235 lines
30 KiB
XML
235 lines
30 KiB
XML
<document id="C5D936CE4262656F6DEDE610D50966D3" ID-DOI="http://dx.doi.org/10.3897/zookeys.577.7254" ID-GBIF-Dataset="bad453df-1230-42fb-98f6-30fdcaa45f51" ID-PMC="PMC4829886" ID-Pensoft-Pub="1313-2970-577-125" ID-PubMed="27110191" ID-ZooBank="327F242512D544E28CD6E5BC91773077" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-577-125" ModsDocOrigin="ZooKeys 577" ModsDocTitle="Description of two new species of the Exocelinabroschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1459885117108" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo & Balke, Michael" docDate="2016" docId="B3D599E47C481D4613E986E5BC58185E" docLanguage="en" docName="ZooKeys 577: 125-148" docOrigin="ZooKeys 577" docSource="http://dx.doi.org/10.3897/zookeys.577.7254" docTitle="Exocelina broschii Balke 1998" docType="treatment" docVersion="5" lastPageNumber="128" masterDocId="FFECB328490DFFA1BA62B2070F4EC00F" masterDocTitle="Description of two new species of the Exocelinabroschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="148" masterPageNumber="125" pageNumber="128" updateTime="1701316882021" updateUser="plazi">
|
||
<mods:mods id="BC7D286F95AE84FA4FB96DE24DFC863E" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="E475231F998A0A555EB78DFBA5680A72">
|
||
<mods:title id="843B7F2923174AB2DABE490CFBCC3C32">Description of two new species of the Exocelinabroschii-group from Papua New Guinea, with revision and key to all representatives of this species group (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="54D182A83DC423DFFA321A13F973B943" type="personal">
|
||
<mods:role id="18F4BAF29DA4AAF162CDE5EBCAAB31F4">
|
||
<mods:roleTerm id="824F8DCE63B5D2E98EDB20CDD443A87E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="77A67F846D53B8622F082B646BBD56FC">Shaverdo, Helena</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="7B3682118E6308ACC906F65D28BD7DE8" type="personal">
|
||
<mods:role id="E4AC2102C3F0ABE2CBA42EA9B7B2E267">
|
||
<mods:roleTerm id="DE7184D3A33D72E011278FD06FC9B0CC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9A3017BB07920B2A50F47CF85090CD1D">Sagata, Katayo</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FCD80D594741E42651E1ACA775EF5886" type="personal">
|
||
<mods:role id="16852DA206AD0241C6BC864183595803">
|
||
<mods:roleTerm id="16F70AFD1647F5D09EDBCBD8014FF5A5">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="139D3A7CA6D64CD23CC54DCC2A9FCB49">Balke, Michael</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E05A3C938574DFEF7550B14D3103029D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="707383C1FF224C4271B3C5CB92F5DBC6" type="host">
|
||
<mods:titleInfo id="B1646E0781CCC4CA8F7180D1B6E8590F">
|
||
<mods:title id="9580152D0F2D27AE21257AE7A2446368">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="109F91241BEAEC92837FB3754CC1B8A6">
|
||
<mods:date id="1D33A70B83C512AF521374BBC30A59BE">2016</mods:date>
|
||
<mods:detail id="7C97E69FD1DCAE7AA7B06C6505C9FC83" type="volume">
|
||
<mods:number id="C96BE7303A3617F93E7470455478398C">577</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="9F3DAC484825041EF3D1F2385DA3CC48" unit="page">
|
||
<mods:start id="0EF839AD733D04C8AD26D94DA073407E">125</mods:start>
|
||
<mods:end id="53BE057382DF043BD9B851A925ACD5A9">148</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="C0204630E69F522BD71D27BD987B35EA">
|
||
<mods:url id="27F2129CF004C1C04D6D6ECEB1B8DB99">http://dx.doi.org/10.3897/zookeys.577.7254</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="9F22E7222863291730CD85EDDA7BA59F">journal article</mods:classification>
|
||
<mods:identifier id="42436D36D3DA4C31761E09C2E52D2810" type="DOI">http://dx.doi.org/10.3897/zookeys.577.7254</mods:identifier>
|
||
<mods:identifier id="E1C74F6C10ADDE58B0E7E4E9BE71BBD7" type="Pensoft-Pub">1313-2970-577-125</mods:identifier>
|
||
<mods:identifier id="D5167CAEE881CE5DC0D57BB801852D54" type="ZooBank">327F242512D544E28CD6E5BC91773077</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="B3D599E47C481D4613E986E5BC58185E" ID-GBIF-Taxon="127880282" LSID="urn:lsid:plazi:treatment:B3D599E47C481D4613E986E5BC58185E" httpUri="http://treatment.plazi.org/id/B3D599E47C481D4613E986E5BC58185E" lastPageNumber="128" pageId="3" pageNumber="128">
|
||
<subSubSection id="CD8736A2A6B3F65CBC9267F834409965" pageId="3" pageNumber="128" type="multiple">
|
||
<paragraph id="E813D45CB42DB1B7554CC83E2A4016C8" pageId="3" pageNumber="128">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="891D9189A98928830F181C2FDE5EE54A" pageId="3" pageNumber="128" type="nomenclature">
|
||
<paragraph id="2EDA67E79FEDA86BB190FD806ED47420" pageId="3" pageNumber="128">
|
||
1.
