497 lines
71 KiB
XML
497 lines
71 KiB
XML
<document id="06EC33C68FC3F27B548B221E533577F4" ID-CLB-Dataset="8049" ID-DOI="10.11646/zootaxa.4869.4.4" ID-GBIF-Dataset="f53aac91-e701-4129-a0dc-a9fcb282fddf" ID-ISSN="1175-5326" ID-Zenodo-Dep="4562510" ID-ZooBank="B88FBB1F-1083-472E-B429-1403BB080E07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610021309880" checkinUser="plazi" docAuthor="Chambrier, Alain De, Brabec, Jan & Scholz, Tomáš" docDate="2020" docId="FE4287ABFF8E3C17FF2CF923FAE9AB90" docLanguage="en" docName="zootaxa.4869.4.4.pdf" docOrigin="Zootaxa 4869 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kapsulotaenia nybelini Chambrier & Brabec & Scholz 2020, n. sp." docType="treatment" docVersion="9" lastPageNumber="552" masterDocId="027BFFD3FF9B3C00FFBBFF81FF8BAC4D" masterDocTitle="Molecular data reveal unexpected species diversity of tapeworms of Australasian reptiles: revision of Kapsulotaenia (Cestoda: Proteocephalidae)" masterLastPageNumber="561" masterPageNumber="529" pageNumber="550" updateTime="1698920232645" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="B8A0BE14BBE7C7D9989F573C4500FEC2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="34CD6CBBE8503E0E0811431ED0E62F38">
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<mods:title id="D7B0EBA99E6829DB33838681075AA244">Molecular data reveal unexpected species diversity of tapeworms of Australasian reptiles: revision of Kapsulotaenia (Cestoda: Proteocephalidae)</mods:title>
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<mods:name id="F3AC2108340172AE2BE8F49C570002E4" type="personal">
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<mods:roleTerm id="760F4973E3F6E05A3C2435782D487DDA">Author</mods:roleTerm>
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<mods:namePart id="CD1EB288DCD917E8B37B8E15209ABE9C">Chambrier, Alain De</mods:namePart>
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<mods:nameIdentifier id="E8BCB4CA58142C9FBB26C8EF7FBA29B1" type="ORCID">0000-0002-2533-2146</mods:nameIdentifier>
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<mods:affiliation id="B2F4E8E74F2F164ABD6BC91C7721A8D9">Department of Invertebrates, Natural History Museum, P. O. Box 6434, CH- 1211 Geneva 6, Switzerland. alain. dechambrier @ ville-ge. ch; https: // orcid. org / 0000 - 0002 - 2533 - 2146</mods:affiliation>
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<mods:nameIdentifier id="CFB9A8DF074671133C536BAAD8021227" type="email">alain.dechambrier@ville-ge.ch</mods:nameIdentifier>
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</mods:name>
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<mods:name id="C3B6F5FFF607915BD76EB101EE8C9863" type="personal">
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<mods:role id="E4070834798C2910C6BFB3BC32154813">
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<mods:roleTerm id="CE38755FBC7865D9F4C68173242E5B8E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="07016D4130EB47B274C71C7998D04CE6">Brabec, Jan</mods:namePart>
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<mods:nameIdentifier id="E5939D22062D8FB0CDBD2796EF2BF080" type="ORCID">0000-0002-2636-414X</mods:nameIdentifier>
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<mods:affiliation id="2080EC64DE240F2175A6FF3477B78FA0">Department of Invertebrates, Natural History Museum, P. O. Box 6434, CH- 1211 Geneva 6, Switzerland. alain. dechambrier @ ville-ge. ch; https: // orcid. org / 0000 - 0002 - 2533 - 2146 & Institute of Parasitology, Biology Centre of the Czech Academy of Sciences, Branišovská 31, 370 05 České Budějovice, Czech Republic. brabcak @ paru. cas. cz; https: // orcid. org / 0000 - 0002 - 2636 - 414 X</mods:affiliation>
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<mods:nameIdentifier id="287F698BD0F04F3FE40B4D9361A3EE50" type="email">alain.dechambrier@ville-ge.ch</mods:nameIdentifier>
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</mods:name>
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<mods:name id="FCA76D747E498215D5F779E70EBE6457" type="personal">
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<mods:roleTerm id="7E6ACDF23E1F88A7B43184B4B5301F3A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D663EFEAF4BFD75678750601B4422E8D">Scholz, Tomáš</mods:namePart>
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<mods:nameIdentifier id="BBC43CEC4059215AC614B753D8A13C1E" type="ORCID">0000-0002-6340-3750</mods:nameIdentifier>
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<mods:affiliation id="57B7E1D23E400B1BA35265745FE0263F">Institute of Parasitology, Biology Centre of the Czech Academy of Sciences, Branišovská 31, 370 05 České Budějovice, Czech Republic. brabcak @ paru. cas. cz; https: // orcid. org / 0000 - 0002 - 2636 - 414 X</mods:affiliation>
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<mods:nameIdentifier id="D8A38171A3A3377954067F63E4F9BB87" type="email">brabcak@paru.cas.cz</mods:nameIdentifier>
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<mods:title id="C6EC78F1FF0FBBC6690257DBD76BA85C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="80B2F99751A4CC9AC2DAA59BD763CEFC">
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<mods:date id="AA219592D876E7094BEE5A37B86AD4F8">2020</mods:date>
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<mods:number id="4D36E282F05A4B646321E8C7F7BA2333">2020-11-04</mods:number>
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</mods:detail>
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<mods:detail id="39416283877625C81E0478319172EDED" type="volume">
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<mods:number id="BE553ABF5405F1334D62826E3D36A5DC">4869</mods:number>
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<mods:number id="F7EA2FB5456A5A22656EBC8515EEFE9D">4</mods:number>
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<mods:identifier id="8E273D908B0B11295BD7363EE3DC1D15" type="DOI">10.11646/zootaxa.4869.4.4</mods:identifier>
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<mods:identifier id="3B4798DA2138630D4270053E632D968A" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="FE4287ABFF8E3C17FF2CF923FAE9AB90" ID-DOI="http://doi.org/10.5281/zenodo.4562558" ID-GBIF-Taxon="179906748" ID-Zenodo-Dep="4562558" LSID="urn:lsid:plazi:treatment:FE4287ABFF8E3C17FF2CF923FAE9AB90" httpUri="http://treatment.plazi.org/id/FE4287ABFF8E3C17FF2CF923FAE9AB90" lastPageId="23" lastPageNumber="552" pageId="21" pageNumber="550">
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<subSubSection id="3EF16536FF8E3C15FF2CF923FD99AAF0" box="[151,530,1698,1725]" pageId="21" pageNumber="550" type="nomenclature">
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<paragraph id="765436BDFF8E3C15FF2CF923FD99AAF0" blockId="21.[151,530,1698,1761]" box="[151,530,1698,1725]" pageId="21" pageNumber="550">
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<heading id="2D1C81D1FF8E3C15FF2CF923FD99AAF0" bold="true" box="[151,530,1698,1725]" fontSize="11" level="1" pageId="21" pageNumber="550" reason="1">
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<emphasis id="449FEAAFFF8E3C15FF2CF923FD99AAF0" bold="true" box="[151,530,1698,1725]" pageId="21" pageNumber="550">
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10.
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<taxonomicName id="B1EB4D3EFF8E3C15FF7AF923FE47AAF0" authority="Chambrier & Brabec & Scholz, 2020" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[193,460,1698,1725]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="21" pageNumber="550" phylum="Platyhelminthes" rank="species" species="nybelini" status="sp. nov.">
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<emphasis id="449FEAAFFF8E3C15FF7AF923FE47AAF0" bold="true" box="[193,460,1698,1725]" italics="true" pageId="21" pageNumber="550">Kapsulotaenia nybelini</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="5FAC57D4FF8E3C15FE68F922FD99AAF0" box="[467,530,1699,1725]" pageId="21" pageNumber="550" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3EF16536FF8E3C15FF2CF946FB87AB6E" pageId="21" pageNumber="550" type="description">
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<paragraph id="765436BDFF8E3C15FF2CF946FEC9AAAC" blockId="21.[151,530,1698,1761]" box="[151,322,1735,1761]" pageId="21" pageNumber="550">
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||
(
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<figureCitation id="EED02A38FF8E3C15FF24F946FF67AAAC" box="[159,236,1735,1761]" captionStart="FIGURE 7" captionStartId="22.[151,250,1168,1192]" captionTargetBox="[155,1429,190,1137]" captionTargetId="figure-24@22.[151,1436,181,1145]" captionTargetPageId="22" captionText="FIGURE 7. Scoleces of Kapsulotaenia cannoni n. sp. (A, B—MHNG-PLAT-36703; C—AHC 35832) and Kapsulotaenia nybelini n. sp. (D–F—AHC 35834), both from Varanus gouldii, Australia." figureDoi="http://doi.org/10.5281/zenodo.4562538" httpUri="https://zenodo.org/record/4562538/files/figure.png" pageId="21" pageNumber="550">Figs. 7</figureCitation>
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D–F, 9)
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</paragraph>
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<paragraph id="765436BDFF8E3C15FF2CF88AFB87AB6E" blockId="21.[151,1036,1803,1827]" box="[151,1036,1803,1827]" pageId="21" pageNumber="550">
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Syn.
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<taxonomicName id="B1EB4D3EFF8E3C15FF70F88AFE11AB6E" authorityName="Freze" authorityYear="1963" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[203,410,1803,1827]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="21" pageNumber="550" phylum="Platyhelminthes" rank="species" species="varia">
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<emphasis id="449FEAAFFF8E3C15FF70F88AFE11AB6E" box="[203,410,1803,1827]" italics="true" pageId="21" pageNumber="550">Kapsulotaenia varia</emphasis>
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</taxonomicName>
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of
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<bibRefCitation id="127A4B4CFF8E3C15FE06F88AFDD0AB6E" author="Nybelin, O." box="[445,603,1803,1827]" pageId="21" pageNumber="550" pagination="1 - 48" refId="ref21965" refString="Nybelin, O. (1917) Australischen Cestoden. Results of D. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. Kungliga Svenska Vetenskapsakademiens Handlingar, 52 (14), 1 - 48." type="journal article" year="1917">Nybelin (1917)</bibRefCitation>
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,
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<emphasis id="449FEAAFFF8E3C15FDDEF88DFD03AB6E" box="[613,648,1804,1827]" italics="true" pageId="21" pageNumber="550">nec</emphasis>
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||
<taxonomicName id="B1EB4D3EFF8E3C15FD35F88DFB87AB6E" authority="(Beddard, 1913) Freze, 1965" authorityName="Freze" authorityYear="1965" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[654,1036,1803,1827]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="21" pageNumber="550" phylum="Platyhelminthes" rank="species" species="varia">
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<emphasis id="449FEAAFFF8E3C15FD35F88DFD6AAB6E" box="[654,737,1804,1827]" italics="true" pageId="21" pageNumber="550">K. varia</emphasis>
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(
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<bibRefCitation id="127A4B4CFF8E3C15FD54F88AFC0CAB6E" author="Beddard, F. E." box="[751,903,1803,1827]" pageId="21" pageNumber="550" pagination="4 - 36" refId="ref20216" refString="Beddard, F. E. (1913) Contributions to the anatomy and systematic arrangement of the Cestoidea. VII. On six species of tapeworms from reptiles, belonging to the genus Ichthyotaenia (s. l.). Proceedings of the Zoological Society of London, Part 1, 4 - 36. https: // doi. org / 10.1111 / j. 1096 - 3642.1913. tb 01980. x" type="book chapter" year="1913">Beddard, 1913</bibRefCitation>
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)
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<bibRefCitation id="127A4B4CFF8E3C15FC2FF88DFB87AB6E" author="Freze, V. I." box="[916,1036,1803,1827]" pageId="21" pageNumber="550" refId="ref21391" refString="Freze, V. I. (1965) [Proteocephalata in Fish, Amphibians and Reptiles]. Essentials of Cestodology. Vol. V. Nauka, Moscow, 538 pp. [in Russian, English translation, Israel Program of Scientific Translation, 1969, Cat. No. 1853, v + 597 pp.]" type="book" year="1965">Freze, 1965</bibRefCitation>
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</taxonomicName>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3EF16536FF8E3C16FF2CF8CDFAEEAA7B" lastPageId="22" lastPageNumber="551" pageId="21" pageNumber="550" type="materials_examined">
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||
<paragraph id="765436BDFF8E3C15FF2CF8CDFBEFAB2A" blockId="21.[151,1124,1868,2003]" box="[151,1124,1868,1895]" pageId="21" pageNumber="550">
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||
<emphasis id="449FEAAFFF8E3C15FF2CF8CDFE53AB2B" bold="true" box="[151,472,1868,1894]" pageId="21" pageNumber="550">
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<typeStatus id="A950881FFF8E3C15FF2CF8CDFF59AB2B" box="[151,210,1868,1894]" pageId="21" pageNumber="550">Type</typeStatus>
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and only known host.
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</emphasis>
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Gould’s monitor,
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<taxonomicName id="B1EB4D3EFF8E3C15FD1FF8CCFCDDAB2B" box="[676,854,1868,1894]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="550" phylum="Chordata" rank="species" species="gouldii">
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<emphasis id="449FEAAFFF8E3C15FD1FF8CCFCDDAB2B" box="[676,854,1868,1894]" italics="true" pageId="21" pageNumber="550">Varanus gouldii</emphasis>
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</taxonomicName>
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||
(
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<taxonomicName id="B1EB4D3EFF8E3C15FCDEF8CDFC52AB2B" box="[869,985,1868,1894]" class="Reptilia" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="550" phylum="Chordata" rank="order">Squamata</taxonomicName>
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||
:
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<taxonomicName id="B1EB4D3EFF8E3C15FC58F8CDFBD2AB2A" box="[995,1113,1868,1895]" class="Reptilia" family="Varanidae" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="550" phylum="Chordata" rank="family">Varanidae</taxonomicName>
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).
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</paragraph>
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||
<paragraph id="765436BDFF8E3C15FF7CF8F1FE73ABC7" blockId="21.[151,1124,1868,2003]" box="[199,504,1904,1930]" pageId="21" pageNumber="550">
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||
<emphasis id="449FEAAFFF8E3C15FF7CF8F1FE02ABC7" bold="true" box="[199,393,1904,1930]" pageId="21" pageNumber="550">Site of infection.</emphasis>
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Intestine.
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</paragraph>
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||
<paragraph id="765436BDFF8E3C15FF7CF815FC81ABE3" blockId="21.[151,1124,1868,2003]" box="[199,778,1940,1967]" pageId="21" pageNumber="550">
|
||
<materialsCitation id="C6833CE0FF8E3C15FF7CF815FC8DABE3" ID-GBIF-Occurrence="3042107314" box="[199,774,1940,1967]" country="Australia" location="Mt. Glorious" pageId="21" pageNumber="550" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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||
<emphasis id="449FEAAFFF8E3C15FF7CF815FEEDABE3" bold="true" box="[199,358,1940,1966]" pageId="21" pageNumber="550">
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||
<typeStatus id="A950881FFF8E3C15FF7CF815FE89ABE3" box="[199,258,1940,1966]" pageId="21" pageNumber="550">Type</typeStatus>
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locality.
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||
</emphasis>
|
||
<location id="73346066FF8E3C15FED7F814FD8BABE3" LSID="urn:lsid:plazi:treatment:FE4287ABFF8E3C17FF2CF923FAE9AB90:73346066FF8E3C15FED7F814FD8BABE3" box="[364,512,1940,1967]" country="Australia" name="Mt. Glorious" pageId="21" pageNumber="550" stateProvince="Queensland">Mt. Glorious</location>
|
||
,
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<collectingRegion id="B42FF85FFF8E3C15FDB0F815FD19ABE3" box="[523,658,1940,1966]" country="Australia" name="Queensland" pageId="21" pageNumber="550">Queensland</collectingRegion>
|
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,
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<collectingCountry id="0EFC762DFF8E3C15FD27F815FC8DABE3" box="[668,774,1940,1966]" name="Australia" pageId="21" pageNumber="550">Australia</collectingCountry>
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</materialsCitation>
|
||
.
|
||
</paragraph>
|
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<paragraph id="765436BDFF8E3C15FF7CF839FD83AB9E" blockId="21.[151,1124,1868,2003]" box="[199,520,1976,2003]" pageId="21" pageNumber="550">
|
||
<emphasis id="449FEAAFFF8E3C15FF7CF839FE36AB9F" bold="true" box="[199,445,1976,2002]" pageId="21" pageNumber="550">Additional localities.</emphasis>
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None.
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||
</paragraph>
|
||
<caption id="22946635FF8D3C16FF2CFB11FCFAA880" ID-DOI="http://doi.org/10.5281/zenodo.4562538" ID-Zenodo-Dep="4562538" httpUri="https://zenodo.org/record/4562538/files/figure.png" pageId="22" pageNumber="551" startId="22.[151,250,1168,1192]" targetBox="[155,1429,190,1137]" targetPageId="22">
|
||
<paragraph id="765436BDFF8D3C16FF2CFB11FCFAA880" blockId="22.[151,1436,1168,1229]" pageId="22" pageNumber="551">
|
||
<emphasis id="449FEAAFFF8D3C16FF2CFB11FE98A8E4" bold="true" box="[151,275,1168,1193]" pageId="22" pageNumber="551">FIGURE 7.</emphasis>
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Scoleces of
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<taxonomicName id="B1EB4D3EFF8D3C16FE2DFB10FD09A8E5" authority="Chambrier & Brabec & Scholz, 2020" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[406,642,1169,1193]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="22" pageNumber="551" phylum="Platyhelminthes" rank="species" species="cannoni" status="sp. nov.">
|
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<emphasis id="449FEAAFFF8D3C16FE2DFB10FD09A8E5" box="[406,642,1169,1193]" italics="true" pageId="22" pageNumber="551">Kapsulotaenia cannoni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="449FEAAFFF8D3C16FD32FB10FD49A8E4" bold="true" box="[649,706,1169,1193]" pageId="22" pageNumber="551">
|
||
<taxonomicNameLabel id="5FAC57D4FF8D3C16FD32FB10FD49A8E4" box="[649,706,1169,1193]" pageId="22" pageNumber="551" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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(A, B—MHNG-PLAT-36703; C—AHC 35832) and
|
||
<taxonomicName id="B1EB4D3EFF8D3C16FB5AFB10FF5AA880" authority="Chambrier & Brabec & Scholz, 2020" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="22" pageNumber="551" phylum="Platyhelminthes" rank="species" species="nybelini" status="sp. nov.">
|
||
<emphasis id="449FEAAFFF8D3C16FB5AFB10FF5AA880" italics="true" pageId="22" pageNumber="551">Kapsulotaenia nybelini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="449FEAAFFF8D3C16FF6CFB34FE9BA880" bold="true" box="[215,272,1205,1229]" pageId="22" pageNumber="551">
|
||
<taxonomicNameLabel id="5FAC57D4FF8D3C16FF6CFB34FE9BA880" box="[215,272,1205,1229]" pageId="22" pageNumber="551" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(D–F—AHC 35834), both from
|
||
<taxonomicName id="B1EB4D3EFF8D3C16FDDAFB34FC89A880" box="[609,770,1205,1229]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="551" phylum="Chordata" rank="species" species="gouldii">
|
||
<emphasis id="449FEAAFFF8D3C16FDDAFB34FC89A880" box="[609,770,1205,1229]" italics="true" pageId="22" pageNumber="551">Varanus gouldii</emphasis>
|
||
</taxonomicName>
|
||
, Australia.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="765436BDFF8D3C16FF7CFA9EFF6CA9CC" blockId="22.[151,1437,1311,2021]" pageId="22" pageNumber="551">
|
||
<emphasis id="449FEAAFFF8D3C16FF7CFA9EFEFEA974" bold="true" box="[199,373,1311,1337]" pageId="22" pageNumber="551">Type material.</emphasis>
|
||
<materialsCitation id="C6833CE0FF8D3C16FEC7FA9EFF6CA9CC" ID-GBIF-Occurrence="3042107310" accessionNumber="AHC 35934, AHC 35832, AHC 35834" collectingDate="1963-08-29" collectionCode="AHC" pageId="22" pageNumber="551" specimenCode="SAM 5714, SAM 5699" specimenCount="4" typeStatus="holotype">
|
||
<typeStatus id="A950881FFF8D3C16FEC7FA9EFE6FA974" box="[380,484,1311,1337]" pageId="22" pageNumber="551" type="holotype">Holotype</typeStatus>
|
||
(complete specimen collected on
|
||
<date id="0255107DFF8D3C16FCDFFA9EFC65A974" box="[868,1006,1311,1337]" pageId="22" pageNumber="551" value="1963-08-29">
|
||
<collectingDate id="1211E995FF8D3C16FCDFFA9EFC65A974" box="[868,1006,1311,1337]" pageId="22" pageNumber="551" value="1963-08-29">29.viii.1963</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="264D9EC6FF8D3C16FC41FA9EFBFEA977" box="[1018,1141,1311,1338]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="22" pageNumber="551">SAM 5714</specimenCode>
|
||
—
|
||
<collectionCode id="10FAAE78FF8D3C16FB29FA9EFB46A977" box="[1170,1229,1311,1338]" pageId="22" pageNumber="551">AHC</collectionCode>
|
||
35934—slide No. 6);
|
||
<specimenCount id="60EDFD34FF8D3C16FF07FAC2FEB0A910" box="[188,315,1347,1373]" pageId="22" pageNumber="551" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
|
||
(
|
||
<specimenCode id="264D9EC6FF8D3C16FEF2FAC2FE4EA913" box="[329,453,1347,1374]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="22" pageNumber="551">SAM 5699</specimenCode>
|
||
—
|
||
<collectionCode id="10FAAE78FF8D3C16FE59FAC2FD96A913" box="[482,541,1347,1374]" pageId="22" pageNumber="551">AHC</collectionCode>
|
||
35832—slides Nos. 2 & 3),
|
||
<specimenCount id="60EDFD34FF8D3C16FCE2FAC2FC52A910" box="[857,985,1347,1373]" pageId="22" pageNumber="551" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
|
||
(
|
||
<specimenCode id="264D9EC6FF8D3C16FC5CFAC2FBE9A913" box="[999,1122,1347,1374]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="22" pageNumber="551">SAM 5714</specimenCode>
|
||
—
|
||
<collectionCode id="10FAAE78FF8D3C16FBC4FAC2FB31A913" box="[1151,1210,1347,1374]" pageId="22" pageNumber="551">AHC</collectionCode>
|
||
35834—slides Nos. 4 & 7).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="765436BDFF8D3C16FF7CFA0AFDA0A9E8" blockId="22.[151,1437,1311,2021]" box="[199,555,1419,1445]" pageId="22" pageNumber="551">
|
||
<emphasis id="449FEAAFFF8D3C16FF7CFA0AFE0CA9E8" bold="true" box="[199,391,1419,1445]" pageId="22" pageNumber="551">DNA sequences.</emphasis>
|
||
Not available.
|
||
</paragraph>
|
||
<paragraph id="765436BDFF8D3C16FF7CFA2EFAEEAA7B" blockId="22.[151,1437,1311,2021]" pageId="22" pageNumber="551">
|
||
<emphasis id="449FEAAFFF8D3C16FF7CFA2EFE1FA984" bold="true" box="[199,404,1455,1481]" pageId="22" pageNumber="551">Material studied.</emphasis>
|
||
Two slides with several whole-mounted specimens (5 scoleces of
|
||
<taxonomicName id="B1EB4D3EFF8D3C16FBCFFA31FB65A984" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[1140,1262,1455,1481]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="22" pageNumber="551" phylum="Platyhelminthes" rank="species" species="nybelini">
|
||
<emphasis id="449FEAAFFF8D3C16FBCFFA31FB0CA984" box="[1140,1159,1456,1481]" italics="true" pageId="22" pageNumber="551">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8D3C16FB2FFA2EFB65A984" box="[1172,1262,1455,1481]" italics="true" pageId="22" pageNumber="551">nybelini</emphasis>
|
||
</taxonomicName>
|
||
and 1 scolex of
|
||
<taxonomicName id="B1EB4D3EFF8D3C16FF2CFA55FE9AA9A0" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[151,273,1491,1517]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="22" pageNumber="551" phylum="Platyhelminthes" rank="species" species="cannoni">
|
||
<emphasis id="449FEAAFFF8D3C16FF2CFA55FF21A9A0" box="[151,170,1492,1517]" italics="true" pageId="22" pageNumber="551">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8D3C16FF0CFA55FE9AA9A0" box="[183,273,1492,1517]" italics="true" pageId="22" pageNumber="551">cannoni</emphasis>
|
||
</taxonomicName>
|
||
), 2 slides with cross sections of scolex and strobila of a specimen from
|
||
<taxonomicName id="B1EB4D3EFF8D3C16FB8AFA55FB15A9A0" box="[1073,1182,1491,1517]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="551" phylum="Chordata" rank="species" species="gouldii">
|
||
<emphasis id="449FEAAFFF8D3C16FB8AFA55FBC9A9A0" box="[1073,1090,1492,1517]" italics="true" pageId="22" pageNumber="551">V</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8D3C16FBF4FA52FB15A9A0" box="[1103,1182,1491,1517]" italics="true" pageId="22" pageNumber="551">gouldii</emphasis>
|
||
</taxonomicName>
|
||
(XP-1105), Mt. Glorious,
|
||
<collectingRegion id="B42FF85FFF8D3C16FF77FA76FED8AA5C" box="[204,339,1527,1553]" country="Australia" name="Queensland" pageId="22" pageNumber="551">Queensland</collectingRegion>
|
||
,
|
||
<collectingCountry id="0EFC762DFF8D3C16FEE7FA76FE4DAA5C" box="[348,454,1527,1553]" name="Australia" pageId="22" pageNumber="551">Australia</collectingCountry>
|
||
,
|
||
<date id="0255107DFF8D3C16FE6AFA76FDC2AA5C" box="[465,585,1527,1553]" pageId="22" pageNumber="551" value="1962-11-26">26.xi.1962</date>
|
||
(
|
||
<collectionCode id="10FAAE78FF8D3C16FDECFA76FD1FAA5C" box="[599,660,1527,1553]" pageId="22" pageNumber="551">AHC</collectionCode>
|
||
35832); 7 slides with whole mounts of several specimens (8 scoleces of
|
||
<taxonomicName id="B1EB4D3EFF8D3C16FF0DF99DFEBBAA78" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[182,304,1563,1589]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="22" pageNumber="551" phylum="Platyhelminthes" rank="species" species="nybelini">
|
||
<emphasis id="449FEAAFFF8D3C16FF0DF99DFF42AA78" box="[182,201,1564,1589]" italics="true" pageId="22" pageNumber="551">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8D3C16FF6DF99AFEBBAA78" box="[214,304,1563,1589]" italics="true" pageId="22" pageNumber="551">nybelini</emphasis>
|
||
</taxonomicName>
|
||
and 2 scoleces of
|
||
<taxonomicName id="B1EB4D3EFF8D3C16FE45F99DFDF2AA78" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[510,633,1563,1589]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="22" pageNumber="551" phylum="Platyhelminthes" rank="species" species="cannoni">
|
||
<emphasis id="449FEAAFFF8D3C16FE45F99DFD9AAA78" box="[510,529,1564,1589]" italics="true" pageId="22" pageNumber="551">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8D3C16FDA4F99DFDF2AA78" box="[543,633,1564,1589]" italics="true" pageId="22" pageNumber="551">cannoni</emphasis>
|
||
</taxonomicName>
|
||
), Mt. Glorious,
|
||
<collectingRegion id="B42FF85FFF8D3C16FC94F99AFC3DAA78" box="[815,950,1563,1589]" country="Australia" name="Queensland" pageId="22" pageNumber="551">Queensland</collectingRegion>
|
||
,
|
||
<collectingCountry id="0EFC762DFF8D3C16FC7BF99AFBA1AA78" box="[960,1066,1563,1589]" name="Australia" pageId="22" pageNumber="551">Australia</collectingCountry>
|
||
,
|
||
<date id="0255107DFF8D3C16FB8EF99AFB37AA78" box="[1077,1212,1563,1589]" pageId="22" pageNumber="551" value="1963-08-29">29.viii.1963</date>
|
||
(
|
||
<collectionCode id="10FAAE78FF8D3C16FB71F99AFA8CAA78" box="[1226,1287,1563,1589]" pageId="22" pageNumber="551">AHC</collectionCode>
|
||
35834).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3EF16536FF8D3C17FF7CF9BEFDECAFCC" lastPageId="23" lastPageNumber="552" pageId="22" pageNumber="551" type="etymology">
|
||
<paragraph id="765436BDFF8D3C16FF7CF9BEFB5FAA30" blockId="22.[151,1437,1311,2021]" pageId="22" pageNumber="551">
|
||
<emphasis id="449FEAAFFF8D3C16FF7CF9BEFEC5AA14" bold="true" box="[199,334,1599,1625]" pageId="22" pageNumber="551">Etymology.</emphasis>
|
||
The new species is named after Orvar Nybelin from
|
||
<collectingCountry id="0EFC762DFF8D3C16FC1FF9BEFC74AA14" box="[932,1023,1599,1625]" name="Sweden" pageId="22" pageNumber="551">Sweden</collectingCountry>
|
||
, who contributed significantly to the knowledge of tapeworms including those parasitising reptiles in the first half of the 20th century.
|
||
</paragraph>
|
||
<paragraph id="765436BDFF8D3C16FF7CF906FBFFAB8C" blockId="22.[151,1437,1311,2021]" pageId="22" pageNumber="551">
|
||
<emphasis id="449FEAAFFF8D3C16FF7CF906FD8EAAEC" bold="true" box="[199,517,1671,1697]" pageId="22" pageNumber="551">Morphological description</emphasis>
|
||
(for measurements—see
|
||
<tableCitation id="3B690306FF8D3C16FC93F906FCF4AAEC" box="[808,895,1671,1697]" captionStart="TABLE 2" captionStartId="10.[151,239,504,530]" captionTargetPageId="10" captionText="TABLE 2. Measurements of species of Kapsulotaenia Freze, 1963 (new measurements of type material are in brackets; measurement of our material in parentheses; measurements are in micrometres unless otherwise stated)." httpUri="http://table.plazi.org/id/22946635FF913C0AFF2CFE79FC80AE62" pageId="22" pageNumber="551" tableUuid="22946635FF913C0AFF2CFE79FC80AE62">Table 2</tableCitation>
|
||
).
|
||
<taxonomicName id="B1EB4D3EFF8D3C16FC2EF906FBD2AAEC" authorityName="La Rue" authorityYear="1911" box="[917,1113,1671,1697]" class="Cestoda" family="Proteocephalidae" kingdom="Animalia" order="Proteocephalidea" pageId="22" pageNumber="551" phylum="Platyhelminthes" rank="family">Proteocephalidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="B1EB4D3EFF8D3C16FBDFF906FAA7AAEC" authorityName="Freze" authorityYear="1963" box="[1124,1324,1671,1697]" class="Cestoda" family="Proteocephalidae" kingdom="Animalia" order="Proteocephalidea" pageId="22" pageNumber="551" phylum="Platyhelminthes" rank="subFamily" subFamily="Acanthotaeniinae">Acanthotaeniinae</taxonomicName>
|
||
. Cestodes up to
|
||
<quantity id="B1139B58FF8D3C16FF62F92AFEBCAA88" box="[217,311,1707,1733]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="22" pageNumber="551" unit="mm" value="103.0">103 mm</quantity>
|
||
in total length; maximum width up to 1,250. Strobila acraspedote, anapolytic. Strobilisation begins about
|
||
<quantity id="B1139B58FF8D3C16FF66F94EFEB8AAA4" box="[221,307,1743,1770]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="551" unit="mm" value="1.5">1.5 mm</quantity>
|
||
posterior to scolex, about 39–
|
||
<specimenCount id="60EDFD34FF8D3C16FD31F94EFC9DAAA4" box="[650,790,1743,1769]" pageId="22" pageNumber="551" type="immature">81 immature</specimenCount>
|
||
proglottids before appearance of spermatozoa in
|
||
<emphasis id="449FEAAFFF8D3C16FAFFF951FF58AB40" italics="true" pageId="22" pageNumber="551">vas deferens</emphasis>
|
||
, 1 mature, 7–12 pregravid proglottids, 8 gravid proglottids and
|
||
<quantity id="B1139B58FF8D3C16FC11F972FB96AB40" box="[938,1053,1779,1806]" metricMagnitude="0" metricUnit="m" metricValue="1.9812" metricValueMax="2.54" metricValueMin="1.4224" pageId="22" pageNumber="551" unit="in" value="78.0" valueMax="100.0" valueMin="56.0">56–100 in</quantity>
|
||
total. Immature proglottids wider than long to longer than wide (length: width ratio 0.6–2.10), mature, pregravid and gravid proglottids longer than wide (ratio 1.49–4.25) (
|
||
<figureCitation id="EED02A38FF8D3C16FE24F8BAFE6BAB18" box="[415,480,1851,1877]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="22" pageNumber="551">Fig. 9</figureCitation>
|
||
A–C). Inner longitudinal musculature present at level of proliferation zone, composed of very few muscle fibres; bundles of muscles not observed in mature and pregravid proglottids (
|
||
<figureCitation id="EED02A38FF8D3C16FB5AF8DEFABCAB34" box="[1249,1335,1887,1913]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="22" pageNumber="551">Fig. 9F</figureCitation>
|
||
). Ventral osmoregulatory canals slightly sinuous,
|
||
<quantity id="B1139B58FF8D3C16FDEDF802FD33ABD0" box="[598,696,1923,1950]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" metricValueMax="8.89" metricValueMin="6.35" pageId="22" pageNumber="551" unit="in" value="30.0" valueMax="35.0" valueMin="25.0">25–35 in</quantity>
|
||
diameter. Dorsal osmoregulatory canals slightly sinuous,
|
||
<quantity id="B1139B58FF8D3C16FA81F802FA17ABD0" box="[1338,1436,1923,1950]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.81" metricValueMin="2.54" pageId="22" pageNumber="551" unit="in" value="12.5" valueMax="15.0" valueMin="10.0">10–15 in</quantity>
|
||
diameter (
|
||
<figureCitation id="EED02A38FF8D3C16FEB3F826FEC1AB8C" box="[264,330,1959,1985]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="22" pageNumber="551">Fig. 9</figureCitation>
|
||
A–E). Canals situated between vitelline follicles and testes (
|
||
<figureCitation id="EED02A38FF8D3C16FC57F826FBECAB8C" box="[1004,1127,1959,1985]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="22" pageNumber="551">Fig. 9A, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="765436BDFF8D3C17FF7CF84AFC0BAC98" blockId="22.[151,1437,1311,2021]" lastBlockId="23.[151,1437,151,2013]" lastPageId="23" lastPageNumber="552" pageId="22" pageNumber="551">
|
||
Scolex wider than neck (
|
||
<figureCitation id="EED02A38FF8D3C16FE61F84AFD90ABA8" box="[474,539,1995,2021]" captionStart="FIGURE 7" captionStartId="22.[151,250,1168,1192]" captionTargetBox="[155,1429,190,1137]" captionTargetId="figure-24@22.[151,1436,181,1145]" captionTargetPageId="22" captionText="FIGURE 7. Scoleces of Kapsulotaenia cannoni n. sp. (A, B—MHNG-PLAT-36703; C—AHC 35832) and Kapsulotaenia nybelini n. sp. (D–F—AHC 35834), both from Varanus gouldii, Australia." figureDoi="http://doi.org/10.5281/zenodo.4562538" httpUri="https://zenodo.org/record/4562538/files/figure.png" pageId="22" pageNumber="551">Fig. 7</figureCitation>
|
||
D–F). Suckers uniloculate, spherical, directed laterally to anterolaterally. Rostel-lum dome-shaped, containing large apical organ, posteriorly surrounded by numerous circular muscular fibres (
|
||
<figureCitation id="EED02A38FF8C3C17FAD4FF16FF57AC98" captionStart="FIGURE 7" captionStartId="22.[151,250,1168,1192]" captionTargetBox="[155,1429,190,1137]" captionTargetId="figure-24@22.[151,1436,181,1145]" captionTargetPageId="22" captionText="FIGURE 7. Scoleces of Kapsulotaenia cannoni n. sp. (A, B—MHNG-PLAT-36703; C—AHC 35832) and Kapsulotaenia nybelini n. sp. (D–F—AHC 35834), both from Varanus gouldii, Australia." figureDoi="http://doi.org/10.5281/zenodo.4562538" httpUri="https://zenodo.org/record/4562538/files/figure.png" pageId="23" pageNumber="552">Fig. 7D, E</figureCitation>
|
||
). Retractor muscles connecting rostellum with neck present.
|
||
</paragraph>
|
||
<paragraph id="765436BDFF8C3C17FF7CFF5EFDD3AD9C" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="552">
|
||
Testes medullary, in 1 layer, rounded to elongate, in 2 wide longitudinal bands on both sides of proglottids, with poral band separated by terminal genitalia into preporal and postporal groups (
|
||
<figureCitation id="EED02A38FF8C3C17FC43FE82FBE4AD50" box="[1016,1135,259,285]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9A, B</figureCitation>
|
||
); testes and ovary degenerate in gravid proglottids. Cirrus-sac elongate, thick-walled (
|
||
<figureCitation id="EED02A38FF8C3C17FC96FEA6FC08AD0C" box="[813,899,295,321]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9E</figureCitation>
|
||
). Cirrus short, covered by spinitriches, its length representing up to 40% of cirrus-sac length (
|
||
<figureCitation id="EED02A38FF8C3C17FD33FECAFD54AD28" box="[648,735,331,357]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9E</figureCitation>
|
||
). Internal sperm duct strongly coiled. External sperm duct (
|
||
<emphasis id="449FEAAFFF8C3C17FACCFECAFF7EADC4" italics="true" pageId="23" pageNumber="552">vas deferens</emphasis>
|
||
) strongly coiled, directed anteriorly, situated between proximal part of cirrus-sac and midline of proglottid, often crossing it (
|
||
<figureCitation id="EED02A38FF8C3C17FE3DFE12FD8BADE0" box="[390,512,403,429]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9A, B</figureCitation>
|
||
). Genital atrium shallow (
|
||
<figureCitation id="EED02A38FF8C3C17FC9EFE12FCF7ADE0" box="[805,892,403,429]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9E</figureCitation>
|
||
); genital pores alternating irregularly, situated in posterior part of proglottid (
|
||
<figureCitation id="EED02A38FF8C3C17FE6BFE36FD98AD9C" box="[464,531,439,465]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9</figureCitation>
|
||
A–C).
|
||
</paragraph>
|
||
<paragraph id="765436BDFF8C3C17FF7CFE5AFE81AEE4" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="552">
|
||
Ovary bilobed, butterfly-shaped (
|
||
<figureCitation id="EED02A38FF8C3C17FDF9FE5AFD0CADB8" box="[578,647,475,501]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9</figureCitation>
|
||
A–D). Mehlis’ gland small (
|
||
<figureCitation id="EED02A38FF8C3C17FC72FE5AFBA8ADB8" box="[969,1059,475,501]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9D</figureCitation>
|
||
). Vaginal canal straight in proximal part, enlarged to form small seminal receptacle anterior to ovarian isthmus (
|
||
<figureCitation id="EED02A38FF8C3C17FB9AFE7EFBF0AE54" box="[1057,1147,511,537]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9B</figureCitation>
|
||
). Terminal (distal) part of vaginal canal (
|
||
<emphasis id="449FEAAFFF8C3C17FE86FDA2FDC0AE71" box="[317,587,547,573]" italics="true" pageId="23" pageNumber="552">pars copulatrix vaginae</emphasis>
|
||
) surrounded by few chromophilic cells (
|
||
<figureCitation id="EED02A38FF8C3C17FBACFDA2FBE4AE70" box="[1047,1135,547,573]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9E</figureCitation>
|
||
). Vagina anterior (28%) or posterior (72%, n = 47) to cirrus-sac (
|
||
<figureCitation id="EED02A38FF8C3C17FD80FDC6FDF7AE2C" box="[571,636,583,609]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9</figureCitation>
|
||
A–C, E). Vitelline follicles arranged in 2 longitudinal bands near lateral margins of proglottids (one to three rows of follicles), interrupted at level of cirrus-sac and vagina ventrally (
|
||
<figureCitation id="EED02A38FF8C3C17FAD4FDEAFF2FAEE4" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9</figureCitation>
|
||
A–C, E).
|
||
</paragraph>
|
||
<paragraph id="765436BDFF8C3C17FF7CFD32FDECAFCC" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="552">
|
||
Primordium of uterine stem ventral, present in immature proglottids (
|
||
<figureCitation id="EED02A38FF8C3C17FC6EFD32FBA4AE80" box="[981,1071,691,717]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9A</figureCitation>
|
||
). Development of uterus of
|
||
<typeStatus id="A950881FFF8C3C17FAD7FD32FA17AE80" box="[1388,1436,691,717]" pageId="23" pageNumber="552">type</typeStatus>
|
||
1 according to
|
||
<bibRefCitation id="127A4B4CFF8C3C17FEFBFD56FDEDAEBC" author="de Chambrier, A. & Zehnder, M. P. & Vaucher, C. & Mariaux, J." box="[320,614,727,753]" pageId="23" pageNumber="552" pagination="159 - 171" refId="ref21085" refString="de Chambrier, A., Zehnder, M. P., Vaucher, C. & Mariaux, J. (2004) The evolution of the Proteocephalidea (Platyhelminthes, Eucestoda) based on an enlarged molecular phylogeny, with comments on their uterine development. Systematic Parasitology, 57, 159 - 171. https: // doi. org / 10.1023 / B: SYPA. 0000019083.26876.34" type="journal article" year="2004">
|
||
de Chambrier
|
||
<emphasis id="449FEAAFFF8C3C17FE59FD56FD9FAEBC" box="[482,532,727,753]" italics="true" pageId="23" pageNumber="552">et al</emphasis>
|
||
. (2004
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="127A4B4CFF8C3C17FDC8FD57FD3BAEBC" author="de Chambrier, A. & Waeschenbach, A. & Fisseha, M. & Scholz, T. & Mariaux, J." box="[627,688,726,753]" pageId="23" pageNumber="552" pagination="25 - 59" refId="ref21013" refString="de Chambrier, A., Waeschenbach, A., Fisseha, M., Scholz, T. & Mariaux, J. (2015) A large 28 S rDNA-based phylogeny confirms the limitations of established morphological characters for classification of proteocephalidean tapeworms (Platyhelminthes, Cestoda). ZooKeys, 500, 25 - 59. https: // doi. org / 10.3897 / zookeys. 500.9360" type="journal article" year="2015">2015</bibRefCitation>
|
||
). In pregravid proglottids, uterus occupies up to 89% of proglottid length, with lateral diverticula on each side, not overlapping ovary. In gravid proglottids, uterus occupies up to 92% of proglottid length, with 30–36 lateral diverticula (
|
||
<figureCitation id="EED02A38FF8C3C17FD58FC9EFCB6AF74" box="[739,829,799,825]" captionStart="FIGURE 9" captionStartId="25.[151,250,1480,1504]" captionTargetBox="[157,1431,188,1448]" captionTargetId="figure-19@25.[151,1436,181,1457]" captionTargetPageId="25" captionText="FIGURE 9. Kapsulotaenia nybelini n. sp. from Varanus varius, Australia (AHC 35834—A, D, E; AHC 35832—B, C, F). A—immature proglottid, ventrally; B—pregravid proglottid, ventrally; C—gravid proglottid, ventrally; D—posterior extent of vitelline follicles, dorsally; E—terminal genitalia, ventrally; F—cross section at level of testes; note absence of inner longitudinal musculature." figureDoi="http://doi.org/10.5281/zenodo.4562548" httpUri="https://zenodo.org/record/4562548/files/figure.png" pageId="23" pageNumber="552">Fig. 9C</figureCitation>
|
||
). Uteroduct enters uterus anterior to ovarian isthmus. Intrauterine eggs spherical, grouped in clusters of eggs (number of eggs not known). Unripe embryophore measured in whole mount slides
|
||
<quantity id="B1139B58FF8C3C17FE2EFCE7FE73AFCC" box="[405,504,870,897]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" metricValueMax="8.382" metricValueMin="6.35" pageId="23" pageNumber="552" unit="in" value="29.0" valueMax="33.0" valueMin="25.0">25–33 in</quantity>
|
||
diameter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3EF16536FF8C3C17FF7CFC0BFCBEA934" pageId="23" pageNumber="552" type="diagnosis">
|
||
<paragraph id="765436BDFF8C3C17FF7CFC0BFCBEA934" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="552">
|
||
<emphasis id="449FEAAFFF8C3C17FF7CFC0BFE46AFE9" bold="true" box="[199,461,906,932]" pageId="23" pageNumber="552">Differential diagnosis.</emphasis>
|
||
The new species is characterized by a large, quadrangular scolex (other species of
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FADFFC0AFE87AF84" authorityName="Freze" authorityYear="1963" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="genus">
|
||
<emphasis id="449FEAAFFF8C3C17FADFFC0AFE87AF84" italics="true" pageId="23" pageNumber="552">Kapsulotaenia</emphasis>
|
||
</taxonomicName>
|
||
from Gould’s monitor possess a much smaller scolex—see above). In the width of the scolex,
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FA99FC2EFA17AF84" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[1314,1436,943,969]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="nybelini">
|
||
<emphasis id="449FEAAFFF8C3C17FA99FC2EFABEAF85" box="[1314,1333,943,968]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FAF9FC2EFA17AF84" box="[1346,1436,943,969]" italics="true" pageId="23" pageNumber="552">nybelini</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FEB6FC52FE20AFA0" box="[269,427,979,1005]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="sangroundi">
|
||
<emphasis id="449FEAAFFF8C3C17FEB6FC52FEABAFA1" box="[269,288,979,1004]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FE97FC52FE20AFA0" box="[300,427,979,1005]" italics="true" pageId="23" pageNumber="552">sangroundi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FE64FC52FDD6AFA0" baseAuthorityName="Wahid" baseAuthorityYear="1968" box="[479,605,979,1005]" class="Cestoda" family="Proteocephalidae" genus="Acanthotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="pythonis">
|
||
<emphasis id="449FEAAFFF8C3C17FE64FC52FE79AFA1" box="[479,498,979,1004]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FE45FC52FDD6AFA0" box="[510,605,979,1005]" italics="true" pageId="23" pageNumber="552">pythonis</emphasis>
|
||
</taxonomicName>
|
||
, but differs from
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FCA5FC52FC40AFA0" baseAuthorityName="Carter" baseAuthorityYear="1943" box="[798,971,979,1005]" class="Cestoda" family="Proteocephalidae" genus="Proteocephalus" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="sandgroundi">
|
||
<emphasis id="449FEAAFFF8C3C17FCA5FC52FCBAAFA1" box="[798,817,979,1004]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FC85FC52FC40AFA0" box="[830,971,979,1005]" italics="true" pageId="23" pageNumber="552">sandgroundi</emphasis>
|
||
</taxonomicName>
|
||
by the position of the vagina to the cirrussac (anterior 28% versus posterior), by the shape of proglottids (longer and narrower versus shorter and larger) and by a smaller scolex (470–645 μm versus 640–900 μm). From
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FCFBFB9AFC35A878" baseAuthorityName="Wahid" baseAuthorityYear="1968" box="[832,958,1051,1077]" class="Cestoda" family="Proteocephalidae" genus="Acanthotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="pythonis">
|
||
<emphasis id="449FEAAFFF8C3C17FCFBFB9AFCD8A879" box="[832,851,1051,1076]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FCE4FB9AFC35A878" box="[863,958,1051,1077]" italics="true" pageId="23" pageNumber="552">pythonis</emphasis>
|
||
</taxonomicName>
|
||
, it differs by the number of eggs in clusters (6–8 versus 19–22) and by the size of the scolex (470–645 μm versus 890 μm).
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FBB7FBBEFA9BA814" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[1036,1296,1087,1113]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="nybelini">
|
||
<emphasis id="449FEAAFFF8C3C17FBB7FBBEFA9BA814" box="[1036,1296,1087,1113]" italics="true" pageId="23" pageNumber="552">Kapsulotaenia nybelini</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FF2CFBE2FEB3A830" authorityName="Jones & de Chambrier" authorityYear="2016" box="[151,312,1123,1149]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="chisholmae">
|
||
<emphasis id="449FEAAFFF8C3C17FF2CFBE2FF21A831" box="[151,170,1123,1148]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FF03FBE2FEB3A830" box="[184,312,1123,1149]" italics="true" pageId="23" pageNumber="552">chisholmae</emphasis>
|
||
</taxonomicName>
|
||
by a smaller cirrus-sac ratio (17% in average versus 24%) and by the position of the vagina in relation to the cirrus-sac (mainly anterior in
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FDE3FB06FD73A8EC" authorityName="Jones & de Chambrier" authorityYear="2016" box="[600,760,1159,1185]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="chisholmae">
|
||
<emphasis id="449FEAAFFF8C3C17FDE3FB06FDE0A8ED" box="[600,619,1159,1184]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FDC3FB06FD73A8EC" box="[632,760,1159,1185]" italics="true" pageId="23" pageNumber="552">chisholmae</emphasis>
|
||
</taxonomicName>
|
||
versus mainly posterior in
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FB9CFB06FB2AA8EC" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[1063,1185,1159,1185]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="nybelini">
|
||
<emphasis id="449FEAAFFF8C3C17FB9CFB06FBB1A8ED" box="[1063,1082,1159,1184]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FBFCFB06FB2AA8EC" box="[1095,1185,1159,1185]" italics="true" pageId="23" pageNumber="552">nybelini</emphasis>
|
||
</taxonomicName>
|
||
). The new species can be distinguished from
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FE2FFB2AFD9FA888" authorityName="Freze" authorityYear="1963" baseAuthorityName="Johnston" baseAuthorityYear="1909" box="[404,532,1195,1221]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="tidswelli">
|
||
<emphasis id="449FEAAFFF8C3C17FE2FFB2AFE2CA889" box="[404,423,1195,1220]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FE0EFB2AFD9FA888" box="[437,532,1195,1221]" italics="true" pageId="23" pageNumber="552">tidswelli</emphasis>
|
||
</taxonomicName>
|
||
by slightly narrower ovary (relative width of the ovary 51–58% versus 63–72% in the latter taxon), and by longer bands of vitelline follicles (relative length of bands of vitelline follicles 90–97% versus 84–90%). From
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FE19FB72FDB2A940" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[418,569,1267,1293]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="beveridgei">
|
||
<emphasis id="449FEAAFFF8C3C17FE19FB72FE3EA941" box="[418,437,1267,1292]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FE78FB72FDB2A940" box="[451,569,1267,1293]" italics="true" pageId="23" pageNumber="552">beveridgei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FDF2FB72FD4EA940" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[585,709,1267,1293]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="nybelini">
|
||
<emphasis id="449FEAAFFF8C3C17FDF2FB72FDD7A941" box="[585,604,1267,1292]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FDD0FB72FD4EA940" box="[619,709,1267,1293]" italics="true" pageId="23" pageNumber="552">nybelini</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by a distinct width of the scolex (470–645 μm versus 335–475 μm), and by a different number of uterine lateral diverticula (30–36 versus 17–20). From
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FA38FA96FF5FA918" authorityName="Schmidt & Kuntz" authorityYear="1974" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="frezei">
|
||
<emphasis id="449FEAAFFF8C3C17FA38FA96FF5FA918" italics="true" pageId="23" pageNumber="552">K. frezei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FF58FABAFED5A918" authorityName="Chambrier & Brabec & Scholz" authorityYear="2020" box="[227,350,1339,1365]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="nybelini">
|
||
<emphasis id="449FEAAFFF8C3C17FF58FABAFED5A918" box="[227,350,1339,1365]" italics="true" pageId="23" pageNumber="552">K. nybelini</emphasis>
|
||
</taxonomicName>
|
||
differs by the shape of the egg clusters (oval versus banana-shaped). From
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FB14FABAFA80A919" authorityName="Freze" authorityYear="1965" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[1199,1291,1339,1365]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="varia">
|
||
<emphasis id="449FEAAFFF8C3C17FB14FABAFB49A919" box="[1199,1218,1339,1364]" italics="true" pageId="23" pageNumber="552">K</emphasis>
|
||
.
|
||
<emphasis id="449FEAAFFF8C3C17FB6BFABAFA80A919" box="[1232,1291,1339,1364]" italics="true" pageId="23" pageNumber="552">varia</emphasis>
|
||
</taxonomicName>
|
||
, it differs by a smaller relative size of the ovary (2.4–4.2% versus 5.2%).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3EF16536FF8C3C17FF7CFA02FAE9AB90" pageId="23" pageNumber="552" type="discussion">
|
||
<paragraph id="765436BDFF8C3C17FF7CFA02FF41AA44" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="552">
|
||
<emphasis id="449FEAAFFF8C3C17FF7CFA02FEB0A9D0" bold="true" box="[199,315,1411,1437]" pageId="23" pageNumber="552">Remarks.</emphasis>
|
||
The new species resembles in its morphology tapeworms found by Mjöberg in
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FB1DFA02FA84A9D0" box="[1190,1295,1411,1437]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="552" phylum="Chordata" rank="species" species="gouldii">
|
||
<emphasis id="449FEAAFFF8C3C17FB1DFA02FA84A9D0" box="[1190,1295,1411,1437]" italics="true" pageId="23" pageNumber="552">V. gouldii</emphasis>
|
||
</taxonomicName>
|
||
from Mount Tambourine (= Tamborine Mountain), South
|
||
<collectingRegion id="B42FF85FFF8C3C17FD2FFA26FC90A98C" box="[660,795,1447,1473]" country="Australia" name="Queensland" pageId="23" pageNumber="552">Queensland</collectingRegion>
|
||
,
|
||
<collectingCountry id="0EFC762DFF8C3C17FC9EFA26FC04A98C" box="[805,911,1447,1473]" name="Australia" pageId="23" pageNumber="552">Australia</collectingCountry>
|
||
.
|
||
<bibRefCitation id="127A4B4CFF8C3C17FC21FA26FBCCA98C" author="Nybelin, O." box="[922,1095,1447,1473]" pageId="23" pageNumber="552" pagination="1 - 48" refId="ref21965" refString="Nybelin, O. (1917) Australischen Cestoden. Results of D. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. Kungliga Svenska Vetenskapsakademiens Handlingar, 52 (14), 1 - 48." type="journal article" year="1917">Nybelin (1917)</bibRefCitation>
|
||
identified these tapeworms as
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FF2CFA4AFEF5A9A9" authorityName="Nybelin" authorityYear="1913" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[151,382,1483,1509]" class="Cestoda" family="Proteocephalidae" genus="Acanthotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="varia">
|
||
<emphasis id="449FEAAFFF8C3C17FF2CFA4AFEF5A9A9" box="[151,382,1483,1509]" italics="true" pageId="23" pageNumber="552">Acanthotaenia varia</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="B1EB4D3EFF8C3C17FE12FA4AFD1BA9A9" authorityName="Freze" authorityYear="1965" baseAuthorityName="Beddard" baseAuthorityYear="1913" box="[425,656,1483,1509]" class="Cestoda" family="Proteocephalidae" genus="Kapsulotaenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Proteocephalidea" pageId="23" pageNumber="552" phylum="Platyhelminthes" rank="species" species="varia">
|
||
<emphasis id="449FEAAFFF8C3C17FE12FA4AFD1BA9A9" box="[425,656,1483,1509]" italics="true" pageId="23" pageNumber="552">Kapsulotaenia varia</emphasis>
|
||
</taxonomicName>
|
||
) based on their presumed conspecificity with
|
||
<bibRefCitation id="127A4B4CFF8C3C17FB1BFA4AFAEEA9A8" author="Beddard, F. E." box="[1184,1381,1483,1509]" pageId="23" pageNumber="552" pagination="4 - 36" refId="ref20216" refString="Beddard, F. E. (1913) Contributions to the anatomy and systematic arrangement of the Cestoidea. VII. On six species of tapeworms from reptiles, belonging to the genus Ichthyotaenia (s. l.). Proceedings of the Zoological Society of London, Part 1, 4 - 36. https: // doi. org / 10.1111 / j. 1096 - 3642.1913. tb 01980. x" type="book chapter" year="1913">Beddard’s (1913)</bibRefCitation>
|
||
species.
|
||
</paragraph>
|
||
<paragraph id="765436BDFF8C3C17FF7CF992FE9AAAF0" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="552">
|
||
<bibRefCitation id="127A4B4CFF8C3C17FF7CF992FEFFAA60" author="Nybelin, O." box="[199,372,1555,1581]" pageId="23" pageNumber="552" pagination="1 - 48" refId="ref21965" refString="Nybelin, O. (1917) Australischen Cestoden. Results of D. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. Kungliga Svenska Vetenskapsakademiens Handlingar, 52 (14), 1 - 48." type="journal article" year="1917">Nybelin (1917)</bibRefCitation>
|
||
provided illustrations of the scolex (his fig. 1), section through the anterior part of the scolex (fig. 2) and mature proglottid (fig. 3). The longest specimen had a total length of
|
||
<quantity id="B1139B58FF8C3C17FB84F9B6FB2BAA1C" box="[1087,1184,1591,1617]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="23" pageNumber="552" unit="mm" value="230.0">230 mm</quantity>
|
||
; maximum width was
|
||
<quantity id="B1139B58FF8C3C17FF2CF9DBFE85AA38" box="[151,270,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="23" pageNumber="552" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
|
||
. The scolex was reported as only weakly separated from the remaining body,
|
||
<quantity id="B1139B58FF8C3C17FB30F9DAFA90AA38" box="[1163,1307,1627,1653]" metricMagnitude="-1" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="23" pageNumber="552" unit="mm" value="410.0" valueMax="420.0" valueMin="400.0">400–420 µm</quantity>
|
||
wide, with spherical suckers
|
||
<quantity id="B1139B58FF8C3C17FEE5F9FEFE66AAD4" box="[350,493,1663,1689]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="23" pageNumber="552" unit="mm" value="210.0" valueMax="220.0" valueMin="200.0">200–220 µm</quantity>
|
||
in diameter. The apical organ measured 120 ×
|
||
<quantity id="B1139B58FF8C3C17FC41F9FEFBDAAAD4" box="[1018,1105,1663,1689]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="552" unit="mm" value="130.0">130 µm</quantity>
|
||
and the cirrus-sac 150–180 ×
|
||
<quantity id="B1139B58FF8C3C17FF2CF923FE86AAF0" box="[151,269,1698,1725]" metricMagnitude="-2" metricUnit="m" metricValue="8.55" metricValueMax="9.6" metricValueMin="7.5" pageId="23" pageNumber="552" unit="mm" value="85.5" valueMax="96.0" valueMin="75.0">75–96 µm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="765436BDFF8C3C17FF7CF946FE06AB3F" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="552">
|
||
<bibRefCitation id="127A4B4CFF8C3C17FF7CF946FEFFAAAC" author="Nybelin, O." box="[199,372,1735,1761]" pageId="23" pageNumber="552" pagination="1 - 48" refId="ref21965" refString="Nybelin, O. (1917) Australischen Cestoden. Results of D. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. Kungliga Svenska Vetenskapsakademiens Handlingar, 52 (14), 1 - 48." type="journal article" year="1917">Nybelin (1917)</bibRefCitation>
|
||
pointed out a high variation in the number of testes (96–158, but usually about 120) and their distribution (sometimes forming two medially well-separated longitudinal fields, sometimes confluent).
|
||
<bibRefCitation id="127A4B4CFF8C3C17FAF9F96AFF69AB64" author="Nybelin, O." pageId="23" pageNumber="552" pagination="1 - 48" refId="ref21965" refString="Nybelin, O. (1917) Australischen Cestoden. Results of D. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. Kungliga Svenska Vetenskapsakademiens Handlingar, 52 (14), 1 - 48." type="journal article" year="1917">Nybelin (1917)</bibRefCitation>
|
||
also observed variation in the extent of
|
||
<emphasis id="449FEAAFFF8C3C17FD1DF88EFCBAAB64" box="[678,817,1807,1833]" italics="true" pageId="23" pageNumber="552">vas deferens</emphasis>
|
||
, the loops of which reach over the mid-line of proglottids to the aporal half, whereas in others not, and relative position of the gonopore (in the posterior fifth to third of the proglottid length).
|
||
</paragraph>
|
||
<paragraph id="765436BDFF8C3C17FF7CF8FAFAE9AB90" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="552">
|
||
Only museum specimens of the new species were available and we were unable to obtain the
|
||
<typeStatus id="A950881FFF8C3C17FB51F8FAFA91ABD8" box="[1258,1306,1915,1941]" pageId="23" pageNumber="552">type</typeStatus>
|
||
material of
|
||
<bibRefCitation id="127A4B4CFF8C3C17FF2CF81EFECEABF4" author="Nybelin, O." box="[151,325,1951,1977]" pageId="23" pageNumber="552" pagination="1 - 48" refId="ref21965" refString="Nybelin, O. (1917) Australischen Cestoden. Results of D. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. Kungliga Svenska Vetenskapsakademiens Handlingar, 52 (14), 1 - 48." type="journal article" year="1917">Nybelin (1917)</bibRefCitation>
|
||
. Therefore, it was uncertain whether Nybelin’s material is conspecific with the new species. Furthermore, no molecular data are available to assess phylogenetic relationships of this parasite of Gould’s monitor.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |