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<document id="8D0C0A8258D9131849E1E5E22DB45E3D" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D58520FF925991FA1654C7F984" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) viridicatum Silva, Louzada &amp; Vaz-De-Mello, 2015, new species" docType="treatment" docVersion="10" lastPageNumber="470" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="469" updateTime="1698565278120" updateUser="plazi">
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<mods:title id="A8EC09DADB9C6EE50E43F1138AA153B5">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="9AE72E24379B404B19CF8923A03EA9D3">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="6178CD2ECACA9A9DEB2607EF75C36157">Louzada, Júlio</mods:namePart>
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<mods:namePart id="840A76823856EBC7218017DC420DA1E2">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:title id="B952507DCA763883AC91C226AB2672F5">Zootaxa</mods:title>
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<mods:part id="C189F1766A9474FF7BCCDAE2FD4A63FF">
<mods:date id="02AE4DD0A05AD2F840AEE8B12F8158F2">2015</mods:date>
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<treatment id="03DE87D58520FF925991FA1654C7F984" ID-DOI="http://doi.org/10.5281/zenodo.6115712" ID-GBIF-Taxon="119598955" ID-Zenodo-Dep="6115712" LSID="urn:lsid:plazi:treatment:03DE87D58520FF925991FA1654C7F984" httpUri="http://treatment.plazi.org/id/03DE87D58520FF925991FA1654C7F984" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469">
<subSubSection id="C36D65488520FF935991FA1657CFF93F" pageId="18" pageNumber="469" type="nomenclature">
<paragraph id="8BC836C38520FF935991FA16565DF9DE" blockId="18.[151,786,1519,1579]" box="[151,786,1519,1546]" pageId="18" pageNumber="469">
<heading id="D08081AF8520FF935991FA16565DF9DE" bold="true" box="[151,786,1519,1546]" fontSize="11" level="1" pageId="18" pageNumber="469" reason="1">
<emphasis id="B903EAD18520FF935991FA16565DF9DE" bold="true" box="[151,786,1519,1546]" pageId="18" pageNumber="469">
<taxonomicName id="4C774D408520FF935991FA165734F9DD" ID-CoL="34KB9" box="[151,635,1519,1546]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="viridicatum" status="sp. nov." subGenus="Aganhyboma">
<emphasis id="B903EAD18520FF935991FA165462F9DD" bold="true" box="[151,301,1519,1545]" italics="true" pageId="18" pageNumber="469">Deltochilum</emphasis>
(
<emphasis id="B903EAD18520FF93583BFA165491F9DD" bold="true" box="[317,478,1519,1545]" italics="true" pageId="18" pageNumber="469">Aganhyboma</emphasis>
)
<emphasis id="B903EAD18520FF9358F6FA165734F9DD" bold="true" box="[496,635,1519,1545]" italics="true" pageId="18" pageNumber="469">viridicatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057AA8520FF935B84FA09565DF9DE" box="[642,786,1520,1546]" pageId="18" pageNumber="469" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836C38520FF935991F9EB54FDF9FF" blockId="18.[151,786,1519,1579]" box="[151,434,1554,1579]" pageId="18" pageNumber="469">
(
<figureCitation id="134C2A468520FF935999F9EB55A8F9FE" box="[159,231,1554,1579]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="18" pageNumber="469">Figs 7</figureCitation>
, 34, 90, 104, 105)
</paragraph>
<paragraph id="8BC836C38520FF935991F9A35647F95B" blockId="18.[151,1254,1626,1771]" pageId="18" pageNumber="469">
<bibRefCitation id="EFE64B328520FF935991F9A354D8F9A4" author="Pereira" box="[151,407,1626,1648]" pageId="18" pageNumber="469" refString="Pereira, F. S. &amp; Martinez, A. (1956) Os generos de Canthonini americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez 1956</bibRefCitation>
: 93 (comment on predation of Diplopoda; referred as
<taxonomicName id="4C774D408520FF935ABEF9A2515AF9A4" box="[952,1045,1626,1648]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="kolbei">
<emphasis id="B903EAD18520FF935ABEF9A2515AF9A4" box="[952,1045,1626,1648]" italics="true" pageId="18" pageNumber="469">D. kolbei</emphasis>
</taxonomicName>
“misidentification”)
<bibRefCitation id="EFE64B328520FF935991F9805460F95B" author="Martinez" box="[151,303,1657,1679]" pageId="18" pageNumber="469" refString="Martinez, A. (1959) Catalogo de los Scarabaeidae argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot;, 5, 1 - 126." type="journal article" year="1959">Martínez 1959</bibRefCitation>
: 52 (catalogue; cites
<bibRefCitation id="EFE64B328520FF935B05F980564EF95B" author="Pereira" box="[515,769,1657,1679]" pageId="18" pageNumber="469" refString="Pereira, F. S. &amp; Martinez, A. (1956) Os generos de Canthonini americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez 1956</bibRefCitation>
)
</paragraph>
<paragraph id="8BC836C38520FF935991F96157CFF93F" blockId="18.[151,1254,1626,1771]" pageId="18" pageNumber="469">
<bibRefCitation id="EFE64B328520FF935991F961546CF97A" author="Halffter" box="[151,291,1688,1710]" pageId="18" pageNumber="469" refString="Halffter, G. (1959) Etologia y paleontologia de Scarabaeinae. Revista Hispano-Americana de Ciencias Puras y Aplicadas, 19, 165 - 168." type="journal article" year="1959">Halffter 1959</bibRefCitation>
: 166 (comment on predation of Diplopoda; cites
<bibRefCitation id="EFE64B328520FF935A12F961515DF97A" author="Pereira" box="[788,1042,1688,1710]" pageId="18" pageNumber="469" refString="Pereira, F. S. &amp; Martinez, A. (1956) Os generos de Canthonini americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez 1956</bibRefCitation>
)
<bibRefCitation id="EFE64B328520FF935991F94F54E8F918" author="Halffter" box="[151,423,1718,1740]" pageId="18" pageNumber="469" refString="Halffter, G. &amp; Matthews, E. G. (1966) The natural history of dung beetles of the subfamily Scarabaeinae. Folia Entomologica Mexicana, 12 - 14, 1 - 312." type="book chapter" year="1966">Halffter &amp; Matthews 1966</bibRefCitation>
: 42 (comment on predation of Diplopoda; cites
<bibRefCitation id="EFE64B328520FF935A8DF94F51C6F918" author="Pereira" box="[907,1161,1718,1740]" pageId="18" pageNumber="469" refString="Pereira, F. S. &amp; Martinez, A. (1956) Os generos de Canthonini americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez 1956</bibRefCitation>
)
<bibRefCitation id="EFE64B328520FF935991F92C5446F93F" author="Cano" box="[151,265,1749,1771]" pageId="18" pageNumber="469" refString="Cano, E. B. (1998) Deltochilum valgum acropyge Bates (Coleoptera: Scarabaeidae: Scarabaeinae): habitats and distribution. The Coleopterists Bulletin, 52, 174 - 178." type="journal article" year="1998">Cano 1998</bibRefCitation>
: 176 (cites
<bibRefCitation id="EFE64B328520FF93587AF92C5735F93F" author="Pereira" box="[380,634,1749,1771]" pageId="18" pageNumber="469" refString="Pereira, F. S. &amp; Martinez, A. (1956) Os generos de Canthonini americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez 1956</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36D65488520FF935991F8EB560DF8FE" box="[151,834,1810,1835]" pageId="18" pageNumber="469" type="etymology">
<paragraph id="8BC836C38520FF935991F8EB560DF8FE" blockId="18.[151,1437,1810,2015]" box="[151,834,1810,1835]" pageId="18" pageNumber="469">
<emphasis id="B903EAD18520FF935991F8EB5451F8FF" bold="true" box="[151,286,1810,1835]" pageId="18" pageNumber="469">Etymology.</emphasis>
The green body color inspired the species name.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488520FF9259C1F8CF572EFCEC" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469" type="description">
<paragraph id="8BC836C38520FF9259C1F8CF572EFCEC" blockId="18.[151,1437,1810,2015]" lastBlockId="19.[151,1437,151,1617]" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469">
<emphasis id="B903EAD18520FF9359C1F8CF54EBF89B" bold="true" box="[199,420,1846,1871]" pageId="18" pageNumber="469">Description. Body.</emphasis>
Head dark green or black-violaceous, with weak sheen or opaque, pronotum dark brown, with dark green spots, elytra dark green (
<figureCitation id="134C2A468520FF935B7BF8A357ABF8A7" box="[637,740,1882,1907]" captionStart="FIGURES 103 – 110" captionStartId="49.[151,264,1951,1974]" captionTargetBox="[197,1421,212,1926]" captionTargetId="figure@49.[165,1421,193,1931]" captionTargetPageId="49" captionText="FIGURES 103 – 110. Species of D. (Aganhyboma). 103. D. viridescens new status. 104. D. viridicatum new species. 105. D. viridicatum new species. 106. D. acropyge new status. 107. D. feeri new species. 108. D. schefflerorum new species. 109. D. streblopodum new species. 110. D. longiceps new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244808/files/figure.png" pageId="18" pageNumber="469">Fig. 104</figureCitation>
). Ventral surface dark green, with a smooth metallic sheen. Length
<quantity id="4C8F9B268520FF9359E9F88654CAF842" box="[239,389,1919,1943]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="18" pageNumber="469" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13 to 14 mm</quantity>
.
