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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.80.13330" ID-PMC="PMC5543270" ID-Pensoft-Pub="1314-2003-80-53" ID-Pensoft-UUID="2404FFC3FFF51978EA29FF84C96CFF79" ID-PubMed="28781558" ID-Zenodo-Dep="816500" ModsDocID="1314-2003-80-53" checkinTime="1495279892315" checkinUser="pensoft" docAuthor="Berger, Andreas" docDate="2017" docId="348F438132DA5FA7892435540D73ABC2" docLanguage="en" docName="PhytoKeys 80: 53-63" docOrigin="PhytoKeys 80" docPubDate="2017-05-19" docSource="http://dx.doi.org/10.3897/phytokeys.80.13330" docTitle="Palicourea tonduzii A. C. Berger 2017, comb. nov." docType="treatment" docVersion="3" id="2404FFC3FFF51978EA29FF84C96CFF79" lastPageNumber="53" masterDocId="2404FFC3FFF51978EA29FF84C96CFF79" masterDocTitle="Two new combinations, lectotypifications and a new name for Costa Rican Palicourea s. l." masterLastPageNumber="63" masterPageNumber="53" pageNumber="53" updateTime="1668141964002" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new combinations, lectotypifications and a new name for Costa Rican Palicourea s. l.</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Berger, Andreas</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3902-3827</mods:nameIdentifier>
<mods:affiliation>Division of Systematic and Evolutionary Botany, Department of Botany and Biodiversity Research, University of Vienna, Rennweg 14, A- 1030 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">andi.berger@univie.ac.at</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-05-19</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>53</mods:start>
<mods:end>63</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.80.13330</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.80.13330</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-80-53</mods:identifier>
<mods:identifier type="Pensoft-UUID">2404FFC3FFF51978EA29FF84C96CFF79</mods:identifier>
<mods:identifier type="Zenodo-Dep">816500</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182226125" LSID="urn:lsid:plazi:treatment:348F438132DA5FA7892435540D73ABC2" httpUri="http://treatment.plazi.org/id/348F438132DA5FA7892435540D73ABC2" lastPageNumber="53" pageId="0" pageNumber="53">
<subSubSection pageId="0" pageNumber="53" type="nomenclature">
<paragraph pageId="0" pageNumber="53">
<taxonomicName LSID="348F4381-32DA-5FA7-8924-35540D73ABC2" authority="(K. Krause) A. C. Berger" authorityName="A. C. Berger" authorityYear="2017" baseAuthorityName="A. C. Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Palicourea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Palicourea tonduzii" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="tonduzii" status="comb. nov.">Palicourea tonduzii (K. Krause) A. C. Berger</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="53">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="reference_group">
<paragraph pageId="0" pageNumber="53">
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="A. C. Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis tonduzii" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="tonduzii" status="comb. nov.">Cephaelis tonduzii</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="53">Basionym.</taxonomicNameLabel>
K. Krause, Bot. Jahrb. Syst. 54(3, Beibl. 119): 45-46, 1916. Non
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="A. C. Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psychotria tonduzii" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="tonduzii" status="comb. nov.">Psychotria tonduzii</taxonomicName>
Standl., J. Wash. Acad. Sci. 15(13): 287, 1925b.-
<emphasis bold="true" pageId="0" pageNumber="53">Type</emphasis>
: COSTA RICA. Cartago: Tuis, 650 m, Nov 1897,
<emphasis italics="true" pageId="0" pageNumber="53">A. Tonduz 11461</emphasis>
(lectotype, designated here: fragm. F barcode V0068631F! ex B; syntype, or possibly holotype: B † [photo: F neg. BN-773!]).
</paragraph>
<paragraph pageId="0" pageNumber="53">
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis discolor" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="discolor" status="comb. nov.">Cephaelis discolor</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="53">=</taxonomicNameLabel>
Pol., Linnaea 41(5-6): 572-573, 1877. ≡
<taxonomicName authorityName="Berger" authorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Uragoga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uragoga angosturensis" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="angosturensis" status="comb. nov.">Uragoga angosturensis</taxonomicName>
Kuntze, Revis. Gen. Pl. 2: 954, 1891, nom. nov., non
<taxonomicName authorityName="Berger" authorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Uragoga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Uragoga discolor" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="discolor" status="comb. nov.">Uragoga discolor</taxonomicName>
(Benth.) Kuntze, Revis. Gen. Pl. 2: 960, 1891. Non
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psychotria discolor" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="discolor" status="comb. nov.">Psychotria discolor</taxonomicName>
(Griseb.) Rolfe, Bull. Misc. Inform. Kew 1893: 258, 1893, nec
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Palicourea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Palicourea discolor" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="discolor" status="comb. nov.">Palicourea discolor</taxonomicName>
K. Krause, Bot. Jahrb. Syst. 54(3, Beibl. 119): 40-41, 1916.-
<emphasis bold="true" pageId="0" pageNumber="53">Type</emphasis>
: COSTA RICA. Cartago: Angostura, Nov 1875,
<emphasis italics="true" pageId="0" pageNumber="53">H. Polakowsky 384</emphasis>
(lectotype, designated here: fragm. F barcode V0068625F! ex B; syntype, or possibly holotype: B † [photo: F neg. BN-722!]).
</paragraph>
<paragraph pageId="0" pageNumber="53">
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Evea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Evea guapilensis" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="guapilensis" status="comb. nov.">Evea guapilensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="53">=</taxonomicNameLabel>
Standl., J. Wash. Acad. Sci. 15(5): 104-105, 1925a. ≡
<taxonomicName authorityName="Berger" authorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis guapilensis" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="guapilensis" status="comb. nov.">Cephaelis guapilensis</taxonomicName>
(Standl.) Standl., Publ. Field Mus. Nat. Hist., Bot. Ser. 4(8): 295, 1929. ≡
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psychotria guapilensis" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="guapilensis" status="comb. nov.">Psychotria guapilensis</taxonomicName>
(Standl.) Hammel, Selbyana 12: 139, 1991. ≡
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Palicourea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Palicourea guapilensis" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="guapilensis" status="comb. nov.">Palicourea guapilensis</taxonomicName>
(Standl.) Borhidi, Acta Bot. Hung. 59(1-2): 17, 2017a.-
<emphasis bold="true" pageId="0" pageNumber="53">Type</emphasis>
: COSTA RICA.
<normalizedToken originalValue="Limón">Limon</normalizedToken>
: Vicinity of
<normalizedToken originalValue="Guápiles">Guapiles</normalizedToken>
, 300-500 m, 12-13 Mar 1924,
<emphasis italics="true" pageId="0" pageNumber="53">P. C. Standley 37025</emphasis>
(holotype: US barcode 00129829!).
</paragraph>
<paragraph pageId="0" pageNumber="53">
<taxonomicName authorityName="Berger" authorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Evea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Evea nana" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="nana" status="comb. nov.">Evea nana</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="53">=</taxonomicNameLabel>
Standl., J. Wash. Acad. Sci. 15(5): 105, 1925a. ≡
<taxonomicName authorityName="Berger" authorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis nana" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="nana" status="comb. nov.">Cephaelis nana</taxonomicName>
(Standl.) Standl., J. Wash. Acad. Sci. 17(7): 171, 1927.
<emphasis bold="true" pageId="0" pageNumber="53">Type</emphasis>
: PANAMA.
<normalizedToken originalValue="Colón">Colon</normalizedToken>
: Hills N of Frijoles, 19 Dec 1923,
<emphasis italics="true" pageId="0" pageNumber="53">P. C. Standley 27550</emphasis>
(holotype: US barcode 1153871!).
</paragraph>
<paragraph pageId="0" pageNumber="53">
<taxonomicName authorityName="Berger" authorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis nicaraguensis" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="nicaraguensis" status="comb. nov.">Cephaelis nicaraguensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="53">=</taxonomicNameLabel>
Standl., Trop. Woods 16: 46, 1928.-
<emphasis bold="true" pageId="0" pageNumber="53">Type</emphasis>
: NICARAGUA.
<normalizedToken originalValue="Atlántico">Atlantico</normalizedToken>
Norte: Puerto Cabezas [
<normalizedToken originalValue="Bragmans">Bragman's</normalizedToken>
Bluff], bank of Kukalaya River, 60 m, 08 Dec 1927,
<emphasis italics="true" pageId="0" pageNumber="53">F. C. Englesing 58</emphasis>
(holotype: F barcode V0068627F!; isotype: fragm. G barcode G00300772! ex F).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="nomenclatural remarks">
<paragraph pageId="0" pageNumber="53">Nomenclatural remarks.</paragraph>
<paragraph pageId="0" pageNumber="53">
The earliest published name for the species is
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis discolor" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis italics="true" pageId="0" pageNumber="53">Cephaelis discolor</emphasis>
</taxonomicName>
but combinations under
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psychotria" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="53">Psychotria</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Palicourea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Palicourea" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="53">Palicourea</emphasis>
</taxonomicName>
are preoccupied by
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psychotria discolor" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis italics="true" pageId="0" pageNumber="53">Psychotria discolor</emphasis>
</taxonomicName>
(Griseb.) Rolfe and
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Palicourea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Palicourea discolor" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis italics="true" pageId="0" pageNumber="53">Palicourea discolor</emphasis>
</taxonomicName>
K. Krause. The next available name
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="A. C. Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis tonduzii" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="tonduzii">
<emphasis italics="true" pageId="0" pageNumber="53">Cephaelis tonduzii</emphasis>
</taxonomicName>
K. Krause cannot be used in
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psychotria" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="53">Psychotria</emphasis>
</taxonomicName>
because of the earlier
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="A. C. Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psychotria tonduzii" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="tonduzii">
<emphasis italics="true" pageId="0" pageNumber="53">Psychotria tonduzii</emphasis>
</taxonomicName>
Standl. Hence, the next name
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Evea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Evea guapilensis" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="guapilensis">
<emphasis italics="true" pageId="0" pageNumber="53">Evea guapilensis</emphasis>
</taxonomicName>
Standl. was adopted and the species became known as
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Psychotria guapilensis" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="guapilensis">
<emphasis italics="true" pageId="0" pageNumber="53">Psychotria guapilensis</emphasis>
</taxonomicName>
(Standl.) Hammel. Recently,
<bibRefCitation DOI="https://doi.org/10.1556/034.59.2017.1-2.3" author="Borhidi, AL" journalOrPublisher="Acta Botanica Hungarica" pageId="0" pageNumber="53" pagination="13 - 23" refId="B7" refString="Borhidi, AL, 2017a. El subgenero Heteropsychotria (Rubiaceae, Palicoureeae) en Mexico y Mesoamerica. Acta Botanica Hungarica 59 (1-2): 13 - 23, DOI: https://doi.org/10.1556/034.59.2017.1-2.3" title="El subgenero Heteropsychotria (Rubiaceae, Palicoureeae) en Mexico y Mesoamerica." url="https://doi.org/10.1556/034.59.2017.1-2.3" volume="59" year="2017 a">Borhidi (2017a)</bibRefCitation>
transferred the species to
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Palicourea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Palicourea" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="53">Palicourea</emphasis>
</taxonomicName>
and proposed the name
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Palicourea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Palicourea guapilensis" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="guapilensis">
<emphasis italics="true" pageId="0" pageNumber="53">Palicourea guapilensis</emphasis>
</taxonomicName>
(Standl.) Borhidi.
</paragraph>
<paragraph pageId="0" pageNumber="53">
However, he overlooked that the earlier published name
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="A. C. Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis tonduzii" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="tonduzii">
<emphasis italics="true" pageId="0" pageNumber="53">Cephaelis tonduzii</emphasis>
</taxonomicName>
K. Krause is still available under
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Palicourea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Palicourea" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="53">Palicourea</emphasis>
</taxonomicName>
and ought to have been adopted. The corresponding new combination is provided here. It is fortunate that the correct name for this showy species honors Swiss botanist Adolphe Tonduz (1862-1921), a long-term employee at the Museo Nacional de Costa Rica (CR) and one of the most prolific plant collectors in the country.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="typification">
<paragraph pageId="0" pageNumber="53">Typification.</paragraph>
<paragraph pageId="0" pageNumber="53">
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="A. C. Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis tonduzii" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="tonduzii">
<emphasis italics="true" pageId="0" pageNumber="53">Cephaelis tonduzii</emphasis>
</taxonomicName>
was described with reference to an entire gathering by Tonduz. Krause did not cite a particular herbarium specimen and all possible duplicates are therefore syntypes (ICN, Art. 40.2 &amp; Note 1; see also
<bibRefCitation DOI="https://doi.org/10.12705/635.7" author="McNeill, J" journalOrPublisher="Taxon" pageId="1" pageNumber="54" pagination="1112 - 1113" refId="B24" refString="McNeill, J, 2014. Holotype specimens and type citations: General issues. Taxon 63 (5): 1112 - 1113, DOI: https://doi.org/10.12705/635.7" title="Holotype specimens and type citations: General issues." url="https://doi.org/10.12705/635.7" volume="63" year="2014">McNeill 2014</bibRefCitation>
). In addition, he was based at B and his working herbarium was destroyed during World War II. Collections made by Tonduz are widely distributed and more or less complete sets are found in CR and US (
<bibRefCitation author="Stafleu, FA" journalOrPublisher="Regnum vegetabile 94, 98, 105, 110, 112, 115, 116, Bohn, Scheltema &amp; Holkema, Utrecht / Antwerpen" pageId="1" pageNumber="54" publicationUrl="http://www.sil.si.edu/DigitalCollections/tl-2" refId="B32" refString="Stafleu, FA, Cowan, RS, 1976-1988. Taxonomic literature: A selective guide to botanical publications and collections with dates, commentaries and types. Regnum vegetabile 94, 98, 105, 110, 112, 115, 116, Bohn, Scheltema &amp; Holkema, Utrecht/Antwerpen 1, http://www.sil.si.edu/DigitalCollections/tl-2" title="Taxonomic literature: A selective guide to botanical publications and collections with dates, commentaries and types" url="http://www.sil.si.edu/DigitalCollections/tl-2" volume="1" year="1976 - 1988">Stafleu and Cowan 1976-1988</bibRefCitation>
). Nevertheless, the only known original material of
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="A. C. Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis tonduzii" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="tonduzii">
<emphasis italics="true" pageId="0" pageNumber="53">Cephaelis tonduzii</emphasis>
</taxonomicName>
is a fragment at F.
</paragraph>
<paragraph pageId="0" pageNumber="53">The fragment originates from the type at B, as shown by information given on the label. It consists of a capsule with a leaf and part of an inflorescence, which is mounted together with a photograph (&quot;Berlin negative&quot;) of the original B specimen. The removed leaf is clearly recognizable on the photograph providing a definite link between both. Although not the best choice, the fragment at F appears to be the only extant original material and is here designated as lectotype.</paragraph>
<paragraph pageId="0" pageNumber="53">
A similar situation is found in
<taxonomicName authorityName="Berger" authorityYear="2017" baseAuthorityName="Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Cephaelis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cephaelis discolor" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis italics="true" pageId="0" pageNumber="53">Cephaelis discolor</emphasis>
</taxonomicName>
, which was based on a gathering of H. Polakowsky. Likewise, the only known original material is a fragment at F that originated from B. It consists of a leaf, some bracts and fruits in a capsule, which is mounted together with a photography of the B specimen, likewise destroyed. The fragment at F is here designated as lectotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="distribution">
<paragraph pageId="0" pageNumber="53">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="53">
<taxonomicName authorityName="A. C. Berger" authorityYear="2017" baseAuthorityName="A. C. Berger" baseAuthorityYear="2017" class="Magnoliopsida" family="Rubiaceae" genus="Palicourea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Palicourea tonduzii" order="Gentianales" pageId="0" pageNumber="53" phylum="Tracheophyta" rank="species" species="tonduzii">
<emphasis italics="true" pageId="0" pageNumber="53">Palicourea tonduzii</emphasis>
</taxonomicName>
is known from Nicaragua, Costa Rica, Panama and Ecuador (
<bibRefCitation author="Taylor, CM" editor="Hammel, BE" journalOrPublisher="Monographs in Systematic Botany from the Missouri Botanical Garden" pageId="2" pageNumber="55" pagination="464 - 779" refId="B46" refString="Taylor, CM, 2014. Rubiaceae. In: Hammel, BE, Grayum, MH, Herrera, C, Zamora, N, Eds., Manual de Plantas de Costa Rica. Vol. VII. Monographs in Systematic Botany from the Missouri Botanical Garden 129: 464 - 779" title="Rubiaceae." volume="129" volumeTitle="Manual de Plantas de Costa Rica. Vol. VII." year="2014">Taylor 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>