|
||
<taxonomicName id="540085C5A22ABF8672E533C23A0E1E49" ID-CoL="3DLPC" authority="Balke, 1998" authorityName="Balke" authorityYear="1998" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii (Balke, 1998)</taxonomicName>
|
||
Figs 1-3, 8, 9
|
||
<normalizedToken id="A02C5614752275F3DF35927325B45266" originalValue="A–E">A-E</normalizedToken>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="538617F7D7868F84F356A6EC245C3C22" pageId="3" pageNumber="128" type="reference_group">
|
||
<paragraph id="B36D11490F2B6AEF625406970B586C92" pageId="3" pageNumber="128">
|
||
<taxonomicName id="61AFF2BAD43BE6170C0FAB44DD2D178A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus (Papuadytes) broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii" subGenus="Papuadytes">Copelatus (Papuadytes) broschii</taxonomicName>
|
||
Balke, 1998: 327;
|
||
<bibRefCitation id="CDB7ED830E998B9844E8CC25E1445956" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="13" pageNumber="138" pagination="1 - 395" title="World catalogue of insects" year="2001">Nilsson 2001</bibRefCitation>
|
||
: 76 (catalogue).
|
||
</paragraph>
|
||
<paragraph id="AC8DF9FA924FDBE8FD282C6A8EBDA480" pageId="3" pageNumber="128">
|
||
<taxonomicName id="18B791411E0FAF18D4B63423FCFF6206" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Papuadytes broschii</taxonomicName>
|
||
(Balke, 1998):
|
||
<bibRefCitation id="FB5E07EAED40D28AD9A6CD88817B1148" author="Shaverdo, HV" journalOrPublisher="Aquatic Insects" pageId="13" pageNumber="138" pagination="269 - 280" title="Five new species of the genus Papuadytes Balke, 1998 from New Guinea (Coleoptera: Dytiscidae)." url="10.1080/01650420500290169" volume="27" year="2005">Shaverdo et al. 2005</bibRefCitation>
|
||
: 270, 271 (notes, illustration).
|
||
</paragraph>
|
||
<paragraph id="BC5CC2CB90A201A09DC44EFB99C760CF" pageId="3" pageNumber="128">
|
||
<taxonomicName id="60834EF22841E825F4439D1DBFD91F46" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Papuadytes broschii</taxonomicName>
|
||
(Balke, 1998):
|
||
<bibRefCitation id="F0B847D579329BE311DA1863DE2E3500" author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="13" pageNumber="138" pagination="55 - 74" title="World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae)." volume="76" year="2006">Nilsson and Fery 2006</bibRefCitation>
|
||
: 56 (addition to catalogue).
|
||
</paragraph>
|
||
<paragraph id="A50BEB871F88C6437637F7792BEAB324" pageId="3" pageNumber="128">
|
||
<taxonomicName id="49416BF3D82A19CFCB67BF1DC98FD8F5" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
|
||
(Balke, 1998):
|
||
<bibRefCitation id="508994F708194FF75DC2377D6B99B1EB" author="Nilsson, AN" journalOrPublisher="Latissimus" pageId="13" pageNumber="138" pagination="33 - 34" title="Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998." volume="23" year="2007">Nilsson 2007</bibRefCitation>
|
||
: 33 (comb. n.).
