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<document id="5FCF3975D675F79E4F5ADF25E7D3EA5C" ID-CLB-Dataset="20138" ID-DOI="10.5852/ejt.2022.806.1697" ID-GBIF-Dataset="28f53295-9155-4b84-ab43-45e852842817" ID-ISSN="2118-9773" ID-Zenodo-Dep="6373945" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1647883015262" checkinUser="felipe" docAuthor="Dang, Minh Quan, Nguyen, Manh Ha, Hoang, Nghia Son, Nguyen, Thi Mai Huong, Truong, Ba Vuong &amp; Dang, Van Son" docDate="2022" docId="0387F954292BFF8AFDD713DBFE83CA2F" docLanguage="en" docName="ejt-806_19-31.pdf" docOrigin="European Journal of Taxonomy 806" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Lasianthus naikii V. S. Dang &amp; Vuong 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="25" masterDocId="FFBE812C292AFF8CFFDD1542FF27C806" masterDocTitle="Two new species of Lasianthus Jack (Rubiaceae) from southern Vietnam" masterLastPageNumber="31" masterPageNumber="19" pageNumber="20" updateTime="1699319045260" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="8AABF45BE0EE8D19C212E6ADCC0419B4">
<mods:title id="2C4CB82D56747C20409C1CA62E2B17C9">Two new species of Lasianthus Jack (Rubiaceae) from southern Vietnam</mods:title>
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<mods:name id="557BA7BEB4E906F3973EB65611EB8173" type="personal">
<mods:role id="658B55D569FAFEE0F44342F1A246F23C">
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<mods:namePart id="C0C1DDAEA1ED97D327D3DE9485B8B972">Dang, Minh Quan</mods:namePart>
<mods:affiliation id="ABD36BD5988D5405F663DD820E85676B">School of Education, Can Tho University, 3 / 2 Street, Ninh Kieu District, Can Tho City, Vietnam. Pedo-Geography and Soil Resources, Institute of Geography (IG), Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan.</mods:affiliation>
<mods:nameIdentifier id="D43AF02E80E46606ED0C4BD512C4C7DC" type="email">dmquan@ctu.edu.vn</mods:nameIdentifier>
</mods:name>
<mods:name id="4EA27DFC4FC13EDD3B0AC081758B08E3" type="personal">
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<mods:namePart id="686DBA4074A742FBECDAD82AF8958AE0">Nguyen, Manh Ha</mods:namePart>
<mods:nameIdentifier id="93F358D7D49F603AA186CC092F82D320" type="email">havdl72@ig.vast.vn</mods:nameIdentifier>
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<mods:namePart id="410A4E4B3AA371322569162B1CC007D1">Hoang, Nghia Son</mods:namePart>
<mods:nameIdentifier id="61EFF1FF2734D78837BAAF0BACDC096B" type="email">hoangnghiason@yahoo.com&amp;tho1010@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="EA848F23277231F4E95E241D1DE560DA" type="personal">
<mods:role id="F2CFD6D81EBB4D400862C86609CA9F59">
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</mods:role>
<mods:namePart id="B4C4E8E2A1DD688F4DC3862DA596B424">Nguyen, Thi Mai Huong</mods:namePart>
<mods:nameIdentifier id="C714E5A4565A8417BD7D9DAE07D3C95C" type="email">ngmaihuong2008@gmail.com&amp;quynhchihonguyen@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="19D17E233E5DCE27671B00E25320DCFE">Author</mods:roleTerm>
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<mods:namePart id="C203D384903D75C8CC41436656818A29">Truong, Ba Vuong</mods:namePart>
<mods:nameIdentifier id="B985EBE26EC8FF4707EB0B7FC3A79776" type="email">bavuong2019@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="D617A792727388B74F5FA65B09A54BE2">Dang, Van Son</mods:namePart>
<mods:nameIdentifier id="2CFF990DD948F5F99473439842A40846" type="email">dvsonitb@gmail.com</mods:nameIdentifier>
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<mods:title id="D769136F4DE2A05532030722D62025D8">European Journal of Taxonomy</mods:title>
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<mods:part id="F56E7ECCC074F8B21D77E4CE926F65FE">
<mods:date id="946013C0BB0B7ACE4B35EFE3E8B7A18D">2022</mods:date>
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<mods:number id="4F7C6A8BE9C7DA862C09FB7AE4558C85">2022-03-18</mods:number>
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<mods:number id="153C62C73348EB7FBB9B66D0887D0F7F">806</mods:number>
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<mods:classification id="9951A0C4D94226BD9569B18A09C89518">journal article</mods:classification>
<mods:identifier id="DF369EBE20E46ABD5F63E74C67D0D61E" type="CLB-Dataset">20138</mods:identifier>
<mods:identifier id="B2CA274D244C7E58A35ECDEC8639686F" type="DOI">10.5852/ejt.2022.806.1697</mods:identifier>
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<treatment id="0387F954292BFF8AFDD713DBFE83CA2F" ID-DOI="http://doi.org/10.5281/zenodo.6379958" ID-GBIF-Taxon="194208393" ID-Zenodo-Dep="6379958" LSID="urn:lsid:plazi:treatment:0387F954292BFF8AFDD713DBFE83CA2F" httpUri="http://treatment.plazi.org/id/0387F954292BFF8AFDD713DBFE83CA2F" lastPageId="6" lastPageNumber="25" pageId="1" pageNumber="20">
<subSubSection id="C3341BC9292BFF8DFDD713DBFB0DCEB2" box="[522,1066,1689,1716]" pageId="1" pageNumber="20" type="nomenclature">
<paragraph id="8B914842292BFF8DFDD713DBFB0DCEB2" blockId="1.[522,1066,1689,1792]" box="[522,1066,1689,1716]" pageId="1" pageNumber="20">
<heading id="D0D9FF2E292BFF8DFDD713DBFB0DCEB2" box="[522,1066,1689,1716]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="20" reason="2">
<taxonomicName id="4C2E33C1292BFF8DFDD713DBFCEFCEB2" authority="V.S.Dang &amp; Vuong" authorityName="V. S. Dang &amp; Vuong" authorityYear="2022" box="[522,968,1689,1716]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="20" phylum="Tracheophyta" rank="species" species="naikii" status="sp. nov.">
<emphasis id="B95A9450292BFF8DFDD713DBFDF9CEB2" bold="true" box="[522,734,1689,1716]" italics="true" pageId="1" pageNumber="20">Lasianthus naikii</emphasis>
V.S.Dang &amp; Vuong
</taxonomicName>
<taxonomicNameLabel id="A269292B292BFF8DFC0D13D8FB0DCEB2" box="[976,1066,1690,1716]" pageId="1" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292BFF8DFD9A13FDFCA7CF06" pageId="1" pageNumber="20" type="description">
<paragraph id="8B914842292BFF8DFD9A13FDFCCACEDC" blockId="1.[522,1066,1689,1792]" box="[583,1005,1727,1754]" pageId="1" pageNumber="20">urn:lsid:ipni.org:names:77295607-1</paragraph>
<paragraph id="8B914842292BFF8DFD6913A4FCA7CF06" blockId="1.[522,1066,1689,1792]" box="[692,896,1766,1792]" pageId="1" pageNumber="20">
<figureCitation id="131554C7292BFF8DFD6913A4FC3ACF06" box="[692,797,1766,1792]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[189,232,1844,1870]" captionStartId-1="3.[189,232,1962,1988]" captionTargetBox-0="[189,1398,265,1805]" captionTargetBox-1="[233,1359,288,1884]" captionTargetId-0="figure-16@2.[189,1398,265,1805]" captionTargetId-1="figure-12@3.[189,1399,262,1924]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Lasianthus naikii V.S.Dang &amp; Vuong sp. nov. A. Branch. B. Stipule. C. Abaxial leaf surfaces. D. Calyces. E–F. Close-up of flowers. G–H. Mature fruits. I. Seeds. Photos by Van-Son Dang on 22 Apr. 2021." captionText-1="Fig. 2. Lasianthus naikii V.S.Dang &amp; Vuong sp. nov. A. Branch. B. Abaxial leaf surfaces. C. Stipule. D. Close-up of flower. E. Fruit. Line drawings from the holotype (Dang 453) by Ba-Vuong Truong." figureDoi-0="http://doi.org/10.5281/zenodo.6373947" figureDoi-1="http://doi.org/10.5281/zenodo.6373949" httpUri-0="https://zenodo.org/record/6373947/files/figure.png" httpUri-1="https://zenodo.org/record/6373949/files/figure.png" pageId="1" pageNumber="20">Figs 1–2</figureCitation>
;
<tableCitation id="C6AC7DF9292BFF8DFCF413A4FCA7CF06" box="[809,896,1766,1792]" captionStart="Table 1" captionStartId="5.[189,258,265,291]" captionTargetPageId="5" captionText="Table 1. Comparison of Lasianthus naikii V.S.Dang &amp; Vuong sp. nov. and L. formosensis Matsumura. * = from Zhu &amp; Taylor 2011." httpUri="http://table.plazi.org/id/DF5118CA292FFF89FF60144BFD32C940" pageId="1" pageNumber="20" tableUuid="DF5118CA292FFF89FF60144BFD32C940">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292BFF8DFF601268FBA3CFEE" pageId="1" pageNumber="20" type="diagnosis">
<paragraph id="8B914842292BFF8DFF601268FE11CF42" blockId="1.[189,310,1834,1860]" box="[189,310,1834,1860]" pageId="1" pageNumber="20">
<heading id="D0D9FF2E292BFF8DFF601268FE11CF42" bold="true" box="[189,310,1834,1860]" fontSize="11" level="3" pageId="1" pageNumber="20" reason="3">
<emphasis id="B95A9450292BFF8DFF601268FE11CF42" bold="true" box="[189,310,1834,1860]" pageId="1" pageNumber="20">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292BFF8DFF60121EFBA3CFEE" blockId="1.[189,1399,1884,2025]" pageId="1" pageNumber="20">
<taxonomicName id="4C2E33C1292BFF8DFF60121EFEAECF70" authority="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang, 2022" authorityName="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang" authorityYear="2022" box="[189,393,1884,1910]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="20" phylum="Tracheophyta" rank="species" species="naikii" status="sp. nov.">
<emphasis id="B95A9450292BFF8DFF60121EFEAECF70" box="[189,393,1884,1910]" italics="true" pageId="1" pageNumber="20">Lasianthus naikii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269292B292BFF8DFE4D121EFECBCF70" box="[400,492,1884,1910]" pageId="1" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="4C2E33C1292BFF8DFD5A121FFCE7CF70" authority="Matsumura" authorityName="Matsumura" box="[647,960,1884,1910]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="20" phylum="Tracheophyta" rank="species" species="formosensis">
<emphasis id="B95A9450292BFF8DFD5A121FFC15CF70" box="[647,818,1884,1910]" italics="true" pageId="1" pageNumber="20">L. formosensis</emphasis>
Matsumura
</taxonomicName>
(
<figureCitation id="131554C7292BFF8DFC0D121EFB30CF70" box="[976,1047,1884,1910]" captionStart="Fig" captionStartId="4.[189,232,1845,1871]" captionTargetBox="[189,1399,265,1806]" captionTargetId="figure-15@4.[189,1399,265,1806]" captionTargetPageId="4" captionText="Fig. 3. Lasianthus formosensis Matsumura. A. Fruiting branch. B. Stipule. C. Abaxial leaf surfaces. D. Close-up of inflorescences and flowers. E. Fruits. Photos by Van-Son Dang on 17 Nov. 2018." figureDoi="http://doi.org/10.5281/zenodo.6373951" httpUri="https://zenodo.org/record/6373951/files/figure.png" pageId="1" pageNumber="20">Fig. 3</figureCitation>
) in leaf morphology and blue fruits, but differs from the latter by having secondary veins 5–6 pairs (vs 6–7 pairs), bracts present
<quantity id="4CD6E5A7292BFF8DFF6012EAFE10CFC5" box="[189,311,1960,1987]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="20" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long (vs absent), calyx tube
<quantity id="4CD6E5A7292BFF8DFD4B12EAFDDECFC5" box="[662,761,1960,1987]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="20" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
long (vs ca
<quantity id="4CD6E5A7292BFF8DFC4C12EAFCF1CFC5" box="[913,982,1960,1987]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="20" unit="mm" value="1.0">1 mm</quantity>
long), calyx lobes
<quantity id="4CD6E5A7292BFF8DFB6212EAFA1ECFC5" box="[1215,1337,1960,1987]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="20" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long (vs
<quantity id="4CD6E5A7292BFF8DFF37128CFE6CCFEF" box="[234,331,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="20" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long) and corolla lobes 6 (vs 5). For a further comparison see
<tableCitation id="C6AC7DF9292BFF8DFBFB128CFB59CFEE" box="[1062,1150,1998,2024]" captionStart="Table 1" captionStartId="5.[189,258,265,291]" captionTargetPageId="5" captionText="Table 1. Comparison of Lasianthus naikii V.S.Dang &amp; Vuong sp. nov. and L. formosensis Matsumura. * = from Zhu &amp; Taylor 2011." httpUri="http://table.plazi.org/id/DF5118CA292FFF89FF60144BFD32C940" pageId="1" pageNumber="20" tableUuid="DF5118CA292FFF89FF60144BFD32C940">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5118CA2928FF8EFF601276FFD9CF93" ID-DOI="http://doi.org/10.5281/zenodo.6373947" ID-Zenodo-Dep="6373947" httpUri="https://zenodo.org/record/6373947/files/figure.png" pageId="2" pageNumber="21" startId="2.[189,232,1844,1870]" targetBox="[189,1398,265,1805]" targetPageId="2">
<paragraph id="8B9148422928FF8EFF601276FFD9CF93" blockId="2.[188,1399,1844,1941]" pageId="2" pageNumber="21">
<emphasis id="B95A94502928FF8EFF601276FE2BCF49" bold="true" box="[189,268,1844,1871]" pageId="2" pageNumber="21">Fig. 1.</emphasis>
<taxonomicName id="4C2E33C12928FF8EFEC81277FDF6CF49" authority="V.S.Dang &amp; Vuong" authorityName="V. S. Dang &amp; Vuong" authorityYear="2022" box="[277,721,1845,1871]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="naikii" status="sp. nov.">
<emphasis id="B95A94502928FF8EFEC81277FEC5CF49" box="[277,482,1845,1871]" italics="true" pageId="2" pageNumber="21">Lasianthus naikii</emphasis>
V.S.Dang &amp; Vuong
</taxonomicName>
<taxonomicNameLabel id="A269292B2928FF8EFD071277FC11CF49" box="[730,822,1845,1871]" pageId="2" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B95A94502928FF8EFC9D1276FC73CF48" bold="true" box="[832,852,1844,1870]" pageId="2" pageNumber="21">A</emphasis>
. Branch.
<emphasis id="B95A94502928FF8EFC171277FCFBCF49" bold="true" box="[970,988,1845,1871]" pageId="2" pageNumber="21">B</emphasis>
. Stipule.
<emphasis id="B95A94502928FF8EFB8F1276FB41CF48" bold="true" box="[1106,1126,1844,1870]" pageId="2" pageNumber="21">C</emphasis>
. Abaxial leaf surfaces.
<emphasis id="B95A94502928FF8EFF63121AFFF5CF74" bold="true" box="[190,210,1880,1906]" pageId="2" pageNumber="21">D</emphasis>
. Calyces.
<emphasis id="B95A94502928FF8EFE95121AFE5BCF74" bold="true" box="[328,380,1880,1906]" pageId="2" pageNumber="21">E–F</emphasis>
. Close-up of flowers.
<emphasis id="B95A94502928FF8EFDA1121AFD9ECF74" bold="true" box="[636,697,1880,1906]" pageId="2" pageNumber="21">G–H</emphasis>
. Mature fruits.
<emphasis id="B95A94502928FF8EFCB7121AFC52CF74" bold="true" box="[874,885,1880,1906]" pageId="2" pageNumber="21">I</emphasis>
. Seeds. Photos by Van-Son Dang on 22 Apr. 2021.
</paragraph>
</caption>
<caption id="DF5118CA2929FF8FFF6012E8FA69CFEE" ID-DOI="http://doi.org/10.5281/zenodo.6373949" ID-Zenodo-Dep="6373949" httpUri="https://zenodo.org/record/6373949/files/figure.png" pageId="3" pageNumber="22" startId="3.[189,232,1962,1988]" targetBox="[233,1359,288,1884]" targetPageId="3">
<paragraph id="8B9148422929FF8FFF6012E8FA69CFEE" blockId="3.[189,1398,1962,2024]" pageId="3" pageNumber="22">
<emphasis id="B95A94502929FF8FFF6012E8FE2BCFC2" bold="true" box="[189,268,1962,1988]" pageId="3" pageNumber="22">Fig. 2.</emphasis>
<taxonomicName id="4C2E33C12929FF8FFEC812E8FDF5CFC3" authority="V.S.Dang &amp; Vuong" authorityName="V. S. Dang &amp; Vuong" authorityYear="2022" box="[277,722,1962,1989]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="22" phylum="Tracheophyta" rank="species" species="naikii" status="sp. nov.">
<emphasis id="B95A94502929FF8FFEC812E8FEC5CFC2" box="[277,482,1962,1988]" italics="true" pageId="3" pageNumber="22">Lasianthus naikii</emphasis>
V.S.Dang &amp; Vuong
</taxonomicName>
<taxonomicNameLabel id="A269292B2929FF8FFD0612E9FC10CFC3" box="[731,823,1963,1989]" pageId="3" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B95A94502929FF8FFC9D12E8FC73CFC2" bold="true" box="[832,852,1962,1988]" pageId="3" pageNumber="22">A</emphasis>
. Branch.
<emphasis id="B95A94502929FF8FFC1712E8FCFBCFC2" bold="true" box="[970,988,1962,1988]" pageId="3" pageNumber="22">B</emphasis>
. Abaxial leaf surfaces.
<emphasis id="B95A94502929FF8FFB2A12E8FA2CCFC2" bold="true" box="[1271,1291,1962,1988]" pageId="3" pageNumber="22">C</emphasis>
. Stipule.
<emphasis id="B95A94502929FF8FFF63128FFFF5CFE1" bold="true" box="[190,210,1997,2023]" pageId="3" pageNumber="22">D</emphasis>
. Close-up of flower.
<emphasis id="B95A94502929FF8FFE17128FFEF9CFE1" bold="true" box="[458,478,1997,2023]" pageId="3" pageNumber="22">E</emphasis>
. Fruit. Line drawings from the holotype (
<emphasis id="B95A94502929FF8FFC1A128CFB1BCFE1" box="[967,1084,1997,2023]" italics="true" pageId="3" pageNumber="22">Dang 453</emphasis>
) by Ba-Vuong Truong.
</paragraph>
</caption>
<caption id="DF5118CA292EFF88FF601277FA02CF74" ID-DOI="http://doi.org/10.5281/zenodo.6373951" ID-Zenodo-Dep="6373951" httpUri="https://zenodo.org/record/6373951/files/figure.png" pageId="4" pageNumber="23" startId="4.[189,232,1845,1871]" targetBox="[189,1399,265,1806]" targetPageId="4">
<paragraph id="8B914842292EFF88FF601277FA02CF74" blockId="4.[189,1399,1845,1907]" pageId="4" pageNumber="23">
<emphasis id="B95A9450292EFF88FF601277FE2ACF49" bold="true" box="[189,269,1845,1871]" pageId="4" pageNumber="23">Fig. 3.</emphasis>
<taxonomicName id="4C2E33C1292EFF88FECB1277FC7FCF49" authority="Matsumura. A. Fruiting" box="[278,856,1845,1871]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="23" phylum="Tracheophyta" rank="species" species="formosensis">
<emphasis id="B95A9450292EFF88FECB1277FD17CF49" box="[278,560,1845,1871]" italics="true" pageId="4" pageNumber="23">Lasianthus formosensis</emphasis>
Matsumura.
<emphasis id="B95A9450292EFF88FD0F1277FDC0CF49" bold="true" box="[722,743,1845,1871]" pageId="4" pageNumber="23">A</emphasis>
. Fruiting
</taxonomicName>
branch.
<emphasis id="B95A9450292EFF88FC1E1277FCF0CF49" bold="true" box="[963,983,1845,1871]" pageId="4" pageNumber="23">B</emphasis>
. Stipule.
<emphasis id="B95A9450292EFF88FB901277FB46CF49" bold="true" box="[1101,1121,1845,1871]" pageId="4" pageNumber="23">C</emphasis>
. Abaxial leaf surfaces.
<emphasis id="B95A9450292EFF88FF62121AFFF4CF74" bold="true" box="[191,211,1880,1906]" pageId="4" pageNumber="23">D</emphasis>
. Close-up of inflorescences and flowers.
<emphasis id="B95A9450292EFF88FD6B121AFDEDCF74" bold="true" box="[694,714,1880,1906]" pageId="4" pageNumber="23">E</emphasis>
. Fruits. Photos by Van-Son Dang on 17 Nov. 2018.
</paragraph>
</caption>
<caption id="DF5118CA292FFF89FF60144BFD32C940" ID-Table-UUID="DF5118CA292FFF89FF60144BFD32C940" httpUri="http://table.plazi.org/id/DF5118CA292FFF89FF60144BFD32C940" pageId="5" pageNumber="24" startId="5.[189,258,265,291]" targetBox="[197,1358,366,713]" targetIsTable="true" targetPageId="5">
<paragraph id="8B914842292FFF89FF60144BFD32C940" blockId="5.[189,1399,265,327]" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF60144BFE07C925" bold="true" box="[189,288,265,291]" pageId="5" pageNumber="24">Table 1.</emphasis>
Comparison of
<taxonomicName id="4C2E33C1292FFF89FE3C144BFCBFC922" authority="V.S.Dang &amp; Vuong" authorityName="V. S. Dang &amp; Vuong" authorityYear="2022" box="[481,920,265,292]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="naikii" status="sp. nov.">
<emphasis id="B95A9450292FFF89FE3C144BFD8AC925" box="[481,685,265,291]" italics="true" pageId="5" pageNumber="24">Lasianthus naikii</emphasis>
V.S.Dang &amp; Vuong
</taxonomicName>
<taxonomicNameLabel id="A269292B292FFF89FC7D1448FCDCC922" box="[928,1019,266,292]" pageId="5" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C2E33C1292FFF89FBEB1448FA50C922" authority="Matsumura." authorityName="Matsumura." box="[1078,1399,265,292]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="formosensis">
<emphasis id="B95A9450292FFF89FBEB1448FBC6C925" box="[1078,1249,265,291]" italics="true" pageId="5" pageNumber="24">L. formosensis</emphasis>
Matsumura.
</taxonomicName>
* = from
<bibRefCitation id="EFBF35B3292FFF89FEF1146EFD36C940" author="Zhu H. &amp; Taylor C. M." box="[300,529,300,326]" pageId="5" pageNumber="24" pagination="185 - 198" refId="ref5573" refString="Zhu H. &amp; Taylor C. M. 2011. Lasianthus. In: Wu Z. Y., Raven P. H. &amp; Hong D. Y. (eds) Flora of China, Vol. 19: 185 - 198. Science Press, Beijing, and Missouri Botanical Garden Press, St. Louis." type="journal article" year="2011">Zhu &amp; Taylor 2011</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8B914842292FFF89FF18142DFA31CACE" pageId="5" pageNumber="24">
<table id="F92EBAE2292F0073FF18142CFA69CACF" box="[197,1358,366,713]" gridcols="3" gridrows="10" pageId="5" pageNumber="24">
<tr id="351E4A00292F0073FF18142CFA69C98F" box="[197,1358,366,393]" gridrow="0" pageId="5" pageNumber="24">
<th id="76CF237C292F0073FF18142CFEB7C98F" box="[197,400,366,393]" gridcol="0" gridrow="0" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF18142DFE76C98F" bold="true" box="[197,337,367,393]" pageId="5" pageNumber="24">Characters</emphasis>
</th>
<th id="76CF237C292F0073FE38142CFC67C98F" box="[485,832,366,393]" gridcol="1" gridrow="0" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FE38142CFC39C98F" bold="true" box="[485,798,366,393]" pageId="5" pageNumber="24">
<taxonomicName id="4C2E33C1292FFF89FE38142CFD9EC98F" authority="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang, 2022" authorityName="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang" authorityYear="2022" box="[485,697,366,393]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="naikii" status="sp. nov.">
<emphasis id="B95A9450292FFF89FE38142CFD9EC98F" bold="true" box="[485,697,366,393]" italics="true" pageId="5" pageNumber="24">Lasianthus naikii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269292B292FFF89FD1C142DFC39C98F" box="[705,798,367,393]" pageId="5" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76CF237C292F0073FC70142CFA69C98F" box="[941,1358,366,393]" gridcol="2" gridrow="0" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FC70142CFBEBC98F" bold="true" box="[941,1228,366,393]" italics="true" pageId="5" pageNumber="24">Lasianthus formosensis</emphasis>
*
</th>
</tr>
<tr id="351E4A00292F0073FF1814D0FA69C9AB" box="[197,1358,402,429]" gridrow="1" pageId="5" pageNumber="24">
<th id="76CF237C292F0073FF1814D0FEB7C9AB" box="[197,400,402,429]" gridcol="0" gridrow="1" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF1814D0FE72C9AA" bold="true" box="[197,341,402,428]" pageId="5" pageNumber="24">Leaf blades</emphasis>
</th>
<td id="76CF237C292F0073FE3814D0FC67C9AB" box="[485,832,402,429]" gridcol="1" gridrow="1" pageId="5" pageNumber="24">12–14 × 3–5 cm</td>
<td id="76CF237C292F0073FC7014D0FA69C9AB" box="[941,1358,402,429]" gridcol="2" gridrow="1" pageId="5" pageNumber="24">7–12 × 2.5–5 cm</td>
</tr>
<tr id="351E4A00292F0073FF1814F7FA69C9D7" box="[197,1358,437,465]" gridrow="2" pageId="5" pageNumber="24">
<th id="76CF237C292F0073FF1814F7FEB7C9D7" box="[197,400,437,465]" gridcol="0" gridrow="2" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF1814F7FEB7C9C9" bold="true" box="[197,400,437,463]" pageId="5" pageNumber="24">Secondary veins</emphasis>
</th>
<td id="76CF237C292F0073FE3814F7FC67C9D7" box="[485,832,437,465]" gridcol="1" gridrow="2" pageId="5" pageNumber="24">5–6 pairs</td>
<td id="76CF237C292F0073FC7014F7FA69C9D7" box="[941,1358,437,465]" gridcol="2" gridrow="2" pageId="5" pageNumber="24">6–7 pairs</td>
</tr>
<tr id="351E4A00292F0073FF18149BFA69C9F2" box="[197,1358,473,500]" gridrow="3" pageId="5" pageNumber="24">
<th id="76CF237C292F0073FF18149BFEB7C9F2" box="[197,400,473,500]" gridcol="0" gridrow="3" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF18149BFE02C9F5" bold="true" box="[197,293,473,499]" pageId="5" pageNumber="24">Petioles</emphasis>
</th>
<td id="76CF237C292F0073FE38149BFC67C9F2" box="[485,832,473,500]" gridcol="1" gridrow="3" pageId="5" pageNumber="24">densely villous, 5–7 mm long</td>
<td id="76CF237C292F0073FC70149BFA69C9F2" box="[941,1358,473,500]" gridcol="2" gridrow="3" pageId="5" pageNumber="24">villous to villosulous, 5–8 mm long</td>
</tr>
<tr id="351E4A00292F0073FF1814BEFA69CA1E" box="[197,1358,508,536]" gridrow="4" pageId="5" pageNumber="24">
<th id="76CF237C292F0073FF1814BEFEB7CA1E" box="[197,400,508,536]" gridcol="0" gridrow="4" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF1814BEFE0FCA10" bold="true" box="[197,296,508,534]" pageId="5" pageNumber="24">Stipules</emphasis>
</th>
<td id="76CF237C292F0073FE3814BEFC67CA1E" box="[485,832,508,536]" gridcol="1" gridrow="4" pageId="5" pageNumber="24">1.5–2 mm long</td>
<td id="76CF237C292F0073FC7014BEFA69CA1E" box="[941,1358,508,536]" gridcol="2" gridrow="4" pageId="5" pageNumber="24">ca 2 mm long</td>
</tr>
<tr id="351E4A00292F0073FF181762FA69CA3D" box="[197,1358,544,571]" gridrow="5" pageId="5" pageNumber="24">
<th id="76CF237C292F0073FF181762FEB7CA3D" box="[197,400,544,571]" gridcol="0" gridrow="5" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF181762FE31CA3C" bold="true" box="[197,278,544,570]" pageId="5" pageNumber="24">Bracts</emphasis>
</th>
<td id="76CF237C292F0073FE381762FC67CA3D" box="[485,832,544,571]" gridcol="1" gridrow="5" pageId="5" pageNumber="24">0.5–1 mm long</td>
<td id="76CF237C292F0073FC701762FA69CA3D" box="[941,1358,544,571]" gridcol="2" gridrow="5" pageId="5" pageNumber="24">absent</td>
</tr>
<tr id="351E4A00292F0073FF181701FA69CA59" box="[197,1358,579,607]" gridrow="6" pageId="5" pageNumber="24">
<th id="76CF237C292F0073FF181701FEB7CA59" box="[197,400,579,607]" gridcol="0" gridrow="6" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF181701FE6BCA58" bold="true" box="[197,332,579,606]" pageId="5" pageNumber="24">Calyx tube</emphasis>
</th>
<td id="76CF237C292F0073FE381701FC67CA59" box="[485,832,579,607]" gridcol="1" gridrow="6" pageId="5" pageNumber="24">2–5 mm long</td>
<td id="76CF237C292F0073FC701701FA69CA59" box="[941,1358,579,607]" gridcol="2" gridrow="6" pageId="5" pageNumber="24">ca 1 mm long</td>
</tr>
<tr id="351E4A00292F0073FF181725FA69CA84" box="[197,1358,615,642]" gridrow="7" pageId="5" pageNumber="24">
<th id="76CF237C292F0073FF181725FEB7CA84" box="[197,400,615,642]" gridcol="0" gridrow="7" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF181725FE72CA87" bold="true" box="[197,341,615,641]" pageId="5" pageNumber="24">Calyx lobes</emphasis>
</th>
<td id="76CF237C292F0073FE381725FC67CA84" box="[485,832,615,642]" gridcol="1" gridrow="7" pageId="5" pageNumber="24">1–1.5 mm long, 6-lobed</td>
<td id="76CF237C292F0073FC701725FA69CA84" box="[941,1358,615,642]" gridcol="2" gridrow="7" pageId="5" pageNumber="24">3–4 mm long, 5-lobed</td>
</tr>
<tr id="351E4A00292F0073FF1817C8FA69CAA3" box="[197,1358,650,677]" gridrow="8" pageId="5" pageNumber="24">
<th id="76CF237C292F0073FF1817C8FEB7CAA3" box="[197,400,650,677]" gridcol="0" gridrow="8" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF1817C8FE04CAA2" bold="true" box="[197,291,650,676]" pageId="5" pageNumber="24">Corolla</emphasis>
</th>
<td id="76CF237C292F0073FE3817C8FC67CAA3" box="[485,832,650,677]" gridcol="1" gridrow="8" pageId="5" pageNumber="24">6-lobed</td>
<td id="76CF237C292F0073FC7017C8FA69CAA3" box="[941,1358,650,677]" gridcol="2" gridrow="8" pageId="5" pageNumber="24">5-lobed</td>
</tr>
<tr id="351E4A00292F0073FF1817ECFA69CACF" box="[197,1358,686,713]" gridrow="9" pageId="5" pageNumber="24">
<th id="76CF237C292F0073FF1817ECFEB7CACF" box="[197,400,686,713]" gridcol="0" gridrow="9" pageId="5" pageNumber="24">
<emphasis id="B95A9450292FFF89FF1817ECFE35CACE" bold="true" box="[197,274,686,712]" pageId="5" pageNumber="24">Fruits</emphasis>
</th>
<td id="76CF237C292F0073FE3817ECFC67CACF" box="[485,832,686,713]" gridcol="1" gridrow="9" pageId="5" pageNumber="24">strigose, 6–7 mm in diam.</td>
<td id="76CF237C292F0073FC7017ECFA69CACF" box="[941,1358,686,713]" gridcol="2" gridrow="9" pageId="5" pageNumber="24">subglabrous, ca 5 mm in diam.</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3341BC9292FFF89FF601647FC9ACB68" pageId="5" pageNumber="24" type="etymology">
<paragraph id="8B914842292FFF89FF601647FE63CB19" blockId="5.[189,324,773,799]" box="[189,324,773,799]" pageId="5" pageNumber="24">
<heading id="D0D9FF2E292FFF89FF601647FE63CB19" bold="true" box="[189,324,773,799]" fontSize="11" level="3" pageId="5" pageNumber="24" reason="3">
<emphasis id="B95A9450292FFF89FF601647FE63CB19" bold="true" box="[189,324,773,799]" pageId="5" pageNumber="24">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292FFF89FF601670FC9ACB68" blockId="5.[188,1398,818,878]" pageId="5" pageNumber="24">
The new species is named in honor of Associate Professor Dr Akyo Naiki of University of the Ryukyus,
<collectingCountry id="F33908D2292FFF89FF611616FE25CB68" box="[188,258,852,878]" name="Japan" pageId="5" pageNumber="24">Japan</collectingCountry>
, for his contributions to the study of
<taxonomicName id="4C2E33C1292FFF89FD6E1616FC0ACB68" box="[691,813,852,878]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
in
<collectingCountry id="F33908D2292FFF89FC8F1616FC9ECB68" box="[850,953,852,878]" name="Vietnam" pageId="5" pageNumber="24">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292FFF89FF6016D6FEB0CBDD" pageId="5" pageNumber="24" type="vernacular_names">
<paragraph id="8B914842292FFF89FF6016D6FEB3CBA8" blockId="5.[189,404,916,942]" box="[189,404,916,942]" pageId="5" pageNumber="24">
<heading id="D0D9FF2E292FFF89FF6016D6FEB3CBA8" bold="true" box="[189,404,916,942]" fontSize="11" level="3" pageId="5" pageNumber="24" reason="3">
<emphasis id="B95A9450292FFF89FF6016D6FEB3CBA8" bold="true" box="[189,404,916,942]" pageId="5" pageNumber="24">Vernacular name</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292FFF89FF601683FEB0CBDD" blockId="5.[189,407,961,989]" box="[189,407,961,989]" pageId="5" pageNumber="24">Xú hƯƠng ʻNaikiʼ.</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292FFF89FF601143FEE1CCA8" pageId="5" pageNumber="24" type="materials_examined">
<paragraph id="8B914842292FFF89FF601143FE4BCC1D" blockId="5.[189,364,1025,1051]" box="[189,364,1025,1051]" pageId="5" pageNumber="24">
<heading id="D0D9FF2E292FFF89FF601143FE4BCC1D" bold="true" box="[189,364,1025,1051]" fontSize="11" level="3" pageId="5" pageNumber="24" reason="3">
<emphasis id="B95A9450292FFF89FF601143FE4BCC1D" bold="true" box="[189,364,1025,1051]" pageId="5" pageNumber="24">
<typeStatus id="5495F6E0292FFF89FF601143FFDDCC1D" box="[189,250,1025,1051]" pageId="5" pageNumber="24">Type</typeStatus>
material
</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292FFF89FF60116CFEE1CCA8" blockId="5.[189,1399,1070,1198]" pageId="5" pageNumber="24">
<materialsCitation id="3B46421F292FFF89FF60116CFA51CC8A" ID-GBIF-Occurrence="3714053304" collectedFrom="primary evergreen forests" collectingDate="2021-04-22" collectionCode="VNM" collectorName="Van- Son Dang &amp; Ba Vuong Truong &amp; Minh Quan Dang &amp; Nghia Son Hoang Dang" country="Vietnam" county="K'Bang District" elevation="1050" latitude="14.486112" location="Kon Chu Rang Nature Reserve" longLatPrecision="1" longitude="108.53864" municipality="Son Lang Commune" pageId="5" pageNumber="24" specimenCount="1" stateProvince="Gia Lai Province" typeStatus="holotype">
<collectingCountry id="F33908D2292FFF89FF60116CFE63CC4E" box="[189,324,1070,1096]" name="Vietnam" pageId="5" pageNumber="24">VIETNAM</collectingCountry>
•
<collectingRegion id="49EA86A0292FFF89FE80116CFD0ECC4E" box="[349,553,1070,1096]" country="Vietnam" name="Gia Lai" pageId="5" pageNumber="24">Gia Lai Province</collectingRegion>
,
<collectingCounty id="62F030CE292FFF89FDE8116CFDD3CC4E" box="[565,756,1070,1096]" pageId="5" pageNumber="24">K’Bang District</collectingCounty>
,
<collectingMunicipality id="6BF5D238292FFF89FCDD116CFCD1CC4E" box="[768,1014,1070,1097]" pageId="5" pageNumber="24">Son Lang Commune</collectingMunicipality>
,
<location id="8EF11E99292FFF89FBDF116DFA55CC4F" LSID="urn:lsid:plazi:treatment:0387F954292BFF8AFDD713DBFE83CA2F:8EF11E99292FFF89FBDF116DFA55CC4F" box="[1026,1394,1070,1097]" country="Vietnam" county="K'Bang District" latitude="14.486112" longLatPrecision="1" longitude="108.53864" municipality="Son Lang Commune" name="Kon Chu Rang Nature Reserve" pageId="5" pageNumber="24" stateProvince="Gia Lai Province">Kon Chu Rang Nature Reserve</location>
,
<collectedFrom id="38320BBF292FFF89FF601112FECACC6C" box="[189,493,1104,1130]" pageId="5" pageNumber="24">primary evergreen forests</collectedFrom>
;
<geoCoordinate id="EE1A2E85292FFF89FE24110CFD95CC6C" box="[505,690,1102,1130]" degrees="14" direction="north" minutes="29" orientation="latitude" pageId="5" pageNumber="24" precision="1" seconds="10.00" value="14.486112">14°29′10.00″ N</geoCoordinate>
,
<geoCoordinate id="EE1A2E85292FFF89FD63110CFCA3CC6C" box="[702,900,1102,1130]" degrees="108" direction="east" minutes="32" orientation="longitude" pageId="5" pageNumber="24" precision="1" seconds="19.12" value="108.53864">108°32′19.12″ E</geoCoordinate>
; at
<elevation id="0003AF71292FFF89FC711112FBAACC6C" box="[940,1165,1104,1130]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="5" pageNumber="24" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">
<quantity id="4CD6E5A7292FFF89FC711112FB6ACC6C" box="[940,1101,1104,1130]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="5" pageNumber="24" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1000–1100 m</quantity>
a.s.l.
</elevation>
;
<date id="FF906E82292FFF89FB471112FA12CC6C" box="[1178,1333,1104,1130]" pageId="5" pageNumber="24" value="2021-04-22">
<collectingDate id="EFD4976A292FFF89FB471112FA12CC6C" box="[1178,1333,1104,1130]" pageId="5" pageNumber="24" value="2021-04-22">22 Apr. 2021</collectingDate>
</date>
;
<emphasis id="B95A9450292FFF89FA9C1113FBB6CC8A" italics="true" pageId="5" pageNumber="24">
<collectorName id="26DB2D94292FFF89FA9C1113FE1DCC8A" pageId="5" pageNumber="24">Van- Son Dang</collectorName>
,
<collectorName id="26DB2D94292FFF89FE951131FD05CC8A" box="[328,546,1139,1164]" pageId="5" pageNumber="24">Ba Vuong Truong</collectorName>
,
<collectorName id="26DB2D94292FFF89FDEC1130FC2FCC8A" box="[561,776,1138,1164]" pageId="5" pageNumber="24">Minh Quan Dang</collectorName>
&amp;
<collectorName id="26DB2D94292FFF89FCEE1130FB7DCC8A" box="[819,1114,1138,1164]" pageId="5" pageNumber="24">Nghia Son Hoang Dang</collectorName>
453
</emphasis>
;
<typeStatus id="5495F6E0292FFF89FB791130FA35CC8A" box="[1188,1298,1138,1164]" pageId="5" pageNumber="24" type="holotype">holotype</typeStatus>
:
<collectionCode id="ED3FD087292FFF89FAC31130FA4ACC8A" box="[1310,1389,1138,1164]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="5" pageNumber="24" type="Herbarium">VNM</collectionCode>
!;
</materialsCitation>
<materialsCitation id="3B46421F292FFF89FF6211D1FE98CCA8" ID-GBIF-Occurrence="3714053301" box="[191,447,1171,1198]" collectedFrom="primary evergreen forests" collectingDate="2021-04-22" collectionCode="HN, VNM" collectorName="Van- Son Dang &amp; Ba Vuong Truong &amp; Minh Quan Dang &amp; Nghia Son Hoang Dang" country="Vietnam" county="K'Bang District" elevation="1050" latitude="14.486112" location="Kon Chu Rang Nature Reserve" longLatPrecision="1" longitude="108.53864" municipality="Son Lang Commune" pageId="5" pageNumber="24" specimenCount="1" stateProvince="Gia Lai Province" typeStatus="isotype">
<typeStatus id="5495F6E0292FFF89FF6211D1FE03CCAB" box="[191,292,1171,1197]" pageId="5" pageNumber="24" type="isotype">isotypes</typeStatus>
:
<collectionCode id="ED3FD087292FFF89FEF311D6FE46CCA8" box="[302,353,1172,1198]" pageId="5" pageNumber="24">HN</collectionCode>
!,
<collectionCode id="ED3FD087292FFF89FEAC11D6FE98CCA8" box="[369,447,1172,1198]" pageId="5" pageNumber="24">VNM</collectionCode>
</materialsCitation>
!.
</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292FFF89FF601191FE15CEF2" pageId="5" pageNumber="24" type="description">
<paragraph id="8B914842292FFF89FF601191FE69CCEB" blockId="5.[189,334,1235,1261]" box="[189,334,1235,1261]" pageId="5" pageNumber="24">
<heading id="D0D9FF2E292FFF89FF601191FE69CCEB" bold="true" box="[189,334,1235,1261]" fontSize="11" level="3" pageId="5" pageNumber="24" reason="3">
<emphasis id="B95A9450292FFF89FF601191FE69CCEB" bold="true" box="[189,334,1235,1261]" pageId="5" pageNumber="24">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292FFF89FF601043FE15CEF2" blockId="5.[187,1399,1280,1780]" pageId="5" pageNumber="24">
Shrubs, evergreen,
<quantity id="4CD6E5A7292FFF89FE451043FEC7CD1D" box="[408,480,1281,1307]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="24" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
tall; branches and branchlets terete,
<quantity id="4CD6E5A7292FFF89FCA01042FCD7CD1D" box="[893,1008,1280,1307]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="5" pageNumber="24" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 mm</quantity>
diam., densely villous, internodes
<quantity id="4CD6E5A7292FFF89FF661061FE33CD3B" box="[187,276,1315,1341]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="24" unit="cm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 cm</quantity>
long. Leaves opposite, blades oblong or elliptic-ovate, 12–14 ×
<quantity id="4CD6E5A7292FFF89FBD21060FB4DCD3B" box="[1039,1130,1314,1341]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="24" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 cm</quantity>
, coriaceous, yellowish brown when dry, adaxially glabrous, abaxially villous especially on the midrib and secondary veins, apex acuminate or cuspidate, aristate
<quantity id="4CD6E5A7292FFF89FDF11024FDADCD87" box="[556,650,1382,1409]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="5" pageNumber="24" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long, villous, base acute or obtuse, margin entire; midrib slightly prominent adaxially, prominent abaxially; secondary veins 5–6 pairs, ascending at an angle of 40–60° from midrib, curved to the margin, flat adaxially, distinct abaxially; tertiary veins reticulate; petioles
<quantity id="4CD6E5A7292FFF89FAC910E8FA50CDC2" box="[1300,1399,1450,1477]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="5" pageNumber="24" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
long, densely villous. Stipules triangular,
<quantity id="4CD6E5A7292FFF89FD691089FC09CDE0" box="[692,814,1483,1510]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="5" pageNumber="24" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
long, persistent, villous. Inflorescences cymose, sessile, 1–3-flowered; bracts small,
<quantity id="4CD6E5A7292FFF89FD8410AFFDE8CE0E" box="[601,719,1517,1544]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="5" pageNumber="24" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long, villous. Flowers sessile; calyx campanulate,
<quantity id="4CD6E5A7292FFF89FACA10AFFA50CE0E" box="[1303,1399,1517,1544]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="24" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long, yellowish brown, densely villous outside, calyx tube
<quantity id="4CD6E5A7292FFF89FC84134DFCEACE2C" box="[857,973,1551,1578]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="5" pageNumber="24" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
long, calyx lobes 6, linear-triangular,
<quantity id="4CD6E5A7292FFF89FF661373FE14CE4D" box="[187,307,1585,1612]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="24" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long, tips acute; corolla cylindrical,
<quantity id="4CD6E5A7292FFF89FD0A1373FC10CE4D" box="[727,823,1585,1611]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="24" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long, white, corolla tube
<quantity id="4CD6E5A7292FFF89FB861373FB9CCE4D" box="[1115,1211,1585,1612]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="24" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
long, pubescent outside, corolla lobes 6, triangular,
<quantity id="4CD6E5A7292FFF89FD961311FD8CCE6B" box="[587,683,1619,1645]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="24" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, puberulous outside, floccose inside; stamens 6, filaments short, anthers oblong,
<quantity id="4CD6E5A7292FFF89FE661336FD08CE89" box="[443,559,1652,1679]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="5" pageNumber="24" unit="mm" value="0.8" valueMax="1.0" valueMin="0.6">0.6–1 mm</quantity>
long; ovary with 5 locules, style linear,
<quantity id="4CD6E5A7292FFF89FC291336FB75CE89" box="[1012,1106,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="24" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
long, stigma
<quantity id="4CD6E5A7292FFF89FB311336FA51CE89" box="[1260,1398,1652,1679]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="24" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
long. Fruit drupaceous, ovoid-globose,
<quantity id="4CD6E5A7292FFF89FD5F13D4FDC6CEB7" box="[642,737,1686,1713]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="5" pageNumber="24" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
long,
<quantity id="4CD6E5A7292FFF89FCF513D4FCA1CEB7" box="[808,902,1686,1713]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="24" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
in diam., strigose, crowned by calyx lobes, blue when ripe; pyrenes 5. Seeds ovoid,
<quantity id="4CD6E5A7292FFF89FD4313FAFC27CED5" box="[670,768,1720,1747]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="24" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, smooth to subwarty on the abaxial face, brown when dry.
</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292FFF8AFF601258FE83CA2F" lastPageId="6" lastPageNumber="25" pageId="5" pageNumber="24" type="distribution">
<subSubSection id="C3341BC9292FFF8AFF601258FBE5C96A" lastPageId="6" lastPageNumber="25" pageId="5" pageNumber="24" type="distribution">
<paragraph id="8B914842292FFF89FF601258FD5ACF32" blockId="5.[189,637,1818,1844]" box="[189,637,1818,1844]" pageId="5" pageNumber="24">
<heading id="D0D9FF2E292FFF89FF601258FD5ACF32" bold="true" box="[189,637,1818,1844]" fontSize="11" level="3" pageId="5" pageNumber="24" reason="3">
<emphasis id="B95A9450292FFF89FF601258FD5ACF32" bold="true" box="[189,637,1818,1844]" pageId="5" pageNumber="24">Distribution, habitat and phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292FFF8AFF601205FBE5C96A" blockId="5.[189,1400,1863,2025]" lastBlockId="6.[189,1398,265,365]" lastPageId="6" lastPageNumber="25" pageId="5" pageNumber="24">
<taxonomicName id="4C2E33C1292FFF89FF601205FEA9CF67" authority="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang, 2022" authorityName="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang" authorityYear="2022" box="[189,398,1863,1889]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="naikii" status="sp. nov.">
<emphasis id="B95A9450292FFF89FF601205FEA9CF67" box="[189,398,1863,1889]" italics="true" pageId="5" pageNumber="24">Lasianthus naikii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269292B292FFF89FE45120AFED0CF64" box="[408,503,1864,1890]" pageId="5" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
is only known from its
<typeStatus id="5495F6E0292FFF89FCFC120AFC72CF64" box="[801,853,1864,1890]" pageId="5" pageNumber="24">type</typeStatus>
locality at
<quantity id="4CD6E5A7292FFF89FC3C1205FBAFCF64" box="[993,1160,1863,1890]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="5" pageNumber="24" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1000–1100 m</quantity>
elevation. It grows under the shade of primary evergreen forest on clay, which is dominated by
<taxonomicName id="4C2E33C1292FFF89FBAF122BFA50CF85" authority="Roxb." authorityName="Roxb." box="[1138,1399,1897,1923]" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="cowa">
<emphasis id="B95A9450292FFF89FBAF122BFA03CF85" box="[1138,1316,1897,1923]" italics="true" pageId="5" pageNumber="24">Garcinia cowa</emphasis>
Roxb.
</taxonomicName>
(
<taxonomicName id="4C2E33C1292FFF89FF1812C9FEBFCFA0" authority="Lindl." authorityName="Lindl." box="[197,408,1931,1958]" class="Magnoliopsida" family="Clusiaceae" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="family">Clusiaceae Lindl.</taxonomicName>
),
<taxonomicName id="4C2E33C1292FFF89FE7112C9FDA1CFA3" box="[428,646,1931,1957]" class="Magnoliopsida" family="Bignoniaceae" genus="Oroxylum" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="indicum">
<emphasis id="B95A9450292FFF89FE7112C9FDA1CFA3" box="[428,646,1931,1957]" italics="true" pageId="5" pageNumber="24">Oroxylum indicum</emphasis>
</taxonomicName>
(L.) Kurz (
<taxonomicName id="4C2E33C1292FFF89FCD412C9FCCACFA3" authority="Juss." authorityName="Juss." box="[777,1005,1931,1957]" class="Magnoliopsida" family="Bignoniaceae" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="family">Bignoniaceae Juss.</taxonomicName>
),
<taxonomicName id="4C2E33C1292FFF89FBDF12C9FFDBCFC0" authority="Wall." authorityName="Wall." class="Magnoliopsida" family="Apocynaceae" genus="Tabernaemontana" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="peduncularis">
<emphasis id="B95A9450292FFF89FBDF12C9FA51CFA3" box="[1026,1398,1931,1957]" italics="true" pageId="5" pageNumber="24">Tabernaemontana peduncularis</emphasis>
Wall.
</taxonomicName>
(
<taxonomicName id="4C2E33C1292FFF89FED412EFFECCCFC1" authority="Juss." authorityName="Juss." box="[265,491,1965,1991]" class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="family">Apocynaceae Juss.</taxonomicName>
),
<taxonomicName id="4C2E33C1292FFF89FE2212EEFC4FCFC0" authority="Pierre" authorityName="Pierre" box="[511,872,1964,1990]" class="Magnoliopsida" family="Fabaceae" genus="Dialium" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="cochinchinense">
<emphasis id="B95A9450292FFF89FE2212EEFC3CCFC0" box="[511,795,1964,1990]" italics="true" pageId="5" pageNumber="24">Dialium cochinchinense</emphasis>
Pierre
</taxonomicName>
(
<taxonomicName id="4C2E33C1292FFF89FCAB12EEFB10CFC1" authority="Lindl." authorityName="Lindl." box="[886,1079,1964,1991]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="family">Fabaceae Lindl.</taxonomicName>
),
<taxonomicName id="4C2E33C1292FFF89FB9612EEFE06CFEE" authority="Gagnep." authorityName="Gagnep." class="Magnoliopsida" family="Elaeocarpaceae" genus="Elaeocarpus" kingdom="Plantae" order="Oxalidales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="kontumensis">
<emphasis id="B95A9450292FFF89FB9612EEFA51CFC0" box="[1099,1398,1964,1990]" italics="true" pageId="5" pageNumber="24">Elaeocarpus kontumensis</emphasis>
Gagnep.
</taxonomicName>
(
<taxonomicName id="4C2E33C1292FFF89FEEF128CFD15CFEF" authority="Juss." authorityName="Juss." box="[306,562,1998,2025]" class="Magnoliopsida" family="Elaeocarpaceae" kingdom="Plantae" order="Oxalidales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="family">Elaeocarpaceae Juss.</taxonomicName>
),
<taxonomicName id="4C2E33C1292FFF89FD96128DFC95CFEE" authority="(Nees) Hook." authorityName="Hook." baseAuthorityName="Nees" box="[587,946,1998,2024]" class="Magnoliopsida" family="Lauraceae" genus="Litsea" kingdom="Plantae" order="Laurales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="elongata">
<emphasis id="B95A9450292FFF89FD96128DFC22CFEE" box="[587,773,1998,2024]" italics="true" pageId="5" pageNumber="24">Litsea elongata</emphasis>
(Nees) Hook.
</taxonomicName>
f. (
<taxonomicName id="4C2E33C1292FFF89FC0F128CFBB3CFEF" authority="Juss." authorityName="Juss." box="[978,1172,1998,2025]" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="family">Lauraceae Juss.</taxonomicName>
),
<taxonomicName id="4C2E33C1292FFF8AFB76128CFE2FC922" authority="Hance" authorityName="Hance" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" lastPageId="6" lastPageNumber="25" order="Gentianales" pageId="5" pageNumber="24" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis id="B95A9450292FFF89FB76128CFA5FCFEE" box="[1195,1400,1998,2024]" italics="true" pageId="5" pageNumber="24">Lasianthus fordii</emphasis>
Hance
</taxonomicName>
(
<taxonomicName id="4C2E33C1292CFF8AFECB144BFEF3C922" authority="Juss." authorityName="Juss." box="[278,468,265,292]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="25" phylum="Tracheophyta" rank="family">Rubiaceae Juss.</taxonomicName>
),
<taxonomicName id="4C2E33C1292CFF8AFE3A144BFCC8C922" authority="(Dandy) Figlar &amp; Noot." authorityName="Figlar &amp; Noot." baseAuthorityName="Dandy" box="[487,1007,265,292]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="6" pageNumber="25" phylum="Tracheophyta" rank="species" species="praecalva">
<emphasis id="B95A9450292CFF8AFE3A144BFDF3C925" box="[487,724,265,291]" italics="true" pageId="6" pageNumber="25">Magnolia praecalva</emphasis>
(Dandy) Figlar &amp; Noot.
</taxonomicName>
(
<taxonomicName id="4C2E33C1292CFF8AFC20144BFBC2C922" authority="Juss." authorityName="Juss." box="[1021,1253,265,292]" class="Magnoliopsida" family="Magnoliaceae" kingdom="Plantae" order="Magnoliales" pageId="6" pageNumber="25" phylum="Tracheophyta" rank="family">Magnoliaceae Juss.</taxonomicName>
),
<taxonomicName id="4C2E33C1292CFF8AFB24144BFD76C94E" authority="(Lour.) S. Moore" authorityName="S. Moore" baseAuthorityName="Lour." class="Magnoliopsida" family="Symplocaceae" genus="Symplocos" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="25" phylum="Tracheophyta" rank="species" species="cochinchinensis">
<emphasis id="B95A9450292CFF8AFB24144BFE59C94E" italics="true" pageId="6" pageNumber="25">Symplocos cochinchinensis</emphasis>
(Lour.) S.Moore
</taxonomicName>
(
<taxonomicName id="4C2E33C1292CFF8AFDBE146CFC47C94F" authority="Desf." authorityName="Desf." box="[611,864,302,329]" class="Magnoliopsida" family="Symplocaceae" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="25" phylum="Tracheophyta" rank="family">Symplocaceae Desf.</taxonomicName>
) and
<taxonomicName id="4C2E33C1292CFF8AFC74146CFE10C96B" authority="(Gagnep.) Merr. &amp; L. M. Perry" authorityName="Merr. &amp; L. M. Perry" baseAuthorityName="Gagnep." class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="25" phylum="Tracheophyta" rank="species" species="chanlos">
<emphasis id="B95A9450292CFF8AFC74146CFBA3C94E" box="[937,1156,302,328]" italics="true" pageId="6" pageNumber="25">Syzygium chanlos</emphasis>
(Gagnep.) Merr. &amp; L.M.Perry
</taxonomicName>
(
<taxonomicName id="4C2E33C1292CFF8AFE9B1410FD2FC96B" authority="Juss." authorityName="Juss." box="[326,520,338,365]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="25" phylum="Tracheophyta" rank="family">Myrtaceae Juss.</taxonomicName>
). Flowering and fruiting specimens were collected in April.
</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292CFF8AFF6014D7FE83CA2F" pageId="6" pageNumber="25" type="description">
<paragraph id="8B914842292CFF8AFF6014D7FEFCC9A9" blockId="6.[189,475,405,431]" box="[189,475,405,431]" pageId="6" pageNumber="25">
<heading id="D0D9FF2E292CFF8AFF6014D7FEFCC9A9" bold="true" box="[189,475,405,431]" fontSize="11" level="3" pageId="6" pageNumber="25" reason="3">
<emphasis id="B95A9450292CFF8AFF6014D7FEFCC9A9" bold="true" box="[189,475,405,431]" pageId="6" pageNumber="25">Population and threats</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292CFF8AFF601487FE83CA2F" blockId="6.[189,1399,453,553]" pageId="6" pageNumber="25">
The new species is so far only known from Kon Chu Rang Nature Reserve,
<collectingRegion id="49EA86A0292CFF8AFBE71487FA22C9D9" box="[1082,1285,453,479]" country="Vietnam" name="Gia Lai" pageId="6" pageNumber="25">Gia Lai Province</collectingRegion>
, southern
<collectingCountry id="F33908D2292CFF8AFF6014A8FE03CA02" box="[189,292,490,516]" name="Vietnam" pageId="6" pageNumber="25">Vietnam</collectingCountry>
. It was collected in a single population containing less than 50 individuals, occupying an area of less than
<quantity id="4CD6E5A7292CFF8AFE97174CFEB3CA2F" box="[330,404,526,553]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="25" unit="km" value="15.0">15 km</quantity>
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.
</paragraph>
</subSubSection>
</subSubSection>
</treatment>
</document>
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