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<document id="F2C3458B63E6C5152ED9BF02FEC3049B" ID-DOI="10.1649/0010-065x-69.3.463" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106050" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630175225603" checkinUser="diego" docAuthor="Filippini, V., Galante, E. &amp; Micó, E." docDate="2015" docId="03CFA3486B61FFCBFF1CFC965863FDA1" docLanguage="en" docName="0010-065X-69.3.463.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-69.3.463" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Anomala divisa Filippini, Galante, and Mico, new species" docType="treatment" docVersion="4" lastPageNumber="471" masterDocId="FFF6DB306B66FFC3FFBEFFE95A73FFC7" masterDocTitle="Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica" masterLastPageNumber="476" masterPageNumber="463" pageNumber="470" updateTime="1699636354092" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E7AF23A0726DBBD4442A677DCA2D2C52" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="63ABF36E77EDFE3695E7DCFFE9D63D66">
<mods:title id="3B4E76CF474BBB556A46EA73C889A1CB">Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="EA0F955072D945A1850B9D862CCF607B" type="personal">
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<mods:namePart id="84B597002F2A40AFF07A06395F3E1951">Filippini, V.</mods:namePart>
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<mods:name id="E41B17A38D37507BDB01EFDE5D06BE09" type="personal">
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<mods:roleTerm id="CCEF4BFA0B075DDC1D077CA46E2FFACF">Author</mods:roleTerm>
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<mods:namePart id="B2C79528B63595B37A6753B9448AD207">Galante, E.</mods:namePart>
</mods:name>
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<mods:role id="6D302C7254F973738CF67C9BB3C53728">
<mods:roleTerm id="EC67E978F7B049F8A095A879B7BF7FD5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6BC23106E69ABF4130BDE48DC5F87A9C">Micó, E.</mods:namePart>
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<mods:typeOfResource id="2F36AED6B483C9CFA9C4FAC58C38E920">text</mods:typeOfResource>
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<mods:title id="DD6C661B4A3373A3C77C254AD1E95481">The Coleopterists Bulletin</mods:title>
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<mods:part id="7CA484AB1F4E1162EAA18E4DAC7D4A70">
<mods:date id="FD126B49416454F15191D26EBBAA0FC0">2015</mods:date>
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<mods:number id="E61BAD89D055A0F5F51322FA288776EF">2015-09-30</mods:number>
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<mods:number id="3FFD83BE006A54DA3DB9397983DF5E20">69</mods:number>
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<mods:number id="E6B234E804A8708D22801593EDB95EF6">3</mods:number>
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<mods:url id="7A02DE6DF91918692106F1B4627A703A">http://dx.doi.org/10.1649/0010-065x-69.3.463</mods:url>
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<mods:classification id="56ECACE8BDC2E9743D1457633D4A23CD">journal article</mods:classification>
<mods:identifier id="A307BC6D2FA63EB18718F85B358F5370" type="DOI">10.1649/0010-065x-69.3.463</mods:identifier>
<mods:identifier id="6AF43C5B33E843A34D72B1DA58C4EC63" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03CFA3486B61FFCBFF1CFC965863FDA1" ID-DOI="http://doi.org/10.5281/zenodo.5413167" ID-Zenodo-Dep="5413167" LSID="urn:lsid:plazi:treatment:03CFA3486B61FFCBFF1CFC965863FDA1" httpUri="http://treatment.plazi.org/id/03CFA3486B61FFCBFF1CFC965863FDA1" lastPageId="8" lastPageNumber="471" pageId="7" pageNumber="470">
<subSubSection id="C37C41D56B61FFC4FF1CFC965BB6FC68" pageId="7" pageNumber="470" type="nomenclature">
<paragraph id="8BD9125E6B61FFC4FF1CFC965BB6FC68" blockId="7.[162,628,895,970]" pageId="7" pageNumber="470">
<heading id="D091A5326B61FFC4FF1CFC965BB6FC68" centered="true" fontSize="10" level="2" pageId="7" pageNumber="470" reason="2">
<emphasis id="B912CE4C6B61FFC4FF1CFC965BB6FC68" bold="true" pageId="7" pageNumber="470">
<taxonomicName id="4C6669DD6B61FFC4FF1CFC96581DFC53" authority="Filippini, Galante, and Mico" authorityName="Filippini, Galante, and Mico" box="[162,622,895,917]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="divisa" status="sp. nov.">
<emphasis id="B912CE4C6B61FFC4FF1CFC965B4FFC52" bold="true" box="[162,316,895,917]" italics="true" pageId="7" pageNumber="470">Anomala divisa</emphasis>
Filippini, Galante, and Micó
</taxonomicName>
,
<taxonomicNameLabel id="A22173376B61FFC4FEF1FC735BB6FC68" box="[335,453,922,943]" pageId="7" pageNumber="470" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B61FFC4FEABFC5C5873FC0D" box="[277,512,949,970]" pageId="7" pageNumber="470" type="description">
<paragraph id="8BD9125E6B61FFC4FEABFC5C5873FC0D" blockId="7.[162,628,895,970]" box="[277,512,949,970]" pageId="7" pageNumber="470">
(
<figureCitation id="135D0EDB6B61FFC4FEA3FC5C5B11FC0D" box="[285,354,949,970]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="7" pageNumber="470">Figs. 3</figureCitation>
,
<figureCitation id="135D0EDB6B61FFC4FED1FC5C5BFBFC0D" box="[367,392,949,970]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="7" pageNumber="470">11</figureCitation>
,
<figureCitation id="135D0EDB6B61FFC4FE2AFC5C5BDDFC0D" box="[404,430,949,970]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="7" pageNumber="470">19</figureCitation>
,
<figureCitation id="135D0EDB6B61FFC4FE04FC5C5BA7FC0D" box="[442,468,949,970]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="7" pageNumber="470">27</figureCitation>
,
<figureCitation id="135D0EDB6B61FFC4FE5EFC5C5B8AFC0D" box="[480,505,949,970]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="7" pageNumber="470">36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B61FFC4FF11FC36595BFADF" pageId="7" pageNumber="470" type="materials_examined">
<paragraph id="8BD9125E6B61FFC4FF11FC365826FA8A" blockId="7.[151,639,991,1729]" pageId="7" pageNumber="470">
<emphasis id="B912CE4C6B61FFC4FF11FC365B35FC33" bold="true" box="[175,326,991,1012]" pageId="7" pageNumber="470">Type Material.</emphasis>
<materialsCitation id="3B0E18036B61FFC4FEEAFC365B13FB83" collectionCode="MNCR" collectorName="G. Barrantes" elevation="1800" location="Costa Rica Cinco Esquinas de Carrizal" pageId="7" pageNumber="470" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="holotype">
<typeStatus id="54DDACFC6B61FFC4FEEAFC365BC0FC33" box="[340,435,991,1012]" pageId="7" pageNumber="470" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D60D9D76B61FFC4FE01FC365BA2FC32" box="[447,465,991,1013]" pageId="7" pageNumber="470" type="male">♂</specimenCount>
&quot;
<collectingRegion id="49A2DCBC6B61FFC4FE5CFC365849FC33" box="[482,570,991,1012]" country="Costa Rica" name="Alajuela" pageId="7" pageNumber="470">Alajuela</collectingRegion>
.
<location id="8EB944856B61FFC4FDF8FC365BA9FBC8" LSID="urn:lsid:plazi:treatment:03CFA3486B61FFCBFF1CFC965863FDA1:8EB944856B61FFC4FDF8FC365BA9FBC8" name="Costa Rica Cinco Esquinas de Carrizal" pageId="7" pageNumber="470" stateProvince="Alajuela">Costa Rica Cinco Esquinas de Carrizal</location>
,
<quantity id="4C9EBFBB6B61FFC4FE58FC13585BFBC8" box="[486,552,1018,1039]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="470" unit="m" value="1800.0">
<elevation id="004BF56D6B61FFC4FE58FC13585BFBC8" box="[486,552,1018,1039]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="470" unit="m" value="1800.0">1800m</elevation>
</quantity>
16 abril de 1988 Col:
<collectorName id="269377886B61FFC4FEA4FBFD5BFCFBEE" box="[282,399,1044,1065]" pageId="7" pageNumber="470">G. Barrantes</collectorName>
/ INBIOCRI002517128&quot; deposited at
<collectionCode id="ED778A9B6B61FFC4FEA8FBC65B13FB83" box="[278,352,1071,1092]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="7" pageNumber="470">MNCR</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B0E18036B61FFC4FED2FBC65B4DFB69" collectingDate="1989-04" collectingDateMax="1989-05-31" collectingDateMin="1989-04" collectionCode="CEUA" collectorName="A. Del Valle &amp; A. Solis" country="Costa Rica" county="Pr." elevation="1700" location="Zarcero" municipality="Alfaro Ruiz" pageId="7" pageNumber="470" specimenCode="CEUA00106199" specimenCount="2" specimenCount-male="2" stateProvince="Alajuela" typeStatus="paratype">
<typeStatus id="54DDACFC6B61FFC4FED2FBC65BBBFB83" box="[364,456,1071,1092]" pageId="7" pageNumber="470" type="paratype">Paratypes</typeStatus>
(6):
<specimenCount id="9D60D9D76B61FFC4FE43FBC65868FB83" box="[509,539,1070,1092]" pageId="7" pageNumber="470" type="male">2♂</specimenCount>
&quot;
<location id="8EB944856B61FFC4FD92FBC65809FB83" LSID="urn:lsid:plazi:treatment:03CFA3486B61FFCBFF1CFC965863FDA1:8EB944856B61FFC4FD92FBC65809FB83" box="[556,634,1071,1092]" country="Costa Rica" county="Pr." municipality="Alfaro Ruiz" name="Zarcero" pageId="7" pageNumber="470" stateProvince="Alajuela">Zarcero</location>
,
<collectingMunicipality id="6BBD88246B61FFC4FF29FBA05B7CFB99" box="[151,271,1097,1118]" pageId="7" pageNumber="470">Alfaro Ruiz</collectingMunicipality>
,
<quantity id="4C9EBFBB6B61FFC4FEA3FBA05B2CFB99" box="[285,351,1097,1118]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="470" unit="m" value="1700.0">
<elevation id="004BF56D6B61FFC4FEA3FBA05B2CFB99" box="[285,351,1097,1118]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="470" unit="m" value="1700.0">1700m</elevation>
</quantity>
<collectingRegion id="49A2DCBC6B61FFC4FED7FBA05BC9FB99" box="[361,442,1097,1118]" country="Costa Rica" name="Alajuela" pageId="7" pageNumber="470">Alajuela</collectingRegion>
<collectingCounty id="62B86AD26B61FFC4FE7AFBA05BACFB99" box="[452,479,1097,1118]" pageId="7" pageNumber="470">Pr.</collectingCounty>
<collectingCountry id="F37152CE6B61FFC4FE56FBA05808FB99" box="[488,635,1097,1118]" name="Costa Rica" pageId="7" pageNumber="470">COSTA RICA</collectingCountry>
.
<date id="FFD8349E6B61FFC4FF29FB8D5B4CFBBE" box="[151,319,1124,1145]" pageId="7" pageNumber="470" value="1989-04" valueMax="1989-05-31" valueMin="1989-04-00">
<collectingDate id="EF9CCD766B61FFC4FF29FB8D5B4CFBBE" box="[151,319,1124,1145]" pageId="7" pageNumber="470" value="1989-04" valueMax="1989-05-31" valueMin="1989-04-00">Apr - May 1989</collectingDate>
</date>
.
<collectorName id="269377886B61FFC4FEF5FB8D5BBBFBBE" box="[331,456,1124,1145]" pageId="7" pageNumber="470">A. Del Valle</collectorName>
&amp;
<collectorName id="269377886B61FFC4FE53FB8D5833FBBE" box="[493,576,1124,1145]" pageId="7" pageNumber="470">A. Solis</collectorName>
, L- N 240500, 493500&quot; / INBIOCRI000022463 and
<specimenCode id="DBC0BA256B61FFC4FF29FB705B4DFB69" box="[151,318,1177,1198]" collectionCode="CEUA" pageId="7" pageNumber="470">CEUA00106199</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B0E18036B61FFC4FEF4FB705B32FB39" collectingDate="1992-03-23" collectingDateMax="1992-05-02" collectingDateMin="1992-03-23" collectionCode="CEUA" country="Costa Rica" county="Araya" elevation="1500" location="Est. Biol. Las Alturas" municipality="Costa Rica" pageId="7" pageNumber="470" specimenCode="CEUA00106198" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="9D60D9D76B61FFC4FEF4FB705B1AFB69" box="[330,361,1176,1198]" pageId="7" pageNumber="470" type="male">3♂</specimenCount>
<specimenCount id="9D60D9D76B61FFC4FECCFB705BE3FB69" box="[370,400,1176,1198]" pageId="7" pageNumber="470" type="female">1♀</specimenCount>
&quot;
<location id="8EB944856B61FFC4FE1CFB70580DFB69" LSID="urn:lsid:plazi:treatment:03CFA3486B61FFCBFF1CFC965863FDA1:8EB944856B61FFC4FE1CFB70580DFB69" box="[418,638,1177,1198]" country="Costa Rica" county="Araya" municipality="Costa Rica" name="Est. Biol. Las Alturas" pageId="7" pageNumber="470" stateProvince="Puntarenas">Est. Biol. Las Alturas</location>
<quantity id="4C9EBFBB6B61FFC4FF29FB5D5A97FB0E" box="[151,228,1204,1225]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="470" unit="m" value="1500.0">
<elevation id="004BF56D6B61FFC4FF29FB5D5A97FB0E" box="[151,228,1204,1225]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="470" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<location id="8EB944856B61FFC4FF4EFB5D5B26FB0E" LSID="urn:lsid:plazi:treatment:03CFA3486B61FFCBFF1CFC965863FDA1:8EB944856B61FFC4FF4EFB5D5B26FB0E" box="[240,341,1204,1225]" country="Costa Rica" county="Araya" municipality="Costa Rica" name="Coto Brus" pageId="7" pageNumber="470" stateProvince="Puntarenas">Coto Brus</location>
,
<location id="8EB944856B61FFC4FEDFFB5D5BE0FB0E" LSID="urn:lsid:plazi:treatment:03CFA3486B61FFCBFF1CFC965863FDA1:8EB944856B61FFC4FEDFFB5D5BE0FB0E" box="[353,403,1204,1225]" country="Costa Rica" county="Araya" municipality="Costa Rica" name="Prov." pageId="7" pageNumber="470" stateProvince="Puntarenas">Prov.</location>
<collectingRegion id="49A2DCBC6B61FFC4FE25FB5D5870FB0E" box="[411,515,1204,1225]" country="Costa Rica" name="Puntarenas" pageId="7" pageNumber="470">Puntarenas</collectingRegion>
,
<collectingMunicipality id="6BBD88246B61FFC4FDB1FB5D5809FB0E" box="[527,634,1204,1225]" pageId="7" pageNumber="470">Costa Rica</collectingMunicipality>
, F.
<collectingCounty id="62B86AD26B61FFC4FF12FB275A96FB24" box="[172,229,1230,1251]" pageId="7" pageNumber="470">Araya</collectingCounty>
,
<date id="FFD8349E6B61FFC4FF53FB275BDAFB24" box="[237,425,1230,1251]" pageId="7" pageNumber="470" value="1992-03-23" valueMax="1992-05-02" valueMin="1992-03-23">
<collectingDate id="EF9CCD766B61FFC4FF53FB275BDAFB24" box="[237,425,1230,1251]" pageId="7" pageNumber="470" value="1992-03-23" valueMax="1992-05-02" valueMin="1992-03-23">23 mar a 2 may 1992</collectingDate>
</date>
, L- S 322500,591300&quot; /
<specimenCode id="DBC0BA256B61FFC4FF29FB005B32FB39" box="[151,321,1257,1278]" collectionCode="CEUA" pageId="7" pageNumber="470">CEUA00106198</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B0E18036B61FFC4FEF3FB005B9AFAF4" collectionCode="MNCR" collectorName="Four" country="Costa Rica" pageId="7" pageNumber="470" specimenCount="1" typeStatus="paratype">
INBIOCRI000909434, INBIO CRI000916198 and INBIOCRI000909406.
<collectorName id="269377886B61FFC4FDEEFAEA580DFADF" box="[592,638,1283,1304]" pageId="7" pageNumber="470">Four</collectorName>
<typeStatus id="54DDACFC6B61FFC4FF29FAF75A80FAF4" box="[151,243,1310,1331]" pageId="7" pageNumber="470" type="paratype">paratypes</typeStatus>
are deposited at
<collectionCode id="ED778A9B6B61FFC4FE1EFAF75B9AFAF4" box="[416,489,1310,1331]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="7" pageNumber="470">MNCR</collectionCode>
</materialsCitation>
, and
<materialsCitation id="3B0E18036B61FFC4FD9EFAF75822FA8A" country="Costa Rica" location="Universidad de Alicante" pageId="7" pageNumber="470" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D60D9D76B61FFC4FD9EFAF75ABBFA8A" pageId="7" pageNumber="470" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
are deposited at
<location id="8EB944856B61FFC4FED4FAD15822FA8A" LSID="urn:lsid:plazi:treatment:03CFA3486B61FFCBFF1CFC965863FDA1:8EB944856B61FFC4FED4FAD15822FA8A" box="[362,593,1336,1357]" country="Costa Rica" name="Universidad de Alicante" pageId="7" pageNumber="470">Universidad de Alicante</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD9125E6B61FFC4FF11FABA590FFB69" blockId="7.[151,639,991,1729]" lastBlockId="7.[685,1173,194,1729]" pageId="7" pageNumber="470">
<emphasis id="B912CE4C6B61FFC4FF11FABA5B60FAAF" bold="true" box="[175,275,1363,1384]" pageId="7" pageNumber="470">
<typeStatus id="54DDACFC6B61FFC4FF11FABA5B7DFAAF" box="[175,270,1363,1384]" pageId="7" pageNumber="470" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="135D0EDB6B61FFC4FEDAFABA5BD6FAAF" box="[356,421,1363,1384]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="7" pageNumber="470">Fig. 3</figureCitation>
). Body oval. Length
<quantity id="4C9EBFBB6B61FFC4FF29FA875A8CFA44" box="[151,255,1390,1411]" metricMagnitude="-2" metricUnit="m" metricValue="1.287" pageId="7" pageNumber="470" unit="mm" value="12.87">12.87 mm</quantity>
, width
<quantity id="4C9EBFBB6B61FFC4FEF2FA875BD4FA44" box="[332,423,1390,1411]" metricMagnitude="-3" metricUnit="m" metricValue="6.47" pageId="7" pageNumber="470" unit="mm" value="6.47">6.47 mm</quantity>
. Head reddish brown with ochre apical fringe and clypeus. Pronotum ochre with 2 large, trapezoidal, reddish brown maculae with bronze luster touching anterior and posterior margins. Scutellum ochre with thin, reddish sides. Elytra ochre with reddish suture. Pygidium ochre. Legs ochre with light to reddish brown apical part of tibia and tarsus. Venter reddish to dark brown, sternum with sides ochre.
<emphasis id="B912CE4C6B61FFC4FF29F9B45AA6F9B5" bold="true" box="[151,213,1629,1650]" pageId="7" pageNumber="470">Head:</emphasis>
Clypeus trapezoidal, anterior angles widely curved, surface densely punctate-reticulate, convex; anterior margin straight, thin in frontal view, ratio width/length 1.99. Frons densely punctate, flat. Ocular canthus long, thin, with acute apex. Ratio of interocular width/width of eye 3.52. Ratio antennal funiculus/club 0.79.
<emphasis id="B912CE4C6B61FFC4FC2FFF1E598FFECB" bold="true" box="[913,1020,247,268]" pageId="7" pageNumber="470">Pronotum:</emphasis>
Subtrapezoidal. Ratio width/length 1.81. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles quadrate. Basal margin sinuate, subapical bead complete. Surface with deep, dense punctures.
<emphasis id="B912CE4C6B61FFC4FB9DFE955EE0FE56" bold="true" box="[1059,1171,380,401]" pageId="7" pageNumber="470">Scutellum:</emphasis>
Subtriangular, with rounded sides and acute apex. Ratio width/length 1.56. Surface sparsely punctate.
<emphasis id="B912CE4C6B61FFC4FD13FE255885FE26" bold="true" box="[685,758,460,481]" pageId="7" pageNumber="470">Elytra:</emphasis>
Costae defined by regular rows of punctures. Subsutural interstice with 2–4 rows of sparse punctures, 2
<superScript id="7C13BF166B61FFC4FCA1FE155943FDCD" attach="none" box="[799,816,508,522]" fontSize="6" pageId="7" pageNumber="470">nd</superScript>
–4
<superScript id="7C13BF166B61FFC4FCF6FE155925FDCD" attach="left" box="[840,854,508,522]" fontSize="6" pageId="7" pageNumber="470">th</superScript>
interstices with 1–2 irregular, secondary stria(e). Surface covered with secondary punctures.
<emphasis id="B912CE4C6B61FFC4FCA9FDDF590CFD8C" bold="true" box="[791,895,566,587]" pageId="7" pageNumber="470">Pygidium:</emphasis>
Finely granulated with long setae on posterior margin. Ratio width/length 1.57.
<emphasis id="B912CE4C6B61FFC4FD13FD825886FD47" bold="true" box="[685,757,619,640]" pageId="7" pageNumber="470">Venter:</emphasis>
Space between mesocoxae narrow, slightly convex. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae (2 rows on second sternite) and 5–7 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane wide.
<emphasis id="B912CE4C6B61FFC4FD58FCE3596FFCD8" bold="true" box="[742,796,778,799]" pageId="7" pageNumber="470">Legs:</emphasis>
Protibia (
<figureCitation id="135D0EDB6B61FFC4FCC6FCE359C9FCD8" box="[888,954,778,799]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="7" pageNumber="470">Fig. 11</figureCitation>
) bidentate, apical tooth long and curved, basal tooth below internal apex of protibia triangular, acute. Metatibia slender, fusiform, length/width ratio 3.39. First external carina with a row of sparse setae, surface punctate. Internal protarsal claw bifurcate, with upper branch of same length and half width of the lower one; inferior margin with a sharp angle at half length.
<emphasis id="B912CE4C6B61FFC4FBE0FC2D5961FC33" bold="true" pageId="7" pageNumber="470">Male genitalia:</emphasis>
Parameres with blunt apex in lateral view (
<figureCitation id="135D0EDB6B61FFC4FD57FC135943FBC8" box="[745,816,1018,1039]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="7" pageNumber="470">Fig. 19</figureCitation>
), ventral angle defined. Endophallus (
<figureCitation id="135D0EDB6B61FFC4FD0BFBFD588FFBEE" box="[693,764,1044,1065]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="7" pageNumber="470">Fig. 27</figureCitation>
) with 1 ventral and 1 dorsal inflation. On dorsal inflation, a long curved, claw and a patch of sclerotized, thick setae; ejaculatory duct opening located on ventral inflation. Median lobes very thin and long, a dorsally located sclerotized plate between their apices.
</paragraph>
<paragraph id="8BD9125E6B61FFC4FD7BFB5D595BFADF" blockId="7.[685,1173,194,1729]" pageId="7" pageNumber="470">
<emphasis id="B912CE4C6B61FFC4FD7BFB5D5965FB0E" bold="true" box="[709,790,1204,1225]" pageId="7" pageNumber="470">Female.</emphasis>
Similar to male except protibia (
<figureCitation id="135D0EDB6B61FFC4FBF5FB5D5EFEFB0E" box="[1099,1165,1204,1225]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="7" pageNumber="470">Fig. 11</figureCitation>
) with wider teeth; internal protarsal claw narrower with curved inferior margin; last sternite with curved apex.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B61FFC4FD7BFAF759F2F990" pageId="7" pageNumber="470" type="description">
<paragraph id="8BD9125E6B61FFC4FD7BFAF759F2F990" blockId="7.[685,1173,194,1729]" pageId="7" pageNumber="470">
<emphasis id="B912CE4C6B61FFC4FD7BFAF75959FAF4" bold="true" box="[709,810,1310,1331]" pageId="7" pageNumber="470">Variation.</emphasis>
Six
<typeStatus id="54DDACFC6B61FFC4FCE2FAF759C9FAF4" box="[860,954,1310,1331]" pageId="7" pageNumber="470" type="paratype">paratypes</typeStatus>
. Maculae on pronotum can fuse into one large macula with a bronze or green luster. Venter of female specimen is ochre. Claw on endophallus can have a bifurcated tip. Body length
<quantity id="4C9EBFBB6B61FFC4FCFAFA615E76FA5A" box="[836,1029,1416,1438]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" metricValueMax="1.356" metricValueMin="1.204" pageId="7" pageNumber="470" unit="mm" value="12.8" valueMax="13.56" valueMin="12.04">12.04–13.56 mm</quantity>
, body width
<quantity id="4C9EBFBB6B61FFC4FD13FA4A5926FA7F" box="[685,853,1442,1464]" metricMagnitude="-3" metricUnit="m" metricValue="6.824999999999999" metricValueMax="7.18" metricValueMin="6.47" pageId="7" pageNumber="470" unit="mm" value="6.824999999999999" valueMax="7.18" valueMin="6.47">6.47–7.18 mm</quantity>
. Clypeal width/length ratio 1.85–2.01. Ratio of interocular width/width of eye 3.42–3.93. Ratio of antennal funiculus/club 0.70–0.79. Pronotal width/length ratio 1.79–1.87. Scutellar width/length ratio 1.22–1.56. Pygidial width/length ratio 1.57–1.66. Metatibial length/ width ratio 3.16–3.61.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B61FFC4FD7BF9B45E1BF906" pageId="7" pageNumber="470" type="etymology">
<paragraph id="8BD9125E6B61FFC4FD7BF9B45E1BF906" blockId="7.[685,1173,194,1729]" pageId="7" pageNumber="470">
<emphasis id="B912CE4C6B61FFC4FD7BF9B4594BF9B5" bold="true" box="[709,824,1629,1650]" pageId="7" pageNumber="470">Etymology.</emphasis>
From the past participle of the Latin verb
<emphasis id="B912CE4C6B61FFC4FCA8F99E5923F94A" box="[790,848,1655,1677]" italics="true" pageId="7" pageNumber="470">divido</emphasis>
, to divide. The ochre median line on the pronotum and dark suture on the elytra visually divide in half the dorsal aspect of this species.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6EFFCBFF7BFF2B58F5FECB" pageId="8" pageNumber="471" type="distribution">
<paragraph id="8BD9125E6B6EFFCBFF7BFF2B58F5FECB" blockId="8.[173,660,194,614]" pageId="8" pageNumber="471">
<emphasis id="B912CE4C6B6EFFCBFF7BFF2B5B34FF10" bold="true" box="[197,327,194,215]" pageId="8" pageNumber="471">Distribution.</emphasis>
<taxonomicName id="4C6669DD6B6EFFCBFEEBFF2B5B88FF1F" box="[341,507,194,216]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="divisa">
<emphasis id="B912CE4C6B6EFFCBFEEBFF2B5B88FF1F" box="[341,507,194,216]" italics="true" pageId="8" pageNumber="471">Anomala divisa</emphasis>
</taxonomicName>
occurs in the Central and Talamanca mountain ranges of
<collectingCountry id="F37152CE6B6EFFCBFDE3FF345AA8FECB" name="Costa Rica" pageId="8" pageNumber="471">Costa Rica</collectingCountry>
, from
<quantity id="4C9EBFBB6B6EFFCBFEA2FF1E5BB7FECB" box="[284,452,247,268]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="8" pageNumber="471" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">1,500 to 1,800 m</quantity>
elevation (
<figureCitation id="135D0EDB6B6EFFCBFD8CFF1E5808FECB" box="[562,635,247,268]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="8" pageNumber="471">Fig. 36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6EFFCBFF7BFEFB5863FDA1" pageId="8" pageNumber="471" type="diagnosis">
<paragraph id="8BD9125E6B6EFFCBFF7BFEFB5863FDA1" blockId="8.[173,660,194,614]" pageId="8" pageNumber="471">
<emphasis id="B912CE4C6B6EFFCBFF7BFEFB5B5EFEE0" bold="true" box="[197,301,274,295]" pageId="8" pageNumber="471">Diagnosis.</emphasis>
This species is recognized by the presence of two large, trapezoidal, dark maculae on the pronotum and the parameres with blunt apex and defined ventral angle. Similar species, also present in
<collectingCountry id="F37152CE6B6EFFCBFEF7FE955BC4FE56" box="[329,439,380,401]" name="Costa Rica" pageId="8" pageNumber="471">Costa Rica</collectingCountry>
, are
<taxonomicName id="4C6669DD6B6EFFCBFE52FE925B58FE6B" authority="Ohaus, 1897" authorityName="Ohaus" authorityYear="1897" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="jansoni">
<emphasis id="B912CE4C6B6EFFCBFE52FE9258E0FE56" box="[492,659,379,401]" italics="true" pageId="8" pageNumber="471">Anomala jansoni</emphasis>
Ohaus, 1897
</taxonomicName>
and
<taxonomicName id="4C6669DD6B6EFFCBFEDDFE7F58FCFE6B" authority="Ohaus, 1897" authorityName="Ohaus" authorityYear="1897" box="[355,655,406,428]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="quiche">
<emphasis id="B912CE4C6B6EFFCBFEDDFE7F5877FE6B" box="[355,516,406,428]" italics="true" pageId="8" pageNumber="471">Anomala quiche</emphasis>
Ohaus, 1897
</taxonomicName>
. The former species is slightly larger, with smaller maculae on the pronotum and long, slender, curved parameres in lateral view. The latter species has smaller and more separated maculae on the pronotum, maculae on the elytral calli, and parameres with a wider and blunter apex (aedeagi illustrated in
<bibRefCitation id="EFF76FAF6B6EFFCBFE8EFDB85876FDA1" author="Filippini" box="[304,517,592,614]" pageId="8" pageNumber="471" pagination="1" refId="ref10277" refString="Filippini, V., E. Galante, and E. Mico. 2015 a. Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 3948 (1): 024 - 040." type="journal article" year="2015">
Filippini
<emphasis id="B912CE4C6B6EFFCBFE35FDB95BCDFDA1" box="[395,446,592,614]" italics="true" pageId="8" pageNumber="471">et al.</emphasis>
2015a
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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