|
||
</paragraph>
|
||
<paragraph id="7E43EEBFDF69C05EEF4244A46216E52C" pageId="3" pageNumber="128">
|
||
<taxonomicName id="14B8604DC1ED4AE62BBBCC619405A91B" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
|
||
undescribed sp. MB1520:
|
||
<bibRefCitation id="E821094868DD26C50F1E88717DBA11B8" author="Toussaint, EFA" journalOrPublisher="Nature Communications" pageId="13" pageNumber="138" pagination="1 - 10" title="The towering orogeny of New Guinea as a trigger for arthropod megadiversity." url="10.1038/ncomms5001" volume="1" year="2014">Toussaint et al. 2014</bibRefCitation>
|
||
: Supplementary figs 1-4, tab. 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DC5711B8BBF1C3764AB8AC3E9DAEF1AA" pageId="3" pageNumber="128" type="type locality">
|
||
<paragraph id="97A7DE1CE39F92EB0C9C95F1C4DD2C36" pageId="3" pageNumber="128">Type locality.</paragraph>
|
||
<paragraph id="4A6FB032BD179EAD72C37D2F8103FF46" pageId="3" pageNumber="128">
|
||
Papua New Guinea: Madang Province, Finisterre Range, Moro, approximately
|
||
<geoCoordinate id="B4967A942C9F8F9C1A2004B77CBE4D36" direction="south" orientation="latitude" precision="1" value="-5.713222">5°42'47.6"S</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="453DA99F87B87C3E5C23771EE81199C6" direction="east" orientation="longitude" precision="1" value="146.06114">146°03'40.1"E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EC1695BA8FE6E2E5889C00FE29AB8CE9" pageId="3" pageNumber="128" type="type material">
|
||
<paragraph id="C237F21CC870E958F94C833348F441C7" pageId="3" pageNumber="128">Type material studied.</paragraph>
|
||
<paragraph id="0F7E06C36B07410BE4789A1DDCF6A72E" pageId="3" pageNumber="128">
|
||
<typeStatus id="983EC6ED214F435C23D22164D3564DC8" pageId="3" pageNumber="128">Holotype</typeStatus>
|
||
: male "Stn. No. 82", "NEW GUINEA: Madang Dist., Finisterre Mts. Moro.C.5550ft. 30.x.-15.xi.1964.", "M.E. Bacchus. B.M. 1965-120",
|
||
<normalizedToken id="9F44CF77EC1EE070ABB7DC8DA7D0B3F3" originalValue="“Holotypus”">"Holotypus"</normalizedToken>
|
||
[red], "
|
||
<taxonomicName id="5A52539923FFB2E7CDE4B2162656007A" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Copelatus broschii</taxonomicName>
|
||
sp.n. Balke des. 1997" (BMNH).
|
||
<typeStatus id="E4DAC060D740418A0CF8C6D1A8F0C2EC" pageId="3" pageNumber="128">Paratypes</typeStatus>
|
||
: 4 males, 1 female with the same labels as the
|
||
<typeStatus id="F358C96956BCBD69DCE488203E2B5A46" pageId="3" pageNumber="128">holotype</typeStatus>
|
||
, except for "Paratypus
|
||
<taxonomicName id="4114F8D9DBEDF990804B19BDBD857219" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Copelatus broschii</taxonomicName>
|
||
sp.n. Balke des. 1997" (BMNH, NHMW, ZSM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DE2EE7B5CBABE3467895DA56FD1EB5CD" pageId="3" pageNumber="128" type="materials_examined">
|
||
<paragraph id="92C22147E32560AE5F6A9E7E539E22A1" pageId="3" pageNumber="128">Additional material.</paragraph>
|
||
<paragraph id="930BB3B79D45D61F41E5390FAF4D693D" pageId="3" pageNumber="128">
|
||
Madang: 1 male "Stn. No. 82", "NEW GUINEA: Madang Dist., Finisterre Mts. Moro.C.5550ft. 30.x.-15.xi.1964.", "M.E. Bacchus. B.M. 1965-120", "Paratypus
|
||
<taxonomicName id="B0D985B9AFA8DD74E81EE638A2823758" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Copelatus broschii</taxonomicName>
|
||
sp.n. Balke des. 1997" (BMNH) - although this specimen is with the
|
||
<typeStatus id="9C48C1915CC2085EA60EE6655B7C8729" pageId="3" pageNumber="128">paratype</typeStatus>
|
||
label, it is not included in the type material of the original description in
|
||
<bibRefCitation id="B98C20807190CB105495A154C03A8AEE" author="Balke, M" journalOrPublisher="Annalen des Naturhistorischen Museum Wien" pageId="12" pageNumber="137" pagination="301 - 341" title="Revision of New Guinea Copelatus Erichson, 1832 (Insecta: Coleoptera: Dytiscidae): The running water species, Part I." volume="100 B" year="1998">Balke (1998)</bibRefCitation>
|
||
. 17 males, 8 females "Papua New Guinea: Madang, Adalbert Mts., Sewan - Keki, 700m, 4.v.2006, 04.42.215S 145.25.154E, Balke & Manaono (PNG 51)" (NARI, NHMW, ZSM). 7 males "Papua New Guinea: Madang, Adalbert Mts., Keki, 850m, 4.v.2006, nr 04.42.300S 145.25.089E, Balke & Manaono (PNG 52)", one male with an additional green label "DNA M.Balke 1300" (NHMW, ZSM). 34 males, 35 females "Papua New Guinea: Madang, Adalbert Mts., below Keki, 790m, 5.v.2006, 04.42.300S 145.25.089E, Balke & Manaono (PNG 53)" (NARI, NHMW, ZSM). 14 males, 13 females "Papua New Guinea: Madang, Adalbert Mts., creek nr Keki, 790m, 28.xi.2006, 04.42.300S 145.25.089E, Binatang Boys leg. (PNG 53a)" (NHMW, ZSM). 4 males, 5 females "Papua New Guinea: Madang, Keki, Adalbert Mts., 400m, 29.xi.2006, 04.43.058S 145.24.437E, Binatang Boys, (PNG 119)" (NHMW, ZSM). 7 males, 6 females "Papua New Guinea: Madang, Keki-Sewan, Adalbert Mts., 700m, 30.xi.2006, nr 04.41.802S 145.25.460E Binatang Boys (PNG 120)" (NHMW, ZSM). 1 male "Papua New Guinea: Madang, Simbai area, 1800-2400m, 8.iii.2007, 05.12.693S 144.35.521E, Kinibel (PNG 151)" (ZSM). 24 males "Papua New Guinea: Madang, Simbai area, 1200m, 10.iii.2007, 05.13.389S 144.37.285E, Kinibel (PNG 152)" (NARI, NHMW, ZSM). 39 males "Papua New Guinea: Madang, Simbai area, 1200m, 11.iii.2007, 05.13.333S 144.37.611E, Kinibel (PNG 153)" (NARI, NHMW, ZSM). 10 males, 2 females "Papua New Guinea: Madang, Simbai - Mombeen, 1100m, 11.iii.2007, 05.12.876S 144.41.759E, Kinibel (PNG 154)" (NHMW, ZSM). Eastern Highlands: 4 males, 6 females "Papua New Guinea: Eastern Highlands, Bena Bridge, 1400m, 8.xii.2007, 06.10.781S 145.26.034E, Balke & Sagata (PNG 164)" (ZSM). 2 males "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 700m, 24.xi.2006, 05.52.754S 145.23.209E, Balke & Kinibel (PNG 109)", one of them with an additional green label "DNA M.Balke 1520" (ZSM). 1 male, 2 females "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 1200m, 24.xi.2006, nr 05.52.754S 145.23.209E, Balke & Kinibel (PNG 110)", one female with an additional green label "DNA M.Balke 1522" (ZSM). 8 males "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 800m, 24.xi.2006, 05.50.021S 145.24.664E, Balke & Kinibel (PNG 112)" (NHMW, ZSM). 18 males "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 750m, 25.xi.2006, 05.49.892S 145.24.491E, Balke & Kinibel (PNG 113)" (NARI, NHMW, ZSM). 3 males "Papua New Guinea: Madang, Akameku - Brahmin, Bismarck Range, 750m, 25.xi.2006, nr 05.49.307S 145.24.389E, Balke & Kinibel (PNG 114)" (NHMW, ZSM).
|
||
</paragraph>
|
||
<paragraph id="1899F245BE3839A3D19C2B6B24174EA6" pageId="3" pageNumber="128">
|
||
Females of doubtful identity. Eastern Highlands: 12 females "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 700m, 24.xi.2006, 05.52.754S 145.23.209E, Balke & Kinibel (PNG 109)" (ZSM). 24 females "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 800m, 24.xi.2006, 05.50.021S 145.24.664E, Balke & Kinibel (PNG 112)" (NARI, NHMW, ZSM). 15 females "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 750m, 25.xi.2006, 05.49.892S 145.24.491E, Balke & Kinibel (PNG 113)" (NHMW, ZSM). 25 females "Papua New Guinea: Madang, Akameku - Brahmin, Bismarck Range, 750m, 25.xi.2006, nr 05.49.307S 145.24.389E, Balke & Kinibel (PNG 114)" (NARI, NHMW, ZSM). These females are a mixture of two species:
|
||
<taxonomicName id="2796A8C5A480CF64943E88B9DCC2451F" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
|
||
and
|
||
<taxonomicName id="416FDEA99E52402A609B62379DE2D717" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina damantiensis" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="damantiensis">Exocelina damantiensis</taxonomicName>
|
||
(Balke, 1998). 19 males "Papua New Guinea: Madang, Simbai area, 1200m, 10.iii.2007, 05.13.389S 144.37.285E, Kinibel (PNG 152)" (NHMW, ZSM). These females are a mixture of two species:
|
||
<taxonomicName id="B2AF9A53C56EB5BA449F0D499BE53CDD" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
|
||
and
|
||
<taxonomicName id="C6CC1A70A412B9E9FB90B7739F1C8AC6" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina damantiensis" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="damantiensis">Exocelina? damantiensis</taxonomicName>
|
||
(Balke, 1998). 53 females "Papua New Guinea: Madang, Simbai area, 1200m, 11.iii.2007, 05.13.333S 144.37.611E, Kinibel (PNG 153)" (NARI, NHMW, ZSM). These females are a mixture of three species:
|
||
<taxonomicName id="6B26B46FE1247D75F5023BCDAFC9358A" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
|
||
,
|
||
<taxonomicName id="D265B20E400C8ABDFF5D21ECC564A77A" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina simbaiarea" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="simbaiarea">Exocelina simbaiarea</taxonomicName>
|
||
Shaverdo & Balke, 2014, and
|
||
<taxonomicName id="542FC7A5B73B6755F84E3E2FA9FA1BA3" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina damantiensis" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="damantiensis">Exocelina? damantiensis</taxonomicName>
|
||
(Balke, 1998).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8DF2F20275912E02E3F0021CA01EC38E" pageId="3" pageNumber="128" type="additions to the description">
|
||
<paragraph id="9A0B8D1372B70EFD5AE827DC48944E9B" pageId="3" pageNumber="128">Additions to the description</paragraph>
|
||
<paragraph id="1854A7717D1C1BA4CE5863C350F7843A" pageId="3" pageNumber="128">
|
||
(original description in
|
||
<bibRefCitation id="93B757FFC7CBF3F49DAFAA9168297219" author="Balke, M" journalOrPublisher="Annalen des Naturhistorischen Museum Wien" pageId="12" pageNumber="137" pagination="301 - 341" title="Revision of New Guinea Copelatus Erichson, 1832 (Insecta: Coleoptera: Dytiscidae): The running water species, Part I." volume="100 B" year="1998">Balke 1998</bibRefCitation>
|
||
, p. 327). Size and shape: Beetles small to medium-sized (TL-H 3.2-4.0 mm, TL 3.7-4.4 mm, MW 1.75-2.15 mm; holotype: TL-H 3.55 mm, TL 3.9 mm, MW 1.9 mm). Male: Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 14 and posterior row of 5 short setae (Fig. 8A). Abdominal ventrite 6 with 3-6 lateral striae on each side. Median lobe with slightly curved apex in lateral view and more or less rounded in ventral view. Its ventral sclerite with almost equal, short apical lobes: left lobe broad or narrow (sometimes with broken apex, e.g., Fig. 9A) and right lobe short, relatively broad (Fig. 8B, C).
|
||
</paragraph>
|
||
<caption id="FE1A4A87BC55D3D0EC3F9B48CFEE3FC7" pageId="3" pageNumber="128">
|
||
<paragraph id="59FF9AC8ABAEC82270258998FDB5EEC4" pageId="3" pageNumber="128">
|
||
Figures 1-3. Habitus and coloration of
|
||
<taxonomicName id="4DFCDF89B2239F23118C42C63CF4884E" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
|
||
(Balke, 1998). 1 holotype 2 Madang, Simbai area, PNG 152, specimen with finer dorsal punctation 3 Madang, Simbai area, PNG 152, specimen with coarser dorsal punctation.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D9381C17323EBA5E5487D31C724999D0" pageId="3" pageNumber="128">
|
||
<paragraph id="5436D668AD5276CBC28D945C7187FA73" pageId="3" pageNumber="128">
|
||
Figures 4-7. Habitus and coloration of the holotypes. 4
|
||
<taxonomicName id="A43E013C7E9B91BB88319F3B8CFEFF7B" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina hintelmannae" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="hintelmannae">Exocelina hintelmannae</taxonomicName>
|
||
(Shaverdo, Sagata & Balke, 2005) 5
|
||
<taxonomicName id="0195020A255AA89C40473BD9D8ACE473" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina marinae" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="marinae">Exocelina marinae</taxonomicName>
|
||
(Shaverdo, Sagata & Balke, 2005) 6
|
||
<taxonomicName id="69FB226A68A24A890FA7A2BBA676B737" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina mondmillensis" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="mondmillensis">Exocelina mondmillensis</taxonomicName>
|
||
sp. n. 7
|
||
<taxonomicName id="280A915F6863A2EC7C012807FC74EC80" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina pseudomarinae" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="pseudomarinae">Exocelina pseudomarinae</taxonomicName>
|
||
sp. n.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3B33FBA0A4EA81498F5040691A6BFD31" pageId="3" pageNumber="128">
|
||
<paragraph id="0B5E41E422E17D19DA5F0B4D25A63457" pageId="3" pageNumber="128">
|
||
Figure 8.
|
||
<taxonomicName id="ACF75E0EDBB8EF1CF4C732F94B60115F" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
|
||
(Balke, 1998). A male protarsomeres 4-5 in ventral view (Madang, Simbai area, PNG 153) B median lobe in ventral view (paratype) C median lobe in lateral view (paratype) D paramere in external view (Madang, Simbai area, PNG 153).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="224F7E4D08AC0DE9A62CC882279CD4BA" pageId="3" pageNumber="128">
|
||
<paragraph id="4AE1C424840CAEED3E3BBF7CB7CD17E8" pageId="3" pageNumber="128">
|
||
Figure 9. Lateral view of median lobe of (
|
||
<normalizedToken id="0C4ABC4447C8DE7930FA7E3687BBBBF2" originalValue="A–E">A-E</normalizedToken>
|
||
)
|
||
<taxonomicName id="D50C507D84405DCBDAB5A145296FE046" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
|
||
(Balke, 1998) and (F, G)
|
||
<taxonomicName id="F1622FB9152871A6197CE54CB1A1774D" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina hintelmannae" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="hintelmannae">Exocelina hintelmannae</taxonomicName>
|
||
(Shaverdo, Sagata & Balke, 2005). A, B Madang, Simbai area, PNG 153 C Madang, Adalbert Mts., Keki, PNG 52 D Madang, Adalbert Mts., below Keki, PNG 53 E Eastern Highlands, Bena Bridge, PNG 164 F, G Simbu/ Eastern Highlands, Crater Mountain, Sera-Herowana, Hulene River, PNG 017. Setae not shown.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E595AA637D659FB0023E3CA0D5F8990B" pageId="3" pageNumber="128">
|
||
Variability: Beetles vary in size, kind of dorsal punctation (Figs 1-3), and shape of ventral sclerite of the median lobe (Fig. 9
|
||
<normalizedToken id="27DF54914D5EE65DB4BB2035344DCD09" originalValue="A–E">A-E</normalizedToken>
|
||
). Dorsal punctation in great majority of the specimens is fine (Fig. 3), but some specimens have slightly coarser punctation, and very few (the types and one specimen from Simbai area, Madang, see Figs 1, 2) have distinct punctation, similar to that of
|
||
<taxonomicName id="BAB787DF8AE96B16E158A2F6AD9DCDAE" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina pseudomarinae" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="pseudomarinae">Exocelina pseudomarinae</taxonomicName>
|
||
sp. n. There are populations (e.g., from Simbai area, Madang, and Akameku-Brahmin, Eastern Highlands) with the left lobe of the ventral sclerite of the median lobe very narrow (Fig. 9A, B). However, this character is not very stable even in one population. Taking into consideration the other different shapes of the ventral sclerite observed (Figs 8B, 9
|
||
<normalizedToken id="05D65A32DD14F5B80F264263FE0BD3CF" originalValue="C–E">C-E</normalizedToken>
|
||
) as well as the fact that it is differently sclerotized in different specimens and because of that is variable in shape, we treated all the material as
|
||
<taxonomicName id="FE0ED2A0C9A1CBAA503962A415252975" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
|
||
. Such variability in shape of the ventral sclerite of the median lobe is also characteristic for other species of this group (Figs 9, 10, 12).
|
||
</paragraph>
|
||
<caption id="1531AA7A8BD87C9940B08147C4873C90" pageId="3" pageNumber="128">
|
||
<paragraph id="98EE925F1A49C6CF4D94AB4D01257DE6" pageId="3" pageNumber="128">
|
||
Figure 10.
|
||
<taxonomicName id="D5A5A344C14B6BBA8C6D5BF81087499A" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina marinae" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="marinae">Exocelina marinae</taxonomicName>
|
||
(Shaverdo, Sagata & Balke, 2005), lateral (
|
||
<normalizedToken id="FAE5D6E6CF0EE0C3A03DA365CCD17AEB" originalValue="A–C">A-C</normalizedToken>
|
||
) and ventral (
|
||
<normalizedToken id="1B765B85727F8650987EADF2EE0E6973" originalValue="D–F">D-F</normalizedToken>
|
||
) views of median lobe. A, D Sandaun, Mianmin, PNG 196 B, E Sandaun, Mianmin area, PNG 234 C, F Southern Highlands, Tari to Koroba, PNG 65. Setae not shown.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="15C5824E3C539129476CA6F29EB0F5C5" pageId="3" pageNumber="128">
|
||
<paragraph id="D447358144D928AA4958000C083088CC" pageId="3" pageNumber="128">
|
||
Figure 11.
|
||
<taxonomicName id="EFB0206007441737BA23E1C4016016AD" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina mondmillensis" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="mondmillensis">Exocelina mondmillensis</taxonomicName>
|
||
sp. n., Western Highlands, Mondmill, PNG 77 A male protarsomeres 4-5 in ventral view B median lobe in ventral view C median lobe in lateral view D paramere in external view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="66528CCB316BEA4C0CC097F771F724D0" pageId="3" pageNumber="128">
|
||
<paragraph id="8BFA2B7AF9B46EA6953C3BEE02E8708F" pageId="3" pageNumber="128">
|
||
Figure 12.
|
||
<taxonomicName id="D1BE4B81B5B91F84155CF110548286CF" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina mondmillensis" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="mondmillensis">Exocelina mondmillensis</taxonomicName>
|
||
sp. n., ventral view of median lobe. A, B Western Highlands, Mt. Hagen Town area, PNG 131 C Western Highlands, Gonzsidai-Sarup, PNG 144 D Enga, Anji, PNG 129. Setae not shown.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="8B7B20F52FF8EBDC9ACA969B85124F03" pageId="3" pageNumber="128" type="distribution">
|
||
<paragraph id="56919796E75D7073E682F54F05EB54B6" pageId="3" pageNumber="128">Distribution.</paragraph>
|
||
<paragraph id="5003E983D9A6F422A3F0810F930A744F" pageId="3" pageNumber="128">Papua New Guinea: Madang and Eastern Highlands Provinces (Fig. 14).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |