177 lines
19 KiB
XML
177 lines
19 KiB
XML
<document id="555DBEFB0F63E3E83FC9291D21EA6C5B" ID-CLB-Dataset="30381" ID-DOI="10.11646/zootaxa.4402.3.12" ID-GBIF-Dataset="c5e3ce0c-4b99-4eef-a3b7-28fd57ac1472" ID-ISSN="1175-5326" ID-Zenodo-Dep="1209670" ID-ZooBank="FEC3A7C5-03BD-44B1-B318-AB5B8498FCA1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1522317782500" checkinUser="plazi" docAuthor="Nakládal, Oto & Hlaváč, Peter" docDate="2018" docId="9943875CFFF0FFBF3886DFC9FB98F95D" docLanguage="en" docName="zootaxa.4402.3.12.pdf" docOrigin="Zootaxa 4402 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monodiger ivoricus Nakládal & Hlaváč 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="596" masterDocId="657AFF24FFF1FFBE3811DE13FF9BFFE0" masterDocTitle="New species and records of Clavigeritae (Coleoptera: Staphylinidae: Pselaphinae) from Africa and Asia" masterLastPageNumber="600" masterPageNumber="595" pageNumber="596" updateTime="1698534358692" updateUser="ExternalLinkService">
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<mods:titleInfo id="355BDF94AD7713BE6E181D08D18D81A1">
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<mods:title id="ADD5A009E25C83C81B7196D6E819580D">New species and records of Clavigeritae (Coleoptera: Staphylinidae: Pselaphinae) from Africa and Asia</mods:title>
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<mods:name id="13E3A6992784BF63F1ED83E2E665553F" type="personal">
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<mods:namePart id="F4542200670FE7C98629118E40C9ED03">Nakládal, Oto</mods:namePart>
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<mods:namePart id="77C8731C165DB3B4A1FF7AC11B39F70D">Hlaváč, Peter</mods:namePart>
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<mods:title id="7F95B50DEDA0C16BCC4D5C7EFB470C84">Zootaxa</mods:title>
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<mods:part id="F5152C3458CEF5DC14FE7154D2595278">
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<mods:date id="ACE0EDC79DF0BE1466FFD54232A81F13">2018</mods:date>
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<mods:number id="91E2A022C3176449BE25503BC7A1850B">2018-03-29</mods:number>
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<mods:number id="950CA4A7F24A66EFBF26363DC8FD9EFD">4402</mods:number>
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<mods:identifier id="27F7F1365C5043907449A9AF21C57E30" type="DOI">10.11646/zootaxa.4402.3.12</mods:identifier>
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<treatment id="9943875CFFF0FFBF3886DFC9FB98F95D" ID-DOI="http://doi.org/10.5281/zenodo.5983602" ID-GBIF-Taxon="142669787" ID-Zenodo-Dep="5983602" LSID="urn:lsid:plazi:treatment:9943875CFFF0FFBF3886DFC9FB98F95D" httpUri="http://treatment.plazi.org/id/9943875CFFF0FFBF3886DFC9FB98F95D" lastPageNumber="596" pageId="1" pageNumber="596">
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<subSubSection id="59F065C1FFF0FFBF3886DFC9FE91FDF6" pageId="1" pageNumber="596" type="nomenclature">
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<paragraph id="1155364AFFF0FFBF3886DFC9FE56FE14" blockId="1.[151,461,474,534]" box="[151,461,474,500]" pageId="1" pageNumber="596">
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<heading id="4A1D8126FFF0FFBF3886DFC9FE56FE14" bold="true" box="[151,461,474,500]" fontSize="11" level="1" pageId="1" pageNumber="596" reason="1">
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<emphasis id="239EEA58FFF0FFBF3886DFC9FE56FE14" bold="true" box="[151,461,474,500]" pageId="1" pageNumber="596">
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<taxonomicName id="D6EA4DC9FFF0FFBF3886DFC9FE1FFE14" ID-CoL="446S8" authority="Nakládal & Hlaváč, 2018" authorityName="Nakládal & Hlaváč" authorityYear="2018" box="[151,388,474,500]" class="Insecta" family="Staphylinidae" genus="Monodiger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="ivoricus" status="sp. nov.">
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<emphasis id="239EEA58FFF0FFBF3886DFC9FE1FFE14" bold="true" box="[151,388,474,500]" italics="true" pageId="1" pageNumber="596">Monodiger ivoricus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="38AD5723FFF0FFBF399ADFC9FE56FE14" box="[395,461,474,500]" pageId="1" pageNumber="596" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="1155364AFFF0FFBF3886DFEEFE91FDF6" blockId="1.[151,461,474,534]" box="[151,266,509,534]" pageId="1" pageNumber="596">
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(
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<figureCitation id="89D12ACFFFF0FFBF388EDFEEFE9AFDF6" box="[159,257,509,534]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[311,1281,192,1937]" captionTargetId="figure@3.[307,1281,185,1937]" captionTargetPageId="3" captionText="FIGURES 1–9. Monodiger ivoricus, 1—habitus; 2—aedeagus, dorsal; 3—aedeagus, lateral; Disarthricerus bruneicus, 4— habitus; 5—aedeagus, dorsal; 6—aedeagus, lateral; Fustigerinus formicarius, 7—habitus; 8—aedeagus, dorsal; 9—aedeagus, lateral." httpUri="https://zenodo.org/record/1209672/files/figure.png" pageId="1" pageNumber="596">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="59F065C1FFF0FFBF3886DC5BFE74FD48" pageId="1" pageNumber="596" type="materials_examined">
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<paragraph id="1155364AFFF0FFBF3886DC5BFE74FD48" blockId="1.[151,1437,581,1725]" pageId="1" pageNumber="596">
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<materialsCitation id="A1823C17FFF0FFBF3886DC5BFE77FD48" ID-GBIF-Occurrence="1836936117" collectingDate="2015-07-12" collectingDateMax="2015-07-18" collectingDateMin="2015-07-12" collectionCode="BMNH" collectorName="Ruzzier, E." country="Ivory Coast" county="Moretto" elevation="1171" location="Material" longitude="0.0019444445" municipality="General Collecting Aristophanous" pageId="1" pageNumber="596" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="239EEA58FFF0FFBF3886DC5BFEC5FD81" bold="true" box="[151,350,584,609]" pageId="1" pageNumber="596">
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<location id="14356091FFF0FFBF3886DC5BFF64FD81" LSID="urn:lsid:plazi:treatment:9943875CFFF0FFBF3886DFC9FB98F95D:14356091FFF0FFBF3886DC5BFF64FD81" box="[151,255,584,609]" country="Ivory Coast" county="Moretto" longitude="0.0019444445" municipality="General Collecting Aristophanous" name="Material" pageId="1" pageNumber="596">Material</location>
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studied
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</emphasis>
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:
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<typeStatus id="CE5188E8FFF0FFBF397CDC5BFD9FFD80" box="[365,516,584,608]" pageId="1" pageNumber="596" type="holotype">HOLOTYPE</typeStatus>
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,
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<specimenCount id="07ECFDC3FFF0FFBF3A01DC5AFDB3FD80" box="[528,552,585,608]" pageId="1" pageNumber="596" type="male">♂</specimenCount>
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:
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<collectingCountry id="69FD76DAFFF0FFBF3A25DC5BFD6BFD80" box="[564,752,584,608]" name="Ivory Coast" pageId="1" pageNumber="596">IVORY COAST</collectingCountry>
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,
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<elevation id="9AC7D179FFF0FFBF3AEADC5BFCC9FD80" box="[763,850,584,608]" metricMagnitude="3" metricUnit="m" metricValue="1.1709999999999998" pageId="1" pageNumber="596" unit="m" value="1171.0">
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<quantity id="D6129BAFFFF0FFBF3AEADC5BFCC9FD80" box="[763,850,584,608]" metricMagnitude="3" metricUnit="m" metricValue="1.1709999999999998" pageId="1" pageNumber="596" unit="m" value="1171.0">1171 m</quantity>
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</elevation>
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,
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||
<location id="14356091FFF0FFBF3B4FDC5AFBB0FD81" LSID="urn:lsid:plazi:treatment:9943875CFFF0FFBF3886DFC9FB98F95D:14356091FFF0FFBF3B4FDC5AFBB0FD81" box="[862,1067,584,609]" country="Ivory Coast" county="Moretto" longitude="0.0019444445" municipality="General Collecting Aristophanous" name="Mt. Tonkoui Peak" pageId="1" pageNumber="596">Mt. Tonkoui Peak</location>
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,
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<geoCoordinate id="74DE508DFFF0FFBF3C26DC5BFBC1FDB3" box="[1079,1114,581,608]" direction="east" orientation="longitude" pageId="1" pageNumber="596" precision="925" value="0.11666667">0 7o</geoCoordinate>
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27’15.2’’N,
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<geoCoordinate id="74DE508DFFF0FFBF3CF5DC5BFA8EFDB3" box="[1252,1301,581,608]" direction="east" orientation="longitude" pageId="1" pageNumber="596" precision="15" value="0.0019444445">0 0 7o</geoCoordinate>
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38’12.5’’W,
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<collectingDate id="7510E962FFF0FFBF3886DC7FFED2FD65" box="[151,329,620,645]" pageId="1" pageNumber="596" value="2015-07-12" valueMax="2015-07-18" valueMin="2015-07-12">
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<date id="6554108AFFF0FFBF3886DC7FFED2FD65" box="[151,329,620,645]" pageId="1" pageNumber="596" value="2015-07-12" valueMax="2015-07-18" valueMin="2015-07-12">12-18.VII.2015</date>
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||
</collectingDate>
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||
.
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<collectingMunicipality id="F131AC30FFF0FFBF3947DC7FFD44FD65" box="[342,735,620,645]" pageId="1" pageNumber="596">General Collecting Aristophanous</collectingMunicipality>
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, M.,
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<collectingCounty id="F8344EC6FFF0FFBF3B0FDC7EFCE0FD64" box="[798,891,621,644]" pageId="1" pageNumber="596">Moretto</collectingCounty>
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, P.,
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<collectorName id="BC1F539CFFF0FFBF3BBFDC7FFBB6FD64" box="[942,1069,620,645]" pageId="1" pageNumber="596">Ruzzier, E.</collectorName>
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leg. /
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<collectionCode id="77FBAE8FFFF0FFBF3C6DDC7EFB4BFD64" box="[1148,1232,621,644]" pageId="1" pageNumber="596">BMNH</collectionCode>
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{E}, 2015-177. /
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<collectionCode id="77FBAE8FFFF0FFBF3886DC82FF70FD48" box="[151,235,657,680]" pageId="1" pageNumber="596">BMNH</collectionCode>
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(E), 1460885.
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<collectionCode id="77FBAE8FFFF0FFBF3985DC82FE77FD48" box="[404,492,657,680]" pageId="1" pageNumber="596">BMNH</collectionCode>
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</materialsCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59F065C1FFF0FFBF38D6DCA7FD09FA21" pageId="1" pageNumber="596" type="description">
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<paragraph id="1155364AFFF0FFBF38D6DCA7FE35FCF4" blockId="1.[151,1437,581,1725]" pageId="1" pageNumber="596">
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||
<emphasis id="239EEA58FFF0FFBF38D6DCA7FECAFD2D" bold="true" box="[199,337,692,717]" pageId="1" pageNumber="596">Description</emphasis>
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: Body (
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||
<figureCitation id="89D12ACFFFF0FFBF39BCDCA7FE75FD2C" box="[429,494,692,717]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[311,1281,192,1937]" captionTargetId="figure@3.[307,1281,185,1937]" captionTargetPageId="3" captionText="FIGURES 1–9. Monodiger ivoricus, 1—habitus; 2—aedeagus, dorsal; 3—aedeagus, lateral; Disarthricerus bruneicus, 4— habitus; 5—aedeagus, dorsal; 6—aedeagus, lateral; Fustigerinus formicarius, 7—habitus; 8—aedeagus, dorsal; 9—aedeagus, lateral." httpUri="https://zenodo.org/record/1209672/files/figure.png" pageId="1" pageNumber="596">Fig. 1</figureCitation>
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) entirely reddish-brown, head, pronotum, abdomen and antennae darker, elytra and legs more reddish, head, pronotum and elytra rugose, composite tergite smooth, shiny. Length
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<quantity id="D6129BAFFFF0FFBF3CA8DCCBFA84FD10" box="[1209,1311,728,753]" metricMagnitude="-3" metricUnit="m" metricValue="2.36" pageId="1" pageNumber="596" unit="mm" value="2.36">2.36 mm</quantity>
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, maximum width of elytra
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<quantity id="D6129BAFFFF0FFBF3952DCEFFE31FCF4" box="[323,426,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="596" unit="mm" value="1.2">1.20 mm</quantity>
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.