<emphasis id="B903EAD18520FF935897F8875497F843" bold="true" box="[401,472,1918,1943]" pageId="18" pageNumber="469">Head.</emphasis>
Surface with dense foveiform punctures, not confluent, separated at center of dorsal interocular surface by approximately one or two times their diameters. Punctures of variable size; largest around eyes and smallest closer to clypeus. Anterior margin of each puncture not defined (as in Fig. 60). Margin between clypeal teeth and genae curved outward approximately at halfway of its length.
<emphasis id="B903EAD18521FF925D12FF6E513BFF64" bold="true" box="[1044,1140,151,176]" pageId="19" pageNumber="470">Thorax.</emphasis>
Surface of pronotum with small punctures, apparently simple and uniformly distributed. Punctures located on anterior angles larger, with a typical foveiform shape. Pronotum with a narrow greenish band extending the entire edge (
<figureCitation id="134C2A468521FF925DB6FF26505BFF2C" box="[1200,1300,223,248]" captionStart="FIGURES 103 – 110" captionStartId="49.[151,264,1951,1974]" captionTargetBox="[197,1421,212,1926]" captionTargetId="figure@49.[165,1421,193,1931]" captionTargetPageId="49" captionText="FIGURES 103 – 110. Species of D. (Aganhyboma). 103. D. viridescens new status. 104. D. viridicatum new species. 105. D. viridicatum new species. 106. D. acropyge new status. 107. D. feeri new species. 108. D. schefflerorum new species. 109. D. streblopodum new species. 110. D. longiceps new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244808/files/figure.png" pageId="19" pageNumber="470">Fig. 104</figureCitation>
). Margin of pronotum (close to lateral angles) straight and emarginated, without depression. Margin between lateral and posterior angles approximately straight.
<emphasis id="B903EAD18521FF925B77FEDE578BFE94" bold="true" box="[625,708,295,320]" pageId="19" pageNumber="470">Elytra.</emphasis>
Slightly elongate (
<figureCitation id="134C2A468521FF925AAFFEDE511CFE94" box="[937,1107,295,320]" captionStart="FIGURES 103 – 110" captionStartId="49.[151,264,1951,1974]" captionTargetBox="[197,1421,212,1926]" captionTargetId="figure@49.[165,1421,193,1931]" captionTargetPageId="49" captionText="FIGURES 103 – 110. Species of D. (Aganhyboma). 103. D. viridescens new status. 104. D. viridicatum new species. 105. D. viridicatum new species. 106. D. acropyge new status. 107. D. feeri new species. 108. D. schefflerorum new species. 109. D. streblopodum new species. 110. D. longiceps new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244808/files/figure.png" pageId="19" pageNumber="470">Figs 104, 105</figureCitation>
). Striae well impressed and punctate. Interstrial surface with small, dispersed foveiform punctures, amid small bright ocellate punctures, randomly distributed. First interstria brighter than rest of elytra. Basal carina of seventh interstria prominent and elongate, extending a little further than humeral callus portion (as in Fig. 71). Length of apical carina of seventh interstria not surpassing three times that of apical carina of sixth interstria.
<emphasis id="B903EAD18521FF925D2CFE4E51E9FE04" bold="true" box="[1066,1190,439,464]" pageId="19" pageNumber="470">Abdomen.</emphasis>
Punctures large and conspicuous, concentrated anterolaterally on each sternite, reduced to a narrow belt along midline of abdomen (as in Fig. 73). Pygidium with foveiform punctures denser on medial portion, near the base. Apex of pygidium weakly acuminate or rounded.
<emphasis id="B903EAD18521FF92589DFDDD5495FDE9" bold="true" box="[411,474,548,573]" pageId="19" pageNumber="470">Legs.</emphasis>
Protibia not abruptly widened. Middle protibial tooth equidistant between apical and basal teeth. Protibial spur acuminate at apex. Apical third of metatibia only slightly curved inward (as in Fig. 68), with five clearly visible longitudinal carinae. Inner margin of metatibia without longitudinal row of small tubercles. Anterior edge of profemur completely emarginate, with abundant setae in two-thirds of its length. Foveiform punctures of profemur with transverse elliptical shape. Metatarsus longer than two-thirds of metatibia length.
<emphasis id="B903EAD18521FF925991FD2E54B1FD24" bold="true" box="[151,510,727,752]" pageId="19" pageNumber="470">Secondary sexual characters.</emphasis>
<collectingCountry id="F36076538521FF925B0DFD2E570AFD24" box="[523,581,727,752]" name="Malta" pageId="19" pageNumber="470">Male</collectingCountry>
abdomen more longitudinally flattened than that of female. Males may possess medially between first and second ventrites, a small tubercle. Apical margin of pygidium longer in male. Protibial spur more acuminate in female.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488521FF9259C1FCBD5754FBE1" pageId="19" pageNumber="470" type="materials_examined">
<paragraph id="8BC836C38521FF9259C1FCBD519AFC88" blockId="19.[151,1437,151,1617]" box="[199,1237,836,861]" pageId="19" pageNumber="470">
<emphasis id="B903EAD18521FF9259C1FCBD578CFC89" bold="true" box="[199,707,836,861]" pageId="19" pageNumber="470">
Material studied.
<typeStatus id="54CC88618521FF92589DFCBD5743FC89" box="[411,524,836,861]" pageId="19" pageNumber="470" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F36076538521FF925B1EFCBD578CFC89" box="[536,707,836,861]" name="Argentina" pageId="19" pageNumber="470">ARGENTINA</collectingCountry>
</emphasis>
: MISIONES:
<collectingCountry id="F36076538521FF925A63FCBD5695FC89" box="[869,986,836,861]" name="Saint Pierre and Miquelon" pageId="19" pageNumber="470">San Pedro</collectingCountry>
,
<date id="FFC910038521FF925AE3FCBC517BFC89" box="[997,1076,836,861]" pageId="19" pageNumber="470" value="1956-01">I. 1956</date>
(
<specimenCount id="9D71FD4A8521FF925D41FCBD5120FC88" box="[1095,1135,836,860]" pageId="19" pageNumber="470" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068521FF925D70FCBD5186FC88" box="[1142,1225,836,860]" pageId="19" pageNumber="470">CEMT</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38521FF9259C1FC9E5190FC38" blockId="19.[151,1437,151,1617]" pageId="19" pageNumber="470">
<emphasis id="B903EAD18521FF9259C1FC9E5761FC54" bold="true" box="[199,558,871,896]" pageId="19" pageNumber="470">
<typeStatus id="54CC88618521FF9259C1FC9E5470FC54" box="[199,319,871,896]" pageId="19" pageNumber="470" type="paratype">Paratypes</typeStatus>
[7]:
<collectingCountry id="F36076538521FF925885FC9E5761FC54" box="[387,558,871,896]" name="Argentina" pageId="19" pageNumber="470">ARGENTINA</collectingCountry>
</emphasis>
: MISIONES: Loreto, 1955, F. H. Walz (1 ♀
<collectionCode id="ED66AE068521FF925D58FC91518FFC54" box="[1118,1216,872,896]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="470">MZUSP</collectionCode>
); Loreto,
<date id="FFC910038521FF925C3EFC9E50D7FC54" box="[1336,1432,871,896]" pageId="19" pageNumber="470" value="1958-11">XI.1958</date>
, Martínez (1 ♀
<collectionCode id="ED66AE068521FF92583AFC7554C0FC70" box="[316,399,908,932]" pageId="19" pageNumber="470">CEMT</collectionCode>
); Loreto, A. Ogloblin (
<specimenCount id="9D71FD4A8521FF925B97FC7557F6FC70" box="[657,697,908,932]" pageId="19" pageNumber="470" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068521FF925BC6FC75566DFC70" box="[704,802,908,932]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="470">MZUSP</collectionCode>
); [no indication of city],
<date id="FFC910038521FF925D30FC7551C5FC71" box="[1078,1162,908,933]" pageId="19" pageNumber="470" value="1954-02">II.1954</date>
, F. H. Walz (
<specimenCount id="9D71FD4A8521FF925C1BFC75500AFC70" box="[1309,1349,908,932]" pageId="19" pageNumber="470" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068521FF925C4AFC7550DEFC70" LSID="urn:lsid:biocol.org:col:34935" box="[1356,1425,908,932]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="19" pageNumber="470">CMN</collectionCode>
).
<emphasis id="B903EAD18521FF925991FC56544DFC1C" bold="true" box="[151,258,943,968]" pageId="19" pageNumber="470">
<collectingCountry id="F36076538521FF925991FC56544DFC1C" box="[151,258,943,968]" name="Brazil" pageId="19" pageNumber="470">BRAZIL</collectingCountry>
</emphasis>
: RIO GRANDE DO SUL: Neu Wurttemberg [currently Panambi], 1915, E. Garbe (
<specimenCount id="9D71FD4A8521FF925DF5FC49506EFC13" box="[1267,1313,944,968]" pageId="19" pageNumber="470" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068521FF925C28FC4950DFFC1C" box="[1326,1424,944,968]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="470">MZUSP</collectionCode>
); Guaíba, 1976, M.H. Galileo (1 ♀
<collectionCode id="ED66AE068521FF925B14FC2D572AFC38" box="[530,613,980,1004]" pageId="19" pageNumber="470">CEMT</collectionCode>
).
<emphasis id="B903EAD18521FF925B70FC2D564CFC39" bold="true" box="[630,771,980,1005]" pageId="19" pageNumber="470">
<collectingCountry id="F36076538521FF925B70FC2D564CFC39" box="[630,771,980,1005]" name="Uruguay" pageId="19" pageNumber="470">URUGUAY</collectingCountry>
</emphasis>
[doubtful]: Puntas, Arenal (
<specimenCount id="9D71FD4A8521FF925D47FC2D5126FC38" box="[1089,1129,980,1004]" pageId="19" pageNumber="470" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068521FF925D77FC2D519CFC38" box="[1137,1235,980,1004]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="470">MZUSP</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38521FF9259C1FC0E5754FBE1" blockId="19.[151,1437,151,1617]" pageId="19" pageNumber="470">
<emphasis id="B903EAD18521FF9259C1FC0E5465FBC4" bold="true" box="[199,298,1015,1040]" pageId="19" pageNumber="470">Habitat.</emphasis>
<collectingCountry id="F36076538521FF925832FC0E54E9FBC4" box="[308,422,1015,1040]" name="Argentina" pageId="19" pageNumber="470">Argentina</collectingCountry>
, southern
<collectingCountry id="F36076538521FF925B27FC0E572AFBC4" box="[545,613,1015,1040]" name="Brazil" pageId="19" pageNumber="470">Brazil</collectingCountry>
and
<collectingCountry id="F36076538521FF925BA2FC015649FBC4" box="[676,774,1016,1040]" name="Uruguay" pageId="19" pageNumber="470">Uruguay</collectingCountry>
[doubtful] (
<figureCitation id="134C2A468521FF925A93FC0E56A1FBC4" box="[917,1006,1015,1040]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="19" pageNumber="470">Fig. 90</figureCitation>
). Only one specimen with label data &quot;
<collectingCountry id="F36076538521FF925999FBE55593FBE0" box="[159,220,1052,1076]" name="Malta" pageId="19" pageNumber="470">Mata</collectingCountry>
de galeria&quot; [riparian forest].
</paragraph>
</subSubSection>
<subSubSection id="C36D65488521FF9259C1FBC654C7F984" pageId="19" pageNumber="470" type="diagnosis">
<paragraph id="8BC836C38521FF9259C1FBC6577EFAAC" blockId="19.[151,1437,151,1617]" pageId="19" pageNumber="470">
<emphasis id="B903EAD18521FF9259C1FBC65492FB8C" bold="true" box="[199,477,1087,1112]" pageId="19" pageNumber="470">Diagnosis and remarks</emphasis>
(within the
<taxonomicName id="4C774D408521FF925B6EFBB957AAFB8C" box="[616,741,1088,1112]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="cupreicolle">
<emphasis id="B903EAD18521FF925B6EFBB957AAFB8C" box="[616,741,1088,1112]" italics="true" pageId="19" pageNumber="470">cupreicolle</emphasis>
</taxonomicName>
complex): head dark green or black-violaceous, with a weak sheen or opaque; apex of pygidium weakly acuminate or rounded; basal carina of seventh interstria prominent and elongate, extending a little further than humeral callus portion (as in Fig. 71), though somewhat shorter and inconspicuous on the basis than in
<taxonomicName id="4C774D408521FF925B4DFB5457BCFB10" box="[587,755,1196,1220]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="viridescens">
<emphasis id="B903EAD18521FF925B4DFB5457BCFB10" box="[587,755,1196,1220]" italics="true" pageId="19" pageNumber="470">D. viridescens</emphasis>
</taxonomicName>
. The following aedeagus characters are also diagnostic: parameres shorter than phallobase, rounded at apex, in lateral view forming an angle of approximately 30° with phallobase (
<figureCitation id="134C2A468521FF925826FB0D5429FAD8" box="[288,358,1268,1293]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="19" pageNumber="470">Fig. 7</figureCitation>
); parameres more enlarged at the base in lateral view (
<figureCitation id="134C2A468521FF925AD9FB0D516AFAD8" box="[991,1061,1268,1293]" captionStart="FIGURES 4 – 11" captionStartId="38.[151,264,1745,1768]" captionTargetBox="[181,1434,225,1688]" captionTargetId="figure@38.[151,1435,195,1688]" captionTargetPageId="38" captionText="FIGURES 4 – 11. Aedeagus in Deltochilum (Aganhyboma). 4. D. titovidaurrei new species. 5. D. cupreicolle. 6. D. viridescens new status. 7. D. viridicatum new species. 8. D. amandaarcanjoae new species. 9. D. trisignatum. 10. D. cangalha new species. and D. alpercata new species. 11. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244797/files/figure.png" pageId="19" pageNumber="470">Fig. 7</figureCitation>
) than in
<taxonomicName id="4C774D408521FF925D88FB0C507FFAD8" box="[1166,1328,1268,1292]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="viridescens">
<emphasis id="B903EAD18521FF925D88FB0C507FFAD8" box="[1166,1328,1268,1292]" italics="true" pageId="19" pageNumber="470">D. viridescens</emphasis>
</taxonomicName>
. Lamella copulatrix (LC): left lobe “J”-shaped (Fig. 34); right lobe with shape similar to a human leg (Fig. 34). Frontolateral peripheral sclerite (FLP) darkly sclerotized (as in Fig. 51). Center of genital segment with two wellsclerotized portions with oval shape.
</paragraph>
<paragraph id="8BC836C38521FF9259C1FA7C5597FA31" blockId="19.[151,1437,151,1617]" pageId="19" pageNumber="470">
We examined a specimen with label data &quot;Puntas, Arenal&quot; and we believe this location is Puntas del Arenal,
<collectingCountry id="F36076538521FF925991FA5055B4FA15" box="[151,251,1449,1473]" name="Uruguay" pageId="19" pageNumber="470">Uruguay</collectingCountry>
. This specimen is black (
<figureCitation id="134C2A468521FF925B1EFA5E5735FA14" box="[536,634,1447,1472]" captionStart="FIGURES 103 – 110" captionStartId="49.[151,264,1951,1974]" captionTargetBox="[197,1421,212,1926]" captionTargetId="figure@49.[165,1421,193,1931]" captionTargetPageId="49" captionText="FIGURES 103 – 110. Species of D. (Aganhyboma). 103. D. viridescens new status. 104. D. viridicatum new species. 105. D. viridicatum new species. 106. D. acropyge new status. 107. D. feeri new species. 108. D. schefflerorum new species. 109. D. streblopodum new species. 110. D. longiceps new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244808/files/figure.png" pageId="19" pageNumber="470">Fig. 105</figureCitation>
), and the foveiform punctures on central portion of head are extremely close.
</paragraph>
<paragraph id="8BC836C38521FF9259C1FA1654C7F984" blockId="19.[151,1437,151,1617]" pageId="19" pageNumber="470">The species can show other variations in pattern of colors. Pronotum of some individuals can vary between dark and light brown or reddish. Elytra can vary between dark green and black. Ventral surface may also vary from dark green and black.</paragraph>
</subSubSection>
</treatment>
</document>
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