208 lines
22 KiB
XML
208 lines
22 KiB
XML
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<mods:title id="EA25ECEB70BFBBE968F167EF68935161">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="759E98B6174027C522B814DCBEAF641E">Assing, Volker</mods:namePart>
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<mods:title id="1F6F00EED194C035DB657426334EA18E">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="BA6A1C8712BE23C852D747A26F5F8BF7">2017</mods:date>
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<mods:number id="41899983F95BA283B3476ED01AF0CAAD">67</mods:number>
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<treatment id="03A787BAFFF2E506531A851B61DDFCD5" ID-DOI="http://doi.org/10.5281/zenodo.5888537" ID-GBIF-Taxon="192498145" ID-Zenodo-Dep="5888537" LSID="urn:lsid:plazi:treatment:03A787BAFFF2E506531A851B61DDFCD5" httpUri="http://treatment.plazi.org/id/03A787BAFFF2E506531A851B61DDFCD5" lastPageId="52" lastPageNumber="169" pageId="51" pageNumber="168">
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<subSubSection id="C3146527FFF2E501531A851B6627FD9A" box="[831,1324,531,558]" pageId="51" pageNumber="168" type="nomenclature">
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<paragraph id="8BB136ACFFF2E501531A851B6627FD9A" blockId="51.[831,1324,531,558]" box="[831,1324,531,558]" pageId="51" pageNumber="168">
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<heading id="D0F981C0FFF2E501531A851B6627FD9A" box="[831,1324,531,558]" centered="true" fontSize="12" level="2" pageId="51" pageNumber="168" reason="2">
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<taxonomicName id="4C0E4D2FFFF2E501531A851B6627FD9A" ID-CoL="8VS6V" authority="CAMERON, 1939" authorityName="CAMERON" authorityYear="1939" box="[831,1324,531,558]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="168" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEABEFFF2E501531A851B607EFD9A" box="[831,885,531,558]" italics="true" pageId="51" pageNumber="168">Zyras</emphasis>
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(incertae sedis)
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<emphasis id="B97AEABEFFF2E5015406851B678AFD9A" box="[1059,1153,531,558]" italics="true" pageId="51" pageNumber="168">coloratus</emphasis>
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CAMERON, 1939
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</taxonomicName>
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</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C3146527FFF2E501533F854467EBFDD5" box="[794,1248,587,609]" pageId="51" pageNumber="168" type="reference_group">
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<paragraph id="8BB136ACFFF2E501533F854467EBFDD5" blockId="51.[794,1248,587,609]" box="[794,1248,587,609]" pageId="51" pageNumber="168">
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<treatmentCitationGroup id="AB1E1182FFF2E501533F854467EBFDD5" box="[794,1248,587,609]" pageId="51" pageNumber="168">
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<taxonomicName id="4C0E4D2FFFF2E501533F854467C5FDD5" ID-CoL="8VS6V" authority="CAMERON, 1939 a: 545" authorityName="CAMERON" authorityPageNumber="545" authorityYear="1939" box="[794,1230,587,609]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="168" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEABEFFF2E501533F85446045FDD5" box="[794,846,588,609]" italics="true" pageId="51" pageNumber="168">Zyras</emphasis>
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(
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<emphasis id="B97AEABEFFF2E501537985436082FDD4" box="[860,905,587,608]" italics="true" pageId="51" pageNumber="168">Pella</emphasis>
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?)
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<treatmentCitation id="0AAF10BDFFF2E5015385854367C5FDD5" author="CAMERON, M." box="[928,1230,587,609]" page="545" pageId="51" pageNumber="168" year="1939">
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<emphasis id="B97AEABEFFF2E5015385854360FEFDD4" box="[928,1013,587,608]" italics="true" pageId="51" pageNumber="168">coloratus</emphasis>
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<bibRefCitation id="EF9F4B5DFFF2E50153DF854367C5FDD5" author="CAMERON, M." box="[1018,1230,587,609]" pageId="51" pageNumber="168" refId="ref57482" refString="CAMERON, M. 1939 a: Coleoptera: Staphylinidae. Volume IV Parts I-II. - Sewell, R. B. S. (ed): The Fauna of British India, including Ceylon and Burma. - London: 691 pp." type="book" year="1939">
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CAMERON, 1939a:
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<specimenCount id="9D08FD25FFF2E501548E854367C5FDD5" box="[1195,1230,587,609]" pageId="51" pageNumber="168" type="generic">545</specimenCount>
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</bibRefCitation>
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||
</treatmentCitation>
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</taxonomicName>
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f.
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||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146527FFF2E501533F85816756FCF5" pageId="51" pageNumber="168" type="materials_examined">
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<paragraph id="8BB136ACFFF2E501533F85816756FCF5" blockId="51.[794,1399,649,833]" pageId="51" pageNumber="168">
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<emphasis id="B97AEABEFFF2E501533F85816726FD15" bold="true" box="[794,1069,649,673]" pageId="51" pageNumber="168">Type material examined</emphasis>
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:
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<materialsCitation id="3B663CF1FFF2E501541A85816609FC95" ID-GBIF-Occurrence="3455147319" collectingDate="1921-08-08" collectionCode="BMNH" collectorName="Dr. M. Cameron & M. Cameron. Bequest." determinerName="Det. Maruyama" location="Zyras" pageId="51" pageNumber="168" specimenCount="4" stateProvince="B" typeStatus="syntype">
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<typeStatus id="54B5880EFFF2E501541A858167ABFD15" box="[1087,1184,649,673]" pageId="51" pageNumber="168" type="syntype">Syntypes</typeStatus>
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: 1 ♀: “Dehra Dun,
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<collectorName id="26FB537AFFF2E501533F85A260C4FD75" box="[794,975,681,705]" pageId="51" pageNumber="168">Dr. M. Cameron</collectorName>
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,
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<date id="FFB0106CFFF2E50153FE85A16730FD75" box="[987,1083,681,705]" pageId="51" pageNumber="168" value="1921-08-08">
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<collectingDate id="EFF4E984FFF2E50153FE85A16730FD75" box="[987,1083,681,705]" pageId="51" pageNumber="168" value="1921-08-08">8-8-1921</collectingDate>
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</date>
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/ at light [overleaf] /
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<taxonomicName id="4C0E4D2FFFF2E501550D85A26081FD55" authority="Cam" authorityName="CAMERON" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="168" phylum="Arthropoda" rank="species" species="coloratus">
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<emphasis id="B97AEABEFFF2E501550D85A26044FD55" italics="true" pageId="51" pageNumber="168">Z. coloratus</emphasis>
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Cam
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</taxonomicName>
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<typeStatus id="54B5880EFFF2E50153B785C260CDFD56" box="[914,966,714,738]" pageId="51" pageNumber="168">Type</typeStatus>
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/
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<collectorName id="26FB537AFFF2E50153FA85C267DAFD55" box="[991,1233,713,737]" pageId="51" pageNumber="168">M. Cameron. Bequest.</collectorName>
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<collectingRegion id="49CAF84EFFF2E50154FD85C267E1FD55" box="[1240,1258,714,737]" pageId="51" pageNumber="168">B</collectingRegion>
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.M. 1955-147 /
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<typeStatus id="54B5880EFFF2E501530B85E16088FCB5" box="[814,899,745,769]" pageId="51" pageNumber="168" type="syntype">Syntype</typeStatus>
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/
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<typeStatus id="54B5880EFFF2E501538685E160F3FCB5" box="[931,1016,745,769]" pageId="51" pageNumber="168" type="syntype">Syntype</typeStatus>
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<taxonomicName id="4C0E4D2FFFF2E501542685E26736FCB5" box="[1027,1085,746,769]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="168" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B97AEABEFFF2E501542685E26736FCB5" box="[1027,1085,746,769]" italics="true" pageId="51" pageNumber="168">
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||
<location id="8ED16077FFF2E501542685E26736FCB5" LSID="urn:lsid:plazi:treatment:03A787BAFFF2E506531A851B61DDFCD5:8ED16077FFF2E501542685E26736FCB5" box="[1027,1085,746,769]" name="Zyras" pageId="51" pageNumber="168" stateProvince="B">Zyras</location>
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</emphasis>
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||
</taxonomicName>
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(
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<emphasis id="B97AEABEFFF2E501547485E1678FFCB5" box="[1105,1156,745,769]" italics="true" pageId="51" pageNumber="168">
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||
<location id="8ED16077FFF2E501547485E1678FFCB5" LSID="urn:lsid:plazi:treatment:03A787BAFFF2E506531A851B61DDFCD5:8ED16077FFF2E501547485E1678FFCB5" box="[1105,1156,745,769]" name="Pella" pageId="51" pageNumber="168" stateProvince="B">Pella</location>
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</emphasis>
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?)
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<taxonomicName id="4C0E4D2FFFF2E501548685E16044FC95" authority="Cameron, 1939" authorityName="CAMERON" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="168" phylum="Arthropoda" rank="species" species="coloratus">
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<emphasis id="B97AEABEFFF2E501548685E1660AFCB5" box="[1187,1281,745,769]" italics="true" pageId="51" pageNumber="168">coloratus</emphasis>
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<location id="8ED16077FFF2E501552985E16679FCB5" LSID="urn:lsid:plazi:treatment:03A787BAFFF2E506531A851B61DDFCD5:8ED16077FFF2E501552985E16679FCB5" box="[1292,1394,745,769]" name="Cameron" pageId="51" pageNumber="168" stateProvince="B">Cameron</location>
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, 1939
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</taxonomicName>
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,
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<determinerName id="7E6DDA68FFF2E501537884026703FC96" box="[861,1032,778,802]" pageId="51" pageNumber="168">Det. Maruyama</determinerName>
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2002 / 3-A” (
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<collectionCode id="ED1FAE69FFF2E5015484840167FDFC95" box="[1185,1270,777,801]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="51" pageNumber="168">BMNH</collectionCode>
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);
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</materialsCitation>
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<materialsCitation id="3B663CF1FFF2E501552E84016751FCF5" ID-GBIF-Occurrence="3455147313" collectingDate="1921-08-08" collectionCode="BMNH" collectorName="Dr. M. Cameron & M. Cameron. Bequest." determinerName="Det. Maruyama" location="Zyras" pageId="51" pageNumber="168" specimenCount="4" stateProvince="B" typeStatus="syntype">
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1 ♀: same data, but “9-9.1922” (
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<collectionCode id="ED1FAE69FFF2E50153D984216759FCF5" box="[1020,1106,809,833]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="51" pageNumber="168">BMNH</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146527FFF2E501533F84616618FB75" pageId="51" pageNumber="168" type="discussion">
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<paragraph id="8BB136ACFFF2E501533F84616618FB75" blockId="51.[794,1399,873,1217]" pageId="51" pageNumber="168">
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<emphasis id="B97AEABEFFF2E501533F84616082FC35" bold="true" box="[794,905,873,897]" pageId="51" pageNumber="168">Comment</emphasis>
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||
: The original description is based on an unspecified number of
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<typeStatus id="54B5880EFFF2E501540A84826781FC16" box="[1071,1162,906,930]" pageId="51" pageNumber="168" type="syntype">syntypes</typeStatus>
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from “Dehra Dun” (
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<bibRefCitation id="EF9F4B5DFFF2E501530684A160D1FC75" author="CAMERON, M." box="[803,986,937,961]" pageId="51" pageNumber="168" refId="ref57482" refString="CAMERON, M. 1939 a: Coleoptera: Staphylinidae. Volume IV Parts I-II. - Sewell, R. B. S. (ed): The Fauna of British India, including Ceylon and Burma. - London: 691 pp." type="book" year="1939">CAMERON 1939a</bibRefCitation>
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||
). Two
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<typeStatus id="54B5880EFFF2E501540684A2678BFC76" box="[1059,1152,938,962]" pageId="51" pageNumber="168" type="syntype">syntypes</typeStatus>
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, both of them females, were found in the Cameron collection at the BMNH. While
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<bibRefCitation id="EF9F4B5DFFF2E501534784E1672DFBB5" author="CAMERON, M." box="[866,1062,1001,1025]" pageId="51" pageNumber="168" refId="ref57482" refString="CAMERON, M. 1939 a: Coleoptera: Staphylinidae. Volume IV Parts I-II. - Sewell, R. B. S. (ed): The Fauna of British India, including Ceylon and Burma. - London: 691 pp." type="book" year="1939">CAMERON (1939a)</bibRefCitation>
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tentatively assigned the species to
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<emphasis id="B97AEABEFFF2E501531C83016060FB95" box="[825,875,1033,1057]" italics="true" pageId="51" pageNumber="168">Pella</emphasis>
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||
STEPHENS, 1835,
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<bibRefCitation id="EF9F4B5DFFF2E50154168302661FFB95" author="HLAVAC, P. & NEWTON, A. F. & MARUYAMA, M." box="[1075,1300,1033,1061]" pageId="51" pageNumber="168" pagination="1 - 151" refId="ref57922" refString="HLAVAC, P.; NEWTON, A. F. & MARUYAMA, M. 2011: World catalogue of the species of the tribe Lomechusini (Staphylinidae: Aleocharinae). - Zootaxa 3075: 1 - 151." type="journal article" year="2011">HLAVÁČ et al. (2011)</bibRefCitation>
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list it as
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<taxonomicName id="4C0E4D2FFFF2E501533F8322605FFBF5" box="[794,852,1066,1089]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="168" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B97AEABEFFF2E501533F8322605FFBF5" box="[794,852,1066,1089]" italics="true" pageId="51" pageNumber="168">Zyras</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24957C5FFF2E501537283226086FBF5" box="[855,909,1066,1089]" pageId="51" pageNumber="168" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
An examination of the type specimens revealed that they significantly differ in so many respects (habitus, morphology of antennae, punctation pattern, absence of non-setiferous punctation, short legs, etc.) from
|
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<taxonomicName id="4C0E4D2FFFF2E50155188382667CFB15" box="[1341,1399,1162,1185]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="168" phylum="Arthropoda" rank="genus" sensu="stricto">
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||
<emphasis id="B97AEABEFFF2E50155188382667CFB15" box="[1341,1399,1162,1185]" italics="true" pageId="51" pageNumber="168">Zyras</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24957C5FFF2E501533F83A2605BFB75" box="[794,848,1194,1217]" pageId="51" pageNumber="168" sensu="stricto">s. str.</taxonomicNameLabel>
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||
that they are treated as incertae sedis here.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFF2E506533F83E16165FCB5" lastPageId="52" lastPageNumber="169" pageId="51" pageNumber="168" type="description">
|
||
<paragraph id="8BB136ACFFF2E501533F83E160B7F9F5" blockId="51.[794,1399,1257,2049]" pageId="51" pageNumber="168">
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||
<emphasis id="B97AEABEFFF2E501533F83E160BCFAB5" bold="true" box="[794,951,1257,1281]" pageId="51" pageNumber="168">Redescription</emphasis>
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: Body subparallel and somewhat depressed, in habitus and punctation somewhat resembling a species of
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<taxonomicName id="4C0E4D2FFFF2E50153D982216620FAF5" authority="KRAATZ, 1856" authorityName="KRAATZ" authorityYear="1856" box="[1020,1323,1321,1345]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="168" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEABEFFF2E50153D98221677DFAF5" box="[1020,1142,1321,1345]" italics="true" pageId="51" pageNumber="168">Tomoglossa</emphasis>
|
||
KRAATZ, 1856
|
||
</taxonomicName>
|
||
. Body length
|
||
<quantity id="4CF69B49FFF2E501534B824160FAFAD5" box="[878,1009,1353,1377]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="51" pageNumber="168" unit="mm" value="4.6" valueMax="4.7" valueMin="4.5">4.5–4.7 mm</quantity>
|
||
; length of forebody 1.7–2.0 mm. Coloration: head reddish-brown to brown; pronotum reddish-yellow; elytra pale-reddish, noticeably darker than pronotum; abdomen: tergites II–V uniformly palereddish or weakly infuscate in the middle, VI–VIII brown; legs pale-yellowish; antennae pale reddishbrown; maxillary palpi yellowish, with palpomere III slightly darker.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFF2E501533F8141606BF895" blockId="51.[794,1399,1257,2049]" pageId="51" pageNumber="168">
|
||
Head distinctly transverse; punctation coarse and dense, somewhat sparser in median portion. Eyes much longer than postocular region in dorsal view. Antenna short and strongly incrassate apically,
|
||
<quantity id="4CF69B49FFF2E501546D81A16791F975" box="[1096,1178,1705,1729]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="51" pageNumber="168" unit="mm" value="1.2">1.2 mm</quantity>
|
||
long; antennomeres IV–X disc-shaped, approximately three times as broad as long, XI large and longer than the combined length of IX and X.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFF2E501533F80226701F835" blockId="51.[794,1399,1257,2049]" pageId="51" pageNumber="168">Pronotum approximately 1.3 times as broad as long and approximately 1.2 times as broad as head; punctation coarse and very dense.</paragraph>
|
||
<paragraph id="8BB136ACFFF2E501533F8081675CF7B5" blockId="51.[794,1399,1257,2049]" pageId="51" pageNumber="168">Elytra approximately 0.9 times as long as pronotum; punctation similar to that of pronotum. Hind wings fully developed. Metatarsomere I approximately as long as the combined length of II and III.</paragraph>
|
||
<paragraph id="8BB136ACFFF5E506509887E16118FDF5" blockId="52.[189,794,233,609]" pageId="52" pageNumber="169">Abdomen approximately as broad as elytra, not distinctly tapering posteriad, segments III–VII of subequal width; non-setiferous punctation absent; tergites III–V with very shallow anterior impressions, very sparsely and finely punctate in anterior half and less finely and less sparsely punctate in posterior half; tergites VI–VII sparsely and very finely punctate anteriorly and less finely punctate and more densely punctate in posterior portion; tergite VIII with more distinct and denser punctation than tergites III–VII; posterior margin of tergite VII with palisade fringe.</paragraph>
|
||
<paragraph id="8BB136ACFFF5E506509885426240FDD5" blockId="52.[189,794,233,609]" box="[189,331,585,609]" pageId="52" pageNumber="169">♂: unknown.</paragraph>
|
||
<paragraph id="8BB136ACFFF5E506509885816165FCB5" blockId="52.[189,794,649,769]" pageId="52" pageNumber="169">
|
||
<emphasis id="B97AEABEFFF5E506509885816285FD15" bold="true" box="[189,398,649,673]" pageId="52" pageNumber="169">Comparative notes</emphasis>
|
||
: In body shape,
|
||
<taxonomicName id="4C0E4D2FFFF5E506521A858261B2FD15" authorityName="CAMERON" authorityYear="1939" box="[575,697,649,673]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="169" phylum="Arthropoda" rank="species" species="coloratus">
|
||
<emphasis id="B97AEABEFFF5E506521A858261B2FD15" box="[575,697,649,673]" italics="true" pageId="52" pageNumber="169">Z. coloratus</emphasis>
|
||
</taxonomicName>
|
||
is somewhat similar to
|
||
<taxonomicName id="4C0E4D2FFFF5E506514B85A26108FD75" authorityName="PACE" authorityYear="2004" box="[366,515,681,705]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="169" phylum="Arthropoda" rank="species" species="angkoricola">
|
||
<emphasis id="B97AEABEFFF5E506514B85A26108FD75" box="[366,515,681,705]" italics="true" pageId="52" pageNumber="169">Z. angkoricola</emphasis>
|
||
</taxonomicName>
|
||
(see below), from which
|
||
<taxonomicName id="4C0E4D2FFFF5E506509885C2623CFD55" authorityName="CAMERON" authorityYear="1939" box="[189,311,713,737]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="169" phylum="Arthropoda" rank="species" species="coloratus">
|
||
<emphasis id="B97AEABEFFF5E506509885C2623CFD55" box="[189,311,713,737]" italics="true" pageId="52" pageNumber="169">Z. coloratus</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguished by much coarser and denser punctation of the forebody alone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFF5E5065098842161DDFCD5" pageId="52" pageNumber="169" type="distribution">
|
||
<paragraph id="8BB136ACFFF5E5065098842161DDFCD5" blockId="52.[189,794,809,866]" pageId="52" pageNumber="169">
|
||
<emphasis id="B97AEABEFFF5E50650988421624DFCF5" bold="true" box="[189,326,809,833]" pageId="52" pageNumber="169">Distribution</emphasis>
|
||
: This species is currently known only from its
|
||
<typeStatus id="54B5880EFFF5E50650F984426201FCD6" box="[220,266,842,866]" pageId="52" pageNumber="169">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="49CAF84EFFF5E506515A8441614DFCD5" box="[383,582,841,865]" country="India" name="Himachal Pradesh" pageId="52" pageNumber="169">Himachal Pradesh</collectingRegion>
|
||
, North
|
||
<collectingCountry id="F319763CFFF5E50652B2844161D9FCD5" box="[663,722,841,865]" name="India" pageId="52" pageNumber="169">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |