150 lines
17 KiB
XML
150 lines
17 KiB
XML
<document id="0DD7305EC090B261BF9DA55416C32330" ID-DOI="10.5281/zenodo.282595" ID-GBIF-Dataset="3c0b0572-9d83-492a-8656-229c8b2043a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="282595" ID-ZooBank="F832C768-A8CA-4FEE-8C3B-BD933247FA6E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460543541282" checkinUser="plazi" docAuthor="Yamamoto, Shûhei & Maruyama, Munetoshi" docDate="2012" docId="03DE87D8082FFFF2FF28FB25299F3CE4" docLanguage="en" docName="zt03517p052.pdf" docOrigin="Zootaxa 3517" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Aleocharina Fleming 1821" docType="treatment" docVersion="9" lastPageNumber="4" masterDocId="FFE7FFA0082CFFF1FFBFFFF628053B11" masterDocTitle="Revision of the Seashore-dwelling Subgenera Emplenota Casey and Triochara Bernhauer (Coleoptera: Staphylinidae: genus Aleochara) from Japan" masterLastPageNumber="52" masterPageNumber="1" pageNumber="4" updateTime="1698317426905" updateUser="plazi">
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<mods:mods id="794C81BAEFBCDB11F19E95D72F5FEB7F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="06B38DD40D4C2F3FE9C3390A85C5FA3B">
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<mods:title id="C73E85E69BFBA8898257524D6DA7E45A">Revision of the Seashore-dwelling Subgenera Emplenota Casey and Triochara Bernhauer (Coleoptera: Staphylinidae: genus Aleochara) from Japan</mods:title>
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<mods:namePart id="1F25399D323F72ABC7BCA4FCE5F97310">Yamamoto, Shûhei</mods:namePart>
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<mods:name id="73E9F27D46BF10D8A0ABD25C024037AC" type="personal">
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<mods:roleTerm id="54CDD8919CB91D50C855FDE28942C370">Author</mods:roleTerm>
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<mods:namePart id="A64A5CF17DF2F8A2E721903005590CC0">Maruyama, Munetoshi</mods:namePart>
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<mods:typeOfResource id="B790C112F545615BB0CC8A35C3BDF3E5">text</mods:typeOfResource>
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<mods:title id="444B0F7C28037B707BAF938CFA40732B">Zootaxa</mods:title>
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<mods:part id="69D212E3D6DF4EE893DFD47265F87275">
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<mods:date id="CCA9EBA2D0301823C3438AE6423F7CF9">2012</mods:date>
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<mods:detail id="BFD762D84BF984406C14BE40C7B31304" type="volume">
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<mods:number id="029A6BD0ABCB612DFF50C231875B45E6">3517</mods:number>
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<mods:start id="651B498CD19ED5FB68D76924661DAFCB">1</mods:start>
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<mods:classification id="5E1EA20AD335CEE85BF1B471D85D185B">journal article</mods:classification>
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<mods:identifier id="F967F008990F37F0D123F287858E299B" type="DOI">10.5281/zenodo.282595</mods:identifier>
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<mods:identifier id="FE9E6F06BE80A56CF2D4034AEE8B8E3B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03DE87D8082FFFF2FF28FB25299F3CE4" ID-DOI="http://doi.org/10.5281/zenodo.6175362" ID-GBIF-Taxon="119528580" ID-Zenodo-Dep="6175362" LSID="urn:lsid:plazi:treatment:03DE87D8082FFFF2FF28FB25299F3CE4" httpUri="http://treatment.plazi.org/id/03DE87D8082FFFF2FF28FB25299F3CE4" lastPageNumber="4" pageId="3" pageNumber="4">
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<subSubSection id="C36D6545082FFFF2FF28FB252BA13E5E" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="8BC836CE082FFFF2FF28FB252A593FFC" blockId="3.[151,604,1235,1261]" box="[151,604,1235,1261]" pageId="3" pageNumber="4">
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<heading id="D08081A2082FFFF2FF28FB252A593FFC" bold="true" box="[151,604,1235,1261]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<emphasis id="B903EADC082FFFF2FF28FB252A593FFC" bold="true" box="[151,604,1235,1261]" pageId="3" pageNumber="4">
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Subtribe
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<taxonomicName id="4C774D4D082FFFF2FEB4FB252A593FFC" ID-CoL="8VRBX" authority="Fleming, 1821" authorityName="Fleming" authorityYear="1821" box="[267,604,1235,1261]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subTribe" subTribe="Aleocharina">
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Aleocharina
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<bibRefCitation id="EFE64B3F082FFFF2FE14FB252A593FFC" author="Fleming" box="[427,604,1235,1261]" pageId="3" pageNumber="4" refString="Fleming, J. (1821) Insecta. Supplement to the fourth, fifth and sixth editions of the Encyclopedia Britannica, with preliminary dissertations on the history of sciences. Volume Fifth [Part 1]. A. Constable and Company, Edinburgh, pp. 41 - 56 + pl. 85." type="book chapter" year="1821">Fleming, 1821</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC836CE082FFFF2FF28FAE22BA13E5E" blockId="3.[151,1006,1300,1360]" pageId="3" pageNumber="4">
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Aleocharidae
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<bibRefCitation id="EFE64B3F082FFFF2FE8CFAE229DF3E3D" author="Fleming" box="[307,474,1300,1324]" pageId="3" pageNumber="4" refString="Fleming, J. (1821) Insecta. Supplement to the fourth, fifth and sixth editions of the Encyclopedia Britannica, with preliminary dissertations on the history of sciences. Volume Fifth [Part 1]. A. Constable and Company, Edinburgh, pp. 41 - 56 + pl. 85." type="book chapter" year="1821">Fleming, 1821</bibRefCitation>
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: 49 (
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<typeStatus id="54CC886C082FFFF2FDAFFAE22A473E3D" box="[528,578,1300,1324]" pageId="3" pageNumber="4">type</typeStatus>
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genus:
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<taxonomicName id="4C774D4D082FFFF2FD26FAE22BE33E3D" authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" box="[665,998,1300,1324]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EADC082FFFF2FD26FAE22B0B3E3D" box="[665,782,1300,1324]" italics="true" pageId="3" pageNumber="4">Aleochara</emphasis>
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<bibRefCitation id="EFE64B3F082FFFF2FCABFAE22BE33E3D" author="Gravenhorst" box="[788,998,1300,1324]" pageId="3" pageNumber="4" refString="Gravenhorst, J. L. C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. Brunsuigae, Carolus Reichard, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
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</taxonomicName>
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) Piochardiae
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<bibRefCitation id="EFE64B3F082FFFF2FE9DFACE29BE3E5E" author="Fenyes" box="[290,443,1335,1360]" pageId="3" pageNumber="4" refString="Fenyes, A. (1918 - 1921) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 a - c. M. Nijhoff, The Hague and L. Desmet-Verteneuil, Brussels, 453 pp. + 7 pls." type="book" year="1918">Fenyes, 1918</bibRefCitation>
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: 20 (
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<typeStatus id="54CC886C082FFFF2FE4EFAC12A263E5E" box="[497,547,1335,1359]" pageId="3" pageNumber="4">type</typeStatus>
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genus:
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<taxonomicName id="4C774D4D082FFFF2FDC5FACE2B9E3E5E" authority="Heyden, 1870" authorityName="Heyden" authorityYear="1870" box="[634,923,1335,1360]" class="Insecta" family="Staphylinidae" genus="Piochardia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EADC082FFFF2FDC5FACE2AF23E41" box="[634,759,1336,1360]" italics="true" pageId="3" pageNumber="4">Piochardia</emphasis>
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Heyden, 1870
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</taxonomicName>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6545082FFFF2FF28FA8F2D103D54" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="8BC836CE082FFFF2FF28FA8F2D103D54" blockId="3.[151,1436,1401,2037]" pageId="3" pageNumber="4">
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<emphasis id="B903EADC082FFFF2FF28FA8F29143E83" bold="true" box="[151,273,1401,1426]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
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The subtribe is distinguished by the following characteristics: 5-5-5 tarsal formula (4-5-
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<quantity id="4C8F9B2B082FFFF2FAADFA8F2D453E80" box="[1298,1344,1401,1425]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="4" unit="in" value="5.0">5 in</quantity>
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<taxonomicName id="4C774D4D082FFFF2FAF6FA8C291B3EA4" authority="Sharp, 1833" authorityName="Sharp" authorityYear="1833" class="Insecta" family="Staphylinidae" genus="Tinotus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EADC082FFFF2FAF6FA8C2D9F3E80" box="[1353,1434,1402,1425]" italics="true" pageId="3" pageNumber="4">Tinotus</emphasis>
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Sharp, 1833
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</taxonomicName>
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) (
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<bibRefCitation id="EFE64B3F082FFFF2FE8AFA6B29CB3EA4" author="Hanley" box="[309,462,1437,1461]" pageId="3" pageNumber="4" refString="Hanley, R. S. (2002) A new species of Mexican Tinotus from the refuse piles of Atta ants, including an annotated world catalog of Tinotus (Coleoptera: Staphylinidae: Aleocharinae: Aleocharini). The Coleopterists Bulletin, 56 (4): 453 - 471." type="journal article" year="2002">Hanley, 2002</bibRefCitation>
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;
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<bibRefCitation id="EFE64B3F082FFFF2FE66FA6B2B043EA4" author="Klimaszewski" box="[473,769,1437,1461]" pageId="3" pageNumber="4" refString="Klimaszewski, J., Pelletier, G. & Sweeney, J. (2002) Genus Tinotus (Coleoptera: Staphylinidae, Aleocharinae) from America north of Mexico: review of the types, distribution records, and key to species. The Canadian Entomologist, 134, 281 - 298." type="journal article" year="2002">
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Klimaszewski
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<emphasis id="B903EADC082FFFF2FDC1FA682AAA3EA4" box="[638,687,1437,1461]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
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., 2002
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</bibRefCitation>
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); maxillary palpus 4-segmented, plus apical pseudosegment (looks 5-segmented); labial palpus 3-segmented, plus apical pseudosegment (looks 4-segmented); ligula bilobed apically, shorter than the segment I of the labial palpus; lateral projections of labial apodeme elongated (Maruyama, 2004); median lobe of the aedeagus without dorsal apodeme (Maruyama, 2004); at least some species have a reticulated velum on the paramerite of the male aedeagus (
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<bibRefCitation id="EFE64B3F082FFFF2FCCAF9DB2C123D54" author="Seevers" box="[885,1047,1581,1605]" pageId="3" pageNumber="4" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana Zoology, 71, 1 - 289." type="journal article" year="1978">Seevers, 1978</bibRefCitation>
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; Ashe & Maus, 1998).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6545082FFFF2FF78F9A7299F3CE4" pageId="3" pageNumber="4" type="discussion">
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<paragraph id="8BC836CE082FFFF2FF78F9A72C593DE8" blockId="3.[151,1436,1401,2037]" pageId="3" pageNumber="4">
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<emphasis id="B903EADC082FFFF2FF78F9A7294A3D7B" bold="true" box="[199,335,1617,1642]" pageId="3" pageNumber="4">Comments.</emphasis>
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This subtribe is composed of 21 genera and is distributed throughout all zoogeographic regions. The subtribe
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<taxonomicName id="4C774D4D082FFFF2FE92F98329B33D9C" box="[301,438,1653,1677]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subTribe" subTribe="Aleocharina">Aleocharina</taxonomicName>
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is easily distinguished from the other two subtribes comprised of termitophiles, namely Compactopediina Kistner, 1970, and Hodoxenina Kistner, 1970, in the tribe
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<taxonomicName id="4C774D4D082FFFF2FBBDF96F2C8C3DA0" box="[1026,1161,1689,1713]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Aleocharini">Aleocharini</taxonomicName>
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, by a rather generalized body shape (with some exceptions). Recently, however, a myrmecoid genus of the subtribe
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<taxonomicName id="4C774D4D082FFFF2FAB3F94B2D9D3DC4" box="[1292,1432,1725,1749]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subTribe" subTribe="Aleocharina">Aleocharina</taxonomicName>
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,
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<emphasis id="B903EADC082FFFF2FF28F91429333DE8" box="[151,310,1762,1785]" italics="true" pageId="3" pageNumber="4">Myrmecostica</emphasis>
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Maruyama, 2011, was described from Borneo (Maruyama
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<emphasis id="B903EADC082FFFF2FC71F9142BFA3DE8" box="[974,1023,1761,1785]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
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., 2011).
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</paragraph>
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<paragraph id="8BC836CE082FFFF2FF78F8F32C5A3CBC" blockId="3.[151,1436,1401,2037]" pageId="3" pageNumber="4">
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The phylogenetic relationships within the subtribe remain uncertain. The phylogenetic concept of this subtribe is very unstable due to uncertain limits of the genus
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<taxonomicName id="4C774D4D082FFFF2FCADF8DF2B823C50" box="[786,903,1833,1857]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EADC082FFFF2FCADF8DF2B823C50" box="[786,903,1833,1857]" italics="true" pageId="3" pageNumber="4">Aleochara</emphasis>
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</taxonomicName>
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and related taxa (Ashe & Maus, 1998). The monophyly of
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<taxonomicName id="4C774D4D082FFFF2FE81F8BB29C23C74" box="[318,455,1869,1893]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subTribe" subTribe="Aleocharina">Aleocharina</taxonomicName>
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is still not established (Ashe & Maus, 1998), and no autapomorphy has been detected for it.
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<bibRefCitation id="EFE64B3F082FFFF2FF5FF88429BA3C98" box="[224,447,1905,1929]" pageId="3" pageNumber="4" refString="Kanao, T., Maruyama, M. & Hashim, R. (2011) A new genus of Compactopediina (Coleoptera: Staphylinidae) associated with Longipeditermes (Isoptera: Termitidae) in Peninsular Malaysia with a key to and phylogenetic analysis of all genera of the subtribe. Insect Systematics & Evolution, 42, 349 - 364." type="journal article">
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Kanao
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<emphasis id="B903EADC082FFFF2FE8CF88429633C98" box="[307,358,1905,1929]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
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. (2011)
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</bibRefCitation>
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showed that Compactopediina belongs to the
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<taxonomicName id="4C774D4D082FFFF2FC6BF8842CE33C98" authority="Kraatz, 1857" authorityName="Kraatz" authorityYear="1857" box="[980,1254,1905,1929]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EADC082FFFF2FC6BF8842C4D3C98" box="[980,1096,1906,1929]" italics="true" pageId="3" pageNumber="4">Tetrasticta</emphasis>
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Kraatz, 1857
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</taxonomicName>
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genus group of
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<taxonomicName id="4C774D4D082FFFF2FF28F86329253CBC" box="[151,288,1941,1965]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subTribe" subTribe="Aleocharina">Aleocharina</taxonomicName>
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(sensu
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<bibRefCitation id="EFE64B3F082FFFF2FECAF8602A463CBC" author="Maruyama" box="[373,579,1941,1966]" pageId="3" pageNumber="4" refString="Maruyama, M. (2004 a) Redescription of the genus Creochara (Coleoptera: Staphylinidae: Aleocharinae: Aleocharini) and its systematic position. The Canadian Entomologist, 136, 621 - 637." type="journal article" year="2004" yearSuffix="a">Maruyama, 2004a</bibRefCitation>
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) and suggested non-monophyly of
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<taxonomicName id="4C774D4D082FFFF2FC71F8632C5E3CBC" box="[974,1115,1941,1965]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subTribe" subTribe="Aleocharina">Aleocharina</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BC836CE082FFFF2FF78F84C299F3CE4" blockId="3.[151,1436,1401,2037]" pageId="3" pageNumber="4">
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In
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<collectingCountry id="F360765E082FFFF2FF56F84C29293CC0" box="[233,300,1978,2001]" name="Japan" pageId="3" pageNumber="4">Japan</collectingCountry>
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, 5 genera, 6 subgenera (all subgenera belong to
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<taxonomicName id="4C774D4D082FFFF2FCD7F84F2BD93CC0" box="[872,988,1977,2001]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EADC082FFFF2FCD7F84F2BD93CC0" box="[872,988,1977,2001]" italics="true" pageId="3" pageNumber="4">Aleochara</emphasis>
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</taxonomicName>
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), and 27 species of this subtribe have hitherto been recorded.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |