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<document ID-CLB-Dataset="2474" ID-DOI="http://dx.doi.org/10.3897/afrinvertebr.63.76934" ID-GBIF-Dataset="4ec9e932-443a-46e3-a337-cf6ab4cb368d" ID-Pensoft-Pub="2305-2562-1-1" ID-Pensoft-UUID="A8A8A48233C35D628C500C4B33DD7260" ID-ZooBank="CB17C6B2CF804B11A100E340E5CB66A2" ModsDocID="2305-2562-63-1-1" checkinTime="1642775392551" checkinUser="pensoft" docAuthor="Bossert, Silas &amp; Noort, Simon van" docDate="2022" docId="417FB9D69E2B5D2A8029E05F19AE61B2" docLanguage="en" docName="AfrInvert 63(1): 1-18" docOrigin="African Invertebrates 63 (1)" docPubDate="2022-01-21" docSource="http://dx.doi.org/10.3897/afrinvertebr.63.76934" docTitle="Scrapter heterodoxus" docType="treatment" docVersion="5" id="A8A8A48233C35D628C500C4B33DD7260" lastPageNumber="1" masterDocId="A8A8A48233C35D628C500C4B33DD7260" masterDocTitle="A group of two: Scrapter peringueyi is not a synonym of Scrapter heterodoxus (Hymenoptera, Colletidae, Scraptrinae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1732938959083" updateUser="ExternalLinkService">
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<mods:title id="6E786287608A7990B53117CCEA4B0857">A group of two: Scrapter peringueyi is not a synonym of Scrapter heterodoxus (Hymenoptera, Colletidae, Scraptrinae)</mods:title>
</mods:titleInfo>
<mods:name id="EBC8DB75744CA1ADA10C5FF03229DA45" type="personal">
<mods:role id="CA7BDDFBDE721FE10BFE61A5DA076580">
<mods:roleTerm id="616AE8628FCC5D1C9C7AE4422529CBEC">Author</mods:roleTerm>
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<mods:namePart id="3A4C76EFA22FEFE4A04546040CA75227">Bossert, Silas</mods:namePart>
<mods:nameIdentifier id="7143105AF86FB2A12C0532FFAB75A943" type="ORCID">https://orcid.org/0000-0002-3620-5468</mods:nameIdentifier>
<mods:affiliation id="E255830394A8E19988B85971C17A7215">Department of Entomology, Washington State University, Pullman, WA, 99164, USA &amp; Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA</mods:affiliation>
<mods:nameIdentifier id="079E5ECF8D9A4163F483A5A4AA19F5FC" type="email">sb2346@cornell.edu</mods:nameIdentifier>
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<mods:namePart id="5F73ABA4D7E6090FD749A27F668B4D98">Noort, Simon van</mods:namePart>
<mods:nameIdentifier id="03B6704C568D712FDAEF37CD17430E14" type="ORCID">https://orcid.org/0000-0001-6930-9741</mods:nameIdentifier>
<mods:affiliation id="9976E35BEFF9F11C701BFFB93B1C4628">Research and Exhibitions Department, South African Museum, Iziko Museums of South Africa, Cape Town, 8000, South Africa &amp; Department of Biological Sciences, University of Cape Town, Private Bag, Rondebosch, 7701, South Africa</mods:affiliation>
</mods:name>
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<mods:titleInfo id="C0136C24956898CD2637A4C8C93A5AF6">
<mods:title id="DF809F5318F9A45D51094563246F39C6">African Invertebrates</mods:title>
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<mods:date id="0CE2983F3E87BF5F9F6A1A9FA04DA65B">2022</mods:date>
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<mods:classification id="89199BFEF0928B3B17E2CE2004358E9B">journal article</mods:classification>
<mods:identifier id="6B5C3C01C33B083BBBEFEEAF34456832" type="DOI">http://dx.doi.org/10.3897/afrinvertebr.63.76934</mods:identifier>
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<treatment id="417FB9D69E2B5D2A8029E05F19AE61B2" ID-GBIF-Taxon="192496488" LSID="urn:lsid:plazi:treatment:417FB9D69E2B5D2A8029E05F19AE61B2" httpUri="http://treatment.plazi.org/id/417FB9D69E2B5D2A8029E05F19AE61B2" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Colletidae" scope_order="Hymenoptera">
<subSubSection id="E383FD11A31094812EE9A09356B44869" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="FF4C7144C5D9BD3E4E86A993EF00010F" pageId="0" pageNumber="1">
<taxonomicName id="9A7BA47C1FF674A2275B2AC219269F29" ID-CoL="4VYL5" LSID="417FB9D6-9E2B-5D2A-8029-E05F19AE61B2" authority="(Cockerell, 1921)" baseAuthorityName="Cockerell" baseAuthorityYear="1921" class="Insecta" family="Colletidae" genus="Scrapter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scrapter heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">Scrapter heterodoxus (Cockerell, 1921)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="632BF4C840BED3DCFA6D1B37696AD5E1" pageId="0" pageNumber="1" type="description">
<paragraph id="C17DE59D0246730ACD6A885D0517C635" pageId="0" pageNumber="1">
<figureCitation id="2746E4AC72A03044941210454F57CD38" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male lectotype of Scrapter heterodoxus (Cockerell, 1921). A habitus, dorsolateral view, and labels B habitus, lateral view C head, frontal view D propodeum E hind leg tibia F fore wing." figureDoi="10.3897/afrinvertebr.63.76934.figure1" httpUri="https://binary.pensoft.net/fig/636741" pageId="0" pageNumber="1">Figures 1</figureCitation>
<figureCitation id="519960AB685CA23535CD8EFE1767A940" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Female non-type specimen of Scrapter heterodoxus (Cockerell, 1921), (SAM-HYM-B 007786). A habitus, dorsal view B habitus, lateral view C head, frontal view D propodeum E labels F fore wing." figureDoi="10.3897/afrinvertebr.63.76934.figure2" httpUri="https://binary.pensoft.net/fig/636742" pageId="0" pageNumber="1">, 2</figureCitation>
<figureCitation id="18354F5386030C4E85C29A96BB5F0B65" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of the shape of the male hind leg tibia and propodeal sculpture. A Scrapter peringueyi (Cockerell, 1921), stat. rev. hind leg tibial apex, non-type male specimen (deposited in CUIC) B Scrapter heterodoxus (Cockerell, 1921) hind leg tibial apex, non-type male specimen (deposited in CUIC) C Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, non-type male specimen (SAM-HYM-B 007139) D Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, lectotype male (SAM-HYM-B 000145) E Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, holotype female (SAM-HYM-B 000144) F Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, non-type female (SAM-HYM-B 007786)." figureDoi="10.3897/afrinvertebr.63.76934.figure5" httpUri="https://binary.pensoft.net/fig/636745" pageId="0" pageNumber="1">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="1D672DF9D26E6024FF8A459B66306C9F" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="3E0103D08F8ABF14177249ECB053738A" pageId="0" pageNumber="1">
<taxonomicName id="B1BC6791551AC693919F6BD3D413FD18" authorityName="Cockerell" authorityYear="1921" class="Insecta" family="Colletidae" genus="Polyglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyglossa heterodoxa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxa">Polyglossa heterodoxa</taxonomicName>
Cockerell, 1921: 204.
</paragraph>
</subSubSection>
<subSubSection id="FF2BFD243D65A7A41C40CDC5B737E273" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="17EB7858CADE48FBD271D69EFC384156" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="4C10FFCE71177F43C4F8F6658AD27BBD" pageId="0" pageNumber="1">
<materialsCitation id="AA4FF49D22A1559177064418CDB832AE" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="F. Foly" country="South Africa" location="Cape" specimenCount="♂" stateProvince="Cape" typeStatus="Lectotype">
<emphasis id="EF3F14D085E2274C9B5A34DFB961283A" bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus id="4CB89769D8558ECAD9870509F4546569">Lectotype</typeStatus>
</emphasis>
:
<collectingCountry id="1F221FEE4DB509EFDEEDABDDC65B0E16" name="South Africa">South Africa</collectingCountry>
:
<collectingRegion id="BCBA182861C678A88C4730BD4990F703" country="South Africa" name="Western Cape">Cape</collectingRegion>
Town, leg.
<collectorName id="E61064D12A80C982848CF511FAFA39C8">F. Foly</collectorName>
,
<specimenCount id="C365CB5DA0EEF8C787ED5A2DC23A8163"></specimenCount>
, SAMC, catalogue no. SAM-HYM-B000145. Labels associated with this specimen are shown in Fig.
<figureCitation id="B46025DDEA4AEF5F1A4533ACE836F373" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male lectotype of Scrapter heterodoxus (Cockerell, 1921). A habitus, dorsolateral view, and labels B habitus, lateral view C head, frontal view D propodeum E hind leg tibia F fore wing." figureDoi="10.3897/afrinvertebr.63.76934.figure1" httpUri="https://binary.pensoft.net/fig/636741" pageId="0" pageNumber="1">1A</figureCitation>
. According to
<bibRefCitation id="306E975AA1FD9863F9A7ECD8A66A6F06" DOI="https://doi.org/10.1080/00222932108632512" author="Cockerell, TDA" journalOrPublisher="Evolution" pageId="0" pageNumber="1" refId="B13" refString="Cockerell, TDA, 1921. XXV.-Descriptions and Records of Bees.-XC. Annals and Magazine of Natural History (Ser 9) 7: 201-212. https://doi.org/10.1080/00222932108632512" title="XXV. - Descriptions and Records of Bees. - XC. Annals and Magazine of Natural History (Ser 9) 7: 201 - 212." url="https://doi.org/10.1080/00222932108632512" year="1921">Cockerell (1921)</bibRefCitation>
, the specimen was collected in 1914
</materialsCitation>
.
<materialsCitation id="C1A2A7D86F46B7B5C3AEE391F607831D" collectingDate="1925-10-25" collectingDateMax="2007-08-25" collectingDateMin="1925-10-25" collectorName="B. N. Danforth, Danforth, C. D, Eardley, K. L, V. B. Whitehead, Katberg Pass, R, K. Steiner, S. van Noort, H. Brauns" country="South Africa" latitude="-34.1167" location="Elands Bay" longLatPrecision="6" longitude="18.8333" municipality="Clanwilliam" specimenCount="46" specimenCount-female="25" specimenCount-male="21" stateProvince="Cape Province">
Additional material:
<collectingCountry id="5B9CC2CB35134B28D862DAA542E4E124" name="South Africa">South Africa</collectingCountry>
:
<collectingRegion id="365FF0168E9A469D6D57D7D7B1E66BED" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
,
<locationDeviation id="4966ABF8CC75B5B17E2026E3F87904A9" location="Clanwilliam">
<quantity id="A63A5CEBD1ED501AE50FF538E24EF005" metricMagnitude="4" metricUnit="m" metricValue="3.1" unit="km" value="31.0">31 km</quantity>
S of Clanwilliam
</locationDeviation>
,
<geoCoordinate id="8AF45C837036071D9CCA1E7BD2DC03B9" degrees="32" direction="south" minutes="23.1" orientation="latitude" precision="92" value="-32.385">32°23.1'S</geoCoordinate>
,
<geoCoordinate id="DC93F1501FE79E91FAD39705FFAB6664" degrees="18" direction="east" minutes="56.8" orientation="longitude" precision="92" value="18.946667">18°56.8'E</geoCoordinate>
,
<collectingDate id="007F23CE67EAC98216A0AC92F027AB22" value="2001-09-07">7 September 2001</collectingDate>
, leg.
<collectorName id="7A1121B019CB63135187FE685340792C">
B. N.
<collectorName id="9C06C43F2057E8A903C8DE3F29DACBBA">Danforth, C. D</collectorName>
</collectorName>
.
<collectorName id="0F34FB509625FD6B828CB3C14B29D528">Eardley, K. L</collectorName>
. Walker,
<specimenCount id="5AA06498EF8A864ED184C0A0E1B98D95" type="male">6♂</specimenCount>
,
<specimenCount id="24391F8848E63372BCD66136F328BBBA" type="female">15♀</specimenCount>
, CUIC.
<collectingRegion id="8F94CD27B8AE998F964C8CD355DA7052" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
, Sauer, Suurfontein, -32.85, 18.5667,
<collectingDate id="3A34A928CAA21D39E99D21363BCDEF2D" value="1994-08-25">25 August 1994</collectingDate>
, leg.
<collectorName id="00CE990D3326A85CBD6B38160AAC9B62">V. B. Whitehead</collectorName>
,
<specimenCount id="E071A3BFA6729660CFD586101D457F26" type="male">2♂</specimenCount>
, SAMC, cat. no. SAM-HYM-B007774.
<collectingRegion id="87BCBD8C0FBF3F0DA7EA96D89A5D9C40" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
, Holfontein,
<quantity id="6BB0BD9D9497DDBD0AAF26E5A233E80D" metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
S. of
<collectingMunicipality id="FDCA5D1D32723C860D97434F4A106F20">Clanwilliam</collectingMunicipality>
, -32.4333, 18.95,
<collectingDate id="EFED0B209E3962DB3C28EEED8D6EE8B5" value="1984-08-08">8 August 1984</collectingDate>
, leg.
<collectorName id="7964EA82E4A787F921D1133AAA45C230">V. B. Whitehead</collectorName>
,
<specimenCount id="D0696C9D7E7C06E225E7BE986E0EB272" type="male">3♂</specimenCount>
, SAMC, cat. no. SAM-HYM-B007777.
<collectingRegion id="659183C7D5B9274750266D72C8D4DFE2" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
, Piketberg, Wittewater, -32.9167, 18.7,
<collectingDate id="C4A95616EE9B1B7827172BB15CB6AE22" value="1990-09-05">5 September 1990</collectingDate>
, leg.
<collectorName id="8794A95C889A64B393439E4E9BBCEE12">V. B. Whitehead</collectorName>
,
<specimenCount id="B20BBDD85F7F75FED7859EA7D601D5B2" type="female">1♀</specimenCount>
, SAMC, cat. no. SAM-HYM-B007784.
<collectingRegion id="AA5E3100A05570433875800044AAD2F4" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
, Piketberg, farm Hartbeesrivier, Kapteinskloof,
<geoCoordinate id="1BBD06F8892B5A81D09FAEF8D2C16987" degrees="32.875" direction="south" orientation="latitude" precision="55" value="-32.875">-32.875</geoCoordinate>
,
<geoCoordinate id="AF9134E278D0B4DD8BFC35CCFF6BBF9A" degrees="18.625" direction="east" orientation="longitude" precision="55" value="18.625">18.625</geoCoordinate>
,
<collectingDate id="0B6B085C2E842D04D4FDD5A5EC8EFDB7" value="1991-08-23">23 August 1991</collectingDate>
, leg.
<collectorName id="0FB1786F49C6BDBD2433B8CD899BFC05">V. B. Whitehead</collectorName>
,
<specimenCount id="BC09CDC73E552F61D7FB61F7939D252A" type="male">1♂</specimenCount>
, SAMC, cat. no. SAM-HYM-B007785.
<collectingRegion id="2D0B2DD9E268B208E8B4F32AA0F0B343" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
, Piketberg, Banghoek, -32.75, 18.6,
<collectingDate id="E18CB2EF082588C2624E1B55EAF9B5B3" value="1991-09-20">20 September 1991</collectingDate>
, leg.
<collectorName id="DB329870CAFBD5EA00C8A86A17EF6B1B">V. B. Whitehead</collectorName>
,
<specimenCount id="4BD7E1B7D6B2482EF9A435FAD752A633" type="female">1♀</specimenCount>
, SAMC, cat. no. SAM-HYM-B007786.
<collectingRegion id="1A0A853763C70CBD8BD5A58F7A2D59DF" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
, Mamre, Malmesbury Div.
<collectingRegion id="178F79C27AF3D10F29EA15F1A8ADF9F6" country="South Africa" name="Western Cape">Cape</collectingRegion>
,
<geoCoordinate id="7AA1CFCB8CEFC2A6018A9584A3D9A75B" degrees="33.5167" direction="south" orientation="latitude" precision="5" value="-33.5167">-33.5167</geoCoordinate>
,
<geoCoordinate id="3CF74B55CC83C16A735442F99445D8AE" degrees="18.4667" direction="east" orientation="longitude" precision="5" value="18.4667">18.4667</geoCoordinate>
,
<collectingDate id="C806D57ABD301F1637BAA5B33E1091EF" value="1977-08-25">25 August 1977</collectingDate>
, leg.
<collectorName id="9AB0D36AD67B29B5F47DAA06D5D11B26">V. B. Whitehead</collectorName>
,
<specimenCount id="4E4A9546FC54DDA9C987B8A8B37BAB84" type="male">1♂</specimenCount>
, SAMC, cat. no. SAM-HYM-B007788.
<collectingRegion id="97B20C0D50089A5F193BF689D7DB2F6B" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
, Joostenbergkloof, Stellenbosch,
<geoCoordinate id="F1CC0FF91E3571E4669816355D21F164" degrees="33.7667" direction="south" orientation="latitude" precision="5" value="-33.7667">-33.7667</geoCoordinate>
,
<geoCoordinate id="16D321A1A956D3B9FA7EE45500D9A94D" degrees="18.7667" direction="east" orientation="longitude" precision="5" value="18.7667">18.7667</geoCoordinate>
,
<collectingDate id="9BBE54E7BDEA73959AC2BCB32F0EA43C" value="1988-08-14">14 August 1988</collectingDate>
, leg.
<collectorName id="1B9E18F8D75D4F5CF3FB13D3DB28CA52">V. B. Whitehead</collectorName>
,
<specimenCount id="1DBCEB00579D4D98892DB4238C318D4C" type="male">3♂</specimenCount>
, SAMC, cat. no. SAM-HYM-B007789.
<collectingRegion id="34621CA62CA0598C1E785255CF81BEAF" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
, Joostenbergkloof, Stellenbosch,
<geoCoordinate id="D448532DE11512EF01F0630E56D9B03D" degrees="33.7667" direction="south" orientation="latitude" precision="5" value="-33.7667">-33.7667</geoCoordinate>
,
<geoCoordinate id="E81776ACE80ABE614721C72A2B2937EC" degrees="18.7667" direction="east" orientation="longitude" precision="5" value="18.7667">18.7667</geoCoordinate>
,
<collectingDate id="26D3D8ECA34AF34DEEBBA097AC69804F" value="1988-07-31">31 July 1988</collectingDate>
, leg.
<collectorName id="32651C4A336F3BA954122F24DE312AE3">V. B. Whitehead</collectorName>
,
<specimenCount id="4ADB91AAEC81415AAC198041230AB4B5" type="male">1♂</specimenCount>
, SAMC, cat. no. SAM-HYM-B007790.
<collectingRegion id="CB454289E08E1E56F2422A7AC3F26A6D" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
,
<collectorName id="B6C6393E9C0A9CAC4B1829F4B974DE69">Katberg Pass, R</collectorName>
351, -32.4667, 26.65,
<collectingDate id="E75EB254843FA6A366D9DB51AC4DBF0B" value="1985-11-25">25 November 1985</collectingDate>
,
<specimenCount id="877CFF08F706BBFFE448BC6A0B13A5D6" type="female">1♀</specimenCount>
, SAMC, cat. no. SAM-HYM-B007791.
<collectingRegion id="F1E843AA8FB364BAA88D5B14D2876F05" country="South Africa" name="Western Cape">Cape Province</collectingRegion>
, Kakamas,
<locationDeviation id="1603EE4973954BFD4FFD4718C03CB5C1" location="Orange">
<quantity id="4670DF68E3496462985D76160F07AC04" metricMagnitude="4" metricUnit="m" metricValue="1.65" unit="km" value="16.5">16.5 km</quantity>
N of Orange
</locationDeviation>
R. bridge, Rd to
<collectingCountry id="714EABD47753E955C6E788E8E78C8863" name="Namibia">Namibia</collectingCountry>
,
<geoCoordinate id="60CF7731CEA23DABED83A0A41633BECE" degrees="28.6000" direction="south" orientation="latitude" precision="5" value="-28.6">-28.6000</geoCoordinate>
,
<geoCoordinate id="673B2E864B130474FD24703DA4776C65" degrees="20.5667" direction="east" orientation="longitude" precision="5" value="20.5667">20.5667</geoCoordinate>
,
<collectingDate id="124DA8F7B2ABF73AB7EF9E8CD1125062" value="1993-07-22">22 July 1993</collectingDate>
, leg.
<collectorName id="8BC6AA203837ED26ECB069D6AC9EA586">V. B. Whitehead</collectorName>
,
<specimenCount id="B51D9E101CC25AF0F78A4547C9986624" type="female">1♀</specimenCount>
, SAMC, cat. no. SAM-HYM-B007792. Cape Province, Leipoldtville,
<geoCoordinate id="8FB81640EDC03009C0C25392CE5B329D" degrees="32.2333" direction="south" orientation="latitude" precision="5" value="-32.2333">-32.2333</geoCoordinate>
,
<geoCoordinate id="6EC1A8DB81581396C60FEE20F046F32A" degrees="18.4833" direction="east" orientation="longitude" precision="5" value="18.4833">18.4833</geoCoordinate>
,
<collectingDate id="4B2E35A948AB09C5AEEC7C31517C0E2B" value="1984-09-14">14 September 1984</collectingDate>
, leg.
<collectorName id="52A63A43321A2EC98534E063DC643FCD">V. B. Whitehead</collectorName>
,
<specimenCount id="EAF653BCAF059FE94DAF633D289278DE" type="female">2♀</specimenCount>
, SAMC, cat. no. SAM-HYM-B007793. Cape Province, Hetkruis, Groenrivier, - 32.6, 18.75,
<collectingDate id="0410398993299AB56F5FE897CED225AA" value="1991-08-14">14 August 1991</collectingDate>
, leg.
<collectorName id="59A5752E70C07A648D8F8B033882A9D1">V. B. Whitehead</collectorName>
,
<specimenCount id="21B40B03BDA76A5D65D02FB247C5AE0C" type="male">1♂</specimenCount>
, SAMC, cat. no. SAM-HYM-B007794. Cape Province,
<quantity id="15C7D006C192890520E6DDF0CF7007A1" metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
N. of Elandsbaai, -32.25, 18.35,
<collectingDate id="FAE6397A52F664238376FEA03328389B" value="1984-09-21">21 September 1984</collectingDate>
, leg.
<collectorName id="421403E47046026155F78967E65A291F">V. B. Whitehead</collectorName>
,
<specimenCount id="154ECDA706AABD7B27321884C9DF2C18" type="male">1♂</specimenCount>
and
<specimenCount id="D7852CE063278F6D507F3A4A13376F02" type="female">1♀</specimenCount>
, SAMC, cat. no. SAM-HYM-B007795. Cape Province, Doringfontein,
<quantity id="2B732DAA413A16ACD61784A496388E2E" metricMagnitude="4" metricUnit="m" metricValue="3.3" unit="km" value="33.0">33km</quantity>
N. of Piketberg,
<collectingDate id="D3F981B201C1D277D549EF69EA6089FB" value="1987-08-30">30 August 1987</collectingDate>
, -32.6, 18.7667, leg.
<collectorName id="EC4E052C1AD29197AD3721F28069F253">V. B. Whitehead</collectorName>
,
<specimenCount id="7FFC62082E28A5921E68033CEFEF73C7" type="male">1♂</specimenCount>
, SAMC, cat. no. SAM-HYM-B007796. Cape Province, Tygerberg, -33.8833, 18.6,
<collectingDate id="63665AA0C4BD348E1F676069C8C225FD" value="1990-09-14">14 September 1990</collectingDate>
, leg.
<collectorName id="576BF0B11A923F3C0BD30A866B768616">K. Steiner</collectorName>
,
<specimenCount id="610D9CDB467770C9A0E031D01F8FE97A" type="female">1♀</specimenCount>
, SAMC, cat. no. SAM-HYM-B009506. Cape Province,
<location id="79B846246433076E02BEE1A74C4EF3D8" LSID="urn:lsid:plazi:treatment:417FB9D69E2B5D2A8029E05F19AE61B2:79B846246433076E02BEE1A74C4EF3D8" country="South Africa" latitude="-34.1167" longLatPrecision="6" longitude="18.8333" municipality="Clanwilliam" name="Elands Bay" stateProvince="Cape Province">Elands Bay</location>
, -32.3, 18.35,
<collectingDate id="D1ADA5A87ACEA37660EBBC10BA38B7BF" value="1985-09-26">26 September 1985</collectingDate>
, leg.
<collectorName id="F8D185B0804DEC4638887A487B7ED165">V. B. Whitehead</collectorName>
,
<specimenCount id="2AB269F7B88174031C7CCB7634760D1A" type="male">1♂</specimenCount>
, SAMC, cat. no. SAM-HYM-B009507. Cape Province, Sevilla,
<normalizedToken id="E99F8154E66783CE0D154B6EBEA19343" originalValue="Travellers">Traveller's</normalizedToken>
Rest,
<geoCoordinate id="764364E93A05B69AFDEA66CD6BBB1B16" degrees="32.07278" direction="south" orientation="latitude" precision="1" value="-32.07278">-32.07278</geoCoordinate>
,
<geoCoordinate id="7397E98693757042B08FEAC4DA2881DB" degrees="19.08056" direction="east" orientation="longitude" precision="1" value="19.08056">19.08056</geoCoordinate>
,
<collectingDate id="69CECD656083950A679F224EE57AAABF" value="2007-08-25">25 August 2007</collectingDate>
, leg.
<collectorName id="7E3AA04E96F71EBDA0E554DD96002440">S. van Noort</collectorName>
,
<specimenCount id="6F29F991D1D0FEBD43B81900EDA0C83A" type="female">1♀</specimenCount>
, SAMC, cat. no. SAM-HYM-B010372. Cape Province, Somerset (W.) Strand,
<collectingDate id="8981F269183E6BE790EA378546C34935" value="1925-10-25">25 October 1925</collectingDate>
,
<geoCoordinate id="60569089419D99D1B7FFA7CC428EDCC1" degrees="34.1167" direction="south" orientation="latitude" precision="5" value="-34.1167">-34.1167</geoCoordinate>
,
<geoCoordinate id="9A5A4032647BC94BB920996D134DC261" degrees="18.8333" direction="east" orientation="longitude" precision="5" value="18.8333">18.8333</geoCoordinate>
, leg.
<collectorName id="DF0475A9990316CF770C79205F6F8D43">H. Brauns</collectorName>
,
<specimenCount id="0F3A53AB5BB9EC190A230F9BB1A2C5F5" type="female">1♀</specimenCount>
, SAMC, cat. no. SAM-HYM-B007776a
</materialsCitation>
.
</paragraph>
<caption id="AEAC8B7D2A9B15127FFF169291B02BD2" doi="10.3897/afrinvertebr.63.76934.figure1" httpUri="https://binary.pensoft.net/fig/636741" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
<paragraph id="39DB41B4248D34646FD1E9941F722627" pageId="0" pageNumber="1">
<emphasis id="28319082760B6A9268E3F553DE83820E" bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
Male
<typeStatus id="1376D360A05EF2CB318DA1E0AD5203EA">lectotype</typeStatus>
of
<taxonomicName id="243A262136AEF717CD94A39F2418B12C" baseAuthorityName="Cockerell" baseAuthorityYear="1921" class="Insecta" family="Colletidae" genus="Scrapter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scrapter heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="83327F4D7B13CBC948C136B4A26D07BE" italics="true" pageId="0" pageNumber="1">Scrapter heterodoxus</emphasis>
</taxonomicName>
(Cockerell, 1921).
<emphasis id="60BB98CBD5E0557406DFC6D83548BBBD" bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus, dorsolateral view, and labels
<emphasis id="0304768265744BFBB6AFD2B7D48891B2" bold="true" pageId="0" pageNumber="1">B</emphasis>
habitus, lateral view
<emphasis id="D72928565633B05604940DE45D220859" bold="true" pageId="0" pageNumber="1">C</emphasis>
head, frontal view
<emphasis id="1623CC04138DFC84E613235AC552BE40" bold="true" pageId="0" pageNumber="1">D</emphasis>
propodeum
<emphasis id="E0FFE4957314A92D826A92F291224034" bold="true" pageId="0" pageNumber="1">E</emphasis>
hind leg tibia
<emphasis id="2E811E0CC22696CA54E25E4A2DC453D2" bold="true" pageId="0" pageNumber="1">F</emphasis>
fore wing.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="744D752809D677EED18F20F632BB49EC" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="7F01EFFEB045D8E194029259B1182FB1" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="36CF315892D2E03BCDB020C1B16BE8BD" pageId="0" pageNumber="1">
♂: the male of
<taxonomicName id="C560794C0310F764CCAFFBE9F9003C7E" class="Insecta" kingdom="Animalia" lsidName="S. heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="6090B621F0DD0B24E010EE9EC13A9778" italics="true" pageId="0" pageNumber="1">S. heterodoxus</emphasis>
</taxonomicName>
differs from all other
<taxonomicName id="0F40ECD3BFC11CEC1089F08F2469CCA5" authorityName="Lepeletier &amp; Serville" authorityYear="1828" class="Insecta" family="Colletidae" genus="Scrapter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scrapter" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="6ACF9ECE307E7A99547DD2ED09C9750D" italics="true" pageId="0" pageNumber="1">Scrapter</emphasis>
</taxonomicName>
species except
<taxonomicName id="A5B9DC4643A2EFB27213FC7EB1DE7F1B" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="C882941DEA9A2A4C9E8870A3EFEA62D0" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
in possessing the unique combination of the following characters: body size of ≥ 12 mm, hind femur greatly enlarged, hind tibia strongly broadened apically (Figs
<figureCitation id="7BDAE8C4C22DED1056A218E47252E231" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male lectotype of Scrapter heterodoxus (Cockerell, 1921). A habitus, dorsolateral view, and labels B habitus, lateral view C head, frontal view D propodeum E hind leg tibia F fore wing." figureDoi="10.3897/afrinvertebr.63.76934.figure1" httpUri="https://binary.pensoft.net/fig/636741" pageId="0" pageNumber="1">1E</figureCitation>
,
<figureCitation id="98CA3957BA9195FD84A28EC04A7FE0FF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of the shape of the male hind leg tibia and propodeal sculpture. A Scrapter peringueyi (Cockerell, 1921), stat. rev. hind leg tibial apex, non-type male specimen (deposited in CUIC) B Scrapter heterodoxus (Cockerell, 1921) hind leg tibial apex, non-type male specimen (deposited in CUIC) C Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, non-type male specimen (SAM-HYM-B 007139) D Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, lectotype male (SAM-HYM-B 000145) E Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, holotype female (SAM-HYM-B 000144) F Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, non-type female (SAM-HYM-B 007786)." figureDoi="10.3897/afrinvertebr.63.76934.figure5" httpUri="https://binary.pensoft.net/fig/636745" pageId="0" pageNumber="1">5B</figureCitation>
), hind basitarsus unmodified (enlarged in
<taxonomicName id="091FE005716C9A421A2AA1CFBB41BDDD" class="Insecta" kingdom="Animalia" lsidName="S. amplitarsus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="amplitarsus">
<emphasis id="44157C79B1CE781F989CAB195868D86F" italics="true" pageId="0" pageNumber="1">S. amplitarsus</emphasis>
</taxonomicName>
Eardley, 1996) and midleg basitarsus unmodified (enlarged in
<taxonomicName id="64B768BAAA656A9222AF0C1585E21FEA" class="Insecta" kingdom="Animalia" lsidName="S. armatipes" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="armatipes">
<emphasis id="941D8F179D47BA2FA25B32908CC5685D" italics="true" pageId="0" pageNumber="1">S. armatipes</emphasis>
</taxonomicName>
(Friese, 1913)).
<taxonomicName id="C806255E63B1B6781B31EAB32F154D25" baseAuthorityName="Cockerell" baseAuthorityYear="1921" class="Insecta" family="Colletidae" genus="Scrapter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scrapter heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="46DC1C86E7D3C0AED7E66895BF7A094B" italics="true" pageId="0" pageNumber="1">Scrapter heterodoxus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="5D056A99FF31EF0C831FA59AF32002E9" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="5BD1BE8F57DDBA8AEC39F2056CBFC36B" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
in the shape of the apical section of the hind tibia: the shape of the projecting apical portion is tapering in
<taxonomicName id="1AA84CB83190FB794A5A3B3C0B141040" class="Insecta" kingdom="Animalia" lsidName="S. heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="9EF524E6CCE48FE64ABA0B9D3D201097" italics="true" pageId="0" pageNumber="1">S. heterodoxus</emphasis>
</taxonomicName>
, resembling a triangular shape, whereas the projecting apical portion of the tibia in
<taxonomicName id="96806DA051B87887DAF4B41D4161C405" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="B01B45E81AB22268F2B5CC5121CA973D" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
is parallel-sided, resembling a rectangular shape (Figs
<figureCitation id="6354744F658E7A7D3BE49CA601DDA249" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male lectotype of Scrapter heterodoxus (Cockerell, 1921). A habitus, dorsolateral view, and labels B habitus, lateral view C head, frontal view D propodeum E hind leg tibia F fore wing." figureDoi="10.3897/afrinvertebr.63.76934.figure1" httpUri="https://binary.pensoft.net/fig/636741" pageId="0" pageNumber="1">1E</figureCitation>
,
<figureCitation id="4F594FD14946851F2BF36B743767B818" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Non-type male specimen of Scrapter peringueyi (Cockerell, 1921), stat. rev. A habitus, dorsal view B habitus, lateral view C head, frontal view D propodeum E hind leg tibia F fore wing." figureDoi="10.3897/afrinvertebr.63.76934.figure4" httpUri="https://binary.pensoft.net/fig/636744" pageId="0" pageNumber="1">4E</figureCitation>
,
<figureCitation id="232286B732135549FE82BE884B568E4F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of the shape of the male hind leg tibia and propodeal sculpture. A Scrapter peringueyi (Cockerell, 1921), stat. rev. hind leg tibial apex, non-type male specimen (deposited in CUIC) B Scrapter heterodoxus (Cockerell, 1921) hind leg tibial apex, non-type male specimen (deposited in CUIC) C Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, non-type male specimen (SAM-HYM-B 007139) D Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, lectotype male (SAM-HYM-B 000145) E Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, holotype female (SAM-HYM-B 000144) F Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, non-type female (SAM-HYM-B 007786)." figureDoi="10.3897/afrinvertebr.63.76934.figure5" httpUri="https://binary.pensoft.net/fig/636745" pageId="0" pageNumber="1">5A</figureCitation>
,
<figureCitation id="8F47F71381BC913358CAA0E3C6A1C060" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of the shape of the male hind leg tibia and propodeal sculpture. A Scrapter peringueyi (Cockerell, 1921), stat. rev. hind leg tibial apex, non-type male specimen (deposited in CUIC) B Scrapter heterodoxus (Cockerell, 1921) hind leg tibial apex, non-type male specimen (deposited in CUIC) C Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, non-type male specimen (SAM-HYM-B 007139) D Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, lectotype male (SAM-HYM-B 000145) E Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, holotype female (SAM-HYM-B 000144) F Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, non-type female (SAM-HYM-B 007786)." figureDoi="10.3897/afrinvertebr.63.76934.figure5" httpUri="https://binary.pensoft.net/fig/636745" pageId="0" pageNumber="1">5B</figureCitation>
). As in the female sex, the surface sculpturing of the basal zone of the propodeum is rugulose in
<taxonomicName id="F0E1AE5BA5F49BDB5D13ACC0DC0A7981" class="Insecta" kingdom="Animalia" lsidName="S. heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="FA50D9F17F6C61A903ED630D78C0D610" italics="true" pageId="0" pageNumber="1">S. heterodoxus</emphasis>
</taxonomicName>
, whereas it is substrigulate in
<taxonomicName id="104659D74C16F98F5B88266593428D9B" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="B6BFD778C61EE9CEC4E5E53C2572A236" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
(Figs
<figureCitation id="45B6C91D028A617614D4DE890ADCF09D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of the shape of the male hind leg tibia and propodeal sculpture. A Scrapter peringueyi (Cockerell, 1921), stat. rev. hind leg tibial apex, non-type male specimen (deposited in CUIC) B Scrapter heterodoxus (Cockerell, 1921) hind leg tibial apex, non-type male specimen (deposited in CUIC) C Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, non-type male specimen (SAM-HYM-B 007139) D Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, lectotype male (SAM-HYM-B 000145) E Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, holotype female (SAM-HYM-B 000144) F Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, non-type female (SAM-HYM-B 007786)." figureDoi="10.3897/afrinvertebr.63.76934.figure5" httpUri="https://binary.pensoft.net/fig/636745" pageId="0" pageNumber="1">5C</figureCitation>
,
<figureCitation id="E1D91A654A0210D10FC0B1468EAC5443" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of the shape of the male hind leg tibia and propodeal sculpture. A Scrapter peringueyi (Cockerell, 1921), stat. rev. hind leg tibial apex, non-type male specimen (deposited in CUIC) B Scrapter heterodoxus (Cockerell, 1921) hind leg tibial apex, non-type male specimen (deposited in CUIC) C Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, non-type male specimen (SAM-HYM-B 007139) D Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, lectotype male (SAM-HYM-B 000145) E Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, holotype female (SAM-HYM-B 000144) F Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, non-type female (SAM-HYM-B 007786)." figureDoi="10.3897/afrinvertebr.63.76934.figure5" httpUri="https://binary.pensoft.net/fig/636745" pageId="0" pageNumber="1">5D</figureCitation>
). The integument between the punctation on the mesoscutum is polished in
<taxonomicName id="9C814998EE2851768E14452D8D1A6691" class="Insecta" kingdom="Animalia" lsidName="S. heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="2CC97A574CAB33C27D8F74B54C6AB24A" italics="true" pageId="0" pageNumber="1">S. heterodoxus</emphasis>
</taxonomicName>
and shagreened (dull) in
<taxonomicName id="F9FC65F4A4A5BFE6DC25CDB0E1B0896E" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="E00A40C455EDCCC144C5CF19905AA570" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="B8FD962B8586F896C8270DEC0921D644" doi="10.3897/afrinvertebr.63.76934.figure2" httpUri="https://binary.pensoft.net/fig/636742" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph id="4C9F404D18D39331B3FDF0F9E468561D" pageId="0" pageNumber="1">
<emphasis id="7A6BDD8A0444FAD6CF7FEC40C9EE3AD5" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
Female non-type specimen of
<taxonomicName id="FD0DC406C4189A60D61E358F18246325" baseAuthorityName="Cockerell" baseAuthorityYear="1921" class="Insecta" family="Colletidae" genus="Scrapter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scrapter heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="044441FB4CA9367EF8915B24B7001CE7" italics="true" pageId="0" pageNumber="1">Scrapter heterodoxus</emphasis>
</taxonomicName>
(Cockerell, 1921), (SAM-HYM-B007786).
<emphasis id="65B87B95A23E84E87229D7AE9858C1E7" bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus, dorsal view
<emphasis id="EFD031B5724F6A732B23BA288ADD5397" bold="true" pageId="0" pageNumber="1">B</emphasis>
habitus, lateral view
<emphasis id="13D83D363053EC83BE85D4C0D8729634" bold="true" pageId="0" pageNumber="1">C</emphasis>
head, frontal view
<emphasis id="4992E13F015FB9CA0FE9F887EADAA1B6" bold="true" pageId="0" pageNumber="1">D</emphasis>
propodeum
<emphasis id="BD3376D2FC474A179B94A74E44B11FDB" bold="true" pageId="0" pageNumber="1">E</emphasis>
labels
<emphasis id="9C482A0DFE8A7C2DD9290802596AC521" bold="true" pageId="0" pageNumber="1">F</emphasis>
fore wing.
</paragraph>
</caption>
<paragraph id="EF9E4EC43CE6A518D3AAAF7B00D21AE8" pageId="0" pageNumber="1">
♀: The female differs from most species of
<taxonomicName id="58256A71F59B8D85F983997DBB8A9B07" authorityName="Lepeletier &amp; Serville" authorityYear="1828" class="Insecta" family="Colletidae" genus="Scrapter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scrapter" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="EE4862E6BD760D1E42FF01CE61360D57" italics="true" pageId="0" pageNumber="1">Scrapter</emphasis>
</taxonomicName>
, except
<taxonomicName id="1E1248CEB1E6AA72264C4E8B9F282DB0" class="Insecta" kingdom="Animalia" lsidName="S. caesariatus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="caesariatus">
<emphasis id="E40849934C0C45B183DB156A1A2BAD1C" italics="true" pageId="0" pageNumber="1">S. caesariatus</emphasis>
</taxonomicName>
Eardley, 1996,
<taxonomicName id="B2E5CDCBDB2937F955AB43B74A71EF69" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="54EF70D7710FCE1865AF1B1DA89A3689" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
and those of the
<taxonomicName id="5D8C53C82690D82EF97856174C9C189E" class="Insecta" kingdom="Animalia" lsidName="S. nitidus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="BA9151F58AE41ED1F349B01E711DC0AA" italics="true" pageId="0" pageNumber="1">S. nitidus</emphasis>
</taxonomicName>
and
<taxonomicName id="0B089A84DB6FE014E018B2935D56D9CB" class="Insecta" kingdom="Animalia" lsidName="S. basutorum" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="basutorum">
<emphasis id="84B30601C12698203D983E0F93A0BA68" italics="true" pageId="0" pageNumber="1">S. basutorum</emphasis>
</taxonomicName>
species groups, in having a medio-longitudinally depressed clypeus (&quot;mediolongitudinal sulcus&quot; in
<bibRefCitation id="36C2B83061FF0615689ADCBE16165AC1" author="Eardley, CD" journalOrPublisher="African Entomology" pageId="0" pageNumber="1" pagination="37 - 92" refId="B17" refString="Eardley, CD, 1996. The genus Scrapter Lepeletier &amp; Serville (Hymenoptera: Colletidae). African Entomology 4: 37 - 92" title="The genus Scrapter Lepeletier &amp; Serville (Hymenoptera: Colletidae)." volume="4" year="1996">Eardley 1996</bibRefCitation>
), but the depression is shallower than in the species of the
<taxonomicName id="34CC6FC6CFEF1D93F510E12F40927E31" class="Insecta" kingdom="Animalia" lsidName="S. nitidus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="A2F4DB4885537A8780751B901DBA1BB6" italics="true" pageId="0" pageNumber="1">S. nitidus</emphasis>
</taxonomicName>
and
<taxonomicName id="A114518089C92121C195531A47C23D76" class="Insecta" kingdom="Animalia" lsidName="S. basutorum" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="basutorum">
<emphasis id="9EE549A4DF33B88265827B4B44A30A59" italics="true" pageId="0" pageNumber="1">S. basutorum</emphasis>
</taxonomicName>
groups. With 6.8-9.3 mm body length,
<taxonomicName id="08509D6DB1B7926BAED10342F9DA5CE4" class="Insecta" kingdom="Animalia" lsidName="S. heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="FACE79293FA08E6C82BEAB335C045CCB" italics="true" pageId="0" pageNumber="1">S. heterodoxus</emphasis>
</taxonomicName>
is on average larger than
<taxonomicName id="8238447A8C2BA84AD6EFEB984B766278" class="Insecta" kingdom="Animalia" lsidName="S. caesariatus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="caesariatus">
<emphasis id="FE62282305FA3E5EF1B7F0428F33DA2C" italics="true" pageId="0" pageNumber="1">S. caesariatus</emphasis>
</taxonomicName>
(at most 7.3 mm) and the species of the
<taxonomicName id="2224C4BE634BAC7135B2BA9127658721" class="Insecta" kingdom="Animalia" lsidName="S. nitidus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="5F73EF5088A2AF729887AC580A3F9B8B" italics="true" pageId="0" pageNumber="1">S. nitidus</emphasis>
</taxonomicName>
group (at most 7.7 mm long). It differs from all aforementioned
<taxonomicName id="3CB15B766A623EF56634EAC0D73F68ED" authorityName="Lepeletier &amp; Serville" authorityYear="1828" class="Insecta" family="Colletidae" genus="Scrapter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scrapter" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="3D59569F042A320A395DC4936A428AD6" italics="true" pageId="0" pageNumber="1">Scrapter</emphasis>
</taxonomicName>
species except
<taxonomicName id="754DCEE50B8531186AF93CAEBEB09EBE" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="65DB072283EDAD5C295314B559E7C04E" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
in having a declivous propodeal surface, without a nearly horizontal basal zone.
<taxonomicName id="CFD3109BF940506EFA3187539BAC53A5" baseAuthorityName="Cockerell" baseAuthorityYear="1921" class="Insecta" family="Colletidae" genus="Scrapter" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scrapter heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="348C30DF629912D156178E199E48DFEF" italics="true" pageId="0" pageNumber="1">Scrapter heterodoxus</emphasis>
</taxonomicName>
is very similar and clearly closely related to
<taxonomicName id="87175F94E1AD780234730B7EBD5AE187" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="743A3AFE9D63DCCF8A89D388A514231B" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
. It differs in having a rugulose surface sculpturing of the basal zone of the propodeum, particularly of the anterior portion, whereas the sculpturing of
<taxonomicName id="A65033BE86194C3C9B465ED0BC2EAEE3" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="883A907EE62F40557AC8CE0A51855286" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
is substrigulate (Figs
<figureCitation id="3FDCC24597E7A192F79620DC228E149C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of the shape of the male hind leg tibia and propodeal sculpture. A Scrapter peringueyi (Cockerell, 1921), stat. rev. hind leg tibial apex, non-type male specimen (deposited in CUIC) B Scrapter heterodoxus (Cockerell, 1921) hind leg tibial apex, non-type male specimen (deposited in CUIC) C Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, non-type male specimen (SAM-HYM-B 007139) D Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, lectotype male (SAM-HYM-B 000145) E Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, holotype female (SAM-HYM-B 000144) F Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, non-type female (SAM-HYM-B 007786)." figureDoi="10.3897/afrinvertebr.63.76934.figure5" httpUri="https://binary.pensoft.net/fig/636745" pageId="0" pageNumber="1">5E</figureCitation>
,
<figureCitation id="C221C6D2C8FC933C80CBB4ED524280A6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of the shape of the male hind leg tibia and propodeal sculpture. A Scrapter peringueyi (Cockerell, 1921), stat. rev. hind leg tibial apex, non-type male specimen (deposited in CUIC) B Scrapter heterodoxus (Cockerell, 1921) hind leg tibial apex, non-type male specimen (deposited in CUIC) C Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, non-type male specimen (SAM-HYM-B 007139) D Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, lectotype male (SAM-HYM-B 000145) E Scrapter peringueyi (Cockerell, 1921), stat. rev. propodeal sculpture, holotype female (SAM-HYM-B 000144) F Scrapter heterodoxus (Cockerell, 1921) propodeal sculpture, non-type female (SAM-HYM-B 007786)." figureDoi="10.3897/afrinvertebr.63.76934.figure5" httpUri="https://binary.pensoft.net/fig/636745" pageId="0" pageNumber="1">5F</figureCitation>
). As in the male, the integument between the punctation on the mesoscutum is polished in
<taxonomicName id="764D59973574271F3CEA634A6273D848" class="Insecta" kingdom="Animalia" lsidName="S. heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="E27AF4312544DCCD31E477FE54E923CB" italics="true" pageId="0" pageNumber="1">S. heterodoxus</emphasis>
</taxonomicName>
and shagreened (dull) in
<taxonomicName id="ED6419D24FC4D4466315D16BBF693DDC" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="76286CACEFDAD6C72359EDE674DDC6D5" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="134B1050851205CC37CFEADE4B774879" pageId="0" pageNumber="1" type="comments">
<paragraph id="F683952F91E547DC4AFC7B3258DCE6A6" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="766011107A10D7F91DD4C0B17CECD716" pageId="0" pageNumber="1">
The females of
<taxonomicName id="9B6674A9D5715471EC787411C508159F" class="Insecta" kingdom="Animalia" lsidName="S. heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="B8FE4256B6D00B39EAAAFCD8694DE43A" italics="true" pageId="0" pageNumber="1">S. heterodoxus</emphasis>
</taxonomicName>
and
<taxonomicName id="E8EF112F06A52C869EB6A033F6CB9CC6" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="FF86AF1AFE852E55AA3C8BEA7461BE6B" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
are difficult to distinguish at times, whereas the males are easily recognized. The published sequence data of
<taxonomicName id="4EF6234520F8BCA623D3811AF1762230" class="Insecta" kingdom="Animalia" lsidName="S. heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="C2A655156B7243EDD80E195DD205390F" italics="true" pageId="0" pageNumber="1">S. heterodoxus</emphasis>
</taxonomicName>
from previous molecular-phylogenetic treatments (
<bibRefCitation id="679BB125672F03917890473E6DE395D7" DOI="https://doi.org/10.1016/j.ympev.2008.09.028" author="Almeida, EAB" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="1" pagination="290 - 309" refId="B2" refString="Almeida, EAB, Danforth, BN, 2009. Phylogeny of colletid bees (Hymenoptera: Colletidae) inferred from four nuclear genes. Molecular Phylogenetics and Evolution 50 (2): 290 - 309, DOI: https://doi.org/10.1016/j.ympev.2008.09.028" title="Phylogeny of colletid bees (Hymenoptera: Colletidae) inferred from four nuclear genes." url="https://doi.org/10.1016/j.ympev.2008.09.028" volume="50" year="2009">Almeida and Danforth 2009</bibRefCitation>
;
<bibRefCitation id="91C217325DBDCAB969CDE25844CBF10C" DOI="https://doi.org/10.1016/j.cub.2017.03.027" author="Branstetter, MG" journalOrPublisher="Current Biology" pageId="0" pageNumber="1" pagination="1019 - 1025" refId="B8" refString="Branstetter, MG, Danforth, BN, Pitts, JP, Faircloth, BC, Ward, PS, Buffington, ML, Gates, MW, Kula, RR, Brady, SG, 2017. Phylogenomic insights into the evolution of stinging wasps and the origins of ants and bees. Current Biology 27 (7): 1019 - 1025, DOI: https://doi.org/10.1016/j.cub.2017.03.027" title="Phylogenomic insights into the evolution of stinging wasps and the origins of ants and bees." url="https://doi.org/10.1016/j.cub.2017.03.027" volume="27" year="2017">Branstetter et al. 2017</bibRefCitation>
;
<bibRefCitation id="1F385771370AD2B8CAE1B4248448B9D3" DOI="https://doi.org/10.1111/zsc.12333" author="Almeida, EAB" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="1" pagination="226 - 242" refId="B4" refString="Almeida, EAB, Packer, L, Melo, GAR, Danforth, BN, Cardinal, SC, Quinteiro, FB, Pie, MR, 2019. The diversification of neopasiphaeine bees during the Cenozoic (Hymenoptera: Colletidae). Zoologica Scripta 48 (2): 226 - 242, DOI: https://doi.org/10.1111/zsc.12333" title="The diversification of neopasiphaeine bees during the Cenozoic (Hymenoptera: Colletidae)." url="https://doi.org/10.1111/zsc.12333" volume="48" year="2019">Almeida et al. 2019</bibRefCitation>
) is associated with a vouchered specimen deposited in the CUIC. This specimen is part of the examined series listed under additional material, which was collected in the Western Cape Province, 31 km S of Clanwilliam. The voucher corresponds to the type specimen of
<taxonomicName id="A3BCEE10C5E0F794245C2D1316897048" class="Insecta" kingdom="Animalia" lsidName="S. heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="F48AD4A79D8F20A6E4E32A0460FE044B" italics="true" pageId="0" pageNumber="1">S. heterodoxus</emphasis>
</taxonomicName>
, which means that the previously published DNA data refers to the true
<taxonomicName id="FCDF9A3D42624A55ECB76F354AB8D5A5" class="Insecta" kingdom="Animalia" lsidName="S. heterodoxus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="heterodoxus">
<emphasis id="EF005427310CB56D96BE514AB0186E0B" italics="true" pageId="0" pageNumber="1">S. heterodoxus</emphasis>
</taxonomicName>
and not to
<taxonomicName id="22B7C26BFB830084A9150B2717B899D9" class="Insecta" kingdom="Animalia" lsidName="S. peringueyi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="peringueyi">
<emphasis id="8D4943D1E3F7686A741E52BD6D2BD2CA" italics="true" pageId="0" pageNumber="1">S. peringueyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>