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<document id="4886D5DE5CC2598258873C84E755EF20" ID-DOI="10.5281/zenodo.282595" ID-GBIF-Dataset="3c0b0572-9d83-492a-8656-229c8b2043a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="282595" ID-ZooBank="F832C768-A8CA-4FEE-8C3B-BD933247FA6E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460543541282" checkinUser="plazi" docAuthor="Yamamoto, Shûhei &amp; Maruyama, Munetoshi" docDate="2012" docId="03DE87D80835FFEAFF28FC9129133F8E" docLanguage="en" docName="zt03517p052.pdf" docOrigin="Zootaxa 3517" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Aleochara (Emplenota) yamato Yamamoto &amp; Maruyama, n. sp." docType="treatment" docUuid="F5361030-3349-44AE-AF16-455C34BDDBB5" docUuidSource="ZooBank" docVersion="10" lastPageNumber="28" masterDocId="FFE7FFA0082CFFF1FFBFFFF628053B11" masterDocTitle="Revision of the Seashore-dwelling Subgenera Emplenota Casey and Triochara Bernhauer (Coleoptera: Staphylinidae: genus Aleochara) from Japan" masterLastPageNumber="52" masterPageNumber="1" pageNumber="26" updateTime="1698317426905" updateUser="plazi">
<mods:mods id="982350E029F900E8C913D56E83624A08" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FBFE64E88F3F2B6C35398292E5C87A12">
<mods:title id="9002EFD966DB0038855D51C804848AC7">Revision of the Seashore-dwelling Subgenera Emplenota Casey and Triochara Bernhauer (Coleoptera: Staphylinidae: genus Aleochara) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="26F6B352FBFA28D85717291D0903DE29" type="personal">
<mods:role id="672776CA26ED8C8A211EB521E299434A">
<mods:roleTerm id="6F911CC6356B90CF488185CAE8CB602E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="72012A56E09E7CA142E955C9976B0DEE">Yamamoto, Shûhei</mods:namePart>
</mods:name>
<mods:name id="CD87F23EFCD26D0B6CBF30CCA5061DCE" type="personal">
<mods:role id="85CCAD5861F0BBE95F8F894F7B72545A">
<mods:roleTerm id="2D36A9FC8744340A2207F2FF898545BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="78DD64807017F008806422D3FC3D24EA">Maruyama, Munetoshi</mods:namePart>
</mods:name>
<mods:typeOfResource id="BEC0CB2AD2254B7539300A552CE88525">text</mods:typeOfResource>
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<mods:title id="B44C5D5E6B1DE01FF7D1EC7DBAA58C16">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4981BCD31E5B55E838EE7A5ADC0C00C4">
<mods:date id="9328CF734A64BD05235BE89A3B44E7EA">2012</mods:date>
<mods:detail id="BA22551686AC4CE363C3308078CBDECF" type="volume">
<mods:number id="FDACE26ED43CF1FAA91C15D6B3E6812A">3517</mods:number>
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<mods:start id="2C30F8116AA32054F30F98AEA3A86B45">1</mods:start>
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<mods:classification id="BE591FFA39990FB26C17F4D1C66760F5">journal article</mods:classification>
<mods:identifier id="FD03199AF3ADA44F604FF92879BDCDED" type="DOI">10.5281/zenodo.282595</mods:identifier>
<mods:identifier id="962E6C1A0C65B5C601CE2E0DAF9563AB" type="GBIF-Dataset">3c0b0572-9d83-492a-8656-229c8b2043a3</mods:identifier>
<mods:identifier id="02A76D677CC0A7BB63C378FA69257586" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EB0A826550DF0704077336B9FE3C4E21" type="Zenodo-Dep">282595</mods:identifier>
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</mods:mods>
<treatment id="03DE87D80835FFEAFF28FC9129133F8E" ID-DOI="http://doi.org/10.5281/zenodo.6175376" ID-GBIF-Taxon="119528586" ID-Zenodo-Dep="6175376" LSID="urn:lsid:zoobank.org:act:F5361030-3349-44AE-AF16-455C34BDDBB5" httpUri="http://treatment.plazi.org/id/03DE87D80835FFEAFF28FC9129133F8E" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<subSubSection id="C36D65450835FFE8FF28FC9129AE38B3" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BC836CE0835FFE8FF28FC912B983890" blockId="25.[151,925,871,930]" box="[151,925,871,897]" pageId="25" pageNumber="26">
<heading id="D08081A20835FFE8FF28FC912B983890" bold="true" box="[151,925,871,897]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="B903EADC0835FFE8FF28FC912B983890" bold="true" box="[151,925,871,897]" pageId="25" pageNumber="26">
<taxonomicName id="4C774D4D0835FFE8FF28FC912B563890" ID-CoL="BMZ7" authority="Yamamoto &amp; Maruyama" authorityName="Yamamoto &amp; Maruyama" box="[151,851,871,897]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="yamato" status="sp. nov." subGenus="Emplenota">
<emphasis id="B903EADC0835FFE8FF28FC9129113890" bold="true" box="[151,276,871,897]" italics="true" pageId="25" pageNumber="26">Aleochara</emphasis>
(
<emphasis id="B903EADC0835FFE8FE9AFC9129AD3890" bold="true" box="[293,424,871,897]" italics="true" pageId="25" pageNumber="26">Emplenota</emphasis>
)
<emphasis id="B903EADC0835FFE8FE05FC912A173890" bold="true" box="[442,530,871,897]" italics="true" pageId="25" pageNumber="26">yamato</emphasis>
Yamamoto &amp; Maruyama
</taxonomicName>
<taxonomicNameLabel id="A23057A70835FFE8FCE5FC912B983890" box="[858,925,871,897]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836CE0835FFE8FF28FC7C29AE38B3" blockId="25.[151,925,871,930]" box="[151,427,906,930]" pageId="25" pageNumber="26">
(
<figureCitation id="134C2A4B0835FFE8FF20FC7C28EA38B3" box="[159,239,906,930]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1918,1941]" captionTargetBox="[183,1407,300,1871]" captionTargetId="figure@5.[151,1436,279,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Japanese species of Aleochara (Emplenota) (Figs. 1 – 5, habitus). 1. Aleochara (Emplenota) fucicola; 2. A. (E.) puetzi; 3. A. (E.) hayamai; 4. A. (E.) yamato; 5. A. (E.) segregata; 6. dorsal head of A. yamato; 7. mesoventrite and metaventrite of A. fucicola in ventral view." httpUri="https://zenodo.org/record/282596/files/figure.png" pageId="25" pageNumber="26">Figs. 4</figureCitation>
,
<figureCitation id="134C2A4B0835FFE8FF45FC7C294138B3" box="[250,324,906,930]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="25" pageNumber="26">46–52</figureCitation>
,
<figureCitation id="134C2A4B0835FFE8FEF1FC7C296938B3" box="[334,364,906,930]" captionStart="FIGURES 93 – 100" captionStartId="43.[151,264,1848,1871]" captionTargetBox="[201,1397,201,1819]" captionTargetId="figure@43.[186,1410,193,1833]" captionTargetPageId="43" captionText="FIGURES 93 – 100. Collecting sites of the subgenera Emplenota and Triochara of the genus Aleochara in Japan. 93. Kourakaigan (Matsue-shi, Shimane-ken; habitat of A. (E.) fucicola, A. (E.) segregata, and A. (E.) hayamai; arrow); 94. drifted seaweed and flotsams on beach (Koura-kaigan); 95. Mochirippu-numa (Hamanaka-chô, Hokkaidô; A. (E.) puetzi, arrow); 96. Ochiishi-misaki (Nemuro-shi, Hokkaidô; A. puetzi); 97. Kakanokuketo (Shimane-ken; habitat of A. fucicola, A. segregata, A. hayamai, and the type locality of A. (E.) yamato; A. fucicola, arrow); 98. Hamamatsu (Nemuro-shi, Hokkaidô; habitat of A. (T.) nubis, arrow); 99. drifted seaweed on beach (Hamamatsu); 100. mating behaviour of A. nubis at Hamamatsu." httpUri="https://zenodo.org/record/282608/files/figure.png" pageId="25" pageNumber="26">97</figureCitation>
,
<figureCitation id="134C2A4B0835FFE8FEC7FC7C29A638B3" box="[376,419,906,930]" captionStart="FIGURES 101 – 103" captionStartId="44.[151,264,1923,1946]" captionTargetBox="[182,1388,523,1897]" captionTargetId="figure@44.[151,1436,515,1909]" captionTargetPageId="44" captionText="FIGURES 101 – 103. Collecting localities of Aleochara (Emplenota) spp. based on specimens examined. 101. A. (E.) fucicola (Pointed by black circle); 102. A. (E.) segregata (Pointed by black square); 103. A. (E.) puetzi (Pointed by black triangle); A. (E.) hayamai (Pointed by black diamond); A. (E.) yamato (Pointed by black star)." httpUri="https://zenodo.org/record/282609/files/figure.png" pageId="25" pageNumber="26">103</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36D65450835FFE8FF28FC252B0E3F3E" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BC836CE0835FFE8FF28FC252B243F1C" blockId="25.[151,1437,978,2008]" pageId="25" pageNumber="26">
<emphasis id="B903EADC0835FFE8FF28FC25299938FA" bold="true" box="[151,412,978,1003]" pageId="25" pageNumber="26">
<typeStatus id="54CC886C0835FFE8FF28FC2528D738FA" box="[151,210,979,1003]" pageId="25" pageNumber="26">Type</typeStatus>
series.
<typeStatus id="54CC886C0835FFE8FE90FC25299938FA" box="[303,412,979,1003]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
</emphasis>
, 3,
<emphasis id="B903EADC0835FFE8FE6CFC242A2838FA" bold="true" box="[467,557,978,1003]" pageId="25" pageNumber="26">
<collectingCountry id="F360765E0835FFE8FE6CFC242A2838FA" box="[467,557,978,1003]" name="Japan" pageId="25" pageNumber="26">JAPAN</collectingCountry>
</emphasis>
: Kakanokuketo, Shimane-chô, Matsue-shi, Shimane-ken, Honshû (
<geoCoordinate id="EE4350090835FFE8FA8DFC242D9D38FB" box="[1330,1432,978,1002]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="55" value="35.578">35.578N</geoCoordinate>
,
<geoCoordinate id="EE4350090835FFE8FF28FC0329023F1C" box="[151,263,1013,1037]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="55" value="133.05">133.050E</geoCoordinate>
),
<date id="FFC9100E0835FFE8FEA5FC03298A3F1C" box="[282,399,1013,1037]" pageId="25" pageNumber="26" value="2009-04-01">1 IV 2009</date>
, Hayama-T. (shingle beach; KUM).
</paragraph>
<paragraph id="8BC836CE0835FFE8FF78FBEE2B0E3F3E" blockId="25.[151,1437,978,2008]" box="[199,779,1047,1072]" pageId="25" pageNumber="26">
<typeStatus id="54CC886C0835FFE8FF78FBEE29323F21" box="[199,311,1048,1072]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
, 4 3, 4 Ƥ, same data as
<typeStatus id="54CC886C0835FFE8FDF9FBE12AAD3F3E" box="[582,680,1047,1071]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
(KUM).
</paragraph>
</subSubSection>
<subSubSection id="C36D65450835FFEAFF78FBCC2CEE3ADE" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
<paragraph id="8BC836CE0835FFE8FF78FBCC2B6B3DF4" blockId="25.[151,1437,978,2008]" pageId="25" pageNumber="26">
<emphasis id="B903EADC0835FFE8FF78FBCC299B3F42" bold="true" box="[199,414,1082,1107]" pageId="25" pageNumber="26">Description. Body</emphasis>
(
<figureCitation id="134C2A4B0835FFE8FE10FBCC29F33F43" box="[431,502,1082,1106]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1918,1941]" captionTargetBox="[183,1407,300,1871]" captionTargetId="figure@5.[151,1436,279,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Japanese species of Aleochara (Emplenota) (Figs. 1 – 5, habitus). 1. Aleochara (Emplenota) fucicola; 2. A. (E.) puetzi; 3. A. (E.) hayamai; 4. A. (E.) yamato; 5. A. (E.) segregata; 6. dorsal head of A. yamato; 7. mesoventrite and metaventrite of A. fucicola in ventral view." httpUri="https://zenodo.org/record/282596/files/figure.png" pageId="25" pageNumber="26">Fig. 4</figureCitation>
): extremely large; robust; somewhat thickly elongated; entire surface of pronotum and elytra covered with numerous brown hairs somewhat densely, but inconspicuous; somewhat large hexagonal microstructures on dorsal surface of head, pronotum and elytra; distinct and prominent punctures on dorsal surface of head and pronotum; punctures on elytra deep.
<emphasis id="B903EADC0835FFE8FD78FB542B1E3FAA" bold="true" box="[711,795,1186,1211]" pageId="25" pageNumber="26">Colour</emphasis>
(
<figureCitation id="134C2A4B0835FFE8FC94FB542B9F3FAB" box="[811,922,1186,1210]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1918,1941]" captionTargetBox="[183,1407,300,1871]" captionTargetId="figure@5.[151,1436,279,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Japanese species of Aleochara (Emplenota) (Figs. 1 – 5, habitus). 1. Aleochara (Emplenota) fucicola; 2. A. (E.) puetzi; 3. A. (E.) hayamai; 4. A. (E.) yamato; 5. A. (E.) segregata; 6. dorsal head of A. yamato; 7. mesoventrite and metaventrite of A. fucicola in ventral view." httpUri="https://zenodo.org/record/282596/files/figure.png" pageId="25" pageNumber="26">Figs. 4, 6</figureCitation>
): gland colour blackish brown to black; legs, especially tarsal segments, reddish brown: maxillary and labial palpi blackish brown to reddish brown; antennae blackish brown and partly light brown.
<emphasis id="B903EADC0835FFE8FDF0FB1E2A8A3E11" bold="true" box="[591,655,1256,1280]" pageId="25" pageNumber="26">Head</emphasis>
(
<figureCitation id="134C2A4B0835FFE8FD22FB112AE73FEE" box="[669,738,1255,1279]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1918,1941]" captionTargetBox="[183,1407,300,1871]" captionTargetId="figure@5.[151,1436,279,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Japanese species of Aleochara (Emplenota) (Figs. 1 – 5, habitus). 1. Aleochara (Emplenota) fucicola; 2. A. (E.) puetzi; 3. A. (E.) hayamai; 4. A. (E.) yamato; 5. A. (E.) segregata; 6. dorsal head of A. yamato; 7. mesoventrite and metaventrite of A. fucicola in ventral view." httpUri="https://zenodo.org/record/282596/files/figure.png" pageId="25" pageNumber="26">Fig. 6</figureCitation>
): somewhat wide (HW/HL = 1.25), widest at just behind eyes; impunctured area of forepart flattened but of basal part prominently and spherically swollen (not carinate; see,
<figureCitation id="134C2A4B0835FFE8FAD0FAFC28AD3E54" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1918,1941]" captionTargetBox="[183,1407,300,1871]" captionTargetId="figure@5.[151,1436,279,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Japanese species of Aleochara (Emplenota) (Figs. 1 – 5, habitus). 1. Aleochara (Emplenota) fucicola; 2. A. (E.) puetzi; 3. A. (E.) hayamai; 4. A. (E.) yamato; 5. A. (E.) segregata; 6. dorsal head of A. yamato; 7. mesoventrite and metaventrite of A. fucicola in ventral view." httpUri="https://zenodo.org/record/282596/files/figure.png" pageId="25" pageNumber="26">Fig. 6</figureCitation>
: arrow); surface sparsely covered with medium sized setae; uniformly observed huge hexagonal reticulations on dorsal surface with relatively large distinct punctures.
<emphasis id="B903EADC0835FFE8FD43FAB92B6A3E79" bold="true" box="[764,879,1359,1384]" pageId="25" pageNumber="26">Antennae</emphasis>
: thick and robust; segment I, not so thick, nearly 3.0 times as long as broad; segment II clearly shorter than I; segment III clearly longer than II; segment IV short barrel shape, slightly longer than width; segments V and VI barrel like shape; segments VII to X clearly transverse; segment XI, about 2.0 times as long as broad; relative length (width) of segments from basal to apical: 9(3): 4.5(2.5): 6.5(2.5): 3.5(3): 3.5(4): 3.5(4): 3.5(5): 3.5(5): 3.5(5): 3.5(5): 8(4).
<emphasis id="B903EADC0835FFE8FC49FA2D2C4A3EE2" bold="true" box="[1014,1103,1499,1523]" pageId="25" pageNumber="26">Thorax</emphasis>
: pronotum moderately wider than long (PW/PL =1.24), a little broader than head (PW/HW =1.36); strongly constricted near base; surface mat with large hexagonal reticulations and deep distinct punctures. Inter coxal cavities of mesoventrite very narrowly separated. Apex of inter coxal process of mesoventrite extremely sharp with exceedingly short carina (almost nothing) along midline, about 0.11 times as long as mesoventrite. Metaventrite, about 1.8 times as long as mesoventrite. Inter coxal process of metaventrite broad and short, rounded apically.
<emphasis id="B903EADC0835FFE8FBCEF97E2CAC3DB1" bold="true" box="[1137,1193,1672,1696]" pageId="25" pageNumber="26">Legs</emphasis>
: hindtibia somewhat long, nearly same length as elytra; relative lengths of tarsomeres from basal to apical: 7: 4: 4: 3.5:
<quantity id="4C8F9B2B0835FFE8FB59F95C2D1A3DD3" box="[1254,1311,1706,1730]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="25" pageNumber="26" unit="in" value="16.0">16 in</quantity>
foretarsus, 11: 7.5: 7.5: 7.5:
<quantity id="4C8F9B2B0835FFE8FEE9F93B29A13DF4" box="[342,420,1741,1765]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" pageId="25" pageNumber="26" unit="in" value="20.5">20.5 in</quantity>
midtarsus, 12: 11: 8: 8:
<quantity id="4C8F9B2B0835FFE8FD0CF93B2AE83DF4" box="[691,749,1741,1765]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="25" pageNumber="26" unit="in" value="23.0">23 in</quantity>
hindtarsus.
</paragraph>
<paragraph id="8BC836CE0835FFE8FF78F9062BDD3CC9" blockId="25.[151,1437,978,2008]" pageId="25" pageNumber="26">
[
<emphasis id="B903EADC0835FFE8FF6FF90629083C19" bold="true" box="[208,269,1776,1800]" pageId="25" pageNumber="26">Male</emphasis>
]: tergite VIII (
<figureCitation id="134C2A4B0835FFE8FE02F9192A163C16" box="[445,531,1775,1799]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="25" pageNumber="26">Fig. 46</figureCitation>
) clearly emarginated medially at posterior margin, with around 8 macrosetae. Sternite VIII (
<figureCitation id="134C2A4B0835FFE8FE89F8E4298C3C3B" box="[310,393,1810,1834]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="25" pageNumber="26">Fig. 48</figureCitation>
) with about 7 macrosetae and around 7 thin macrosetae; posterior margin strongly rounded. Aedeagus (
<figureCitation id="134C2A4B0835FFE8FEAAF8C329983C5C" box="[277,413,1845,1869]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="25" pageNumber="26">Figs. 50–51</figureCitation>
) extremely large; median lobe elongated and moderately narrowed toward apex, elongated pyriform in ventral view (
<figureCitation id="134C2A4B0835FFE8FE02F8A12A143C7E" box="[445,529,1879,1903]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="25" pageNumber="26">Fig. 51</figureCitation>
); a pair of subapico-ventral projections straight, quite thick and robust in lateral aspect (
<figureCitation id="134C2A4B0835FFE8FF52F88C29473C83" box="[237,322,1914,1938]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="25" pageNumber="26">Fig. 50</figureCitation>
); basal swelling of aedeagus, circularly curved and pointed in lateral view (
<figureCitation id="134C2A4B0835FFE8FB27F88C2CE83C83" box="[1176,1261,1914,1938]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="25" pageNumber="26">Fig. 50</figureCitation>
); apical lobe of median lobe long and moderately curved in lateral view (
<figureCitation id="134C2A4B0835FFE8FCA6F86B2B683CA4" box="[793,877,1949,1973]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="25" pageNumber="26">Fig. 50</figureCitation>
), and clearly isosceles shape in ventral view (
<figureCitation id="134C2A4B0835FFE8FAD1F86B28B33CC9" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="25" pageNumber="26">Fig. 51</figureCitation>
); flagellum shorter than the whole length of median lobe (
<figureCitation id="134C2A4B0835FFE8FCFDF8362BC93CC9" box="[834,972,1984,2008]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="25" pageNumber="26">Figs. 50–51</figureCitation>
).
</paragraph>
<caption id="DF0866460836FFEBFF28F92F29B23C21" httpUri="https://zenodo.org/record/282603/files/figure.png" pageId="26" pageNumber="27" targetBox="[151,1433,193,1728]" targetPageId="26">
<paragraph id="8BC836CE0836FFEBFF28F92F29B23C21" blockId="26.[151,1436,1753,1840]" pageId="26" pageNumber="27">
<emphasis id="B903EADC0836FFEBFF28F92F29513DFF" bold="true" box="[151,340,1753,1776]" pageId="26" pageNumber="27">FIGURES 53–64.</emphasis>
Japanese species of
<taxonomicName id="4C774D4D0836FFEBFD98F92F2B0B3DFE" authority="(Triochara)" baseAuthorityName="Triochara" box="[551,782,1753,1775]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0836FFEBFD98F92F2A953DFE" box="[551,656,1753,1775]" italics="true" pageId="26" pageNumber="27">Aleochara</emphasis>
(
<emphasis id="B903EADC0836FFEBFD20F92F2B063DFE" box="[671,771,1753,1775]" italics="true" pageId="26" pageNumber="27">Triochara</emphasis>
)
</taxonomicName>
. 53-56.
<taxonomicName id="4C774D4D0836FFEBFCDDF92F2CB53DFE" box="[866,1200,1753,1775]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="trisulcata" subGenus="Triochara">
<emphasis id="B903EADC0836FFEBFCDDF92F2BCE3DFE" box="[866,971,1753,1775]" italics="true" pageId="26" pageNumber="27">Aleochara</emphasis>
(
<emphasis id="B903EADC0836FFEBFC64F92F2C3A3DFE" box="[987,1087,1753,1775]" italics="true" pageId="26" pageNumber="27">Triochara</emphasis>
)
<emphasis id="B903EADC0836FFEBFBF0F92F2CB53DFE" box="[1103,1200,1753,1775]" italics="true" pageId="26" pageNumber="27">trisulcata</emphasis>
</taxonomicName>
; 57-60.
<taxonomicName id="4C774D4D0836FFEBFAB9F92C2D913DFE" box="[1286,1428,1753,1775]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zerchei" subGenus="Triochara">
<emphasis id="B903EADC0836FFEBFAB9F92C2D103DFE" box="[1286,1301,1754,1775]" italics="true" pageId="26" pageNumber="27">A</emphasis>
. (
<emphasis id="B903EADC0836FFEBFA94F92C2D323DFE" box="[1323,1335,1754,1775]" italics="true" pageId="26" pageNumber="27">T</emphasis>
.)
<emphasis id="B903EADC0836FFEBFAF3F92F2D913DFE" box="[1356,1428,1753,1775]" italics="true" pageId="26" pageNumber="27">zerchei</emphasis>
</taxonomicName>
; 61-64.
<taxonomicName id="4C774D4D0836FFEBFF60F90C295E3C1E" box="[223,347,1785,1807]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="nubis" subGenus="Triochara">
<emphasis id="B903EADC0836FFEBFF60F90C28EB3C1E" box="[223,238,1786,1807]" italics="true" pageId="26" pageNumber="27">A</emphasis>
. (
<emphasis id="B903EADC0836FFEBFEBBF90C29153C1E" box="[260,272,1786,1807]" italics="true" pageId="26" pageNumber="27">T</emphasis>
.)
<emphasis id="B903EADC0836FFEBFE9AF90F295E3C1E" box="[293,347,1785,1807]" italics="true" pageId="26" pageNumber="27">nubis</emphasis>
</taxonomicName>
. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64).
</paragraph>
</caption>
<paragraph id="8BC836CE0836FFEAFF78F8A92A363BE6" blockId="26.[151,1437,1886,2019]" lastBlockId="27.[151,1437,151,1184]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
[
<emphasis id="B903EADC0836FFEBFF6FF8A929223C66" bold="true" box="[208,295,1887,1911]" pageId="26" pageNumber="27">Female</emphasis>
]: tergite VIII (
<figureCitation id="134C2A4B0836FFEBFE71F8A82A253C67" box="[462,544,1886,1910]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="26" pageNumber="27">Fig. 47</figureCitation>
) strongly emarginated at posterior margin, with around 4 macrosetae. Shape of sternite VIII (
<figureCitation id="134C2A4B0836FFEBFEFFF874299C3C8B" box="[320,409,1922,1946]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="26" pageNumber="27">Fig. 49</figureCitation>
) closely similar to that of male, but pointed somewhat weakly, with 5 macrosetae. Spermatheca (
<figureCitation id="134C2A4B0836FFEBFE87F85029883CAF" box="[312,397,1958,1982]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="26" pageNumber="27">Fig. 52</figureCitation>
): spermathecal head (sh) about twice longer than apical portion of spermathecal stem; neck of spermatheca very long, somewhat longer than (sh); collar thick; basal portion of spermathecal stem (sb) slightly narrowing toward base, with bent at base; sclerotized portion of spermathecal stem short, erect and sclerotized; each part of spermatheca except for membraneous portion of spermathecal duct (sm) entirely and moderately sclerotized; (sm) moderate in length.
</paragraph>
<paragraph id="8BC836CE0837FFEAFF78FEF22CEF3A75" blockId="27.[151,1437,151,1184]" pageId="27" pageNumber="28">
<emphasis id="B903EADC0837FFEAFF78FEF22A1C3A0D" bold="true" box="[199,537,259,284]" pageId="27" pageNumber="28">Measurements (male: n=5)</emphasis>
: BL, 4.80–5.99 (5.63±0.47); FBL, 2.59–2.96 (2.76±0.19); HL, 0.65–0.78 (0.71±0.06); HW, 0.81–0.95 (0.89±0.06); AL, 1.46–1.76 (1.59±0.14); PL, 0.87–1.06 (0.98±0.07); PW, 1.15–1.27 (1.22±0.06); EL, 0.94–1.20 (1.03±0.11); EW, 1.03–1.71 (1.46±0.27); HTL, 0.92–1.13 (1.00±0.09).
</paragraph>
<paragraph id="8BC836CE0837FFEAFF78FE862CEE3ADE" blockId="27.[151,1437,151,1184]" pageId="27" pageNumber="28">
<emphasis id="B903EADC0837FFEAFF78FE862A2E3A99" bold="true" box="[199,555,367,392]" pageId="27" pageNumber="28">Measurements (female: n=3)</emphasis>
: BL, 5.50–6.25 (5.95±0.40); FBL, 2.73–2.89 (2.82±0.08); HL, 0.73–0.79 (0.75±0.03); HW, 0.88–0.96 (0.93±0.04); AL, 1.61–1.69 (1.64±0.04); PL, 0.94–1.06 (1.02±0.07); PW, 1.16–1.32 (1.26±0.09); EL, 1.00–1.15 (1.06±0.08); EW, 1.56–1.66 (1.63±0.06); HTL, 0.91–1.04 (0.98±0.07).
</paragraph>
</subSubSection>
<subSubSection id="C36D65450837FFEAFF78FE2D2B8A3AE5" box="[199,911,475,500]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BC836CE0837FFEAFF78FE2D2B8A3AE5" blockId="27.[151,1437,151,1184]" box="[199,911,475,500]" pageId="27" pageNumber="28">
<emphasis id="B903EADC0837FFEAFF78FE2D29653AE5" bold="true" box="[199,352,475,500]" pageId="27" pageNumber="28">Distribution.</emphasis>
[
<emphasis id="B903EADC0837FFEAFECFFE2D29CF3AE5" bold="true" box="[368,458,475,500]" pageId="27" pageNumber="28">
<collectingCountry id="F360765E0837FFEAFECFFE2D29CF3AE5" box="[368,458,475,500]" name="Japan" pageId="27" pageNumber="28">JAPAN</collectingCountry>
</emphasis>
]: Honshû (Shimane-ken). See,
<figureCitation id="134C2A4B0837FFEAFC94FE2A2B8E3AE5" box="[811,907,476,500]" captionStart="FIGURES 101 – 103" captionStartId="44.[151,264,1923,1946]" captionTargetBox="[182,1388,523,1897]" captionTargetId="figure@44.[151,1436,515,1909]" captionTargetPageId="44" captionText="FIGURES 101 – 103. Collecting localities of Aleochara (Emplenota) spp. based on specimens examined. 101. A. (E.) fucicola (Pointed by black circle); 102. A. (E.) segregata (Pointed by black square); 103. A. (E.) puetzi (Pointed by black triangle); A. (E.) hayamai (Pointed by black diamond); A. (E.) yamato (Pointed by black star)." httpUri="https://zenodo.org/record/282609/files/figure.png" pageId="27" pageNumber="28">Fig. 103</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D65450837FFEAFF78FE092BCD38B5" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8BC836CE0837FFEAFF78FE092BCD38B5" blockId="27.[151,1437,151,1184]" pageId="27" pageNumber="28">
<emphasis id="B903EADC0837FFEAFF78FE0929443909" bold="true" box="[199,321,511,536]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
This species,
<taxonomicName id="4C774D4D0837FFEAFE41FDF62AD23909" box="[510,727,512,536]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="yamato">
<emphasis id="B903EADC0837FFEAFE41FDF62AD23909" box="[510,727,512,536]" italics="true" pageId="27" pageNumber="28">Aleochara yamato</emphasis>
</taxonomicName>
, can be easily differentiated from other congeners by a combination of features as follows: very rare species (only one locality known so far); body extremely large (BL =5.79;
<figureCitation id="134C2A4B0837FFEAFF52FDB1293C394E" box="[237,313,583,607]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1918,1941]" captionTargetBox="[183,1407,300,1871]" captionTargetId="figure@5.[151,1436,279,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Japanese species of Aleochara (Emplenota) (Figs. 1 – 5, habitus). 1. Aleochara (Emplenota) fucicola; 2. A. (E.) puetzi; 3. A. (E.) hayamai; 4. A. (E.) yamato; 5. A. (E.) segregata; 6. dorsal head of A. yamato; 7. mesoventrite and metaventrite of A. fucicola in ventral view." httpUri="https://zenodo.org/record/282596/files/figure.png" pageId="27" pageNumber="28">Fig. 4</figureCitation>
); dorsal surface of forebody with prominently deep and distinct punctures; dorsal surface of impunctured area of head largely flattened but of basal part prominently and spherically swollen (see,
<figureCitation id="134C2A4B0837FFEAFAF0FD9A2D923995" box="[1359,1431,620,644]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1918,1941]" captionTargetBox="[183,1407,300,1871]" captionTargetId="figure@5.[151,1436,279,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Japanese species of Aleochara (Emplenota) (Figs. 1 – 5, habitus). 1. Aleochara (Emplenota) fucicola; 2. A. (E.) puetzi; 3. A. (E.) hayamai; 4. A. (E.) yamato; 5. A. (E.) segregata; 6. dorsal head of A. yamato; 7. mesoventrite and metaventrite of A. fucicola in ventral view." httpUri="https://zenodo.org/record/282596/files/figure.png" pageId="27" pageNumber="28">Fig. 6</figureCitation>
: arrow); head capsule transverse (HW/HL =1.25: other Japanese species ≈1.14); quite short carina on mesoventrite about 0.1 times as long as mesoventrite. [
<emphasis id="B903EADC0837FFEAFD17FD422AE139DD" bold="true" box="[680,740,692,716]" pageId="27" pageNumber="28">Male</emphasis>
]: posterior margin of tergite VIII (
<figureCitation id="134C2A4B0837FFEAFB1FFD422CFE39DD" box="[1184,1275,692,716]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="27" pageNumber="28">Fig. 46</figureCitation>
) prominently emarginated; outer margin of sternite VIII (
<figureCitation id="134C2A4B0837FFEAFD3FFD212AD739FE" box="[640,722,727,751]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="27" pageNumber="28">Fig. 48</figureCitation>
) clearly rounded in the middle; median lobe of aedeagus (
<figureCitation id="134C2A4B0837FFEAFADCFD2128DB3805" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="27" pageNumber="28">Figs. 50–51</figureCitation>
) large and elongated; subapico-ventral projections on median lobe straight, thick and robust in lateral view (
<figureCitation id="134C2A4B0837FFEAFF20FCE928F63826" box="[159,243,799,823]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="27" pageNumber="28">Fig. 50</figureCitation>
); apical lobe of median lobe long and moderately curved in lateral aspect (
<figureCitation id="134C2A4B0837FFEAFB91FCE92C873826" box="[1070,1154,799,823]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="27" pageNumber="28">Fig. 50</figureCitation>
); basal swelling (
<figureCitation id="134C2A4B0837FFEAFAFCFCE92D913826" box="[1347,1428,799,823]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="27" pageNumber="28">Fig. 50</figureCitation>
) of median lobe strongly rounded toward ventral margin as in
<taxonomicName id="4C774D4D0837FFEAFCDFFCB22BC0384D" box="[864,965,836,860]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="puetzi">
<emphasis id="B903EADC0837FFEAFCDFFCB22BC0384D" box="[864,965,836,860]" italics="true" pageId="27" pageNumber="28">A. puetzi</emphasis>
</taxonomicName>
(
<figureCitation id="134C2A4B0837FFEAFC68FCB22C2B384D" box="[983,1070,836,860]" captionStart="FIGURES 23 – 29" captionStartId="11.[151,264,1937,1960]" captionTargetBox="[180,1402,187,1870]" captionTargetId="figure@11.[151,1436,169,1916]" captionTargetPageId="11" captionText="FIGURES 23 – 29. Terminalia of Aleochara (Emplenota) puetzi. 23. tergite VIII of male; 24. tergite VIII of female; 25. sternite VIII of male; 26. sternite VIII of female; 27. male genitalia: median lobe of aedeagus in lateral view; 28. male genitalia: median lobe of aedeagus in ventral view; 29. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282599/files/figure.png" pageId="27" pageNumber="28">Fig. 27</figureCitation>
). [
<emphasis id="B903EADC0837FFEAFBF2FCB22CA1384D" bold="true" box="[1101,1188,836,860]" pageId="27" pageNumber="28">Female</emphasis>
]: posterior margin of tergite VIII (
<figureCitation id="134C2A4B0837FFEAFE96FC912978386E" box="[297,381,871,895]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="27" pageNumber="28">Fig. 47</figureCitation>
) strongly emarginated; spermatheca (
<figureCitation id="134C2A4B0837FFEAFC93FC912B85386E" box="[812,896,871,895]" captionStart="FIGURES 46 – 52" captionStartId="24.[151,264,1810,1833]" captionTargetBox="[189,1408,191,1756]" captionTargetId="figure@24.[158,1428,105,1806]" captionTargetPageId="24" captionText="FIGURES 46 – 52. Terminalia of Aleochara (Emplenota) yamato. 46. tergite VIII of male; 47. tergite VIII of female; 48. sternite VIII of male; 49. sternite VIII of female; 50. male genitalia: median lobe of aedeagus in lateral view; 51. male genitalia: median lobe of aedeagus in ventral view; 52. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282602/files/figure.png" pageId="27" pageNumber="28">Fig. 52</figureCitation>
) unique in shape; combined length of (sh) and (sn) very long, more than 4 times as apical portion of spermathecal stem.
</paragraph>
</subSubSection>
<subSubSection id="C36D65450837FFEAFF78FC462BA438D6" box="[199,929,943,968]" pageId="27" pageNumber="28" type="etymology">
<paragraph id="8BC836CE0837FFEAFF78FC462BA438D6" blockId="27.[151,1437,151,1184]" box="[199,929,943,968]" pageId="27" pageNumber="28">
<emphasis id="B903EADC0837FFEAFF78FC46294B38D9" bold="true" box="[199,334,944,968]" pageId="27" pageNumber="28">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C774D4D0837FFEAFD87FC472A8E38D9" box="[568,651,945,968]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="yamato">
<emphasis id="B903EADC0837FFEAFD87FC472A8E38D9" box="[568,651,945,968]" italics="true" pageId="27" pageNumber="28">yamato</emphasis>
</taxonomicName>
is an old name of
<collectingCountry id="F360765E0837FFEAFCE6FC462B9938D6" box="[857,924,944,967]" name="Japan" pageId="27" pageNumber="28">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D65450837FFEAFF78FC2229133F8E" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BC836CE0837FFEAFF78FC2229133F8E" blockId="27.[151,1437,151,1184]" pageId="27" pageNumber="28">
<emphasis id="B903EADC0837FFEAFF78FC22293E38FD" bold="true" box="[199,315,980,1004]" pageId="27" pageNumber="28">Remarks.</emphasis>
The species clearly differs from the other
<taxonomicName id="4C774D4D0837FFEAFC95FC222BA138FD" box="[810,932,980,1004]" class="Insecta" family="Staphylinidae" genus="Emplenota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0837FFEAFC95FC222BA138FD" box="[810,932,980,1004]" italics="true" pageId="27" pageNumber="28">Emplenota</emphasis>
</taxonomicName>
species by having a larger body, prominent spherical swelling on the dorsal surface of the head base (see,
<figureCitation id="134C2A4B0837FFEAFCCDFC012BBC3F1E" box="[882,953,1015,1039]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1918,1941]" captionTargetBox="[183,1407,300,1871]" captionTargetId="figure@5.[151,1436,279,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Japanese species of Aleochara (Emplenota) (Figs. 1 – 5, habitus). 1. Aleochara (Emplenota) fucicola; 2. A. (E.) puetzi; 3. A. (E.) hayamai; 4. A. (E.) yamato; 5. A. (E.) segregata; 6. dorsal head of A. yamato; 7. mesoventrite and metaventrite of A. fucicola in ventral view." httpUri="https://zenodo.org/record/282596/files/figure.png" pageId="27" pageNumber="28">Fig. 6</figureCitation>
), and deep punctures in the fore body. In addition, tergite VIII and sternite VIII of
<taxonomicName id="4C774D4D0837FFEAFDCEFBEA2B443F25" box="[625,833,1052,1076]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="yamato">
<emphasis id="B903EADC0837FFEAFDCEFBEA2B443F25" box="[625,833,1052,1076]" italics="true" pageId="27" pageNumber="28">Aleochara yamato</emphasis>
</taxonomicName>
are unique in both sexes. Only nine specimens have been obtained to date.
<typeStatus id="54CC886C0837FFEAFE12FBB629E13F49" box="[429,484,1088,1112]" pageId="27" pageNumber="28">Type</typeStatus>
materials were collected by Mr. T. Hayama in Shimane-ken (Honshû) as with
<taxonomicName id="4C774D4D0837FFEAFF28FB9229703F6D" box="[151,373,1124,1148]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="hayamai">
<emphasis id="B903EADC0837FFEAFF28FB9229703F6D" box="[151,373,1124,1148]" italics="true" pageId="27" pageNumber="28">Aleochara hayamai</emphasis>
</taxonomicName>
. Ecological information for this species, including detailed habitats and seasonal patterns, is not known.
</paragraph>
</subSubSection>
</treatment>
</document>
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