289 lines
43 KiB
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289 lines
43 KiB
XML
<document id="95475FE2DDFC573AADAFC0662E5B8AB5" ID-DOI="10.11646/zootaxa.5016.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222275" ID-ZooBank="87DFF908-BB63-47C9-918B-E9D63261B4B1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629380528309" checkinUser="plazi" docAuthor="Fuhrmann, Juares" docDate="2021" docId="03DE8794FF8A8F6FE7E9F9C5FD496089" docLanguage="en" docName="zootaxa.5016.3.5.pdf" docOrigin="Zootaxa 5016 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zagurya Fuhrmann 2021, new genus" docType="treatment" docVersion="4" lastPageNumber="400" masterDocId="FFE7FFECFF8B8F6AE77EFFF2FF84651B" masterDocTitle="Zagurya, a new genus from Brazil with notes on claw movement in Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="406" masterPageNumber="395" pageNumber="396" updateTime="1699070514233" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FFE29C4D3C01F2F97E99B1E63E91846F">Zagurya, a new genus from Brazil with notes on claw movement in Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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<mods:name id="FE487CCB3D4DFC011CC6DA04165D51FC" type="personal">
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<mods:namePart id="AD30C90BBAE2FB9E7D51F69D5C1AED43">Fuhrmann, Juares</mods:namePart>
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<mods:title id="EBBEB19624704E9042032AD16A451BC1">Zootaxa</mods:title>
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<mods:part id="F99B0896C40ABCCD3331FE4173D38763">
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<mods:date id="161AED760E28D091375C159CC2590F15">2021</mods:date>
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<mods:number id="B7E2E31CD07CAAD01ADC7019AE05F17E">2021-08-06</mods:number>
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<mods:number id="94B230E38C9D8FA629CC17BBDB28E520">5016</mods:number>
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<mods:classification id="F64905A116965B7CFF0BC94F3A84225F">journal article</mods:classification>
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<mods:identifier id="3A7A4F29311A905718EE52289BF5FDDF" type="DOI">10.11646/zootaxa.5016.3.5</mods:identifier>
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<mods:identifier id="E7364CA58A5DC19552EAB61C8A328358" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03DE8794FF8A8F6FE7E9F9C5FD496089" ID-DOI="http://doi.org/10.5281/zenodo.5222344" ID-Zenodo-Dep="5222344" LSID="urn:lsid:plazi:treatment:03DE8794FF8A8F6FE7E9F9C5FD496089" httpUri="http://treatment.plazi.org/id/03DE8794FF8A8F6FE7E9F9C5FD496089" lastPageId="5" lastPageNumber="400" pageId="1" pageNumber="396">
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<subSubSection id="C36D6509FF8A8F6BE7E9F9C5FD99634A" box="[151,541,1591,1617]" pageId="1" pageNumber="396" type="nomenclature">
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<paragraph id="8BC83682FF8A8F6BE7E9F9C5FD99634A" blockId="1.[151,541,1591,1617]" box="[151,541,1591,1617]" pageId="1" pageNumber="396">
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<heading id="D08081EEFF8A8F6BE7E9F9C5FD99634A" bold="true" box="[151,541,1591,1617]" fontSize="11" level="1" pageId="1" pageNumber="396" reason="1">
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<emphasis id="B903EA90FF8A8F6BE7E9F9C5FD99634A" bold="true" box="[151,541,1591,1617]" pageId="1" pageNumber="396">
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<taxonomicName id="4C774D01FF8A8F6BE7E9F9C5FE0A634A" ID-CoL="8NCCQ" authority="Fuhrmann" authorityName="Fuhrmann" authorityYear="2021" box="[151,398,1591,1617]" class="Insecta" family="Scarabaeidae" genus="Zagurya" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B903EA90FF8A8F6BE7E9F9C5FF7A634A" bold="true" box="[151,254,1591,1617]" italics="true" pageId="1" pageNumber="396">Zagurya</emphasis>
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Fuhrmann
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23057EBFF8A8F6BE6E5F9C5FD99634A" box="[411,541,1591,1617]" pageId="1" pageNumber="396" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6509FF8A8F6BE7E9F989FCA56388" box="[151,801,1659,1684]" pageId="1" pageNumber="396" type="type_taxon">
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<paragraph id="8BC83682FF8A8F6BE7E9F989FCA56388" blockId="1.[151,801,1659,1684]" box="[151,801,1659,1684]" pageId="1" pageNumber="396">
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||
<emphasis id="B903EA90FF8A8F6BE7E9F989FE9A6388" bold="true" box="[151,286,1659,1683]" pageId="1" pageNumber="396">
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<typeStatus id="54CC8820FF8A8F6BE7E9F989FF486388" box="[151,204,1659,1683]" pageId="1" pageNumber="396">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="4C774D01FF8A8F6BE654F989FDF56388" authority="Frey, 1975" authorityName="Frey" authorityYear="1975" box="[298,625,1659,1684]" class="Insecta" family="Melolonthidae" genus="Plectris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="species" species="eusquamosa">
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<emphasis id="B903EA90FF8A8F6BE654F989FE786388" box="[298,508,1659,1683]" italics="true" pageId="1" pageNumber="396">Plectris eusquamosa</emphasis>
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<bibRefCitation id="EFE64B73FF8A8F6BE57CF98EFDF56388" author="Frey, G." box="[514,625,1659,1684]" pageId="1" pageNumber="396" pagination="252 - 255" refId="ref5689" refString="Frey, G. (1975) Neue Macrodactylini (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum G. Frey, 26, 252 - 255." type="journal article" year="1975">Frey, 1975</bibRefCitation>
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</taxonomicName>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6509FF8A8F6BE7E9F933FB4C628B" pageId="1" pageNumber="396" type="diagnosis">
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<paragraph id="8BC83682FF8A8F6BE7E9F933FB4C628B" blockId="1.[151,1437,1729,2007]" pageId="1" pageNumber="396">
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<emphasis id="B903EA90FF8A8F6BE7E9F933FE8E63C0" bold="true" box="[151,266,1729,1755]" pageId="1" pageNumber="396">Diagnosis</emphasis>
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. Dorsal surface with glabrous and shiny areas mixed with areas covered by scale-like setae, some sparse hair-like setae present (
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<figureCitation id="134C2A07FF8A8F6BE6E5F917FE6F63E4" box="[411,491,1765,1791]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1865,1889]" captionTargetBox="[151,1436,179,1839]" captionTargetId="figure-26@2.[151,1436,179,1839]" captionTargetPageId="2" captionText="FIGURES 1–8. Specimen and habitat of Zagurya eusquamosa (Frey, 1975), new combination. 1, Banho do Belchior, Santuário do Caraça (credits: Santuário do Caraça); 2–3, female holotype (dorsal and labels; credits: Christoph Germann); 4, head of male (left side showing posterior angle of clypeus (red) not in canthus area (green)); 5, scutellum (arrow showing the angulate contact of mesoscutum−scutellum); 6–7, male at Banho do Belchior locality (credits: Fernando Frieiro-Costa); 8, female at Refúgio do Peixe Tolo locality (credits: Frederico Falcão Salles). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222277" httpUri="https://zenodo.org/record/5222277/files/figure.png" pageId="1" pageNumber="396">Figs. 2</figureCitation>
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,
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<figureCitation id="134C2A07FF8A8F6BE689F917FDB663E4" box="[503,562,1765,1791]" captionStart="FIGURES 9–17" captionStartId="3.[151,264,1548,1572]" captionTargetBox="[151,1436,181,1524]" captionTargetId="figure-16@3.[151,1436,181,1524]" captionTargetPageId="3" captionText="FIGURES 9–17. Zagurya eusquamosa (Frey, 1975), new combination, habitus. 9–13, male (dorsal, latero-dorsal, antero-dorsal, lateral, posterior); 14–17, female (dorsal, antero-dorsal, lateral, posterior). Scale = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5222280" httpUri="https://zenodo.org/record/5222280/files/figure.png" pageId="1" pageNumber="396">9–17</figureCitation>
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); posterior angles of clypeus not forming part of canthus area (
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<figureCitation id="134C2A07FF8A8F6BE38FF917FAB263E4" box="[1265,1334,1765,1791]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1865,1889]" captionTargetBox="[151,1436,179,1839]" captionTargetId="figure-26@2.[151,1436,179,1839]" captionTargetPageId="2" captionText="FIGURES 1–8. Specimen and habitat of Zagurya eusquamosa (Frey, 1975), new combination. 1, Banho do Belchior, Santuário do Caraça (credits: Santuário do Caraça); 2–3, female holotype (dorsal and labels; credits: Christoph Germann); 4, head of male (left side showing posterior angle of clypeus (red) not in canthus area (green)); 5, scutellum (arrow showing the angulate contact of mesoscutum−scutellum); 6–7, male at Banho do Belchior locality (credits: Fernando Frieiro-Costa); 8, female at Refúgio do Peixe Tolo locality (credits: Frederico Falcão Salles). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222277" httpUri="https://zenodo.org/record/5222277/files/figure.png" pageId="1" pageNumber="396">Fig. 4</figureCitation>
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); contact of mesoscutum−scutellum angulate (
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<figureCitation id="134C2A07FF8A8F6BE542F8FBFD07623F" box="[572,643,1801,1828]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1865,1889]" captionTargetBox="[151,1436,179,1839]" captionTargetId="figure-26@2.[151,1436,179,1839]" captionTargetPageId="2" captionText="FIGURES 1–8. Specimen and habitat of Zagurya eusquamosa (Frey, 1975), new combination. 1, Banho do Belchior, Santuário do Caraça (credits: Santuário do Caraça); 2–3, female holotype (dorsal and labels; credits: Christoph Germann); 4, head of male (left side showing posterior angle of clypeus (red) not in canthus area (green)); 5, scutellum (arrow showing the angulate contact of mesoscutum−scutellum); 6–7, male at Banho do Belchior locality (credits: Fernando Frieiro-Costa); 8, female at Refúgio do Peixe Tolo locality (credits: Frederico Falcão Salles). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222277" httpUri="https://zenodo.org/record/5222277/files/figure.png" pageId="1" pageNumber="396">Fig. 5</figureCitation>
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); protibiae and mesotibiae of males wide and flat (
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<figureCitation id="134C2A07FF8A8F6BE3AAF8FBFAB06238" box="[1236,1332,1801,1827]" captionStart="FIGURES 9–17" captionStartId="3.[151,264,1548,1572]" captionTargetBox="[151,1436,181,1524]" captionTargetId="figure-16@3.[151,1436,181,1524]" captionTargetPageId="3" captionText="FIGURES 9–17. Zagurya eusquamosa (Frey, 1975), new combination, habitus. 9–13, male (dorsal, latero-dorsal, antero-dorsal, lateral, posterior); 14–17, female (dorsal, antero-dorsal, lateral, posterior). Scale = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5222280" httpUri="https://zenodo.org/record/5222280/files/figure.png" pageId="1" pageNumber="396">Figs. 11</figureCitation>
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,
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<figureCitation id="134C2A07FF8A8F6BE23CF8FBFA0A6238" box="[1346,1422,1801,1827]" captionStart="FIGURES 18–27" captionStartId="4.[151,264,1861,1885]" captionTargetBox="[161,1424,191,1828]" captionTargetId="figure-26@4.[151,1436,181,1837]" captionTargetPageId="4" captionText="FIGURES 18–27. Zagurya eusquamosa (Frey, 1975), new combination, appendages. 18–19, antennae (male, female); 20, labium, ventral (left side with setae omitted); 21–22, protibia–tarsus, anterior (male, female); 23–24, male mesotibia–tarsus (anterior, posterior; some setae omitted); 25, female mesotibia–tarsus, anterior; 26–27, metatibia–tarsus, anterior (male, female). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222282" httpUri="https://zenodo.org/record/5222282/files/figure.png" pageId="1" pageNumber="396">21, 23</figureCitation>
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); protibia with two outer teeth and with a spur (
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<figureCitation id="134C2A07FF8A8F6BE5D0F8DFFC87625C" box="[686,771,1837,1863]" captionStart="FIGURES 18–27" captionStartId="4.[151,264,1861,1885]" captionTargetBox="[161,1424,191,1828]" captionTargetId="figure-26@4.[151,1436,181,1837]" captionTargetPageId="4" captionText="FIGURES 18–27. Zagurya eusquamosa (Frey, 1975), new combination, appendages. 18–19, antennae (male, female); 20, labium, ventral (left side with setae omitted); 21–22, protibia–tarsus, anterior (male, female); 23–24, male mesotibia–tarsus (anterior, posterior; some setae omitted); 25, female mesotibia–tarsus, anterior; 26–27, metatibia–tarsus, anterior (male, female). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222282" httpUri="https://zenodo.org/record/5222282/files/figure.png" pageId="1" pageNumber="396">Fig. 21</figureCitation>
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); mesotarsomere I of males with an inner hook-like tooth (
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<figureCitation id="134C2A07FF8A8F6BE7E1F8A3FEAE6270" box="[159,298,1873,1899]" captionStart="FIGURES 18–27" captionStartId="4.[151,264,1861,1885]" captionTargetBox="[161,1424,191,1828]" captionTargetId="figure-26@4.[151,1436,181,1837]" captionTargetPageId="4" captionText="FIGURES 18–27. Zagurya eusquamosa (Frey, 1975), new combination, appendages. 18–19, antennae (male, female); 20, labium, ventral (left side with setae omitted); 21–22, protibia–tarsus, anterior (male, female); 23–24, male mesotibia–tarsus (anterior, posterior; some setae omitted); 25, female mesotibia–tarsus, anterior; 26–27, metatibia–tarsus, anterior (male, female). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222282" httpUri="https://zenodo.org/record/5222282/files/figure.png" pageId="1" pageNumber="396">Figs. 23–24</figureCitation>
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); metatarsomere IV with two well-defined groups of inner spiny setae, one of these at apex and other at proximal area (
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<figureCitation id="134C2A07FF8A8F6BE622F887FE356294" box="[348,433,1909,1935]" captionStart="FIGURES 34–52" captionStartId="7.[151,264,1775,1799]" captionTargetBox="[195,1394,191,1746]" captionTargetId="figure-16@7.[185,1402,181,1752]" captionTargetPageId="7" captionText="FIGURES 34–52. Macrodactylini and other phytophagous Scarabaeidae, distal metatarsomeres and claws, some setae omitted. 34–35, Zagurya eusquamosa (Frey, 1975), new combination (anterior, inner); 36–37, Plectris tomentosa LePeletier & Audinet-Serville, 1828 (anterior, inner); 38–39, Mallotarsus spadiceus Blanchard, 1850 (anterior, inner); 40–44, Ancistrosoma klugi Curtis, 1835; 40–41, anterior (with and without some setae); 42, inner; 43–44, internal section (with empodium protracted and retracted); 45–49, Alvarinus pallidipennis Blanchard, 1850; 45, anterior; 46, internal section; 47–49, inner (with empodium protracted, partially retracted, and fully retracted); 50–51, Hoplia argentea (Poda, 1761), Hopliini (anterior, inner); 52, Rutela lineola (Linnaeus, 1767), Rutelini, inner. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222286" httpUri="https://zenodo.org/record/5222286/files/figure.png" pageId="1" pageNumber="396">Fig. 34</figureCitation>
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); inner side of metatarsomere V with a raised carina (
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<figureCitation id="134C2A07FF8A8F6BE376F887FBE16294" box="[1032,1125,1909,1935]" captionStart="FIGURES 18–27" captionStartId="4.[151,264,1861,1885]" captionTargetBox="[161,1424,191,1828]" captionTargetId="figure-26@4.[151,1436,181,1837]" captionTargetPageId="4" captionText="FIGURES 18–27. Zagurya eusquamosa (Frey, 1975), new combination, appendages. 18–19, antennae (male, female); 20, labium, ventral (left side with setae omitted); 21–22, protibia–tarsus, anterior (male, female); 23–24, male mesotibia–tarsus (anterior, posterior; some setae omitted); 25, female mesotibia–tarsus, anterior; 26–27, metatibia–tarsus, anterior (male, female). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222282" httpUri="https://zenodo.org/record/5222282/files/figure.png" pageId="1" pageNumber="396">Figs. 27</figureCitation>
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,
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<figureCitation id="134C2A07FF8A8F6BE30FF887FB38628B" box="[1137,1212,1909,1936]" captionStart="FIGURES 34–52" captionStartId="7.[151,264,1775,1799]" captionTargetBox="[195,1394,191,1746]" captionTargetId="figure-16@7.[185,1402,181,1752]" captionTargetPageId="7" captionText="FIGURES 34–52. Macrodactylini and other phytophagous Scarabaeidae, distal metatarsomeres and claws, some setae omitted. 34–35, Zagurya eusquamosa (Frey, 1975), new combination (anterior, inner); 36–37, Plectris tomentosa LePeletier & Audinet-Serville, 1828 (anterior, inner); 38–39, Mallotarsus spadiceus Blanchard, 1850 (anterior, inner); 40–44, Ancistrosoma klugi Curtis, 1835; 40–41, anterior (with and without some setae); 42, inner; 43–44, internal section (with empodium protracted and retracted); 45–49, Alvarinus pallidipennis Blanchard, 1850; 45, anterior; 46, internal section; 47–49, inner (with empodium protracted, partially retracted, and fully retracted); 50–51, Hoplia argentea (Poda, 1761), Hopliini (anterior, inner); 52, Rutela lineola (Linnaeus, 1767), Rutelini, inner. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222286" httpUri="https://zenodo.org/record/5222286/files/figure.png" pageId="1" pageNumber="396">34–35</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6509FF8A8F6BE7B9F86BFE8262CC" pageId="1" pageNumber="396" type="etymology">
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<paragraph id="8BC83682FF8A8F6BE7B9F86BFE8262CC" blockId="1.[151,1437,1729,2007]" pageId="1" pageNumber="396">
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<emphasis id="B903EA90FF8A8F6BE7B9F86BFECC62A8" bold="true" box="[199,328,1945,1971]" pageId="1" pageNumber="396">Etymology</emphasis>
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.
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<taxonomicName id="4C774D01FF8A8F6BE628F868FE3162A8" authorityName="Fuhrmann" authorityYear="2021" box="[342,437,1946,1971]" class="Insecta" family="Scarabaeidae" genus="Zagurya" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EA90FF8A8F6BE628F868FE3162A8" box="[342,437,1946,1971]" italics="true" pageId="1" pageNumber="396">Zagurya</emphasis>
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</taxonomicName>
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is named in honor of Dr. Fernando Zagury Vaz-de-Mello (CEMT). The name is feminine in gender.
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</paragraph>
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</subSubSection>
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|
||
<paragraph id="8BC83682FF898F68E7E9F8BBFC4762EB" blockId="2.[151,1436,1865,2033]" pageId="2" pageNumber="397">
|
||
<emphasis id="B903EA90FF898F68E7E9F8BBFEB2627A" bold="true" box="[151,310,1865,1889]" pageId="2" pageNumber="397">FIGURES 1–8</emphasis>
|
||
. Specimen and habitat of
|
||
<taxonomicName id="4C774D01FF898F68E536F8B8FC29627A" authority="(Frey, 1975)" authorityName="Fuhrmann" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1975" box="[584,941,1865,1889]" class="Insecta" family="Scarabaeidae" genus="Zagurya" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="eusquamosa" status="comb. nov.">
|
||
<emphasis id="B903EA90FF898F68E536F8B8FCA7627A" box="[584,803,1866,1889]" italics="true" pageId="2" pageNumber="397">Zagurya eusquamosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B73FF898F68E44CF8BBFC21627A" author="Frey, G." box="[818,933,1865,1889]" pageId="2" pageNumber="397" pagination="252 - 255" refId="ref5689" refString="Frey, G. (1975) Neue Macrodactylini (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum G. Frey, 26, 252 - 255." type="journal article" year="1975">Frey, 1975</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B903EA90FF898F68E4C7F8BBFBF7627A" bold="true" box="[953,1139,1865,1889]" pageId="2" pageNumber="397">
|
||
<taxonomicNameLabel id="A23057EBFF898F68E4C7F8BBFBF7627A" box="[953,1139,1865,1889]" pageId="2" pageNumber="397" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B903EA90FF898F68E3FCF8BBFB0B627A" bold="true" box="[1154,1167,1865,1889]" pageId="2" pageNumber="397">1</emphasis>
|
||
, Banho do Belchior, Santuário do Caraça (credits: Santuário do Caraça);
|
||
<emphasis id="B903EA90FF898F68E507F89FFD1B629E" bold="true" box="[633,671,1901,1925]" pageId="2" pageNumber="397">2–3</emphasis>
|
||
, female holotype (dorsal and labels; credits: Christoph Germann);
|
||
<emphasis id="B903EA90FF898F68E244F89FFAC3629E" bold="true" box="[1338,1351,1901,1925]" pageId="2" pageNumber="397">4</emphasis>
|
||
, head of male (left side showing posterior angle of clypeus (
|
||
<emphasis id="B903EA90FF898F68E5E1F863FD4762B2" bold="true" box="[671,707,1937,1961]" pageId="2" pageNumber="397">red</emphasis>
|
||
) not in canthus area (
|
||
<emphasis id="B903EA90FF898F68E4E1F863FC5F62B2" bold="true" box="[927,987,1937,1961]" pageId="2" pageNumber="397">green</emphasis>
|
||
));
|
||
<emphasis id="B903EA90FF898F68E487F863FB8262B2" bold="true" box="[1017,1030,1937,1961]" pageId="2" pageNumber="397">5</emphasis>
|
||
, scutellum (arrow showing the angulate contact of mesoscutum−scutellum);
|
||
<emphasis id="B903EA90FF898F68E571F847FDB162D6" bold="true" box="[527,565,1973,1997]" pageId="2" pageNumber="397">6–7</emphasis>
|
||
, male at Banho do Belchior locality (credits: Fernando Frieiro-Costa);
|
||
<emphasis id="B903EA90FF898F68E25EF847FAA962D6" bold="true" box="[1312,1325,1973,1997]" pageId="2" pageNumber="397">8</emphasis>
|
||
, female at Refúgio do Peixe Tolo locality (credits: Frederico Falcão Salles). Scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF08660AFF888F69E7E9F9FEFBC36353" ID-DOI="http://doi.org/10.5281/zenodo.5222280" ID-Zenodo-Dep="5222280" httpUri="https://zenodo.org/record/5222280/files/figure.png" pageId="3" pageNumber="398" startId="3.[151,264,1548,1572]" targetBox="[151,1436,181,1524]" targetPageId="3">
|
||
<paragraph id="8BC83682FF888F69E7E9F9FEFBC36353" blockId="3.[151,1436,1548,1609]" pageId="3" pageNumber="398">
|
||
<emphasis id="B903EA90FF888F69E7E9F9FEFEC5633F" bold="true" box="[151,321,1548,1572]" pageId="3" pageNumber="398">FIGURES 9–17</emphasis>
|
||
.
|
||
<taxonomicName id="4C774D01FF888F69E630F9FFFD2A633F" authority="(Frey, 1975)" authorityName="Fuhrmann" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1975" box="[334,686,1548,1573]" class="Insecta" family="Scarabaeidae" genus="Zagurya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="eusquamosa" status="comb. nov.">
|
||
<emphasis id="B903EA90FF888F69E630F9FFFDA3633F" box="[334,551,1549,1572]" italics="true" pageId="3" pageNumber="398">Zagurya eusquamosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B73FF888F69E54BF9FFFD22633F" author="Frey, G." box="[565,678,1548,1573]" pageId="3" pageNumber="398" pagination="252 - 255" refId="ref5689" refString="Frey, G. (1975) Neue Macrodactylini (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum G. Frey, 26, 252 - 255." type="journal article" year="1975">Frey, 1975</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B903EA90FF888F69E5C6F9FFFCF5633F" bold="true" box="[696,881,1548,1573]" pageId="3" pageNumber="398">
|
||
<taxonomicNameLabel id="A23057EBFF888F69E5C6F9FFFCF5633F" box="[696,881,1548,1573]" pageId="3" pageNumber="398" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitus.
|
||
<emphasis id="B903EA90FF888F69E4AAF9FEFB83633F" bold="true" box="[980,1031,1548,1572]" pageId="3" pageNumber="398">9–13</emphasis>
|
||
, male (dorsal, latero-dorsal, antero-dorsal, lateral, posterior);
|
||
<emphasis id="B903EA90FF888F69E602F9C2FE3F6353" bold="true" box="[380,443,1584,1608]" pageId="3" pageNumber="398">14–17</emphasis>
|
||
, female (dorsal, antero-dorsal, lateral, posterior). Scale = 4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36D6509FF888F6FE7B9F985FD496089" lastPageId="5" lastPageNumber="400" pageId="3" pageNumber="398" type="discussion">
|
||
<paragraph id="8BC83682FF888F69E7B9F985FDCE63E6" blockId="3.[151,1437,1655,2041]" pageId="3" pageNumber="398">
|
||
<emphasis id="B903EA90FF888F69E7B9F985FEB0638A" bold="true" box="[199,308,1655,1681]" pageId="3" pageNumber="398">Remarks</emphasis>
|
||
. Within all South American Melolonthinae,
|
||
<taxonomicName id="4C774D01FF888F69E453F985FC30638A" authorityName="Blanchard" authorityYear="1850" box="[813,948,1655,1681]" class="Insecta" family="Melolonthidae" genus="Mallotarsus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA90FF888F69E453F985FC30638A" box="[813,948,1655,1681]" italics="true" pageId="3" pageNumber="398">Mallotarsus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C774D01FF888F69E493F985FBC8638B" authorityName="Fuhrmann" authorityYear="2021" box="[1005,1100,1655,1680]" class="Insecta" family="Scarabaeidae" genus="Zagurya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA90FF888F69E493F985FBC8638B" box="[1005,1100,1655,1680]" italics="true" pageId="3" pageNumber="398">Zagurya</emphasis>
|
||
</taxonomicName>
|
||
are the only genera that have mesotarsomere I of males with an inner hook-like tooth, metatarsomere V with an inner raised carina or tubercle, and all tarsi with inner distal sulci (“claws sulci”) long (length of sulci at least one-fourth of length of inner area of metatarsomere V,
|
||
<figureCitation id="134C2A07FF888F69E61FF911FDB963E6" box="[353,573,1762,1789]" captionStart="FIGURES 34–52" captionStartId="7.[151,264,1775,1799]" captionTargetBox="[195,1394,191,1746]" captionTargetId="figure-16@7.[185,1402,181,1752]" captionTargetPageId="7" captionText="FIGURES 34–52. Macrodactylini and other phytophagous Scarabaeidae, distal metatarsomeres and claws, some setae omitted. 34–35, Zagurya eusquamosa (Frey, 1975), new combination (anterior, inner); 36–37, Plectris tomentosa LePeletier & Audinet-Serville, 1828 (anterior, inner); 38–39, Mallotarsus spadiceus Blanchard, 1850 (anterior, inner); 40–44, Ancistrosoma klugi Curtis, 1835; 40–41, anterior (with and without some setae); 42, inner; 43–44, internal section (with empodium protracted and retracted); 45–49, Alvarinus pallidipennis Blanchard, 1850; 45, anterior; 46, internal section; 47–49, inner (with empodium protracted, partially retracted, and fully retracted); 50–51, Hoplia argentea (Poda, 1761), Hopliini (anterior, inner); 52, Rutela lineola (Linnaeus, 1767), Rutelini, inner. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222286" httpUri="https://zenodo.org/record/5222286/files/figure.png" pageId="3" pageNumber="398">Figs. 35, 37, 39, 42</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC83682FF888F69E7B9F8F5FBE262E2" blockId="3.[151,1437,1655,2041]" pageId="3" pageNumber="398">
|
||
<taxonomicName id="4C774D01FF888F69E7B9F8F5FEA2623B" authorityName="Fuhrmann" authorityYear="2021" box="[199,294,1799,1824]" class="Insecta" family="Scarabaeidae" genus="Zagurya" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA90FF888F69E7B9F8F5FEA2623B" box="[199,294,1799,1824]" italics="true" pageId="3" pageNumber="398">Zagurya</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C774D01FF888F69E55FF8F5FD2C623A" authorityName="Blanchard" authorityYear="1850" box="[545,680,1799,1825]" class="Insecta" family="Melolonthidae" genus="Mallotarsus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA90FF888F69E55FF8F5FD2C623A" box="[545,680,1799,1825]" italics="true" pageId="3" pageNumber="398">Mallotarsus</emphasis>
|
||
</taxonomicName>
|
||
by (characters of
|
||
<taxonomicName id="4C774D01FF888F69E40AF8F5FC7F623A" authorityName="Blanchard" authorityYear="1850" box="[884,1019,1799,1825]" class="Insecta" family="Melolonthidae" genus="Mallotarsus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA90FF888F69E40AF8F5FC7F623A" box="[884,1019,1799,1825]" italics="true" pageId="3" pageNumber="398">Mallotarsus</emphasis>
|
||
</taxonomicName>
|
||
given in parentheses): dorsal surface with groups of scale-like setae (never with scale-like setae); protibia with two outer teeth (with three teeth); protibia and mesotibia of males wide and 1.6 or two time longer than wide (long and more than three times longer than wide); mesotibia of males with one spur (with two spurs); metafemur without row of spiny setae (with a longitudinal row of spiny setae); inner area of metatarsomere IV with proximal spiny setae (without proximal spiny setae); inner distal area of metatarsomere IV with spiny setae and without spine (without spiny setae and with a spine,
|
||
<figureCitation id="134C2A07FF888F69E245F849FA0A62CE" box="[1339,1422,1979,2005]" captionStart="FIGURES 34–52" captionStartId="7.[151,264,1775,1799]" captionTargetBox="[195,1394,191,1746]" captionTargetId="figure-16@7.[185,1402,181,1752]" captionTargetPageId="7" captionText="FIGURES 34–52. Macrodactylini and other phytophagous Scarabaeidae, distal metatarsomeres and claws, some setae omitted. 34–35, Zagurya eusquamosa (Frey, 1975), new combination (anterior, inner); 36–37, Plectris tomentosa LePeletier & Audinet-Serville, 1828 (anterior, inner); 38–39, Mallotarsus spadiceus Blanchard, 1850 (anterior, inner); 40–44, Ancistrosoma klugi Curtis, 1835; 40–41, anterior (with and without some setae); 42, inner; 43–44, internal section (with empodium protracted and retracted); 45–49, Alvarinus pallidipennis Blanchard, 1850; 45, anterior; 46, internal section; 47–49, inner (with empodium protracted, partially retracted, and fully retracted); 50–51, Hoplia argentea (Poda, 1761), Hopliini (anterior, inner); 52, Rutela lineola (Linnaeus, 1767), Rutelini, inner. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222286" httpUri="https://zenodo.org/record/5222286/files/figure.png" pageId="3" pageNumber="398">Fig. 38</figureCitation>
|
||
); metatarsus with claws symmetrical (anterior claw smaller than posterior claw,
|
||
<figureCitation id="134C2A07FF888F69E378F82DFBDE62E2" box="[1030,1114,2015,2041]" captionStart="FIGURES 34–52" captionStartId="7.[151,264,1775,1799]" captionTargetBox="[195,1394,191,1746]" captionTargetId="figure-16@7.[185,1402,181,1752]" captionTargetPageId="7" captionText="FIGURES 34–52. Macrodactylini and other phytophagous Scarabaeidae, distal metatarsomeres and claws, some setae omitted. 34–35, Zagurya eusquamosa (Frey, 1975), new combination (anterior, inner); 36–37, Plectris tomentosa LePeletier & Audinet-Serville, 1828 (anterior, inner); 38–39, Mallotarsus spadiceus Blanchard, 1850 (anterior, inner); 40–44, Ancistrosoma klugi Curtis, 1835; 40–41, anterior (with and without some setae); 42, inner; 43–44, internal section (with empodium protracted and retracted); 45–49, Alvarinus pallidipennis Blanchard, 1850; 45, anterior; 46, internal section; 47–49, inner (with empodium protracted, partially retracted, and fully retracted); 50–51, Hoplia argentea (Poda, 1761), Hopliini (anterior, inner); 52, Rutela lineola (Linnaeus, 1767), Rutelini, inner. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222286" httpUri="https://zenodo.org/record/5222286/files/figure.png" pageId="3" pageNumber="398">Fig. 39</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF08660AFF8F8F6EE7E9F8B7FEAF62D2" ID-DOI="http://doi.org/10.5281/zenodo.5222282" ID-Zenodo-Dep="5222282" httpUri="https://zenodo.org/record/5222282/files/figure.png" pageId="4" pageNumber="399" startId="4.[151,264,1861,1885]" targetBox="[161,1424,191,1828]" targetPageId="4">
|
||
<paragraph id="8BC83682FF8F8F6EE7E9F8B7FEAF62D2" blockId="4.[151,1437,1861,1993]" pageId="4" pageNumber="399">
|
||
<emphasis id="B903EA90FF8F8F6EE7E9F8B7FED46246" bold="true" box="[151,336,1861,1885]" pageId="4" pageNumber="399">FIGURES 18–27</emphasis>
|
||
.
|
||
<taxonomicName id="4C774D01FF8F8F6EE61EF8B4FD436246" authority="(Frey, 1975)" authorityName="Fuhrmann" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1975" box="[352,711,1861,1885]" class="Insecta" family="Scarabaeidae" genus="Zagurya" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="399" phylum="Arthropoda" rank="species" species="eusquamosa" status="comb. nov.">
|
||
<emphasis id="B903EA90FF8F8F6EE61EF8B4FDBF6246" box="[352,571,1862,1885]" italics="true" pageId="4" pageNumber="399">Zagurya eusquamosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B73FF8F8F6EE532F8B7FD3B6246" author="Frey, G." box="[588,703,1861,1885]" pageId="4" pageNumber="399" pagination="252 - 255" refId="ref5689" refString="Frey, G. (1975) Neue Macrodactylini (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum G. Frey, 26, 252 - 255." type="journal article" year="1975">Frey, 1975</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B903EA90FF8F8F6EE5AAF8B7FC0B6246" bold="true" box="[724,911,1861,1885]" pageId="4" pageNumber="399">
|
||
<taxonomicNameLabel id="A23057EBFF8F8F6EE5AAF8B7FC0B6246" box="[724,911,1861,1885]" pageId="4" pageNumber="399" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
, appendages.
|
||
<emphasis id="B903EA90FF8F8F6EE35AF8B7FBE06246" bold="true" box="[1060,1124,1861,1885]" pageId="4" pageNumber="399">18–19</emphasis>
|
||
, antennae (male, female);
|
||
<emphasis id="B903EA90FF8F8F6EE203F8B7FA126246" bold="true" box="[1405,1430,1861,1885]" pageId="4" pageNumber="399">20</emphasis>
|
||
, labium, ventral (left side with setae omitted);
|
||
<emphasis id="B903EA90FF8F8F6EE506F89BFD33629A" bold="true" box="[632,695,1897,1921]" pageId="4" pageNumber="399">21–22</emphasis>
|
||
, protibia–tarsus, anterior (male, female);
|
||
<emphasis id="B903EA90FF8F8F6EE317F89BFB2D629A" bold="true" box="[1129,1193,1897,1921]" pageId="4" pageNumber="399">23–24</emphasis>
|
||
, male mesotibia–tarsus (anterior, posterior; some setae omitted);
|
||
<emphasis id="B903EA90FF8F8F6EE54AF87FFDC962BE" bold="true" box="[564,589,1933,1957]" pageId="4" pageNumber="399">25</emphasis>
|
||
, female mesotibia–tarsus, anterior;
|
||
<emphasis id="B903EA90FF8F8F6EE4CDF87FFC7662BE" bold="true" box="[947,1010,1933,1957]" pageId="4" pageNumber="399">26–27</emphasis>
|
||
, metatibia–tarsus, anterior (male, female). Scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF08660AFF8E8F6FE7E9FC1FFEAF61A2" ID-DOI="http://doi.org/10.5281/zenodo.5222284" ID-Zenodo-Dep="5222284" httpUri="https://zenodo.org/record/5222284/files/figure.png" pageId="5" pageNumber="400" startId="5.[151,264,1005,1029]" targetBox="[156,1430,187,976]" targetPageId="5">
|
||
<paragraph id="8BC83682FF8E8F6FE7E9FC1FFEAF61A2" blockId="5.[151,1437,1005,1210]" pageId="5" pageNumber="400">
|
||
<emphasis id="B903EA90FF8E8F6FE7E9FC1FFEC9611E" bold="true" box="[151,333,1005,1029]" pageId="5" pageNumber="400">FIGURES 28–33</emphasis>
|
||
.
|
||
<taxonomicName id="4C774D01FF8E8F6FE624FC1CFD3C611E" authority="(Frey, 1975)" authorityName="Fuhrmann" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1975" box="[346,696,1005,1030]" class="Insecta" family="Scarabaeidae" genus="Zagurya" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="species" species="eusquamosa" status="comb. nov.">
|
||
<emphasis id="B903EA90FF8E8F6FE624FC1CFDB7611E" box="[346,563,1006,1029]" italics="true" pageId="5" pageNumber="400">Zagurya eusquamosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B73FF8E8F6FE53EFC1CFD34611E" author="Frey, G." box="[576,688,1005,1030]" pageId="5" pageNumber="400" pagination="252 - 255" refId="ref5689" refString="Frey, G. (1975) Neue Macrodactylini (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum G. Frey, 26, 252 - 255." type="journal article" year="1975">Frey, 1975</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B903EA90FF8E8F6FE5BCFC1CFCFF611E" bold="true" box="[706,891,1005,1030]" pageId="5" pageNumber="400">
|
||
<taxonomicNameLabel id="A23057EBFF8E8F6FE5BCFC1CFCFF611E" box="[706,891,1005,1030]" pageId="5" pageNumber="400" rank="species">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
, appendages and terminalia.
|
||
<emphasis id="B903EA90FF8E8F6FE3DFFC1FFB64611E" bold="true" box="[1185,1248,1005,1029]" pageId="5" pageNumber="400">28–29</emphasis>
|
||
, apex of metatibia (male, female);
|
||
<emphasis id="B903EA90FF8E8F6FE649FBE3FED46132" bold="true" box="[311,336,1041,1065]" pageId="5" pageNumber="400">30</emphasis>
|
||
, genital ring;
|
||
<emphasis id="B903EA90FF8E8F6FE6A3FBE3FE726132" bold="true" box="[477,502,1041,1065]" pageId="5" pageNumber="400">31</emphasis>
|
||
, aedeagus, lateral;
|
||
<emphasis id="B903EA90FF8E8F6FE5C6FBE3FD556132" bold="true" box="[696,721,1041,1065]" pageId="5" pageNumber="400">32</emphasis>
|
||
, parameres, distal;
|
||
<emphasis id="B903EA90FF8E8F6FE4EBFBE3FC2A6132" bold="true" box="[917,942,1041,1065]" pageId="5" pageNumber="400">33</emphasis>
|
||
, female terminalia, posterior (left side with setae represented by their puncture).
|
||
<emphasis id="B903EA90FF8E8F6FE6A4FBC4FD846155" bold="true" box="[474,512,1078,1102]" pageId="5" pageNumber="400">dgc</emphasis>
|
||
, distal piece of gonocoxite;
|
||
<emphasis id="B903EA90FF8E8F6FE45BFBC4FCCF6155" bold="true" box="[805,843,1078,1102]" pageId="5" pageNumber="400">dsp</emphasis>
|
||
, distal piece of spiculum gastrale;
|
||
<emphasis id="B903EA90FF8E8F6FE3CDFBC4FB5D6155" bold="true" box="[1203,1241,1078,1102]" pageId="5" pageNumber="400">pgc</emphasis>
|
||
, proximal piece of gonocoxite;
|
||
<emphasis id="B903EA90FF8E8F6FE668FBA8FEB86169" bold="true" box="[278,316,1114,1138]" pageId="5" pageNumber="400">psp</emphasis>
|
||
, proximal piece of spiculum gastrale (apodeme);
|
||
<emphasis id="B903EA90FF8E8F6FE445FBA8FCE16169" bold="true" box="[827,869,1114,1138]" pageId="5" pageNumber="400">scm</emphasis>
|
||
, second connective membrane (connecting genital ring to aedeagus);
|
||
<emphasis id="B903EA90FF8E8F6FE65DFB8FFEC6618E" bold="true" box="[291,322,1149,1173]" pageId="5" pageNumber="400">st9</emphasis>
|
||
, abdominal sternite IX;
|
||
<emphasis id="B903EA90FF8E8F6FE549FB8CFDE4618D" bold="true" box="[567,608,1150,1174]" pageId="5" pageNumber="400">tcm</emphasis>
|
||
, third connective membrane (connecting abdominal segment VIII to genital ring). Scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC83682FF8E8F6FE7B9FB1AFD496089" blockId="5.[151,1437,1256,1426]" pageId="5" pageNumber="400">
|
||
The new genus includes a species formerly included in
|
||
<taxonomicName id="4C774D01FF8E8F6FE42FFB1AFC2C6019" authorityName="LePeletier & Audinet-Serville" authorityYear="1828" box="[849,936,1256,1282]" class="Insecta" family="Melolonthidae" genus="Plectris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA90FF8E8F6FE42FFB1AFC2C6019" box="[849,936,1256,1282]" italics="true" pageId="5" pageNumber="400">Plectris</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C774D01FF8E8F6FE4C7FB1AFB9C601A" authorityName="Fuhrmann" authorityYear="2021" box="[953,1048,1256,1281]" class="Insecta" family="Scarabaeidae" genus="Zagurya" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA90FF8E8F6FE4C7FB1AFB9C601A" box="[953,1048,1256,1281]" italics="true" pageId="5" pageNumber="400">Zagurya</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C774D01FF8E8F6FE261FB1AFAF26019" authorityName="LePeletier & Audinet-Serville" authorityYear="1828" box="[1311,1398,1256,1282]" class="Insecta" family="Melolonthidae" genus="Plectris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA90FF8E8F6FE261FB1AFAF26019" box="[1311,1398,1256,1282]" italics="true" pageId="5" pageNumber="400">Plectris</emphasis>
|
||
</taxonomicName>
|
||
by (character of
|
||
<taxonomicName id="4C774D01FF8E8F6FE650FAFEFE01603D" authorityName="LePeletier & Audinet-Serville" authorityYear="1828" box="[302,389,1292,1318]" class="Insecta" family="Melolonthidae" genus="Plectris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="400" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA90FF8E8F6FE650FAFEFE01603D" box="[302,389,1292,1318]" italics="true" pageId="5" pageNumber="400">Plectris</emphasis>
|
||
</taxonomicName>
|
||
given in parentheses): protibia and mesotibia of males wide and 1.6 or two time longer than wide (long and more than three times longer than wide); metatarsomere IV with two well-defined groups of inner spiny setae: proximal and distal (without spiny setae,
|
||
<figureCitation id="134C2A07FF8E8F6FE58CFAA6FCC16074" box="[754,837,1364,1391]" captionStart="FIGURES 34–52" captionStartId="7.[151,264,1775,1799]" captionTargetBox="[195,1394,191,1746]" captionTargetId="figure-16@7.[185,1402,181,1752]" captionTargetPageId="7" captionText="FIGURES 34–52. Macrodactylini and other phytophagous Scarabaeidae, distal metatarsomeres and claws, some setae omitted. 34–35, Zagurya eusquamosa (Frey, 1975), new combination (anterior, inner); 36–37, Plectris tomentosa LePeletier & Audinet-Serville, 1828 (anterior, inner); 38–39, Mallotarsus spadiceus Blanchard, 1850 (anterior, inner); 40–44, Ancistrosoma klugi Curtis, 1835; 40–41, anterior (with and without some setae); 42, inner; 43–44, internal section (with empodium protracted and retracted); 45–49, Alvarinus pallidipennis Blanchard, 1850; 45, anterior; 46, internal section; 47–49, inner (with empodium protracted, partially retracted, and fully retracted); 50–51, Hoplia argentea (Poda, 1761), Hopliini (anterior, inner); 52, Rutela lineola (Linnaeus, 1767), Rutelini, inner. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222286" httpUri="https://zenodo.org/record/5222286/files/figure.png" pageId="5" pageNumber="400">Fig. 36</figureCitation>
|
||
); metatarsus with claws symmetrical (symmetrical or anterior claw smaller than posterior claw,
|
||
<figureCitation id="134C2A07FF8E8F6FE513FA8AFD456089" box="[621,705,1400,1426]" captionStart="FIGURES 34–52" captionStartId="7.[151,264,1775,1799]" captionTargetBox="[195,1394,191,1746]" captionTargetId="figure-16@7.[185,1402,181,1752]" captionTargetPageId="7" captionText="FIGURES 34–52. Macrodactylini and other phytophagous Scarabaeidae, distal metatarsomeres and claws, some setae omitted. 34–35, Zagurya eusquamosa (Frey, 1975), new combination (anterior, inner); 36–37, Plectris tomentosa LePeletier & Audinet-Serville, 1828 (anterior, inner); 38–39, Mallotarsus spadiceus Blanchard, 1850 (anterior, inner); 40–44, Ancistrosoma klugi Curtis, 1835; 40–41, anterior (with and without some setae); 42, inner; 43–44, internal section (with empodium protracted and retracted); 45–49, Alvarinus pallidipennis Blanchard, 1850; 45, anterior; 46, internal section; 47–49, inner (with empodium protracted, partially retracted, and fully retracted); 50–51, Hoplia argentea (Poda, 1761), Hopliini (anterior, inner); 52, Rutela lineola (Linnaeus, 1767), Rutelini, inner. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222286" httpUri="https://zenodo.org/record/5222286/files/figure.png" pageId="5" pageNumber="400">Fig. 37</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |