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<document id="D6581BB853C8B02E2A5A1509F83DAB2C" ID-DOI="10.11646/zootaxa.3716.2.7" ID-GBIF-Dataset="05daac68-c0ad-4762-b67d-39c2cce89eb1" ID-ISSN="1175-5326" ID-Zenodo-Dep="223425" ID-ZooBank="1D8EEE3F-712D-4B06-B107-06CB7B025A1B" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460681091080" checkinUser="plazi" docAuthor="Hayat, Mohammad, Zeya, Shahid Bin &amp; Veenakumari, K." docDate="2013" docId="03DE87C5FFC95E4DFF53C150FF2BB990" docLanguage="en" docName="zt03716p276.pdf" docOrigin="Zootaxa 3716 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cheiloneurus mandyanus Hayat, sp. nov." docType="treatment" docVersion="8" lastPageNumber="262" masterDocId="FFE7FFBDFFCB5E4EFFC4C371FFCEBC1C" masterDocTitle="On some brachypterous Encyrtidae (Hymenoptera: Chalcidoidea) from India, with description of four new species" masterLastPageNumber="276" masterPageNumber="259" pageNumber="261" updateTime="1698350491968" updateUser="plazi">
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<mods:title id="F7DAF4821661D6A45655C85A2A3861E2">On some brachypterous Encyrtidae (Hymenoptera: Chalcidoidea) from India, with description of four new species</mods:title>
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<mods:namePart id="3C18B8182B64D735C6F553DE6E3D888A">Hayat, Mohammad</mods:namePart>
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<mods:namePart id="6CD4E316F8CFFB531AFB0A3AF88A9CF1">Zeya, Shahid Bin</mods:namePart>
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<mods:namePart id="07D1F3BAC37702B07E68D97BCFEC779B">Veenakumari, K.</mods:namePart>
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<mods:title id="9BFC71E7F386B1C46380D16FF731DF14">Zootaxa</mods:title>
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<mods:date id="1302F579572382C9A572279C8B0E6161">2013</mods:date>
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<treatment id="03DE87C5FFC95E4DFF53C150FF2BB990" ID-DOI="http://doi.org/10.5281/zenodo.6153132" ID-GBIF-Taxon="119579862" ID-Zenodo-Dep="6153132" LSID="urn:lsid:plazi:treatment:03DE87C5FFC95E4DFF53C150FF2BB990" httpUri="http://treatment.plazi.org/id/03DE87C5FFC95E4DFF53C150FF2BB990" lastPageId="3" lastPageNumber="262" pageId="2" pageNumber="261">
<subSubSection id="C36D6558FFC95E4CFF53C150FE4BBBA9" pageId="2" pageNumber="261" type="nomenclature">
<paragraph id="8BC836D3FFC95E4CFF53C150FD41BE27" blockId="2.[151,655,545,605]" box="[151,655,545,571]" pageId="2" pageNumber="261">
<heading id="D08081BFFFC95E4CFF53C150FD41BE27" bold="true" box="[151,655,545,571]" fontSize="11" level="1" pageId="2" pageNumber="261" reason="1">
<emphasis id="B903EAC1FFC95E4CFF53C150FD41BE27" bold="true" box="[151,655,545,571]" pageId="2" pageNumber="261">
<taxonomicName id="4C774D50FFC95E4CFF53C150FDEABE27" authority="Hayat" authorityName="Hayat" box="[151,548,545,571]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="mandyanus" status="sp. nov.">
<emphasis id="B903EAC1FFC95E4CFF53C150FE01BE27" bold="true" box="[151,463,545,571]" italics="true" pageId="2" pageNumber="261">Cheiloneurus mandyanus</emphasis>
Hayat
</taxonomicName>
,
<taxonomicNameLabel id="A23057BAFFC95E4CFDF5C150FD41BE27" box="[561,655,545,571]" pageId="2" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836D3FFC95E4CFF53C135FEC4BE40" blockId="2.[151,655,545,605]" box="[151,266,580,605]" pageId="2" pageNumber="261">
(
<figureCitation id="134C2A56FFC95E4CFF5BC135FECFBE40" box="[159,257,580,605]" captionStart="FIGURES 1 3" captionStartId="3.[151,264,1010,1033]" captionTargetBox="[338,1249,282,941]" captionTargetId="figure@3.[313,1273,265,989]" captionTargetPageId="3" captionText="FIGURES 1 3. Cheiloneurus mandyanus Hayat, sp. nov. Holotype female: 1, body dorsal; 2, antenna; 3, fore wing." httpUri="https://zenodo.org/record/223426/files/figure.png" pageId="2" pageNumber="261">Figs 13</figureCitation>
)
</paragraph>
<paragraph id="8BC836D3FFC95E4CFF53C1FDFD24B864" blockId="2.[151,1436,652,2009]" pageId="2" pageNumber="261">
<emphasis id="B903EAC1FFC95E4CFF53C1FDFF3CBEB8" box="[151,242,652,676]" italics="true" pageId="2" pageNumber="261">Female.</emphasis>
<typeStatus id="54CC8871FFC95E4CFF3CC1FDFEAABEB9" box="[248,356,652,677]" pageId="2" pageNumber="261" type="holotype">Holotype</typeStatus>
. Length,
<quantity id="4C8F9B36FFC95E4CFE0FC1FDFDFCBEB8" box="[459,562,652,676]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="2" pageNumber="261" unit="mm" value="0.98">0.98 mm</quantity>
. Body yellow to pale brownish yellow (
<figureCitation id="134C2A56FFC95E4CFC2BC1FDFBFEBEB8" box="[1007,1072,652,677]" captionStart="FIGURES 1 3" captionStartId="3.[151,264,1010,1033]" captionTargetBox="[338,1249,282,941]" captionTargetId="figure@3.[313,1273,265,989]" captionTargetPageId="3" captionText="FIGURES 1 3. Cheiloneurus mandyanus Hayat, sp. nov. Holotype female: 1, body dorsal; 2, antenna; 3, fore wing." httpUri="https://zenodo.org/record/223426/files/figure.png" pageId="2" pageNumber="261">Fig. 1</figureCitation>
); head with vertex yellow, frons and malar space pale brown, shiny; inter-torular region and mouth margin pale brown. Mandible with tip black. Antenna with radicle brownish yellow; scape brownish yellow with dorsal margin reddish brown; pedicel and F1 brown; F26 yellow (appears white on slide); clava dark brown. Mesosoma brownish yellow except as follows: sides and posterior two-fifths of mesoscutum (with silvery white setae) pale yellow; scutellum brownish yellow, but with reddish tinge, and sides and apical margin reddish brown; propodeum pale brownish yellow with reddish tinge on middle third, laterally up to spiracles whitish yellow, and dark brown distal to spiracles; mesopleuron white except apical fifth or so brown. Fore wing in basal third infuscate, and with a darker infuscated streak subapically, otherwise hyaline including apex (
<figureCitation id="134C2A56FFC95E4CFD67C0DDFD28BFD8" box="[675,742,940,965]" captionStart="FIGURES 1 3" captionStartId="3.[151,264,1010,1033]" captionTargetBox="[338,1249,282,941]" captionTargetId="figure@3.[313,1273,265,989]" captionTargetPageId="3" captionText="FIGURES 1 3. Cheiloneurus mandyanus Hayat, sp. nov. Holotype female: 1, body dorsal; 2, antenna; 3, fore wing." httpUri="https://zenodo.org/record/223426/files/figure.png" pageId="2" pageNumber="261">Fig. 3</figureCitation>
).
<collectingCountry id="F3607643FFC95E4CFD38C0DDFCFABFD9" box="[764,820,940,965]" name="India" pageId="2" pageNumber="261">Hind</collectingCountry>
wing hyaline. Legs with coxae, trochanters, basal third of fore femur, and basal three-fourth of mid femur, white; rest of fore and mid femora and hind femur brownish yellow; fore tibia yellow, becoming whitish apically; mid tibia brownish yellow with base brown and apically whitish with a brown streak along basal half of dorsal margin; hind tibia dark brown, except base narrowly and apical two-fifths brownish yellow; tarsi all white, last segment of hind tarsus brown. Gaster pale brown; T1 dark brown, with purple shine; ovipositor sheaths whitish.
</paragraph>
<paragraph id="8BC836D3FFC95E4CFF03C7F5FCD9BA34" blockId="2.[151,1436,652,2009]" pageId="2" pageNumber="261">
Head (
<figureCitation id="134C2A56FFC95E4CFED0C7F5FE96B880" box="[276,344,1156,1181]" captionStart="FIGURES 1 3" captionStartId="3.[151,264,1010,1033]" captionTargetBox="[338,1249,282,941]" captionTargetId="figure@3.[313,1273,265,989]" captionTargetPageId="3" captionText="FIGURES 1 3. Cheiloneurus mandyanus Hayat, sp. nov. Holotype female: 1, body dorsal; 2, antenna; 3, fore wing." httpUri="https://zenodo.org/record/223426/files/figure.png" pageId="2" pageNumber="261">Fig. 1</figureCitation>
). Occipital margin sharp; frontovertex slightly convex, when fully exposed, head dorsal width 1.3× as broad as long (26:20) and 5.77× as broad as frontovertex width; frontovertex 0.17× head width, and 4.44× as long as broad; ocelli with apical angle acute; posterior ocelli touching eye margins, POL:OCL:AOL = 3:4:4; eyes large, antero-posterior length greater (18.5) than dorso-ventral height (14.5); malar space 0.79× eye height, and with a fine sulcus present; distance between toruli greater than width of frontovertex (5.5:4.5); frons with white setae; setae on vertex and those along eye margins brown. Antenna (
<figureCitation id="134C2A56FFC95E4CFC58C646FC2FB94C" box="[924,993,1335,1360]" captionStart="FIGURES 1 3" captionStartId="3.[151,264,1010,1033]" captionTargetBox="[338,1249,282,941]" captionTargetId="figure@3.[313,1273,265,989]" captionTargetPageId="3" captionText="FIGURES 1 3. Cheiloneurus mandyanus Hayat, sp. nov. Holotype female: 1, body dorsal; 2, antenna; 3, fore wing." httpUri="https://zenodo.org/record/223426/files/figure.png" pageId="2" pageNumber="261">Fig. 2</figureCitation>
) with scape slightly flattened, 4.35× as long as broad, and 2.64× as long as pedicel; pedicel 2.24× as long as broad and slightly shorter than F1F3 combined; F1 subquadrate, other segments all broader than long; F6 1.36× as long as and 1.85× as broad as F1; F3 F6 with longitudinal sensilla; clava large, as long as distal half of F1 and F2F6 combined and with third segment obliquely truncate.
<emphasis id="B903EAC1FFC95E4CFE47C6B9FD41B9FC" box="[387,655,1480,1504]" italics="true" pageId="2" pageNumber="261">Relative measurements</emphasis>
—head dorsal width, 26; head dorsal length (frontovertex fully exposed), 20; frontovertex width, 4.5; head frontal height, 24; eye length with head in profile, 18.5; eye height,14.5; malar space, 11.5; antennal scape length, 15.
</paragraph>
<paragraph id="8BC836D3FFC95E4CFF03C544FB57BB54" blockId="2.[151,1436,652,2009]" pageId="2" pageNumber="261">
Mesosoma (
<figureCitation id="134C2A56FFC95E4CFE90C545FE54BA50" box="[340,410,1588,1613]" captionStart="FIGURES 1 3" captionStartId="3.[151,264,1010,1033]" captionTargetBox="[338,1249,282,941]" captionTargetId="figure@3.[313,1273,265,989]" captionTargetPageId="3" captionText="FIGURES 1 3. Cheiloneurus mandyanus Hayat, sp. nov. Holotype female: 1, body dorsal; 2, antenna; 3, fore wing." httpUri="https://zenodo.org/record/223426/files/figure.png" pageId="2" pageNumber="261">Fig. 1</figureCitation>
). Mesoscutum very finely reticulate; scutellum with raised polygonal reticulations, and deeper than that on mesoscutum; collar of pronotum with a line of brown setae; mesoscutum in anterior three-fifth largely with silvery white setae, and a pair of brown setae in middle of posterior margin; axilla with a few brown setae; scutellum with short, largely pale brown setae, and apically with 3 pairs of brown setae of which the last pair longer and recurved; sides of propodeum with silvery white setae. Fore wing (
<figureCitation id="134C2A56FFC95E4CFBCDC5B5FB85BAC0" box="[1033,1099,1732,1757]" captionStart="FIGURES 1 3" captionStartId="3.[151,264,1010,1033]" captionTargetBox="[338,1249,282,941]" captionTargetId="figure@3.[313,1273,265,989]" captionTargetPageId="3" captionText="FIGURES 1 3. Cheiloneurus mandyanus Hayat, sp. nov. Holotype female: 1, body dorsal; 2, antenna; 3, fore wing." httpUri="https://zenodo.org/record/223426/files/figure.png" pageId="2" pageNumber="261">Fig. 3</figureCitation>
) 3.78× as long as broad, with apex rounded, reaching slightly beyond TII of gaster.
<collectingCountry id="F3607643FFC95E4CFD28C596FCEABB1C" box="[748,804,1767,1792]" name="India" pageId="2" pageNumber="261">Hind</collectingCountry>
wing very short, about 0.22× fore wing length and 3× as long as broad.
<emphasis id="B903EAC1FFC95E4CFE8EC47DFD9FBB38" box="[330,593,1804,1828]" italics="true" pageId="2" pageNumber="261">Relative measurements</emphasis>
— mesosoma length, 29; pronotum length (width), 3 (16); mesoscutum length (width), 10 (20.5); scutellum length (width), 12 (11); propodeum median length, 2.5.
</paragraph>
<paragraph id="8BC836D3FFC95E4CFF03C424FEB8BB8C" blockId="2.[151,1436,652,2009]" pageId="2" pageNumber="261">Metasoma slightly longer than mesosoma (35:29), elongated and apically narrowed; exserted part of ovipositor 0.14× gaster length.</paragraph>
<paragraph id="8BC836D3FFC95E4CFF03C4EDFE4BBBA9" blockId="2.[151,1436,652,2009]" box="[199,389,1948,1973]" pageId="2" pageNumber="261">
<emphasis id="B903EAC1FFC95E4CFF03C4EDFEC6BBA8" box="[199,264,1948,1972]" italics="true" pageId="2" pageNumber="261">
<collectingCountry id="F3607643FFC95E4CFF03C4EDFECABBA8" box="[199,260,1948,1972]" name="Malta" pageId="2" pageNumber="261">Male</collectingCountry>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36D6558FFC95E4DFF03C4B1FE94BCC9" lastPageId="3" lastPageNumber="262" pageId="2" pageNumber="261" type="materials_examined">
<paragraph id="8BC836D3FFC95E4DFF03C4B1FE94BCC9" blockId="2.[151,1436,652,2009]" lastBlockId="3.[151,1436,151,213]" lastPageId="3" lastPageNumber="262" pageId="2" pageNumber="261">
<emphasis id="B903EAC1FFC95E4CFF03C4B1FE78BBC5" bold="true" box="[199,438,1984,2009]" pageId="2" pageNumber="261">Material examined.</emphasis>
<typeStatus id="54CC8871FFC95E4CFE07C4B1FDE5BBC5" box="[451,555,1984,2009]" pageId="2" pageNumber="261" type="holotype">Holotype</typeStatus>
(NBAII, Registration No. ICAR/NBAII/EN8), female (on card with right antenna and wings on slide No. EH.1643) labelled “
<collectingCountry id="F3607643FFC85E4DFCC4C3E9FC98BCAC" box="[768,854,152,176]" name="India" pageId="3" pageNumber="262">INDIA</collectingCountry>
: KARNATAKA: Mandya,
<date id="FFC91013FFC85E4DFB56C3E6FAC4BCAC" box="[1170,1290,151,176]" pageId="3" pageNumber="262" value="2012-09-26">26.ix.2012</date>
(MT), Coll. K.Veenakumari”.
</paragraph>
</subSubSection>
<caption id="DF08665BFFC85E4DFF53C083FAFAB814" box="[151,1332,1010,1033]" httpUri="https://zenodo.org/record/223426/files/figure.png" pageId="3" pageNumber="262" targetBox="[338,1249,282,941]" targetPageId="3">
<paragraph id="8BC836D3FFC85E4DFF53C083FAFAB814" blockId="3.[151,1332,1010,1033]" box="[151,1332,1010,1033]" pageId="3" pageNumber="262">
<emphasis id="B903EAC1FFC85E4DFF53C083FEF4B81B" bold="true" box="[151,314,1010,1033]" pageId="3" pageNumber="262">FIGURES 13.</emphasis>
<taxonomicName id="4C774D50FFC85E4DFE84C083FD46B814" authority="Hayat" authorityName="Hayat" box="[320,648,1010,1032]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="mandyanus" status="sp. nov.">
<emphasis id="B903EAC1FFC85E4DFE84C083FD8CB814" box="[320,578,1010,1032]" italics="true" pageId="3" pageNumber="262">Cheiloneurus mandyanus</emphasis>
Hayat
</taxonomicName>
,
<emphasis id="B903EAC1FFC85E4DFD56C082FD2CB814" bold="true" box="[658,738,1011,1032]" pageId="3" pageNumber="262">
<taxonomicNameLabel id="A23057BAFFC85E4DFD56C082FD2CB814" box="[658,738,1011,1032]" pageId="3" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype female: 1, body dorsal; 2, antenna; 3, fore wing.
</paragraph>
</caption>
<subSubSection id="C36D6558FFC85E4DFF03C75EFDE4B854" box="[199,554,1071,1096]" pageId="3" pageNumber="262" type="distribution">
<paragraph id="8BC836D3FFC85E4DFF03C75EFDE4B854" blockId="3.[151,1436,1071,1420]" box="[199,554,1071,1096]" pageId="3" pageNumber="262">
<emphasis id="B903EAC1FFC85E4DFF03C75EFEAEB854" bold="true" box="[199,352,1071,1096]" pageId="3" pageNumber="262">Distribution.</emphasis>
<collectingCountry id="F3607643FFC85E4DFEA3C75EFE6BB854" box="[359,421,1071,1096]" name="India" pageId="3" pageNumber="262">India</collectingCountry>
: Karnataka.
</paragraph>
</subSubSection>
<subSubSection id="C36D6558FFC85E4DFF03C722FA46B870" box="[199,1416,1107,1132]" pageId="3" pageNumber="262" type="etymology">
<paragraph id="8BC836D3FFC85E4DFF03C722FA46B870" blockId="3.[151,1436,1071,1420]" box="[199,1416,1107,1132]" pageId="3" pageNumber="262">
<emphasis id="B903EAC1FFC85E4DFF03C722FE80B870" bold="true" box="[199,334,1107,1132]" pageId="3" pageNumber="262">Etymology.</emphasis>
The species name is derived from the locality (Mandya) from where the
<typeStatus id="54CC8871FFC85E4DFB44C722FB2CB870" box="[1152,1250,1107,1132]" pageId="3" pageNumber="262" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="C36D6558FFC85E4DFF03C706FF2BB990" pageId="3" pageNumber="262" type="discussion">
<paragraph id="8BC836D3FFC85E4DFF03C706FF2BB990" blockId="3.[151,1436,1071,1420]" pageId="3" pageNumber="262">
<emphasis id="B903EAC1FFC85E4DFF03C706FE81B88C" bold="true" box="[199,335,1143,1168]" pageId="3" pageNumber="262">Comments.</emphasis>
The genus
<taxonomicName id="4C774D50FFC85E4DFE0BC709FD2CB88C" authority="Westwood" authorityName="Westwood" box="[463,738,1143,1168]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAC1FFC85E4DFE0BC709FDA8B88C" box="[463,614,1144,1168]" italics="true" pageId="3" pageNumber="262">Cheiloneurus</emphasis>
Westwood
</taxonomicName>
contains 140 world species (Noyes, 2012). Some species exist either only in brachypterous (short winged) forms or both in brachypterous and macropterous forms (Anis &amp; Hayat 2002; Hayat 2006; Trjapitzin &amp; Triapitsyn 2008; Timberlake 1922, under
<taxonomicName id="4C774D50FFC85E4DFBF1C7B0FB21B8C4" box="[1077,1263,1217,1240]" class="Insecta" family="Encyrtidae" genus="Hypergonatopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAC1FFC85E4DFBF1C7B0FB21B8C4" box="[1077,1263,1217,1240]" italics="true" pageId="3" pageNumber="262">Hypergonatopus</emphasis>
</taxonomicName>
). The narrow frontovertex with width less than distance between antennal toruli, and absence of scutellar brush of setae excludes the new species from the brachypterous forms of species known from
<collectingCountry id="F3607643FFC85E4DFC68C676FC28B93C" box="[940,998,1287,1312]" name="India" pageId="3" pageNumber="262">India</collectingCountry>
(
<taxonomicName id="4C774D50FFC85E4DFC32C679FB7BB93C" box="[1014,1205,1288,1312]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="bangalorensis">
<emphasis id="B903EAC1FFC85E4DFC32C679FB7BB93C" box="[1014,1205,1288,1312]" italics="true" pageId="3" pageNumber="262">C. bangalorensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D50FFC85E4DFB28C679FA42B93C" box="[1260,1420,1288,1312]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="quadricolor">
<emphasis id="B903EAC1FFC85E4DFB28C679FA42B93C" box="[1260,1420,1288,1312]" italics="true" pageId="3" pageNumber="262">C quadricolor</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="4C774D50FFC85E4DFE85C65DFD98B958" authority="Timberlake" authorityName="Timberlake" box="[321,598,1323,1348]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="B903EAC1FFC85E4DFE85C65DFE75B958" box="[321,443,1324,1348]" italics="true" pageId="3" pageNumber="262">C. flavipes</emphasis>
(Timberlake)
</taxonomicName>
and
<taxonomicName id="4C774D50FFC85E4DFD54C65DFC04B958" authority="Timberlake" authorityName="Timberlake" box="[656,970,1323,1348]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="hemipterus">
<emphasis id="B903EAC1FFC85E4DFD54C65DFCE1B958" box="[656,815,1324,1348]" italics="true" pageId="3" pageNumber="262">C. hemipterus</emphasis>
(Timberlake)
</taxonomicName>
(Timberlake 1922; Guerrieri &amp; Viggiani 2005) in dimensions of head width, frontovertex width, different dimensions of flagellar segments, and body colour.
</paragraph>
</subSubSection>
</treatment>
</document>