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<mods:title id="2E8DC4716F13E7A698740CF852BE91B8">The cicadas (Hemiptera: Cicadidae) of Ecuador including the description of five new species, a new subtribe, four new synonymies, and fifteen new records</mods:title>
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<mods:namePart id="4D267D0F93BFAB59C73E2B21EBAFEE1C">Sanborn, Allen F.</mods:namePart>
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<treatment id="03DE878CFFD8FFF0FF3B7499D3CBF93B" ID-DOI="http://doi.org/10.5281/zenodo.4573770" ID-GBIF-Taxon="180323592" ID-Zenodo-Dep="4573770" LSID="urn:lsid:plazi:treatment:03DE878CFFD8FFF0FF3B7499D3CBF93B" httpUri="http://treatment.plazi.org/id/03DE878CFFD8FFF0FF3B7499D3CBF93B" lastPageNumber="33" pageId="32" pageNumber="33">
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<paragraph id="8BC8369AFFD8FFF0FF3B7499D200FB2D" blockId="32.[151,528,1225,1251]" box="[151,528,1225,1251]" pageId="32" pageNumber="33">
<emphasis id="B903EA88FFD8FFF0FF3B7499D0F7FB2D" bold="true" box="[151,231,1225,1251]" pageId="32" pageNumber="33">Genus</emphasis>
<taxonomicName id="4C774D19FFD8FFF0FF427499D200FB2D" ID-CoL="8HQDL" ID-ENA="1465168" authority="Distant, 1905" authorityName="Distant" authorityYear="1905" box="[238,528,1225,1251]" class="Insecta" family="Cicadidae" genus="Quesada" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFD8FFF0FF427499D146FB2D" box="[238,342,1225,1251]" italics="true" pageId="32" pageNumber="33">Quesada</emphasis>
<bibRefCitation id="EFE64B6BFFD8FFF0FEF27499D200FB2D" author="Distant, W. L." box="[350,528,1225,1251]" pageId="32" pageNumber="33" pagination="478 - 486" refId="ref69266" refString="Distant, W. L. (1905 c) Rhynchotal notes XXXII. Annals and Magazine of Natural History, Series 7, 15, 478 - 486. https: // doi. org / 10.1080 / 03745480509442837" type="journal article" year="1905">
<emphasis id="B903EA88FFD8FFF0FEF27499D200FB2D" bold="true" box="[350,528,1225,1251]" pageId="32" pageNumber="33">Distant, 1905c</emphasis>
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</taxonomicName>
</paragraph>
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<paragraph id="8BC8369AFFD8FFF0FF3B755ED1D0FAE8" blockId="32.[151,708,1294,1349]" box="[151,448,1294,1318]" pageId="32" pageNumber="33">
<treatmentCitationGroup id="AB6711B4FFD8FFF0FF3B755ED1D0FAE8" box="[151,448,1294,1318]" pageId="32" pageNumber="33">
<treatmentCitation id="0AD6108BFFD8FFF0FF3B755ED1ABFAE8" author="Distant, W. L." box="[151,443,1294,1318]" page="478" pageId="32" pageNumber="33" year="1905">
<taxonomicName id="4C774D19FFD8FFF0FF3B755ED1ABFAE8" ID-CoL="8HQDL" authority="Distant, 1905" authorityName="Distant" authorityPageNumber="478" authorityYear="1905" box="[151,443,1294,1318]" class="Insecta" family="Cicadidae" genus="Quesada" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFD8FFF0FF3B755ED0E1FAE8" box="[151,241,1294,1318]" italics="true" pageId="32" pageNumber="33">Quesada</emphasis>
<bibRefCitation id="EFE64B6BFFD8FFF0FF54755ED1ABFAE8" author="Distant, W. L." box="[248,443,1294,1318]" pageId="32" pageNumber="33" pagination="478 - 486" refId="ref69266" refString="Distant, W. L. (1905 c) Rhynchotal notes XXXII. Annals and Magazine of Natural History, Series 7, 15, 478 - 486. https: // doi. org / 10.1080 / 03745480509442837" type="journal article" year="1905">Distant 1905c: 478</bibRefCitation>
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.
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</paragraph>
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<typeStatus id="54CC8838FFD8FFF0FF3B757DD0D6FA8C" box="[151,198,1325,1348]" pageId="32" pageNumber="33">TYPE</typeStatus>
SPECIES.
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<taxonomicName id="4C774D19FFD8FFF0FE9D757CD265FA8A" ID-CoL="V5RG" authority="Olivier, 1790" authorityName="Olivier" authorityPageNumber="750" authorityYear="1790" box="[305,629,1324,1349]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B903EA88FFD8FFF0FE9D757CD1A8FA8A" box="[305,440,1324,1348]" italics="true" pageId="32" pageNumber="33">Cicada gigas</emphasis>
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<bibRefCitation id="EFE64B6BFFD8FFF0FE12757CD265FA8A" author="Olivier, G. A." box="[446,629,1324,1349]" pageId="32" pageNumber="33" pagination="735 - 760" refId="ref72065" refString="Olivier, G. A. (1790) Cigale, Cicada. Encyclopedie Methodique Histoire Naturelles Insectes, 5, 735 - 760." type="journal article" year="1790">Olivier 1790: 750</bibRefCitation>
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. (Java).
</paragraph>
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REMARKS.—The genus was recently reassigned to
<taxonomicName id="4C774D19FFD8FFF0FD787523D351FA43" authorityName="Distant" authorityYear="1905" box="[724,833,1395,1421]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="tribe" tribe="Fidicinini">Fidicinini</taxonomicName>
with the synonymy of the
<taxonomicName id="4C774D19FFD8FFF0FBD57523D0CCFA7F" authority="(Marshall et al. 2018)" baseAuthorityName="Marshall" baseAuthorityYear="2018" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="tribe" tribe="Hyantiini">
Hyantiini (
<bibRefCitation id="EFE64B6BFFD8FFF0FB587523D0C4FA7F" author="Marshall, D. C. &amp; Moulds, M. S. &amp; Hill, K. B. R. &amp; Price, B. W. &amp; Wade, E. J. &amp; Owen, C. J. &amp; Goemans, G. &amp; Marathe, K. &amp; Sarkar, V. &amp; Cooley, J. R. &amp; Sanborn, A. F. &amp; Kunte, K. &amp; Villet, M. H. &amp; Simon, C." pageId="32" pageNumber="33" pagination="1 - 64" refId="ref71376" refString="Marshall, D. C., Moulds, M. S., Hill, K. B. R., Price, B. W., Wade, E. J., Owen, C. J., Goemans, G., Marathe, K., Sarkar, V., Cooley, J. R., Sanborn, A. F., Kunte, K., Villet, M. H. &amp; Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily level classification. Zootaxa, 4424 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 4424.1.1" type="journal article" year="2018">
Marshall
<emphasis id="B903EA88FFD8FFF0FACE7523D58DFA43" box="[1378,1437,1395,1421]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
. Only
<taxonomicName id="4C774D19FFD8FFF0FE8B75C7D19BFA7F" authorityName="Distant" authorityYear="1905" box="[295,395,1431,1457]" class="Insecta" family="Cicadidae" genus="Quesada" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFD8FFF0FE8B75C7D19BFA7F" box="[295,395,1431,1457]" italics="true" pageId="32" pageNumber="33">Quesada</emphasis>
</taxonomicName>
in the Ecuadorian
<taxonomicName id="4C774D19FFD8FFF0FDCC75C7D2DDFA7F" authorityName="Distant" authorityYear="1905" box="[608,717,1431,1457]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="tribe" tribe="Fidicinini">Fidicinini</taxonomicName>
have small timbal covers that recurve for only part of its length with an anterior triangular extension and do not cover the timbal. Specimens of
<taxonomicName id="4C774D19FFD8FFF0FBB475EBD46CFA1B" authorityName="Distant" authorityYear="1905" box="[1048,1148,1467,1493]" class="Insecta" family="Cicadidae" genus="Quesada" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA88FFD8FFF0FBB475EBD46CFA1B" box="[1048,1148,1467,1493]" italics="true" pageId="32" pageNumber="33">Quesada</emphasis>
</taxonomicName>
can also be distinguished by the larger body size (body lengths greater than
<quantity id="4C8F9B7FFFD8FFF0FD6D758FD303FA37" box="[705,787,1503,1529]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="33" unit="mm" value="30.0">30 mm</quantity>
), the basal cell of the fore wing is only slightly longer than broad, the abdomen is about as long as the distance between the anterior head and posterior of the cruciform elevation, and the distal terminus of the pygofer angles medially.
</paragraph>
</subSubSection>
<subSubSection id="C36D6511FFD8FFF0FF6B761BD3CBF93B" pageId="32" pageNumber="33" type="distribution">
<paragraph id="8BC8369AFFD8FFF0FF6B761BD3CBF93B" blockId="32.[151,1437,1395,1781]" pageId="32" pageNumber="33">
DISTRIBUTION.—The two species of the genus have been reported from the Antilles,
<collectingCountry id="F360760AFFD8FFF0FBC3761BD4F1F9AB" box="[1135,1249,1611,1637]" name="Argentina" pageId="32" pageNumber="33">Argentina</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FB41761BD527F9AB" box="[1261,1335,1611,1637]" name="Belize" pageId="32" pageNumber="33">Belize</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FAEF761BD588F9AB" box="[1347,1432,1611,1637]" name="Bolivia" pageId="32" pageNumber="33">Bolivia</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FF3B763FD0CEF947" box="[151,222,1647,1673]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FF47763FD14CF947" box="[235,348,1647,1673]" name="Colombia" pageId="32" pageNumber="33">Colombia</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FEC4763FD1F6F947" box="[360,486,1647,1673]" name="Costa Rica" pageId="32" pageNumber="33">Costa Rica</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FE5F763FD242F947" box="[499,594,1647,1673]" name="Ecuador" pageId="32" pageNumber="33">Ecuador</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FDF2763FD2F5F947" box="[606,741,1647,1673]" name="El Salvador" pageId="32" pageNumber="33">El Salvador</collectingCountry>
, French Guiana,
<collectingCountry id="F360760AFFD8FFF0FC0B763FD433F947" box="[935,1059,1647,1673]" name="Guatemala" pageId="32" pageNumber="33">Guatemala</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FB9C763FD49AF947" box="[1072,1162,1647,1673]" name="Guyana" pageId="32" pageNumber="33">Guyana</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FB3A763FD516F947" box="[1174,1286,1647,1673]" name="Honduras" pageId="32" pageNumber="33">Honduras</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FABE763FD57AF947" box="[1298,1386,1647,1673]" name="Mexico" pageId="32" pageNumber="33">Mexico</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FADB763FD0E1F963" name="Nicaragua" pageId="32" pageNumber="33">Nicaragua</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FF5076C3D148F963" box="[252,344,1683,1709]" name="Panama" pageId="32" pageNumber="33">Panama</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FECE76C3D1DBF963" box="[354,459,1683,1709]" name="Paraguay" pageId="32" pageNumber="33">Paraguay</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FE7A76C3D21CF963" box="[470,524,1683,1709]" name="Peru" pageId="32" pageNumber="33">Peru</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FDBA76C3D2FEF963" box="[534,750,1683,1709]" name="Trinidad and Tobago" pageId="32" pageNumber="33">Trinidad &amp; Tobago</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FD5576C3D405F963" box="[761,1045,1683,1709]" name="United States of America" pageId="32" pageNumber="33">United States of America</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FB8C76C3D493F963" box="[1056,1155,1683,1709]" name="Uruguay" pageId="32" pageNumber="33">Uruguay</collectingCountry>
,
<collectingCountry id="F360760AFFD8FFF0FB2176C3D513F963" box="[1165,1283,1683,1709]" name="Venezuela" pageId="32" pageNumber="33">Venezuela</collectingCountry>
, and the West Indies (
<bibRefCitation id="EFE64B6BFFD8FFF0FF4776E7D183F91F" author="Metcalf, Z. P." box="[235,403,1719,1745]" pageId="32" pageNumber="33" pagination="1 - 585" refId="ref71633" refString="Metcalf, Z. P. (1963 a) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae. North Carolina State College Contribution, 1502, i-vii + 1 - 585." type="journal article" year="1963">Metcalf 1963a</bibRefCitation>
;
<bibRefCitation id="EFE64B6BFFD8FFF0FE3376E7D2FEF91F" author="Duffels, J. P. &amp; van der Laan, P. A." box="[415,750,1718,1745]" pageId="32" pageNumber="33" refId="ref69806" refString="Duffels, J. P. &amp; van der Laan, P. A. (1985) Catalogue of the Cicadoidea (Homoptera, Auchenorhyncha) 1956 - 1980. Series Entomologica 34. Dr. W. Junk Publishers, Dordrect, 414 pp." type="book" year="1985">Duffels &amp; van der Laan 1985</bibRefCitation>
; Sanborn 2011b; 2013; 2014a; 2018c; 2019b; 2020b, e;
<bibRefCitation id="EFE64B6BFFD8FFF0FAC576E7D164F93B" author="Sanborn, A. F. &amp; Heath, M. S. &amp; Phillips, P. K. &amp; Heath, J. E." pageId="32" pageNumber="33" pagination="923 - 941" refId="ref74754" refString="Sanborn, A. F., Heath, M. S., Phillips, P. K. &amp; Heath, J. E. (2012) The genus Pacarina Distant, 1905 (Hemiptera: Cicadidae) with the description of a new species. Journal of Natural History, 46, 923 - 941. https: // doi. org / 10.1080 / 00222933.2011.651647" type="journal article" year="2012">Sanborn &amp; Heath 2012</bibRefCitation>
; 2014; 2017;
<bibRefCitation id="EFE64B6BFFD8FFF0FDA1768BD2AFF93B" author="Reis, P. R. &amp; Rebelles, P. P. R. &amp; Pereira, M. C. &amp; Liska, G. R. &amp; Morais, A. R. de" box="[525,703,1754,1781]" pageId="32" pageNumber="33" pagination="527 - 536" refId="ref72195" refString="Reis, P. R., Rebelles, P. P. R., Pereira, M. C., Liska, G. R. &amp; Morais, A. R. de (2015) Eficacia do enxofre aplicado via solo no controle da cigarra Quesada gigas (Olivier) em cafeeiro. Coffee Science, Lavras, 10, 527 - 536." type="journal article" year="2015">
Reis
<emphasis id="B903EA88FFD8FFF0FDE9768BD26EF93B" box="[581,638,1755,1781]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="EFE64B6BFFD8FFF0FD67768BD3DFF93B" author="Maccagnan, D. H. B. &amp; Matuo, T. K. &amp; Martinelli, N. M." box="[715,975,1755,1781]" pageId="32" pageNumber="33" pagination="1 - 4" refId="ref71178" refString="Maccagnan, D. H. B., Matuo, T. K. &amp; Martinelli, N. M. (2017) Oviposition of Quesada gigas in weed no hostess: implication in pest management. Ciencias Rurali, 47 (09), e 20101752, 1 - 4. https: // doi. org / 10.1590 / 0103 - 8478 cr 20170152" type="journal article" year="2017">
Maccagnan
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).
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