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<mods:title id="62CA3ACB93CF94DED88ED77EFD7CF37D">Note on Psila helvetica Shatalkin &amp; Merz, 2010: a new rust fly (Diptera: Psilidae) for mainland France, with key to French species and male genitalia illustration</mods:title>
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Replacement for
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identification key provided by
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<emphasis id="D6B6EA83E47BDD50FF4A09F8FE9EE2BB" bold="true" box="[151,327,225,249]" pageId="2" pageNumber="299">English version</emphasis>
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<keyLead id="5F3395A4E47BDD50FF4A0819FA45E371" pageId="2" pageNumber="299">1 Mesonotum and subscutellum yellow. Postpedicel mainly yellow (sometimes with a blackened area at the base of the arista). Aedeagus shorter than the genitalia, not protruding below the sternites........................................... 2</keyLead>
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- Mesonotum with black stripes and subscutellum black. Postpedicel mostly black. Aedeagus longer than the genitalia (
<figureCitation id="7CF92A14E47BDD50FA880822FA56E313" box="[1365,1423,315,337]" captionStart="FIGURES 56" captionStartId="2.[151,264,1960,1984]" captionTargetBox="[199,1389,508,1936]" captionTargetId="figure-372@2.[199,1389,508,1936]" captionTargetPageId="2" captionText="FIGURES 56. Genitalia of Psila helvetica Shatalkin &amp; Merz, 2010. 5: in ventral view. Scale bar = 0.3 mm. 6: left epandrium in lateral view. Scale bars = 0.1 mm. (Drawings: Jocelyn Claude)." figureDoi="http://doi.org/10.5281/zenodo.7091707" httpUri="https://zenodo.org/record/7091707/files/figure.png" pageId="2" pageNumber="299">Fig. 5</figureCitation>
), and often protruding below the sternites (
<figureCitation id="7CF92A14E47BDD50FDB30841FD7EE32F" box="[622,679,344,366]" captionStart="FIGURES 24" captionStartId="1.[151,264,1971,1995]" captionTargetBox="[220,1386,956,1948]" captionTargetId="figure-428@1.[201,1386,956,1948]" captionTargetPageId="1" captionText="FIGURES 24. Habitus of Psila helvetica Shatalkin &amp; Merz, 2010 in dorsal view with aedeagus indicated by arrow (2), ventral view (3) and head in front (4). (Photos: Christophe Lauriaut)." figureDoi="http://doi.org/10.5281/zenodo.7091705" httpUri="https://zenodo.org/record/7091705/files/figure.png" pageId="2" pageNumber="299">Fig. 3</figureCitation>
).............................
<taxonomicName id="23C24D12E47BDD50FBFD0841FA45E32F" ID-CoL="8DJJV" authority="Shatalkin &amp; Merz, 2010" authorityName="Shatalkin &amp; Merz" authorityYear="2010" box="[1056,1436,344,366]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="helvetica">
<emphasis id="D6B6EA83E47BDD50FBFD0841FB76E32F" box="[1056,1199,344,365]" italics="true" pageId="2" pageNumber="299">Psila helvetica</emphasis>
<bibRefCitation id="80534B60E47BDD50FB690841FA45E32F" author="Shatalkin, A. I. &amp; Merz, B." box="[1204,1436,344,366]" pageId="2" pageNumber="299" pagination="771 - 800" refId="ref2102" refString="Shatalkin, A. I. &amp; Merz, B. (2010) The Psilidae (Diptera, Acalyptrata) of Switzerland, with description of two new species from Central Europe. Revue Suisse de Zoologie, 117 (4), 771 - 800. https: // doi. org / 10.5962 / bhl. part. 117602" type="journal article" year="2010">Shatalkin &amp; Merz, 2010</bibRefCitation>
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2 Postpedicel yellow-orange, blackened at the base of the arista. Genitalia as in Fig.
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...................................................................................
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(Linnaeus, 1761)
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- Postpedicel unicolourous (yellow-orange), without black spot at the base of the arista. Genitalia as in Fig.
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.................................................................
<taxonomicName id="23C24D12E47BDD50FB5508D4FA40E3A1" ID-CoL="4PG7W" authority="Collin, 1944" authorityName="Collin" authorityYear="1944" box="[1160,1433,461,483]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="merdaria">
<emphasis id="D6B6EA83E47BDD50FB5508D4FAC1E3A0" box="[1160,1304,461,482]" italics="true" pageId="2" pageNumber="299">Psila merdaria</emphasis>
Collin, 1944
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.
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<emphasis id="D6B6EA83E47BDD50FF4A0EB1FEE3E582" bold="true" box="[151,314,1960,1984]" pageId="2" pageNumber="299">FIGURES 56.</emphasis>
Genitalia of
<taxonomicName id="23C24D12E47BDD50FE620EB1FC8AE582" authority="Shatalkin &amp; Merz, 2010" authorityName="Shatalkin &amp; Merz" authorityYear="2010" box="[447,851,1960,1984]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="helvetica">
<emphasis id="D6B6EA83E47BDD50FE620EB1FD8DE582" box="[447,596,1960,1984]" italics="true" pageId="2" pageNumber="299">Psila helvetica</emphasis>
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.
<emphasis id="D6B6EA83E47BDD50FC810EB1FCABE582" bold="true" box="[860,882,1960,1984]" pageId="2" pageNumber="299">5:</emphasis>
in ventral view. Scale bar = 0.3 mm.
<emphasis id="D6B6EA83E47BDD50FB300EB1FADBE582" bold="true" box="[1261,1282,1960,1984]" pageId="2" pageNumber="299">6:</emphasis>
left epandrium in lateral view. Scale bars = 0.1 mm. (Drawings: Jocelyn Claude).
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<emphasis id="D6B6EA83E47ADD51FF4A098EFE9BE2ED" bold="true" box="[151,322,151,175]" pageId="3" pageNumber="300">French version</emphasis>
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<keyLead id="5F3395A4E47ADD51FF4A09AFFA45E2AB" pageId="3" pageNumber="300">1 Mésonotum jaune et subscutellum jaune. Postpedicel principalement jaune (parfois avec une zone noircie à la base de larista). Lédeage est plus court que le genitalia, et ne dépasse pas sous les sternites........................................ 2</keyLead>
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- Mésonotum avec des rayures noires et subscutellum noir. Postpedicel principalement noir. Lédeage est plus long que les genitalia (
<figureCitation id="7CF92A14E47ADD51FE9F0817FEA5E366" box="[322,380,270,292]" captionStart="FIGURES 56" captionStartId="2.[151,264,1960,1984]" captionTargetBox="[199,1389,508,1936]" captionTargetId="figure-372@2.[199,1389,508,1936]" captionTargetPageId="2" captionText="FIGURES 56. Genitalia of Psila helvetica Shatalkin &amp; Merz, 2010. 5: in ventral view. Scale bar = 0.3 mm. 6: left epandrium in lateral view. Scale bars = 0.1 mm. (Drawings: Jocelyn Claude)." figureDoi="http://doi.org/10.5281/zenodo.7091707" httpUri="https://zenodo.org/record/7091707/files/figure.png" pageId="3" pageNumber="300">Fig. 5</figureCitation>
), et souvent dépasse sous les sternites (
<figureCitation id="7CF92A14E47ADD51FD2B0817FCF7E361" box="[758,814,270,292]" captionStart="FIGURES 24" captionStartId="1.[151,264,1971,1995]" captionTargetBox="[220,1386,956,1948]" captionTargetId="figure-428@1.[201,1386,956,1948]" captionTargetPageId="1" captionText="FIGURES 24. Habitus of Psila helvetica Shatalkin &amp; Merz, 2010 in dorsal view with aedeagus indicated by arrow (2), ventral view (3) and head in front (4). (Photos: Christophe Lauriaut)." figureDoi="http://doi.org/10.5281/zenodo.7091705" httpUri="https://zenodo.org/record/7091705/files/figure.png" pageId="3" pageNumber="300">Fig. 3</figureCitation>
)..................
<taxonomicName id="23C24D12E47ADD51FBFD0817FA45E361" ID-CoL="8DJJV" authority="Shatalkin &amp; Merz, 2010" authorityName="Shatalkin &amp; Merz" authorityYear="2010" box="[1056,1436,270,292]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="helvetica">
<emphasis id="D6B6EA83E47ADD51FBFD0817FB76E361" box="[1056,1199,270,291]" italics="true" pageId="3" pageNumber="300">Psila helvetica</emphasis>
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</paragraph>
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<keyStep id="5F362E34E47ADD51FF4A0835FA44E3DB" pageId="3" pageNumber="300">
<paragraph id="E47D3691E47ADD51FF4A0835FA45E31D" blockId="3.[151,1437,151,410]" pageId="3" pageNumber="300">
<keyLead id="5F3395A4E47ADD51FF4A0835FA45E31D" pageId="3" pageNumber="300">
2 Postpedicel jaune-orange, noirci à la base de larista. Genitalia comme dans la Fig. 43 dans
<bibRefCitation id="80534B60E47ADD51FB8C0835FA9CE303" author="Withers, P. &amp; Claude, J." box="[1105,1349,300,322]" pageId="3" pageNumber="300" pagination="115 - 135" refId="ref2219" refString="Withers, P. &amp; Claude, J. (2021) Les Psilidae de France (Diptera: Acalyptrata): checklist et cles de determination des genres et des especes. Naturae, 9, 115 - 135. https: // doi. org / 10.5852 / naturae 2021 a 9" type="journal article" year="2021">Withers &amp; Claude (2021)</bibRefCitation>
................................................................................
<taxonomicName id="23C24D12E47ADD51FBBE0850FA45E31D" ID-CoL="4PG6P" authority="(Linnaeus, 1761)" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" box="[1123,1436,329,351]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="fimetaria">
<emphasis id="D6B6EA83E47ADD51FBBE0850FB2BE31C" box="[1123,1266,329,350]" italics="true" pageId="3" pageNumber="300">Psila fimetaria</emphasis>
(Linnaeus, 1761)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="E47D3691E47ADD51FF4A087EFA44E3DB" blockId="3.[151,1437,151,410]" pageId="3" pageNumber="300">
<keyLead id="5F3395A4E47ADD51FF4A087EFA44E3DB" pageId="3" pageNumber="300">
- Postpedicel unicolore (jaune-orange), sans tache noire à la base de larista. Genitalia comme dans la Fig. 44 dans
<bibRefCitation id="80534B60E47ADD51FAEA087FFEB5E3DB" author="Withers, P. &amp; Claude, J." pageId="3" pageNumber="300" pagination="115 - 135" refId="ref2219" refString="Withers, P. &amp; Claude, J. (2021) Les Psilidae de France (Diptera: Acalyptrata): checklist et cles de determination des genres et des especes. Naturae, 9, 115 - 135. https: // doi. org / 10.5852 / naturae 2021 a 9" type="journal article" year="2021">Withers &amp; Claude (2021)</bibRefCitation>
.................................................................
<taxonomicName id="23C24D12E47ADD51FB55089DFA40E3DB" ID-CoL="4PG7W" authority="Collin, 1944" authorityName="Collin" authorityYear="1944" box="[1160,1433,388,410]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="merdaria">
<emphasis id="D6B6EA83E47ADD51FB55089DFAC1E3DB" box="[1160,1304,388,409]" italics="true" pageId="3" pageNumber="300">Psila merdaria</emphasis>
Collin, 1944
</taxonomicName>
.
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="ACD8651AE47ADD51FF4A08D3FD05E0D4" pageId="3" pageNumber="300" type="discussion">
<paragraph id="E47D3691E47ADD51FF4A08D3FD05E0D4" blockId="3.[151,1437,458,662]" pageId="3" pageNumber="300">
Specimens caught in
<collectingCountry id="9CD57601E47ADD51FEAB08D3FE66E3A3" box="[374,447,458,481]" name="France" pageId="3" pageNumber="300">France</collectingCountry>
confirm the late flight period (mid-August to mid-September) of
<taxonomicName id="23C24D12E47ADD51FBB108D3FAD1E3A0" authorityName="Shatalkin &amp; Merz" authorityYear="2010" box="[1132,1288,458,482]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="helvetica">
<emphasis id="D6B6EA83E47ADD51FBB108D3FAD1E3A0" box="[1132,1288,458,482]" italics="true" pageId="3" pageNumber="300">Psila helvetica</emphasis>
</taxonomicName>
. The altitudes are also concordant (
<quantity id="233A9B74E47ADD51FEAA08F7FDCEE047" box="[375,535,494,518]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="12.0" metricValueMin="6.5" pageId="3" pageNumber="300" unit="m" value="925.0" valueMax="1200.0" valueMin="650.0">650 to 1200 m</quantity>
a.s.l.). Examination of the genitalia confirms that
<taxonomicName id="23C24D12E47ADD51FBEA08F7FB0CE044" authorityName="Shatalkin &amp; Merz" authorityYear="2010" box="[1079,1237,494,518]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="helvetica">
<emphasis id="D6B6EA83E47ADD51FBEA08F7FB0CE044" box="[1079,1237,494,518]" italics="true" pageId="3" pageNumber="300">Psila helvetica</emphasis>
</taxonomicName>
differs from other European species of
<taxonomicName id="23C24D12E47ADD51FEA80B0BFE75E068" authorityName="Meigen" authorityYear="1803" box="[373,428,530,554]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="genus">
<emphasis id="D6B6EA83E47ADD51FEA80B0BFE75E068" box="[373,428,530,554]" italics="true" pageId="3" pageNumber="300">Psila</emphasis>
</taxonomicName>
. After
<collectingCountry id="9CD57601E47ADD51FE2B0B0BFDACE068" box="[502,629,530,554]" name="Switzerland" pageId="3" pageNumber="300">Switzerland</collectingCountry>
(
<bibRefCitation id="80534B60E47ADD51FD590B0BFC5BE068" author="Shatalkin, A. I. &amp; Merz, B." box="[644,898,530,554]" pageId="3" pageNumber="300" pagination="771 - 800" refId="ref2102" refString="Shatalkin, A. I. &amp; Merz, B. (2010) The Psilidae (Diptera, Acalyptrata) of Switzerland, with description of two new species from Central Europe. Revue Suisse de Zoologie, 117 (4), 771 - 800. https: // doi. org / 10.5962 / bhl. part. 117602" type="journal article" year="2010">Shatalkin &amp; Merz 2010</bibRefCitation>
) and
<collectingCountry id="9CD57601E47ADD51FC630B0BFBD6E068" box="[958,1039,530,554]" name="Finland" pageId="3" pageNumber="300">Finland</collectingCountry>
(
<bibRefCitation id="80534B60E47ADD51FBC00B0BFA98E068" author="Kahanpaa, J. &amp; Salmela, J." box="[1053,1345,530,554]" pageId="3" pageNumber="300" pagination="1 - 20" refId="ref2063" refString="Kahanpaa, J. &amp; Salmela, J. (2014) Checklist of the Diptera of Finland. Zookeys, 441, 1 - 20. https: // doi. org / 10.3897 / zookeys. 441.7620" type="journal article" year="2014">Kahanpää &amp; Salmela 2014</bibRefCitation>
),
<collectingCountry id="9CD57601E47ADD51FA890B0BFA44E06B" box="[1364,1437,530,553]" name="France" pageId="3" pageNumber="300">France</collectingCountry>
is the third country known for
<taxonomicName id="23C24D12E47ADD51FE020B2FFDA4E00C" authorityName="Shatalkin &amp; Merz" authorityYear="2010" box="[479,637,566,590]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="helvetica">
<emphasis id="D6B6EA83E47ADD51FE020B2FFDA4E00C" box="[479,637,566,590]" italics="true" pageId="3" pageNumber="300">Psila helvetica</emphasis>
</taxonomicName>
. The list of
<taxonomicName id="23C24D12E47ADD51FCDD0B2FFC8DE00C" box="[768,852,566,590]" class="Insecta" family="Psilidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="family">Psilidae</taxonomicName>
from
<collectingCountry id="9CD57601E47ADD51FC4B0B2FFC06E00F" box="[918,991,566,589]" name="France" pageId="3" pageNumber="300">France</collectingCountry>
now stands at 38 species, including three
<taxonomicName id="23C24D12E47ADD51FF4A0B43FF2AE033" box="[151,243,602,626]" class="Insecta" family="Psilidae" genus="Psila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D6B6EA83E47ADD51FF4A0B43FF17E030" box="[151,206,602,626]" italics="true" pageId="3" pageNumber="300">Psila</emphasis>
sp.
</taxonomicName>
However, knowledge of the French fauna remains modest, with other species to be described in the near future, in particular for the genus
<taxonomicName id="23C24D12E47ADD51FE6C0B67FD01E0D4" authority="Hendel, 1917" authorityName="Hendel" authorityYear="1917" box="[433,728,638,662]" class="Insecta" family="Psilidae" genus="Chamaepsila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="genus">
<emphasis id="D6B6EA83E47ADD51FE6C0B67FDE7E0D4" box="[433,574,638,662]" italics="true" pageId="3" pageNumber="300">Chamaepsila</emphasis>
Hendel, 1917
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>