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</paragraph>
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<paragraph id="1155364AFFF0FFBF38D6DD33FBBDFC61" blockId="1.[151,1437,581,1725]" pageId="1" pageNumber="596">Head widest across eyes, with elevated carinae in anterior half, 1.5 as long as wide, with lateraly projecting genae, temples about twice as long as diameter of eyes, frontal and vertexal fovea absent, rostrum strongly projecting anteriad, projecting process not pointed. Clypeus well-visible dorsally.</paragraph>
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<paragraph id="1155364AFFF0FFBF38D6DD9FFB79FC0C" blockId="1.[151,1437,581,1725]" pageId="1" pageNumber="596">Antennae short, with three antennomeres, scape minuscule, not visible dorsally, pedicel also small, partly visible dorsally, terminal antennomeres long, slightly curved, about 6.8 times as long as pedicel and about 3.8 times as long as wide in middle, at apex oval in cross section, with 4 ribs and 4 sulcus ending near apex.</paragraph>
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<paragraph id="1155364AFFF0FFBF38D6DDEAFD4FFBD5" blockId="1.[151,1437,581,1725]" pageId="1" pageNumber="596">Pronotum pentagonal with well-defined anterior corners, 1.3 times as wide as long, with well-defined, wide entire median sulcus, surface of sulcus shagreened.</paragraph>
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<paragraph id="1155364AFFF0FFBF38D6DA53FAE1FB41" blockId="1.[151,1437,581,1725]" pageId="1" pageNumber="596">Elytra twice as wide as long, 1.6 times as long as pronotum, lacking basal foveae, with two entire discal carinae, discal carinae oblique, humeri with well-defined, prominent tooth. Elytral posterior edge with small elytral humps located close to posterior corners, elytral trichomes short attached to elytral humps from their inner side.</paragraph>
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<paragraph id="1155364AFFF0FFBF38D6DABFFD6FFAED" blockId="1.[151,1437,581,1725]" pageId="1" pageNumber="596">Abdomen 1.25 times as wide as long, 1.4 times as long as elytra, composite tergite (IV–VI) with fine puncturation, shiny, with large basal depression ending by elevated short elongate carinae, with three visible paratergites, first paratergites bearing large trichomes.</paragraph>
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<paragraph id="1155364AFFF0FFBF38D6DB0AFC4BFAD0" blockId="1.[151,1437,581,1725]" box="[199,976,1304,1329]" pageId="1" pageNumber="596">Legs short and slender, lacking sexual characters like spines or spurs.</paragraph>
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<paragraph id="1155364AFFF0FFBF38D6DB2FFB17FA7D" blockId="1.[151,1437,581,1725]" pageId="1" pageNumber="596">
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Aedeagus (
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<figureCitation id="89D12ACFFFF0FFBF3956DB2FFE35FAB4" box="[327,430,1340,1365]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[311,1281,192,1937]" captionTargetId="figure@3.[307,1281,185,1937]" captionTargetPageId="3" captionText="FIGURES 1–9. Monodiger ivoricus, 1—habitus; 2—aedeagus, dorsal; 3—aedeagus, lateral; Disarthricerus bruneicus, 4— habitus; 5—aedeagus, dorsal; 6—aedeagus, lateral; Fustigerinus formicarius, 7—habitus; 8—aedeagus, dorsal; 9—aedeagus, lateral." httpUri="https://zenodo.org/record/1209672/files/figure.png" pageId="1" pageNumber="596">Figs 2, 3</figureCitation>
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) about
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||
<quantity id="D6129BAFFFF0FFBF3A19DB2FFDF4FAB4" box="[520,623,1340,1364]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="1" pageNumber="596" unit="mm" value="0.49">0.49 mm</quantity>
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long and
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<quantity id="D6129BAFFFF0FFBF3AF4DB2FFCD7FAB4" box="[741,844,1340,1364]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="1" pageNumber="596" unit="mm" value="0.18">0.18 mm</quantity>
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||
wide, about 2,70 times as long as wide, basal bulb large, apical lobe about as long as basal bulb, slender and tapered to apex, at apex slightly truncate, ventrolateral foramen strongly projecting, phallobase diaphragm oval, about 1.6 of length of basal bulb.
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</paragraph>
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||
<paragraph id="1155364AFFF0FFBF38D6DBBBFD09FA21" blockId="1.[151,1437,581,1725]" box="[199,658,1448,1473]" pageId="1" pageNumber="596">
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||
<emphasis id="239EEA58FFF0FFBF38D6DBBBFE2CFA21" bold="true" box="[199,439,1448,1473]" pageId="1" pageNumber="596">Sexual dimorphism.</emphasis>
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Females unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="59F065C1FFF0FFBF38D6DBDFFC3BF9B0" pageId="1" pageNumber="596" type="diagnosis">
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<paragraph id="1155364AFFF0FFBF38D6DBDFFC3BF9B0" blockId="1.[151,1437,581,1725]" pageId="1" pageNumber="596">
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<emphasis id="239EEA58FFF0FFBF38D6DBDFFE4BFA05" bold="true" box="[199,464,1484,1509]" pageId="1" pageNumber="596">Differential diagnosis:</emphasis>
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<taxonomicName id="D6EA4DC9FFF0FFBF39C9DBDFFD2CFA04" authorityName="Nakládal & Hlaváč" authorityYear="2018" box="[472,695,1484,1509]" class="Insecta" family="Staphylinidae" genus="Monodiger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="ivoricus">
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<emphasis id="239EEA58FFF0FFBF39C9DBDFFD2CFA04" box="[472,695,1484,1509]" italics="true" pageId="1" pageNumber="596">Monodiger ivoricus</emphasis>
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</taxonomicName>
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differs from its congener
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<taxonomicName id="D6EA4DC9FFF0FFBF3BCEDBDEFA9EFA04" authority="Paulian, 1948" authorityName="Paulian" authorityYear="1948" box="[991,1285,1484,1509]" class="Insecta" family="Staphylinidae" genus="Monodiger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="sulcatus">
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<emphasis id="239EEA58FFF0FFBF3BCEDBDEFBFAFA05" box="[991,1121,1484,1509]" italics="true" pageId="1" pageNumber="596">M. sulcatus</emphasis>
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Paulian, 1948
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</taxonomicName>
|
||
by having 1) rostrum strongly projecting anteriad, projecting process not pointed, 2) larger eyes, temples only about twice as long as longitudinal diameter of eyes and 3) anterior corner of pronotum less projecting anteriad, corners below level of its anterior margin (for the comparison see Célis, 1969: 419).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59F065C1FFF0FFBF38D6D84FFE3DF995" box="[199,422,1628,1653]" pageId="1" pageNumber="596" type="biology_ecology">
|
||
<paragraph id="1155364AFFF0FFBF38D6D84FFE3DF995" blockId="1.[151,1437,581,1725]" box="[199,422,1628,1653]" pageId="1" pageNumber="596">
|
||
<emphasis id="239EEA58FFF0FFBF38D6D84FFEBAF995" bold="true" box="[199,289,1628,1653]" pageId="1" pageNumber="596">Biology</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59F065C1FFF0FFBF38D6D893FE68F978" box="[199,499,1664,1689]" pageId="1" pageNumber="596" type="distribution">
|
||
<paragraph id="1155364AFFF0FFBF38D6D893FE68F978" blockId="1.[151,1437,581,1725]" box="[199,499,1664,1689]" pageId="1" pageNumber="596">
|
||
<emphasis id="239EEA58FFF0FFBF38D6D893FEC2F979" bold="true" box="[199,345,1664,1689]" pageId="1" pageNumber="596">Distribution</emphasis>
|
||
:
|
||
<collectingCountry id="69FD76DAFFF0FFBF3979D892FE74F978" box="[360,495,1664,1689]" name="Ivory Coast" pageId="1" pageNumber="596">Ivory Coast</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59F065C1FFF0FFBF38D6D8B7FB98F95D" box="[199,1027,1700,1725]" pageId="1" pageNumber="596" type="etymology">
|
||
<paragraph id="1155364AFFF0FFBF38D6D8B7FB98F95D" blockId="1.[151,1437,581,1725]" box="[199,1027,1700,1725]" pageId="1" pageNumber="596">
|
||
<emphasis id="239EEA58FFF0FFBF38D6D8B7FED3F95D" bold="true" box="[199,328,1700,1725]" pageId="1" pageNumber="596">Etymology</emphasis>
|
||
: Locotypic, named after
|
||
<collectingCountry id="69FD76DAFFF0FFBF3A73D8B6FD72F95C" box="[610,745,1700,1725]" name="Ivory Coast" pageId="1" pageNumber="596">Ivory Coast</collectingCountry>
|
||
, country of its discovery.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |