species of Garra (Teleostei: Cyprinidae) from western Iran. Ichthyological Exploration Freshwaters, 30 (3), 249 - 270. https: // doi. org / 10.23788 / IEF- 1137" type="journal article" year="2021">
Zamani-Faradonbe
<emphasis id="6ED48C025108FFC0AD753D69FDCC3280" box="[575,631,619,645]" italics="true" pageId="14" pageNumber="433">et al.</emphasis>
2021a
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>