1881 lines
245 KiB
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1881 lines
245 KiB
XML
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<mods:title id="5A1475BB5D06F956B820FBAF61C532B9">A reappraisal of Notohypshilophodon comodorensis (Ornithischia: Ornithopoda) from the Late Cretaceous of Patagonia, Argentina</mods:title>
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<mods:namePart id="2679234AF8D07D0CC48C15EB7580B31B">Ibiricu, Lucio M.</mods:namePart>
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<mods:namePart id="2DF71B51046C02379A001F4AE2140369">Martínez, Rubén D.</mods:namePart>
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<mods:namePart id="8BF01C017CD2A7B0E388F143B8F9F484">Luna, Marcelo</mods:namePart>
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<mods:namePart id="755D4E7B54533E830E3FE1ADAD9B6960">Casal, Gabriel A.</mods:namePart>
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<treatment id="6B62B54B5933DA5176D39A59232BFE05" ID-DOI="http://doi.org/10.5281/zenodo.6137396" ID-GBIF-Taxon="119422849" ID-Zenodo-Dep="6137396" LSID="urn:lsid:plazi:treatment:6B62B54B5933DA5176D39A59232BFE05" httpUri="http://treatment.plazi.org/id/6B62B54B5933DA5176D39A59232BFE05" lastPageId="13" lastPageNumber="414" pageId="1" pageNumber="402">
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<paragraph id="E374045D5933DA5D76D39A5920C7F89C" blockId="1.[151,729,1819,1880]" box="[151,729,1819,1844]" pageId="1" pageNumber="402">
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<heading id="B83CB3315933DA5D76D39A5920C7F89C" bold="true" box="[151,729,1819,1844]" fontSize="10" level="2" pageId="1" pageNumber="402" reason="2">
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<taxonomicName id="24CB7FDE5933DA5D76D39A5920C7F89C" authority="Martinez, 1998" authorityName="Martinez" authorityYear="1998" box="[151,729,1819,1844]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="402" phylum="Chordata" rank="species" species="comodorensis">
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<emphasis id="D1BFD84F5933DA5D76D39A5920C7F89C" bold="true" box="[151,729,1819,1844]" pageId="1" pageNumber="402">
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<emphasis id="D1BFD84F5933DA5D76D39A592000F89C" bold="true" box="[151,542,1819,1844]" italics="true" pageId="1" pageNumber="402">Notohypsilophodon comodorensis</emphasis>
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<bibRefCitation id="875A79AC5933DA5D74619A5920C7F89C" author="Martinez" box="[549,729,1819,1844]" pageId="1" pageNumber="402" refString="Martinez, R. D. (1998) Notohypsilophodon comodorensis gen. et sp. nov. Un Hypsilophodontidae (Ornithischia: Ornithopoda) del Cretacico Superior de Chubut, Patagonia central, Argentina. Acta Geologica Leopoldensia, 21, 119 - 135." type="journal article" year="1998">Martínez, 1998</bibRefCitation>
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</emphasis>
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</taxonomicName>
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<paragraph id="E374045D5933DA5D76D39A7D2313F8F0" blockId="1.[151,729,1819,1880]" box="[151,269,1855,1880]" pageId="1" pageNumber="402">
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<figureCitation id="7BF018D85933DA5D76D39A7D2313F8F0" box="[151,269,1855,1880]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStart-8="FIGURE 10" captionStartId-0="2.[151,250,1695,1717]" captionStartId-1="3.[151,250,1587,1609]" captionStartId-2="5.[151,250,1786,1808]" captionStartId-3="6.[151,250,1450,1472]" captionStartId-4="7.[151,250,1595,1617]" captionStartId-5="8.[151,250,1165,1187]" captionStartId-6="10.[151,250,1729,1751]" captionStartId-7="11.[151,250,1696,1718]" captionStartId-8="13.[151,250,1327,1349]" captionTargetBox-0="[151,1436,553,1673]" captionTargetBox-1="[340,1247,697,1565]" captionTargetBox-2="[151,1436,193,1764]" captionTargetBox-3="[340,1247,543,1429]" captionTargetBox-4="[340,1247,808,1572]" captionTargetBox-5="[151,1436,193,1143]" captionTargetBox-6="[151,1436,193,1707]" captionTargetBox-7="[151,1436,373,1674]" captionTargetBox-8="[340,1247,483,1305]" captionTargetId-0="figure@2.[151,1436,553,1674]" captionTargetId-1="figure@3.[340,1247,697,1565]" captionTargetId-2="figure@5.[151,1436,193,1765]" captionTargetId-3="figure@6.[340,1248,543,1429]" captionTargetId-4="figure@7.[340,1247,808,1573]" captionTargetId-5="figure@8.[151,1436,193,1144]" captionTargetId-6="figure@10.[151,1436,193,1707]" captionTargetId-7="figure@11.[151,1436,373,1675]" captionTargetId-8="figure@13.[340,1247,483,1305]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="13" captionText-0="FIGURE 2. Notohypsilophodon comodorensis. Middle cervical vertebra (UNPSJB-PV 942 / 1), in anterior (A), in right lateral (B), in dorsal (C) and ventral (D) views. Abbreviations: nc, neural canal; ncs, neural central suture; ns, neural spine; pa, parapophysis; poz, postzygapophysis; prz, prezygapophysis; vk, ventral keel." captionText-1="FIGURE 3. Notohypsilophodon comodorensis. Anterior dorsal vertebrae (UNPSJB-PV 942 / 5,6,7 respectively), in right lateral (A and B) and ventral (C) views. Abbreviations: dp, diapophysis; ns, neural spine; pa, parapophysis; prz, prezygapophysis; rg, rugosities; vk, ventral keel." captionText-2="FIGURE 4. Notohypsilophodon comodorensis. Sacral vertebrae (UNPSJB-PV 942 / 12,13,14,15,16 respectively), in left lateral (A and B), in dorsal (C and D) and ventral (E and F) views. Abbreviations: ns, neural spine; rg, rugosities; tp, transverse process." captionText-3="FIGURE 5. Notohypsilophodon comodorensis. Anterior caudal vertebrae (UNPSJB-PV 942 / 17,18), in right lateral (A and C) and ventral (B and D) views. Abbreviations: cf, chevron facet; lgr, longitudinal groove; ncs, neurocentral suture; str, striations. Scale bar equals 2 cm." captionText-4="FIGURE 6. Notohypsilophodon comodorensis. Left scapula (UNPSJB-PV 942 / 23), in lateral (A) and proximal (B) views. Right coracoids (UNPSJB-PV 942 / 24), in lateral view (C). Abbreviation: cof, coracoids foramen; gl, glenoid; scb, scapular blade; spb, supraglenoidal buttress; spf, supraglenoid fossa; stp, sternal process." captionText-5="FIGURE 7. Notohypsilophodon comodorensis. Right humerus (UNPSJB-PV 942 / 25), in anterior (A), proximal (B) and distal (C) views. Left ulna (UNPSJB-PV 942 / 27), in medial (D), proximal (E) and distal (F) views. Abbreviation: asr, articular surface radius; dc, deltopectoral crest; hh, humeral head; lsh, lateral shoulder; ms, muscle scars; msh, medial shoulder; op, olecranon process; rc, radial condyle; uc, ulnar condyle." captionText-6="FIGURE 8. Notohypsilophodon comodorensis. Left femur (UNPSJB-PV 942 / 28), in medial (A- D), proximal (B), distal (C), posterolateral (E), lateral (F), anterior (G) and posterior (H) views. Abbreviation: fh, femoral head; gth, greater trochanter; ict, intertrochanteric cleft; lc, lateral condyle; lis, ligament sulcus; lth, lesser trochanter; mc, medial condyle; pgr, posterior groove; tub, tubercle. Note: dotted lines delimit the reconstructed femoral shaft." captionText-7="FIGURE 9. Notohypsilophodon comodorensis. Left tibia (UNPSJB-PV 942 / 29), in medial (A), proximal (B) and distal (C) views. Right tibia (UNPSJB-PV 942 / 30), in distal (D) view. Left fibula (UNPSJB-PV 942 / 32), in medial (E) view. Right astragalus (UNPSJB-PV 942 / 33), in posterior (F) and anterior (G) views. Left calcaneum (UNPSJB-PV 942 / 34), in lateral (H) and medial (I) views. Abbreviation: ac, astragalar contact; cc, cnemial crest; dbl, distal bulge; dpe, distal perforation; fc, fibular condyle; fs, fibular surface; ic, inner condyle; it, incisura tibialis; ppr, proximal process; prb, proximal bulge; ts, tibial surface." captionText-8="FIGURE 10. Notohypsilophodon comodorensis. Non-ungual phalanx (UNPSJB-PV 942 / 35), in medial (A), dorsal (B) and proximal (C) views. Non-ungual phalanx (UNPSJB-PV 942 / 37), in medial (B) view. Non-ungual phalanx (UNPSJB-PV 942 / 39), in dorsal (C) view. Ungual phalanges (UNPSJB-PV 942 / 45,46,47), in dorsal view. Abbreviation: clp, collateral ligament pit; ugr, ungula groove." httpUri-0="https://zenodo.org/record/226295/files/figure.png" httpUri-1="https://zenodo.org/record/226296/files/figure.png" httpUri-2="https://zenodo.org/record/226297/files/figure.png" httpUri-3="https://zenodo.org/record/226298/files/figure.png" httpUri-4="https://zenodo.org/record/226299/files/figure.png" httpUri-5="https://zenodo.org/record/226300/files/figure.png" httpUri-6="https://zenodo.org/record/226301/files/figure.png" httpUri-7="https://zenodo.org/record/226302/files/figure.png" httpUri-8="https://zenodo.org/record/226303/files/figure.png" pageId="1" pageNumber="402">Figs. 2–10</figureCitation>
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</paragraph>
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<subSubSection id="ABD157D65933DA5E76D39AC523A9FE5D" lastPageId="2" lastPageNumber="403" pageId="1" pageNumber="402" type="materials_examined">
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<paragraph id="E374045D5933DA5E76D39AC52310FF7D" blockId="1.[151,1436,1927,2024]" lastBlockId="2.[151,1436,151,501]" lastPageId="2" lastPageNumber="403" pageId="1" pageNumber="402">
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<emphasis id="D1BFD84F5933DA5D76D39AC52315F808" bold="true" box="[151,267,1927,1952]" pageId="1" pageNumber="402">
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<typeStatus id="3C70BAFF5933DA5D76D39AC52318F808" box="[151,262,1927,1952]" pageId="1" pageNumber="402" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="85DA9C985933DA5D775D9ACA23BBF808" box="[281,421,1928,1952]" httpUri="http://grbio.org/cool/h86n-gpzd" name="Universidad Nacional de la Patagonia" pageId="1" pageNumber="402">UNPSJB-Pv</collectionCode>
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942 (
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<figureCitation id="7BF018D85933DA5D77B09AC5206EF808" box="[500,624,1927,1952]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStart-8="FIGURE 10" captionStartId-0="2.[151,250,1695,1717]" captionStartId-1="3.[151,250,1587,1609]" captionStartId-2="5.[151,250,1786,1808]" captionStartId-3="6.[151,250,1450,1472]" captionStartId-4="7.[151,250,1595,1617]" captionStartId-5="8.[151,250,1165,1187]" captionStartId-6="10.[151,250,1729,1751]" captionStartId-7="11.[151,250,1696,1718]" captionStartId-8="13.[151,250,1327,1349]" captionTargetBox-0="[151,1436,553,1673]" captionTargetBox-1="[340,1247,697,1565]" captionTargetBox-2="[151,1436,193,1764]" captionTargetBox-3="[340,1247,543,1429]" captionTargetBox-4="[340,1247,808,1572]" captionTargetBox-5="[151,1436,193,1143]" captionTargetBox-6="[151,1436,193,1707]" captionTargetBox-7="[151,1436,373,1674]" captionTargetBox-8="[340,1247,483,1305]" captionTargetId-0="figure@2.[151,1436,553,1674]" captionTargetId-1="figure@3.[340,1247,697,1565]" captionTargetId-2="figure@5.[151,1436,193,1765]" captionTargetId-3="figure@6.[340,1248,543,1429]" captionTargetId-4="figure@7.[340,1247,808,1573]" captionTargetId-5="figure@8.[151,1436,193,1144]" captionTargetId-6="figure@10.[151,1436,193,1707]" captionTargetId-7="figure@11.[151,1436,373,1675]" captionTargetId-8="figure@13.[340,1247,483,1305]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="13" captionText-0="FIGURE 2. Notohypsilophodon comodorensis. Middle cervical vertebra (UNPSJB-PV 942 / 1), in anterior (A), in right lateral (B), in dorsal (C) and ventral (D) views. Abbreviations: nc, neural canal; ncs, neural central suture; ns, neural spine; pa, parapophysis; poz, postzygapophysis; prz, prezygapophysis; vk, ventral keel." captionText-1="FIGURE 3. Notohypsilophodon comodorensis. Anterior dorsal vertebrae (UNPSJB-PV 942 / 5,6,7 respectively), in right lateral (A and B) and ventral (C) views. Abbreviations: dp, diapophysis; ns, neural spine; pa, parapophysis; prz, prezygapophysis; rg, rugosities; vk, ventral keel." captionText-2="FIGURE 4. Notohypsilophodon comodorensis. Sacral vertebrae (UNPSJB-PV 942 / 12,13,14,15,16 respectively), in left lateral (A and B), in dorsal (C and D) and ventral (E and F) views. Abbreviations: ns, neural spine; rg, rugosities; tp, transverse process." captionText-3="FIGURE 5. Notohypsilophodon comodorensis. Anterior caudal vertebrae (UNPSJB-PV 942 / 17,18), in right lateral (A and C) and ventral (B and D) views. Abbreviations: cf, chevron facet; lgr, longitudinal groove; ncs, neurocentral suture; str, striations. Scale bar equals 2 cm." captionText-4="FIGURE 6. Notohypsilophodon comodorensis. Left scapula (UNPSJB-PV 942 / 23), in lateral (A) and proximal (B) views. Right coracoids (UNPSJB-PV 942 / 24), in lateral view (C). Abbreviation: cof, coracoids foramen; gl, glenoid; scb, scapular blade; spb, supraglenoidal buttress; spf, supraglenoid fossa; stp, sternal process." captionText-5="FIGURE 7. Notohypsilophodon comodorensis. Right humerus (UNPSJB-PV 942 / 25), in anterior (A), proximal (B) and distal (C) views. Left ulna (UNPSJB-PV 942 / 27), in medial (D), proximal (E) and distal (F) views. Abbreviation: asr, articular surface radius; dc, deltopectoral crest; hh, humeral head; lsh, lateral shoulder; ms, muscle scars; msh, medial shoulder; op, olecranon process; rc, radial condyle; uc, ulnar condyle." captionText-6="FIGURE 8. Notohypsilophodon comodorensis. Left femur (UNPSJB-PV 942 / 28), in medial (A- D), proximal (B), distal (C), posterolateral (E), lateral (F), anterior (G) and posterior (H) views. Abbreviation: fh, femoral head; gth, greater trochanter; ict, intertrochanteric cleft; lc, lateral condyle; lis, ligament sulcus; lth, lesser trochanter; mc, medial condyle; pgr, posterior groove; tub, tubercle. Note: dotted lines delimit the reconstructed femoral shaft." captionText-7="FIGURE 9. Notohypsilophodon comodorensis. Left tibia (UNPSJB-PV 942 / 29), in medial (A), proximal (B) and distal (C) views. Right tibia (UNPSJB-PV 942 / 30), in distal (D) view. Left fibula (UNPSJB-PV 942 / 32), in medial (E) view. Right astragalus (UNPSJB-PV 942 / 33), in posterior (F) and anterior (G) views. Left calcaneum (UNPSJB-PV 942 / 34), in lateral (H) and medial (I) views. Abbreviation: ac, astragalar contact; cc, cnemial crest; dbl, distal bulge; dpe, distal perforation; fc, fibular condyle; fs, fibular surface; ic, inner condyle; it, incisura tibialis; ppr, proximal process; prb, proximal bulge; ts, tibial surface." captionText-8="FIGURE 10. Notohypsilophodon comodorensis. Non-ungual phalanx (UNPSJB-PV 942 / 35), in medial (A), dorsal (B) and proximal (C) views. Non-ungual phalanx (UNPSJB-PV 942 / 37), in medial (B) view. Non-ungual phalanx (UNPSJB-PV 942 / 39), in dorsal (C) view. Ungual phalanges (UNPSJB-PV 942 / 45,46,47), in dorsal view. Abbreviation: clp, collateral ligament pit; ugr, ungula groove." httpUri-0="https://zenodo.org/record/226295/files/figure.png" httpUri-1="https://zenodo.org/record/226296/files/figure.png" httpUri-2="https://zenodo.org/record/226297/files/figure.png" httpUri-3="https://zenodo.org/record/226298/files/figure.png" httpUri-4="https://zenodo.org/record/226299/files/figure.png" httpUri-5="https://zenodo.org/record/226300/files/figure.png" httpUri-6="https://zenodo.org/record/226301/files/figure.png" httpUri-7="https://zenodo.org/record/226302/files/figure.png" httpUri-8="https://zenodo.org/record/226303/files/figure.png" pageId="1" pageNumber="402">Figs 2–10</figureCitation>
|
||
). A partial skeleton including four cervical vertebrae, seven dorsal vertebrae, four sacral vertebrae, six caudal vertebrae (all of the vertebrae are partially preserved), rib fragment, a portion of the left scapula, right coracoid, right humerus, left and right ulnae, an incomplete left femur, right tibia, an incomplete left tibia, left fibula, an incomplete right fibula; right astragalus, left calcaneum and 13 pedal phalanges.
|
||
</paragraph>
|
||
<paragraph id="E374045D5930DA5E76839D9D2149FE20" blockId="2.[151,1436,151,501]" pageId="2" pageNumber="403">
|
||
<emphasis id="D1BFD84F5930DA5E76839D9D23A5FF50" bold="true" box="[199,443,223,248]" pageId="2" pageNumber="403">Emended diagnosis.</emphasis>
|
||
A small ornithopod dinosaur distinguished from all other basal ornithopods by the following autapomorphies originally proposed by
|
||
<bibRefCitation id="875A79AC5930DA5E74829C462163FEB4" author="Martinez" box="[710,893,260,285]" pageId="2" pageNumber="403" refString="Martinez, R. D. (1998) Notohypsilophodon comodorensis gen. et sp. nov. Un Hypsilophodontidae (Ornithischia: Ornithopoda) del Cretacico Superior de Chubut, Patagonia central, Argentina. Acta Geologica Leopoldensia, 21, 119 - 135." type="journal article" year="1998">Martinez (1998)</bibRefCitation>
|
||
and supported in this study: Anteromedial bulge on the proximal extremity of the tibia, calcaneum with a pronounced posterodistal projection. Additionally, the following new autapomorphies are proposed: Distal tip of the sacral neural spine strongly projected posteriorly, sternal process of the coracoids markedly reduced (see below).
|
||
</paragraph>
|
||
<paragraph id="E374045D5930DA5E76839CD623A9FE5D" blockId="2.[151,1436,151,501]" pageId="2" pageNumber="403">
|
||
<emphasis id="D1BFD84F5930DA5E76839CD623D5FE05" bold="true" box="[199,459,404,429]" pageId="2" pageNumber="403">Locality and horizon.</emphasis>
|
||
<quantity id="2433A9B85930DA5E77979CD62002FE05" box="[467,540,404,429]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="403" unit="km" value="28.0">28 km</quantity>
|
||
northeast of the town of Buen Pasto, in the central-south of Chubut Province, Patagonia,
|
||
<collectingCountry id="9BDC44CD5930DA5E775F9CF52394FE78" box="[283,394,439,464]" name="Argentina" pageId="2" pageNumber="403">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="7BF018D85930DA5E77E49CF523F7FE78" box="[416,489,439,464]" captionStart="FIGURE 1" captionStartId="1.[151,250,1187,1209]" captionTargetBox="[177,1410,197,1163]" captionTargetId="figure@1.[173,1413,193,1166]" captionTargetPageId="1" captionText="FIGURE 1. Site map showing geographical location of Notohypsilophodon comodorensis (UNPSJB-PV 942), denoted by the black arrow." httpUri="https://zenodo.org/record/226294/files/figure.png" pageId="2" pageNumber="403">Fig. 1</figureCitation>
|
||
). Bajo Barreal Formation, Lower Member (Late Cretaceous, Cenomanian, see
|
||
<bibRefCitation id="875A79AC5930DA5E76D39C9E23B5FE5D" author="Martinez" box="[151,427,476,501]" pageId="2" pageNumber="403" refString="Martinez, R. D. & Novas, F. E. (2006) Aniksosaurus darwini gen. et sp. nov., a new coelurosaurian theropod from the Early Late Cretaceous of central Patagonia, Argentina. Revista Museo Argentino de Ciencias Naturales, 8, 243 - 259." type="journal article" year="2006">Martinez & Novas 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="B7B454D55930DA5E76D39BDD21BAF95A" httpUri="https://zenodo.org/record/226295/files/figure.png" pageId="2" pageNumber="403" targetBox="[151,1436,553,1673]" targetPageId="2">
|
||
<paragraph id="E374045D5930DA5E76D39BDD21BAF95A" blockId="2.[151,1436,1695,1778]" pageId="2" pageNumber="403">
|
||
<emphasis id="D1BFD84F5930DA5E76D39BDD230AF91C" bold="true" box="[151,276,1695,1717]" pageId="2" pageNumber="403">FIGURE 2.</emphasis>
|
||
<taxonomicName id="24CB7FDE5930DA5E775F9BDD206BF91D" box="[283,629,1695,1717]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="comodorensis">
|
||
<emphasis id="D1BFD84F5930DA5E775F9BDD206BF91D" box="[283,629,1695,1717]" italics="true" pageId="2" pageNumber="403">Notohypsilophodon comodorensis</emphasis>
|
||
</taxonomicName>
|
||
. Middle cervical vertebra (UNPSJB-PV 942/1), in anterior (A), in right lateral (B), in dorsal (C) and ventral (D) views. Abbreviations: nc, neural canal; ncs, neural central suture; ns, neural spine; pa, parapophysis; poz, postzygapophysis; prz, prezygapophysis; vk, ventral keel.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="ABD157D65930DA5176839A58232BFE05" lastPageId="13" lastPageNumber="414" pageId="2" pageNumber="403" type="description">
|
||
<paragraph id="E374045D5930DA5F76839A58267FFD2D" blockId="2.[151,1436,1818,2023]" lastBlockId="3.[151,1436,151,645]" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
|
||
<emphasis id="D1BFD84F5930DA5E76839A58202CF89B" bold="true" box="[199,562,1818,1843]" pageId="2" pageNumber="403">Description and comparisons.</emphasis>
|
||
<collectingCountry id="9BDC44CD5930DA5E74789A582045F89A" box="[572,603,1818,1842]" name="American Samoa" pageId="2" pageNumber="403">As</collectingCountry>
|
||
mentioned by
|
||
<bibRefCitation id="875A79AC5930DA5E754A9A5821D4F89A" author="Martinez" box="[782,970,1818,1842]" pageId="2" pageNumber="403" refString="Martinez, R. D. (1998) Notohypsilophodon comodorensis gen. et sp. nov. Un Hypsilophodontidae (Ornithischia: Ornithopoda) del Cretacico Superior de Chubut, Patagonia central, Argentina. Acta Geologica Leopoldensia, 21, 119 - 135." type="journal article" year="1998">Martínez (1998)</bibRefCitation>
|
||
, the morphology of the material clearly differs from that seen in the most derived ornithopods (i.e., hadrosaurids). Likewise, the material does not possess synapomorphies or anatomical similarities of marginocephalians (i.e., pachycephalosaurs and ceratopsians). For this reason, members of both groups were excluded from our comparisons. Therefore, in order to ascertain the evolutionary affinities of
|
||
<taxonomicName id="24CB7FDE5930DA5E77859AE82083F86B" box="[449,669,1962,1987]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="403" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5930DA5E77859AE82083F86B" box="[449,669,1962,1987]" italics="true" pageId="2" pageNumber="403">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
within
|
||
<taxonomicName id="24CB7FDE5930DA5E74BF9AE82197F86B" box="[763,905,1962,1987]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="403" phylum="Chordata" rank="order">Ornithischia</taxonomicName>
|
||
, we compared the specimen to the following genera: basal neornithischians -
|
||
<emphasis id="D1BFD84F5930DA5E74409A8F20B9F84E" box="[516,679,1997,2022]" italics="true" pageId="2" pageNumber="403">Hexinlusaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5930DA5E74F39A8C2195F84E" author="Barrett" box="[695,907,1997,2022]" pageId="2" pageNumber="403" refString="Barrett, P. M., Butler, R. J. & Knoll, F. (2005) Small-bodied ornithischian dinosaurs from the Middle Jurassic of Sichuan, China. Journal of Vertebrate Paleontology, 25, 823 - 834. http: // dx. doi. org / 10.1671 / 0272 - 4634 (2005) 025 [0823: sodftm] 2.0. co; 2" type="journal article" year="2005">
|
||
Barrett
|
||
<emphasis id="D1BFD84F5930DA5E754B9A8D2156F84E" box="[783,840,1997,2022]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="24CB7FDE5930DA5E758F9A8F2645F84E" box="[971,1115,1997,2022]" class="Reptilia" family="Hypsilophodontidae" genus="Yandusaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="403" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5930DA5E758F9A8F2645F84E" box="[971,1115,1997,2022]" italics="true" pageId="2" pageNumber="403">Yandusaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5930DA5E722E9A8C26D1F84E" author="He" box="[1130,1231,1998,2022]" pageId="2" pageNumber="403" refString="He, X. L. (1979) A newly discovered ornithopod dinosaur, Yandusaurus from Zigong, Sichuan. Contribution to International Exchange in Geology, 2, 116 - 123." type="journal article" year="1979">He 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5930DA5E72989A8D2791F84E" author="He" box="[1244,1423,1997,2022]" pageId="2" pageNumber="403" refString="He, X. L. & Cai, K. (1984) The Middle Jurassic Dinosaur Fauna from Dashanpu, Zigong, Sichuan. Vol. 1. The Ornithopod Dinosaurs. Sichuan Publishing House of Science and Technology, Chengdu, 71 pp." type="book" year="1984">He & Cai 1984</bibRefCitation>
|
||
); basal (non-iguanodontian) ornithopods -
|
||
<taxonomicName id="24CB7FDE5931DA5F742E9DD52128FF18" box="[618,822,151,176]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F742E9DD52128FF18" box="[618,822,151,176]" italics="true" pageId="3" pageNumber="404">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F75029DDA21E4FF18" author="Zan" box="[838,1018,151,176]" pageId="3" pageNumber="404" refString="Zan, S. Q., Chen, J., Jin, L. Y. & Li, T. (2005) A primitive ornithopod from the Early Cretaceous Quantou Formation of central Jilin, China. Vertebrata PalAsiatica, 43, 182 - 193." type="journal article" year="2005">
|
||
Zan
|
||
<emphasis id="D1BFD84F5931DA5F75389DDB21A8FF18" box="[892,950,151,176]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
; Butler
|
||
<emphasis id="D1BFD84F5931DA5F72139DDB268CFF18" box="[1111,1170,151,176]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2011),
|
||
<taxonomicName id="24CB7FDE5931DA5F72AF9DDA2782FF07" box="[1259,1436,152,175]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F72AF9DDA2782FF07" box="[1259,1436,152,175]" italics="true" pageId="3" pageNumber="404">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F76DA9DFE23BCFF7D" author="Coria" box="[158,418,188,213]" pageId="3" pageNumber="404" refString="Coria, R. A. & Salgado, L. (1996) A basal iguanodontian (Ornithischia: Ornithopoda) from the Late Cretaceous of South America. Journal of Vertebrate Paleontology, 16, 445 - 457. http: // dx. doi. org / 10.1080 / 02724634.1996.10011333" type="journal article" year="1996">Coria & Salgado 1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F77EB9DFE2090FF7C" author="Salgado" box="[431,654,188,213]" pageId="3" pageNumber="404" refString="Salgado, L., Coria, R. A. & Heredia, S. E. (1997) New materials of Gasparinisaura cincosaltensis (Ornithischia, Ornithopoda) from the Upper Cretaceous of Argentina. Journal of Paleontology, 71, 993 - 940." type="journal volume" year="1997">
|
||
Salgado
|
||
<emphasis id="D1BFD84F5931DA5F74559DFF2054FF7D" box="[529,586,188,213]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
1997
|
||
</bibRefCitation>
|
||
),
|
||
<emphasis id="D1BFD84F5931DA5F74E79DFE2141FF7D" box="[675,863,188,213]" italics="true" pageId="3" pageNumber="404">Gideonmantellia</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F752A9DFE2680FF7C" author="Ruiz-Omenaca" box="[878,1182,188,213]" pageId="3" pageNumber="404" refString="Ruiz-Omenaca, J. I., Canudo, J. I., Cuenca-Bescos, G., Cruzado-Cabellero, P., Gasca, J. M. & Moreno-Azanza, M. (2012) A new basal ornithopod dinosaur from the Barremian of Galve, Spain. Comptes Rendus Palevol, 11, 435 - 444. http: // dx. doi. org / 10.1016 / j. crpv. 2012.06.001" type="journal article" year="2012">
|
||
Ruiz-Omeñaca
|
||
<emphasis id="D1BFD84F5931DA5F72649DFF2644FF7D" box="[1056,1114,188,213]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
),
|
||
<emphasis id="D1BFD84F5931DA5F72F69DFF26F1FF7C" box="[1202,1263,189,212]" italics="true" pageId="3" pageNumber="404">Haya</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F72BA9DFE22E9FF50" author="Makovicky" pageId="3" pageNumber="404" refString="Makovicky, P. J., Kilbourne, B. M., Sadleir, R. W. & Norell, M. A. (2011) A new basal ornithopod (Dinosauria, Ornithischia) from the Late Cretaceous of Mongolia. Journal of Vertebrate Paleontology, 31, 626 - 640. http: // dx. doi. org / 10.1080 / 02724634.2011.557114" type="journal article" year="2011">
|
||
Makovicky
|
||
<emphasis id="D1BFD84F5931DA5F73CC9DFF22AAFF50" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="24CB7FDE5931DA5F77499D9D23A6FF50" box="[269,440,223,248]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F77499D9D23A6FF50" box="[269,440,223,248]" italics="true" pageId="3" pageNumber="404">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F778D9D9D2079FF50" author="Galton" box="[457,615,223,248]" pageId="3" pageNumber="404" refString="Galton, P. M. (1974 a) The ornithischian dinosaur Hypsilophodon from the Wealden of the Isle of Wight. Bulletin of the British Museum (Natural History), Geology, 25, 1 - 152." type="journal article" year="1974" yearSuffix="a">Galton 1974a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F74319DA2217BFF50" author="Torres" box="[629,869,223,248]" pageId="3" pageNumber="404" refString="Torres, J. & Viera, L. (1994) Hypsilophodon foxii (Reptilia, Ornithischia) en el Cretacico inferior de Igea (La Rioja, Espana). Munibe, 46, 3 - 41." type="journal article" year="1994">Torres & Viera 1994</bibRefCitation>
|
||
),
|
||
<taxonomicName id="24CB7FDE5931DA5F753E9D9D2611FF50" box="[890,1039,223,248]" class="Reptilia" family="Thescelosauridae" genus="Jeholosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F753E9D9D2611FF50" box="[890,1039,223,248]" italics="true" pageId="3" pageNumber="404">Jeholosaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F72649DA226D5FF50" author="Xu" box="[1056,1227,223,248]" pageId="3" pageNumber="404" refString="Xu, X., Wang, L. X. & You, H. L. (2000) A primitive ornithopod from the Early Cretaceous Yixian Formation of Liaoning. Vertebrata Palasiatica, 38, 318 - 325." type="journal article" year="2000">
|
||
Xu
|
||
<emphasis id="D1BFD84F5931DA5F72099DA32696FF50" box="[1101,1160,223,248]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2000
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F729D9DA2278EFF50" author="Han" box="[1241,1424,223,248]" pageId="3" pageNumber="404" refString="Han, F. L., Barrett, P. M., Butler, R. J. & Xu, X. (2012) Postcranial anatomy of Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Late Cretaceous Yixian Formation of China. Journal of Vertebrate Paleontology, 32, 1370 - 1395. http: // dx. doi. org / 10.1080 / 02724634.2012.694385" type="journal article" year="2012">
|
||
Han
|
||
<emphasis id="D1BFD84F5931DA5F73559DA32755FF50" box="[1297,1355,223,248]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="24CB7FDE5931DA5F76D39C472006FEB4" authority="Huh et al. 2010" authorityName="Huh et al." authorityYear="2010" box="[151,536,260,285]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F76D39C47235DFEB4" box="[151,323,261,284]" italics="true" pageId="3" pageNumber="404">Koreanosaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F77179C46200EFEB4" author="Huh" box="[339,528,260,285]" pageId="3" pageNumber="404" refString="Huh, M., Lee, D. G., Kim, J. K., Lim, J. D. & Godefroit, P. (2010) A new basal ornithopod dinosaur from the Upper Cretaceous of South Korea. Neues Jahrbuch fur Geologie Palaontologie, Abhandlungen, 259, 1 - 24. http: // dx. doi. org / 10.1127 / 0077 - 7749 / 2010 / 0102" type="journal article" year="2010">
|
||
Huh
|
||
<emphasis id="D1BFD84F5931DA5F77CB9C4723D5FEB5" box="[399,459,260,285]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5931DA5F74629C46260AFEB4" authority="Horner & Weishampel 1988" authorityName="Horner & Weishampel" authorityYear="1988" box="[550,1044,260,285]" class="Reptilia" family="Hypsilophodontidae" genus="Orodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F74629C4620AFFEB5" box="[550,689,260,285]" italics="true" pageId="3" pageNumber="404">Orodromeus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F74879C462612FEB4" author="Horner" box="[707,1036,260,285]" pageId="3" pageNumber="404" refString="Horner, J. R. & Weishampel, D. B. (1988) A comparative embryological study of two ornithischian dinosaurs. Nature, 332, 256 - 257. http: // dx. doi. org / 10.1038 / 332256 a 0" type="journal article" year="1988">Horner & Weishampel 1988</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5931DA5F72679C4622C5FEE8" authority="Varricchio et al. 2007" authorityName="Varricchio et al." authorityYear="2007" class="Reptilia" family="Hypsilophodontidae" genus="Oryctodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F72679C4626CEFEB5" box="[1059,1232,260,285]" italics="true" pageId="3" pageNumber="404">Oryctodromeus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F72A59C4622CDFEE8" author="Varricchio" pageId="3" pageNumber="404" refString="Varricchio, D. J., Martin, A. J. & Katsura, Y. (2007) First trace and body fossil evidence of a burrowing, denning dinosaur. Proceedings of the Royal Society, 274, 1361 - 1368. http: // dx. doi. org / 10.1098 / rspb. 2006.0443" type="journal article" year="2007">
|
||
Varricchio
|
||
<emphasis id="D1BFD84F5931DA5F73259C472782FEB5" box="[1377,1436,260,285]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5931DA5F76A29C65239EFEE8" box="[230,384,295,320]" class="Reptilia" family="Hypsilophodontidae" genus="Parksosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F76A29C65239EFEE8" box="[230,384,295,320]" italics="true" pageId="3" pageNumber="404">Parksosaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F77CB9C65200FFEE8" author="Parks" box="[399,529,295,320]" pageId="3" pageNumber="404" refString="Parks, W. A. (1926) Thescelosaurus warreni, a new species of orthopodous dinosaur from the Edmonton Formation of Alberta. University of Toronto Studies in Geology Series, 21, 1 - 42." type="journal article" year="1926">Parks 1926</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F74589C6520B5FEE8" author="Galton" box="[540,683,295,320]" pageId="3" pageNumber="404" refString="Galton, P. M. (1973) Redescription of the skull and mandible of Parksosaurus from the Late Cretaceous with comments on the family Hypsilophodontidae (Ornithischia). Life Science Contributions of the Royal Ontario Museum, 89, 1 - 21." type="journal article" year="1973">Galton 1973</bibRefCitation>
|
||
),
|
||
<taxonomicName id="24CB7FDE5931DA5F74FB9C652173FEE8" box="[703,877,295,320]" class="Reptilia" family="Hypsilophodontidae" genus="Thescelosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F74FB9C652173FEE8" box="[703,877,295,320]" italics="true" pageId="3" pageNumber="404">Thescelosaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F75389C652603FEE8" author="Gilmore" box="[892,1053,295,320]" pageId="3" pageNumber="404" refString="Gilmore, C. W. (1915) Osteology of Thescelosaurus, an orthopodous dinosaur from the Lance Formation of Wyoming. Proceedings of the United States National Museum, 49, 591 - 616." type="journal article" year="1915">Gilmore 1915</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F726D9C6526D8FEE8" author="Galton" box="[1065,1222,295,320]" pageId="3" pageNumber="404" refString="Galton, P. M. (1974 b) Notes on Thescelosaurus, a conservative ornithopod dinosaur from the Upper Cretaceous of North America, with comments on ornithopod classification. Journal of Paleontology, 48, 1048 - 1067." type="journal article" year="1974" yearSuffix="b">Galton 1974b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F72959C65278EFEE8" author="Boyd" box="[1233,1424,295,320]" pageId="3" pageNumber="404" refString="Boyd, C. A., Brown, C. M., Scheetz, R. D. & Clarke, J. A. (2009) Taxonomic revision of the basal neornithischian taxa Thescelosaurus and Bugenasaura. Journal of Vertebrate Paleontology, 29, 758 - 770. http: // dx. doi. org / 10.1671 / 039.029.0328" type="journal article" year="2009">
|
||
Boyd
|
||
<emphasis id="D1BFD84F5931DA5F73509C6B2753FEE8" box="[1300,1357,295,320]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
),
|
||
<emphasis id="D1BFD84F5931DA5F76D39C0F2313FECC" box="[151,269,333,356]" italics="true" pageId="3" pageNumber="404">Trinisaura</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F77599C0E23FAFECC" author="Coria" box="[285,484,332,357]" pageId="3" pageNumber="404" refString="Coria, R. A., Moly, J. J., Reguero, M., Santillana, S. & Marenssi, S. (2013) A new ornithopod (Dinosauria; Ornithischia) from Antarctica. Cretaceous Research, 41, 186 - 193. http: // dx. doi. org / 10.1016 / j. cretres. 2012.12.004" type="journal article" year="2013">
|
||
Coria
|
||
<emphasis id="D1BFD84F5931DA5F77219C0F2381FECD" box="[357,415,332,357]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
), and
|
||
<emphasis id="D1BFD84F5931DA5F746D9C0F20B8FECC" box="[553,678,333,356]" italics="true" pageId="3" pageNumber="404">Yueosaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F74FD9C0E2194FECC" author="Zheng" box="[697,906,332,357]" pageId="3" pageNumber="404" refString="Zheng, W. J., Jin, X. S., Shibata, M., Azuma, Y. & Yu, F. M. (2012) A new ornithischian dinosaur from the Cretaceous Liangtoutang Formation of Tiantai, Zhejiang Province, China. Cretaceous Research, 34, 208 - 219. http: // dx. doi. org / 10.1016 / j. cretres. 2011.11.001" type="journal article" year="2012">
|
||
Zheng
|
||
<emphasis id="D1BFD84F5931DA5F754F9C0F215BFECD" box="[779,837,332,357]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
); non-hadrosaurid iguanodontians -
|
||
<taxonomicName id="24CB7FDE5931DA5F73609C0E238BFE20" authority="Coria & Calvo 2002" authorityName="Coria & Calvo" authorityYear="2002" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F73609C0E2782FECD" box="[1316,1436,332,357]" italics="true" pageId="3" pageNumber="404">Anabisetia</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F76DA9C2D2393FE20" author="Coria" box="[158,397,367,392]" pageId="3" pageNumber="404" refString="Coria, R. A. & Calvo, J. O. (2002) A new iguanodontian ornithopod from Neuquen Basin, Patagonia, Argentina. Journal of Vertebrate Paleontology, 22, 503 - 509. http: // dx. doi. org / 10.1671 / 0272 - 4634 (2002) 022 [0503: aniofn] 2.0. co; 2" type="journal article" year="2002">Coria & Calvo 2002</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5931DA5F77E79C322088FE20" box="[419,662,367,392]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="D1BFD84F5931DA5F77E79C322088FE20" box="[419,662,367,392]" italics="true" pageId="3" pageNumber="404">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="D1BFD84F5931DA5F74E39C3226F6FE20" box="[679,1256,367,392]" italics="true" pageId="3" pageNumber="404">
|
||
<taxonomicName id="24CB7FDE5931DA5F74E39C32267BFE2F" authority="Cumnoria" authorityName="Cumnoria" box="[679,1125,367,392]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="prestwichii">Camptosaurus prestwichii = Cumnoria</taxonomicName>
|
||
<taxonomicName id="24CB7FDE5931DA5F72299C2D26F6FE20" box="[1133,1256,367,392]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5931DA5F72BD9C322048FE05" italics="true" pageId="3" pageNumber="404">
|
||
<taxonomicName id="24CB7FDE5931DA5F72BD9C3223A8FE05" authority="Uteodon" authorityName="Uteodon" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="aphanoecetes">Camptosaurus aphanoecetes = Uteodon</taxonomicName>
|
||
<taxonomicName id="24CB7FDE5931DA5F77FA9CD62048FE05" box="[446,598,404,429]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="aphanoecetes">aphanoecetes</taxonomicName>
|
||
</emphasis>
|
||
[
|
||
<bibRefCitation id="875A79AC5931DA5F74229CD62112FE04" author="Gilmore" box="[614,780,404,429]" pageId="3" pageNumber="404" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 196 - 302. http: // dx. doi. org / 10.5479 / si. 00963801.36 - 1666.197" type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F755D9CD6262CFE04" author="Galton" box="[793,1074,404,429]" pageId="3" pageNumber="404" refString="Galton, P. M. & Powell, H. P. (1980) The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology, 23, 411 - 443." type="journal article" year="1980">Galton and Powell 1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F72049CD62774FE04" author="Carpenter" box="[1088,1386,404,429]" pageId="3" pageNumber="404" refString="Carpenter, K. & Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum, 76, 227 - 263. http: // dx. doi. org / 10.2992 / 0097 - 4463 (2008) 76 [227: ansoco] 2.0. co; 2" type="journal article" year="2008">Carpenter & Wilson 2008</bibRefCitation>
|
||
, see
|
||
<bibRefCitation id="875A79AC5931DA5F76D39CF5234BFE78" author="McDonald" box="[151,341,439,464]" pageId="3" pageNumber="404" refString="McDonald, A. T. (2011) The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda). Zootaxa, 2783, 52 - 68." type="journal article" year="2011">
|
||
<collectingCountry id="9BDC44CD5931DA5F76D39CF52311FE78" box="[151,271,439,464]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="404">McDonald</collectingCountry>
|
||
2011
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="24CB7FDE5931DA5F77299CFB23EFFE78" box="[365,497,441,464]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F77299CFB23EFFE78" box="[365,497,441,464]" italics="true" pageId="3" pageNumber="404">Dryosaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F74419CF520ACFE78" author="Janensch" box="[517,690,439,464]" pageId="3" pageNumber="404" refString="Janensch, W. (1955) Der ornithopode Dysalotosaurus der Tendaguruschichten. Palaeontographica Supplement, 7, 105 - 176." type="journal article" year="1955">Janensch 1955</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F74879CF52148FE78" author="Galton" box="[707,854,439,464]" pageId="3" pageNumber="404" refString="Galton, P. M. (1981) Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa: postcranial skeleton. Palaontologische Zeitschrift, 55, 271 - 312." type="journal article" year="1981">Galton 1981</bibRefCitation>
|
||
),
|
||
<taxonomicName id="24CB7FDE5931DA5F752A9CF526B6FE78" authority="Norman 1980" authorityName="Norman" authorityYear="1980" box="[878,1192,439,464]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F752A9CF521F9FE78" box="[878,999,439,464]" italics="true" pageId="3" pageNumber="404">Iguanodon</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F75BF9CFA26BEFE78" author="Norman" box="[1019,1184,440,464]" pageId="3" pageNumber="404" refString="Norman, D. B. (1980) On the ornithischian dinosaur Iguanodon bernissartensis from the Lower Cretaceous of Bernissart (Belgium). Memoire de l'Institut Royal des Sciences Naturelles de Belgique, 178, 1 - 103." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D1BFD84F5931DA5F72FC9CF52782FE78" box="[1208,1436,439,464]" italics="true" pageId="3" pageNumber="404">Macrogryphosaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F76DA9C9E2376FE5C" author="Calvo" box="[158,360,476,501]" pageId="3" pageNumber="404" refString="Calvo, J. O., Porfiri, J. D. & Novas, F. E. (2007) Discovery of a new ornithopod dinosaur from the Portezuelo Formation (Upper Cretaceous), Neuquen, Patagonia, Argentina. Arquivos do Museu Nacional, Rio de Janeiro, 65, 471 - 483." type="journal article" year="2007">
|
||
Calvo
|
||
<emphasis id="D1BFD84F5931DA5F76AF9C9F233AFE5D" box="[235,292,476,501]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="24CB7FDE5931DA5F77389C9E2038FE5D" box="[380,550,476,501]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F77389C9E2038FE5D" box="[380,550,476,501]" italics="true" pageId="3" pageNumber="404">Mantellisaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F74719C9E20D2FE5D" author="Hooley" box="[565,716,476,501]" pageId="3" pageNumber="404" refString="Hooley, R. W. (1925) On the skeleton of Iguanodon atherfieldensis sp. nov., from the Wealden shales of Atherfield (Isle of Wight). Quarterly Journal of the Geological Society of London, 81, 1 - 61." type="journal article" year="1925">Hooley 1925</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F749D9C9F2169FE5D" author="Norman" box="[729,887,476,501]" pageId="3" pageNumber="404" refString="Norman, D. B. (1986) On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique: Sciences de la Terre, 56, 281 - 372." type="journal article" year="1986">Norman 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F75C09C9E21E7FE5D" author="Paul" box="[900,1017,476,501]" pageId="3" pageNumber="404" refString="Paul, G. S. (2006) Turning the old into the new: a separate genus for the gracile iguanodont from the Wealden of England. In: Carpenter, K. (Ed.), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Indiana University Press, Bloomington, pp. 69 - 77." type="book chapter" year="2006">Paul 2006</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="875A79AC5931DA5F72419C9E265EFE5C" author="Paul" box="[1029,1088,476,500]" pageId="3" pageNumber="404" refString="Paul, G. S. (2008) A revised taxonomy of the iguanodont dinosaur genera and species. Cretaceous Research, 29, 192 - 216. http: // dx. doi. org / 10.1016 / j. cretres. 2007.04.009" type="journal article" year="2008">2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F72099C9E2716FE5C" author="McDonald" box="[1101,1288,476,501]" pageId="3" pageNumber="404" refString="McDonald, A. T. (2012) The status of Dollodon and other basal iguanodonts (Dinosauria: Ornithischia) from the Lower Cretaceous of Europe. Cretaceous Research, 33, 1 - 6. http: // dx. doi. org / 10.1016 / j. cretres. 2011.03.002" type="journal article" year="2012">
|
||
<collectingCountry id="9BDC44CD5931DA5F72099C9E26DBFE5D" box="[1101,1221,476,501]" name="Heard Island and McDonald Islands" pageId="3" pageNumber="404">McDonald</collectingCountry>
|
||
2012
|
||
</bibRefCitation>
|
||
),
|
||
<emphasis id="D1BFD84F5931DA5F73589C9E2785FE5D" box="[1308,1435,476,501]" italics="true" pageId="3" pageNumber="404">Mochlodon</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F76DB9CBD234AFDB0" author="Osi" box="[159,340,511,536]" pageId="3" pageNumber="404" refString="Osi, A., Prondvai, E., Butler, R. J. & Weishampel, D. B. (2012) Phylogeny, histology and inferred body size evolution in a new rhabdodontid dinosaur from the Late Cretaceous of Hungary. PLoS ONE, 7 (9), e 44318. http: // dx. doi. org / 10.1371 / journal. pone. 0044318" type="journal article" year="2012">
|
||
Ösi
|
||
<emphasis id="D1BFD84F5931DA5F76969F432310FDB0" box="[210,270,511,536]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="24CB7FDE5931DA5F772F9F4220A5FDB0" authority="Taquet 1976" authorityName="Taquet" authorityYear="1976" box="[363,699,511,536]" class="Reptilia" family="Iguanodontidae" genus="Ouranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F772F9F422013FDBF" box="[363,525,512,535]" italics="true" pageId="3" pageNumber="404">Ouranosaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F745B9F4220ADFDB0" author="Taquet" box="[543,691,511,536]" pageId="3" pageNumber="404" refString="Taquet, P. (1976) Geologie et paleontologie de gisement de Gadoufaoua (Aptien du Niger). Cahiers de Paleontologie, C. N. R. S, 1 - 191." type="book chapter" year="1976">Taquet 1976</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5931DA5F748E9CBD2625FDB0" authority="Novas et al. 2004" authorityName="Novas et al." authorityYear="2004" box="[714,1083,511,536]" class="Reptilia" family="Hadrosauridae" genus="Talenkauen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F748E9CBD2154FDB0" box="[714,842,511,536]" italics="true" pageId="3" pageNumber="404">Talenkauen</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F75199F42262DFDB0" author="Novas" box="[861,1075,511,536]" pageId="3" pageNumber="404" refString="Novas, F. E., Cambiaso, A. V. & Ambrosio, A. (2004) A new basal iguanodontian (Dinosauria, Ornithischia) from the Upper Cretaceous of Patagonia. Ameghiniana, 41, 75 - 82." type="journal article" year="2004">
|
||
Novas
|
||
<emphasis id="D1BFD84F5931DA5F75F59F4321F3FDB0" box="[945,1005,511,536]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5931DA5F720E9F4326F0FDB0" box="[1098,1262,513,536]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F720E9F4326F0FDB0" box="[1098,1262,513,536]" italics="true" pageId="3" pageNumber="404">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F73449F422789FDB0" author="Forster" box="[1280,1431,512,536]" pageId="3" pageNumber="404" refString="Forster, C. A. (1990) The postcranial skeleton of the ornithopod dinosaur Tenontosaurus tilletti. Journal of Vertebrate Paleontology, 10, 273 - 294." type="journal article" year="1990">Forster 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F76D39F66236FFD94" author="Winkler" box="[151,369,548,573]" pageId="3" pageNumber="404" refString="Winkler, D. A., Murry, P. A. & Jacobs, L. L. (1997) A new species of Tenontosaurus (Dinosauria: Ornithopoda) from the Early Cretaceous of Texas. Journal of Vertebrate Paleontology, 17, 330 - 348. http: // dx. doi. org / 10.1080 / 02724634.1997.10010978" type="journal article" year="1997">
|
||
Winkler
|
||
<emphasis id="D1BFD84F5931DA5F76BC9F67232EFD95" box="[248,304,548,573]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
1997
|
||
</bibRefCitation>
|
||
),
|
||
<taxonomicName id="24CB7FDE5931DA5F77C09F662010FD95" box="[388,526,548,573]" class="Reptilia" family="Dryosauridae" genus="Valdosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F77C09F662010FD95" box="[388,526,548,573]" italics="true" pageId="3" pageNumber="404">Valdosaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F74599F662105FD94" author="Galton" box="[541,795,548,573]" pageId="3" pageNumber="404" refString="Galton, P. M. & Taquet, F. (1982) Valdosaurus, a hypsilophodontid dinosaur from the Lower Cretaceous of Europe and Africa. Geobios, 15, 147 - 158." type="journal article" year="1982">Galton & Taquet 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F75629F6621ADFD94" author="Galton" box="[806,947,548,573]" pageId="3" pageNumber="404" refString="Galton, P. M. (2009) Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England - Hypsilophodon, Valdosaurus, " Camptosaurus, Iguanodon - and referred specimens from Romania and elsewhere. Revue de Paleobiologie, Geneve, 28, 211 - 273." type="journal article" year="2009">Galton 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F75FA9F672695FD94" author="Barrett" box="[958,1163,548,573]" pageId="3" pageNumber="404" refString="Barrett, P. M., Butler, R. J., Twitchett, R. J. & Hutt, S. (2011) New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of Southern England. Special Papers in Palaeontology, 86, 131 - 163." type="journal article" year="2011">
|
||
Barrett
|
||
<emphasis id="D1BFD84F5931DA5F72579F672654FD95" box="[1043,1098,548,573]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
), and
|
||
<taxonomicName id="24CB7FDE5931DA5F72899F662727FD95" box="[1229,1337,548,573]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F72899F662727FD95" box="[1229,1337,548,573]" italics="true" pageId="3" pageNumber="404">Zalmoxes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5931DA5F73029F6622CDFDC8" author="Nopcsa" pageId="3" pageNumber="404" refString="Nopcsa, F. (1900) Dinosaurierreste aus Siebenburgen. Schadel von Limnosaurus transsylvanicus nov. gen. et spec. Denkschriften der kaiserlichen Akademie der Wissenchaften Wien, Mathematisch Naturwissenschaftliche Classe, 68, 555 - 591." type="journal article" year="1900">Nopcsa 1900</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5931DA5F769A9F0523F5FDC8" author="Weishampel" box="[222,491,583,608]" pageId="3" pageNumber="404" refString="Weishampel, D. B., Jianu, C. M., Csiki, Z. & Norman, D. B. (2003) Osteology and phylogeny of Zalmoxes (n. g.), an unusual euornithopod dinosaur from the latest Cretaceous of Romania. Journal of Systematic Palaeontology, 1, 65 - 123. http: // dx. doi. org / 10.1017 / s 1477201903001032" type="journal article" year="2003">
|
||
Weishampel
|
||
<emphasis id="D1BFD84F5931DA5F772B9F0B23B9FDC8" box="[367,423,583,608]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
). Additionally, other specimens (e.g., the Australian ornithopods) as well as isolated and partially persevered materials are also used in the present comparison (see below).
|
||
</paragraph>
|
||
<caption id="B7B454D55931DA5F76D39B7123B4F92E" httpUri="https://zenodo.org/record/226296/files/figure.png" pageId="3" pageNumber="404" targetBox="[340,1247,697,1565]" targetPageId="3">
|
||
<paragraph id="E374045D5931DA5F76D39B7123B4F92E" blockId="3.[151,1435,1587,1670]" pageId="3" pageNumber="404">
|
||
<emphasis id="D1BFD84F5931DA5F76D39B71230DF9E0" bold="true" box="[151,275,1587,1609]" pageId="3" pageNumber="404">FIGURE 3.</emphasis>
|
||
<taxonomicName id="24CB7FDE5931DA5F775D9B71206FF9E1" box="[281,625,1587,1609]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="comodorensis">
|
||
<emphasis id="D1BFD84F5931DA5F775D9B71206FF9E1" box="[281,625,1587,1609]" italics="true" pageId="3" pageNumber="404">Notohypsilophodon comodorensis</emphasis>
|
||
</taxonomicName>
|
||
. Anterior dorsal vertebrae (UNPSJB-PV 942/5,6,7 respectively), in right lateral (A and B) and ventral (C) views. Abbreviations: dp, diapophysis; ns, neural spine; pa, parapophysis; prz, prezygapophysis; rg, rugosities; vk, ventral keel.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E374045D5931DA5876839BF1212AFD0C" blockId="3.[151,1436,1715,2026]" lastBlockId="4.[151,1436,151,2035]" lastPageId="4" lastPageNumber="405" pageId="3" pageNumber="404">
|
||
<emphasis id="D1BFD84F5931DA5F76839BF123AAF964" bold="true" box="[199,436,1715,1740]" pageId="3" pageNumber="404">Cervical vertebrae.</emphasis>
|
||
The cervical vertebrae (
|
||
<figureCitation id="7BF018D85931DA5F74A69BF12132F963" box="[738,812,1715,1740]" captionStart="FIGURE 2" captionStartId="2.[151,250,1695,1717]" captionTargetBox="[151,1436,553,1673]" captionTargetId="figure@2.[151,1436,553,1674]" captionTargetPageId="2" captionText="FIGURE 2. Notohypsilophodon comodorensis. Middle cervical vertebra (UNPSJB-PV 942 / 1), in anterior (A), in right lateral (B), in dorsal (C) and ventral (D) views. Abbreviations: nc, neural canal; ncs, neural central suture; ns, neural spine; pa, parapophysis; poz, postzygapophysis; prz, prezygapophysis; vk, ventral keel." httpUri="https://zenodo.org/record/226295/files/figure.png" pageId="3" pageNumber="404">Fig. 2</figureCitation>
|
||
) of
|
||
<taxonomicName id="24CB7FDE5931DA5F752C9BF126F2F964" box="[872,1260,1715,1740]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="comodorensis">
|
||
<emphasis id="D1BFD84F5931DA5F752C9BF126F2F964" box="[872,1260,1715,1740]" italics="true" pageId="3" pageNumber="404">Notohypsilophodon comodorensis</emphasis>
|
||
</taxonomicName>
|
||
, based on the morphology of the centra and the portions of the neural arches preserved are presumably middle cervicals. The centra are amphyplatian as in
|
||
<emphasis id="D1BFD84F5931DA5F74499BBE2115F8BB" box="[525,779,1786,1811]" italics="true" pageId="3" pageNumber="404">
|
||
Haya,
|
||
<taxonomicName id="24CB7FDE5931DA5F74249BB82115F8BB" box="[608,779,1786,1811]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">Hypsilophodon</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="24CB7FDE5931DA5F75169BB821F9F8BB" box="[850,999,1786,1811]" class="Reptilia" family="Thescelosauridae" genus="Jeholosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F75169BB821F9F8BB" box="[850,999,1786,1811]" italics="true" pageId="3" pageNumber="404">Jeholosaurus</emphasis>
|
||
</taxonomicName>
|
||
. In more derived ornithopods (e.g.,
|
||
<taxonomicName id="24CB7FDE5931DA5F76D39A5D23A7F89E" authority="Cumnoria" authorityName="Cumnoria" box="[151,441,1823,1846]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F76D39A5D2324F89E" box="[151,314,1823,1846]" italics="true" pageId="3" pageNumber="404">Camptosaurus</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5931DA5F770C9A5D23A7F89E" box="[328,441,1823,1846]" italics="true" pageId="3" pageNumber="404">Cumnoria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5931DA5F77829A5D2054F89E" box="[454,586,1823,1846]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F77829A5D2054F89E" box="[454,586,1823,1846]" italics="true" pageId="3" pageNumber="404">Dryosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5931DA5F741C9A5D20E2F89E" box="[600,764,1823,1846]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F741C9A5D20E2F89E" box="[600,764,1823,1846]" italics="true" pageId="3" pageNumber="404">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5931DA5F754E9A5C218BF89F" box="[778,917,1822,1847]" class="Reptilia" family="Dryosauridae" genus="Valdosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F754E9A5C218BF89F" box="[778,917,1822,1847]" italics="true" pageId="3" pageNumber="404">Valdosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE5931DA5F758D9A5C262BF89F" box="[969,1077,1822,1847]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F758D9A5C262BF89F" box="[969,1077,1822,1847]" italics="true" pageId="3" pageNumber="404">Zalmoxes</emphasis>
|
||
</taxonomicName>
|
||
), on the other hand, the cervical centra are slightly opisthocoelous or amphicoelous, as in
|
||
<emphasis id="D1BFD84F5931DA5F756E9A002610F8F3" box="[810,1038,1858,1883]" italics="true" pageId="3" pageNumber="404">Macrogryphosaurus</emphasis>
|
||
. The centra of
|
||
<taxonomicName id="24CB7FDE5931DA5F72859A002782F8F3" box="[1217,1436,1858,1883]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F72859A002782F8F3" box="[1217,1436,1858,1883]" italics="true" pageId="3" pageNumber="404">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
are slightly wider than tall, as is the case in most basal neornithischians and basal ornithopods (e.g.,
|
||
<taxonomicName id="24CB7FDE5931DA5F76D39ACB237DF80A" box="[151,355,1929,1954]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F76D39ACB237DF80A" box="[151,355,1929,1954]" italics="true" pageId="3" pageNumber="404">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5931DA5F77319ACB20AAF809" authority="Yueosaurus" authorityName="Yueosaurus" box="[373,692,1929,1954]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F77319ACB203EF80A" box="[373,544,1929,1954]" italics="true" pageId="3" pageNumber="404">Hypsilophodon</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5931DA5F74709AC820AAF809" box="[564,692,1930,1953]" italics="true" pageId="3" pageNumber="404">Yueosaurus</emphasis>
|
||
</taxonomicName>
|
||
). In ornithopods more derived than
|
||
<taxonomicName id="24CB7FDE5931DA5F72239AC82715F809" box="[1127,1291,1930,1953]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F72239AC82715F809" box="[1127,1291,1930,1953]" italics="true" pageId="3" pageNumber="404">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="875A79AC5931DA5F73149ACB22CAF86E" author="Galton" pageId="3" pageNumber="404" refString="Galton, P. M. (2009) Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England - Hypsilophodon, Valdosaurus, " Camptosaurus, Iguanodon - and referred specimens from Romania and elsewhere. Revue de Paleobiologie, Geneve, 28, 211 - 273." type="journal article" year="2009">Galton 2009</bibRefCitation>
|
||
: fig. 20),
|
||
<emphasis id="D1BFD84F5931DA5F77019AEF2037F86E" box="[325,553,1965,1990]" italics="true" pageId="3" pageNumber="404">Macrogryphosaurus</emphasis>
|
||
and
|
||
<taxonomicName id="24CB7FDE5931DA5F74209AEF20FAF86E" box="[612,740,1965,1990]" class="Reptilia" family="Hadrosauridae" genus="Talenkauen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="3" pageNumber="404" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5931DA5F74209AEF20FAF86E" box="[612,740,1965,1990]" italics="true" pageId="3" pageNumber="404">Talenkauen</emphasis>
|
||
</taxonomicName>
|
||
, the cervical centra are anteroposteriorly longer than tall. In lateral view, the centra are “divided” in two slightly concave surfaces as in basal ornithopods due to the presence of the longitudinal ridge at approximately centrum midheight. Ventrally, the cervical vertebrae of
|
||
<taxonomicName id="24CB7FDE5936DA5872859DD52782FF18" box="[1217,1436,151,176]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5872859DD52782FF18" box="[1217,1436,151,176]" italics="true" pageId="4" pageNumber="405">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
exhibit a thin midline keel similar to those seen in
|
||
<taxonomicName id="24CB7FDE5936DA58748F9DF9215DFF7C" box="[715,835,187,212]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA58748F9DF9215DFF7C" box="[715,835,187,212]" italics="true" pageId="4" pageNumber="405">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5875159DF92603FF7C" box="[849,1053,187,212]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5875159DF92603FF7C" box="[849,1053,187,212]" italics="true" pageId="4" pageNumber="405">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA58726F9DFE26C2FF7B" box="[1067,1244,188,211]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA58726F9DFE26C2FF7B" box="[1067,1244,188,211]" italics="true" pageId="4" pageNumber="405">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5872AE9DF92365FF50" authority="Macrogryphosaurus" authorityName="Macrogryphosaurus" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5872AE9DF9278BFF7C" box="[1258,1429,187,212]" italics="true" pageId="4" pageNumber="405">Hypsilophodon</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5936DA5876D39D9D2365FF50" box="[151,379,223,248]" italics="true" pageId="4" pageNumber="405">Macrogryphosaurus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="24CB7FDE5936DA5877FE9D9D205AFF50" box="[442,580,223,248]" class="Reptilia" family="Dryosauridae" genus="Valdosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5877FE9D9D205AFF50" box="[442,580,223,248]" italics="true" pageId="4" pageNumber="405">Valdosaurus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="875A79AC5936DA5874C69DA2214AFF50" author="Barrett" box="[642,852,223,248]" pageId="4" pageNumber="405" refString="Barrett, P. M., Butler, R. J., Twitchett, R. J. & Hutt, S. (2011) New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of Southern England. Special Papers in Palaeontology, 86, 131 - 163." type="journal article" year="2011">
|
||
Barrett
|
||
<emphasis id="D1BFD84F5936DA58749C9DA22114FF50" box="[728,778,223,248]" italics="true" pageId="4" pageNumber="405">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
: fig. 2) and unlike the thicker, rugose ventral keels, present in some basal iguanodontids (e.g.,
|
||
<taxonomicName id="24CB7FDE5936DA5874EB9C4621C1FEB3" authority="Cumnoria" authorityName="Cumnoria" box="[687,991,260,283]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5874EB9C46214CFEB3" box="[687,850,260,283]" italics="true" pageId="4" pageNumber="405">Camptosaurus</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5936DA58752F9C4621C1FEB3" box="[875,991,260,283]" italics="true" pageId="4" pageNumber="405">Cumnoria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5875B09C46270EFEB4" authority="Uteodon" authorityName="Uteodon" box="[1012,1296,259,284]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
Tenontosaurus,
|
||
<emphasis id="D1BFD84F5936DA5872F49C41270EFEB4" box="[1200,1296,259,284]" italics="true" pageId="4" pageNumber="405">Uteodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA58736D9C4123D2FE97" authority="Carpenter & Wilson 2008" authorityName="Carpenter & Wilson" authorityYear="2008" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA58736D9C41278BFEB4" box="[1321,1429,259,284]" italics="true" pageId="4" pageNumber="405">Zalmoxes</emphasis>
|
||
:
|
||
<bibRefCitation id="875A79AC5936DA5876D39C6523D2FE97" author="Carpenter" box="[151,460,294,319]" pageId="4" pageNumber="405" refString="Carpenter, K. & Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum, 76, 227 - 263. http: // dx. doi. org / 10.2992 / 0097 - 4463 (2008) 76 [227: ansoco] 2.0. co; 2" type="journal article" year="2008">Carpenter & Wilson 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
; Norman 2004). In anterior view, the cervical vertebrae have pentagonal articular surfaces with an acute ventral apex as the rhabdodontid ornithopod
|
||
<taxonomicName id="24CB7FDE5936DA5875CE9C0821E8FECB" box="[906,1014,330,355]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5875CE9C0821E8FECB" box="[906,1014,330,355]" italics="true" pageId="4" pageNumber="405">Zalmoxes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5936DA5872479C08270EFECB" author="Weishampel" box="[1027,1296,330,355]" pageId="4" pageNumber="405" refString="Weishampel, D. B., Jianu, C. M., Csiki, Z. & Norman, D. B. (2003) Osteology and phylogeny of Zalmoxes (n. g.), an unusual euornithopod dinosaur from the latest Cretaceous of Romania. Journal of Systematic Palaeontology, 1, 65 - 123. http: // dx. doi. org / 10.1017 / s 1477201903001032" type="journal article" year="2003">
|
||
Weishampel
|
||
<emphasis id="D1BFD84F5936DA5872D19C0E26D3FECB" box="[1173,1229,330,355]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
: fig. 15) and the basal ornithopod
|
||
<emphasis id="D1BFD84F5936DA5877D69C2D2011FE2E" box="[402,527,367,390]" italics="true" pageId="4" pageNumber="405">Yueosaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5936DA5874619C2C211FFE2F" author="Zheng" box="[549,769,366,391]" pageId="4" pageNumber="405" refString="Zheng, W. J., Jin, X. S., Shibata, M., Azuma, Y. & Yu, F. M. (2012) A new ornithischian dinosaur from the Cretaceous Liangtoutang Formation of Tiantai, Zhejiang Province, China. Cretaceous Research, 34, 208 - 219. http: // dx. doi. org / 10.1016 / j. cretres. 2011.11.001" type="journal article" year="2012">
|
||
Zheng
|
||
<emphasis id="D1BFD84F5936DA58743F9C2D20A6FE2F" box="[635,696,366,391]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
: fig. 3); however, in the latter, the ventral apex is more pronounced than in
|
||
<taxonomicName id="24CB7FDE5936DA5877309CD0204EFE03" box="[372,592,402,427]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5877309CD0204EFE03" box="[372,592,402,427]" italics="true" pageId="4" pageNumber="405">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE5936DA5874C19CD020EEFE03" box="[645,752,402,427]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5874C19CD020EEFE03" box="[645,752,402,427]" italics="true" pageId="4" pageNumber="405">Zalmoxes</emphasis>
|
||
</taxonomicName>
|
||
. The neural canal is proportionally large and ovoid in outline (a feature which could be related to the juvenile status of
|
||
<emphasis id="D1BFD84F5936DA5875689CF72610FE66" box="[812,1038,437,462]" italics="true" pageId="4" pageNumber="405">
|
||
<taxonomicName id="24CB7FDE5936DA5875689CF72614FE66" box="[812,1034,437,462]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">Notohypsilophodon</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
see below). The articular facets of the prezygapophyses are subcircular in outline and project strongly anterodorsally as in
|
||
<taxonomicName id="24CB7FDE5936DA58723E9C9B273BFE5A" box="[1146,1317,473,498]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA58723E9C9B273BFE5A" box="[1146,1317,473,498]" italics="true" pageId="4" pageNumber="405">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
. Based on the preserved portion of the cervical neural spines, they appear to be notably reduced, a condition also reported in the basal ornithopods
|
||
<taxonomicName id="24CB7FDE5936DA5877C99F632026FD92" box="[397,568,545,570]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5877C99F632026FD92" box="[397,568,545,570]" italics="true" pageId="4" pageNumber="405">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5874029F6020E9FD91" box="[582,759,546,569]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5874029F6020E9FD91" box="[582,759,546,569]" italics="true" pageId="4" pageNumber="405">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
and the basal iguanodontian
|
||
<taxonomicName id="24CB7FDE5936DA58727A9F6326A0FD92" box="[1086,1214,545,570]" class="Reptilia" family="Hadrosauridae" genus="Talenkauen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA58727A9F6326A0FD92" box="[1086,1214,545,570]" italics="true" pageId="4" pageNumber="405">Talenkauen</emphasis>
|
||
</taxonomicName>
|
||
. In lateral view, the centra exhibit an open neurocentral suture, indicating that
|
||
<taxonomicName id="24CB7FDE5936DA58750C9F06263AFDF5" box="[840,1060,580,605]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA58750C9F06263AFDF5" box="[840,1060,580,605]" italics="true" pageId="4" pageNumber="405">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
was not somatically mature (as
|
||
<bibRefCitation id="875A79AC5936DA5876D39F2A235EFD29" author="Martinez" box="[151,320,616,641]" pageId="4" pageNumber="405" refString="Martinez, R. D. (1998) Notohypsilophodon comodorensis gen. et sp. nov. Un Hypsilophodontidae (Ornithischia: Ornithopoda) del Cretacico Superior de Chubut, Patagonia central, Argentina. Acta Geologica Leopoldensia, 21, 119 - 135." type="journal article" year="1998">Martinez 1998</bibRefCitation>
|
||
, pointed out in his original study, see below). A parapophysis located just below the neurocentral suture is also present in
|
||
<taxonomicName id="24CB7FDE5936DA5877E09FCF2056FD0C" box="[420,584,653,676]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5877E09FCF2056FD0C" box="[420,584,653,676]" italics="true" pageId="4" pageNumber="405">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE5936DA58743B9FCE2133FD0D" box="[639,813,652,677]" class="Reptilia" family="Hypsilophodontidae" genus="Thescelosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA58743B9FCE2133FD0D" box="[639,813,652,677]" italics="true" pageId="4" pageNumber="405">Thescelosaurus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E374045D5936DA5876839FED22FEFAE3" blockId="4.[151,1436,151,2035]" pageId="4" pageNumber="405">
|
||
<emphasis id="D1BFD84F5936DA5876839FED2385FD60" bold="true" box="[199,411,687,712]" pageId="4" pageNumber="405">Dorsal vertebrae.</emphasis>
|
||
The dorsal vertebrae (
|
||
<figureCitation id="7BF018D85936DA5874E09FED20F4FD60" box="[676,746,687,712]" captionStart="FIGURE 3" captionStartId="3.[151,250,1587,1609]" captionTargetBox="[340,1247,697,1565]" captionTargetId="figure@3.[340,1247,697,1565]" captionTargetPageId="3" captionText="FIGURE 3. Notohypsilophodon comodorensis. Anterior dorsal vertebrae (UNPSJB-PV 942 / 5,6,7 respectively), in right lateral (A and B) and ventral (C) views. Abbreviations: dp, diapophysis; ns, neural spine; pa, parapophysis; prz, prezygapophysis; rg, rugosities; vk, ventral keel." httpUri="https://zenodo.org/record/226296/files/figure.png" pageId="4" pageNumber="405">Fig. 3</figureCitation>
|
||
) are slightly amphicoelous and hourglass-shaped in ventral view as a consequence of their lateral compression. There are strongly-developed longitudinal striations around the margins of each of the articular surfaces, probably related to the insertion of soft tissues (i.e., the dorsal hypaxial musculature:
|
||
<bibRefCitation id="875A79AC5936DA58776A9E5923EBFC9B" author="Butler" box="[302,501,795,820]" pageId="4" pageNumber="405" refString="Butler, R. J., Galton, P. M., Porro, L. B., Chiappe, L. M., Henderson, D. M. & Erickson, M. (2010) Lower limits of ornithischian dinosaur body size inferred from a new Upper Jurassic heterodontosaurid from North America. Proceedings of the Royal Society, 277, 375 - 381. http: // dx. doi. org / 10.1098 / rspb. 2009.1494" type="journal article" year="2010">
|
||
Butler
|
||
<emphasis id="D1BFD84F5936DA58773F9E5E23ACFC9C" box="[379,434,795,820]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
; Butler
|
||
<emphasis id="D1BFD84F5936DA5874089E5E209AFC9C" box="[588,644,795,820]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
|
||
2011;
|
||
<bibRefCitation id="875A79AC5936DA5874949E592187FC9B" author="Zheng" box="[720,921,795,820]" pageId="4" pageNumber="405" refString="Zheng, W. J., Jin, X. S., Shibata, M., Azuma, Y. & Yu, F. M. (2012) A new ornithischian dinosaur from the Cretaceous Liangtoutang Formation of Tiantai, Zhejiang Province, China. Cretaceous Research, 34, 208 - 219. http: // dx. doi. org / 10.1016 / j. cretres. 2011.11.001" type="journal article" year="2012">
|
||
Zheng
|
||
<emphasis id="D1BFD84F5936DA58755A9E5E2148FC9C" box="[798,854,795,820]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
). These longitudinal striations are comparable to those observed in many ornithopods, such as
|
||
<taxonomicName id="24CB7FDE5936DA5874FD9E7D2142FCFE" box="[697,860,831,854]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5874FD9E7D2142FCFE" box="[697,860,831,854]" italics="true" pageId="4" pageNumber="405">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA58752F9E0226FCFCFF" authority="Macrogryphosaurus" authorityName="Macrogryphosaurus" box="[875,1250,830,855]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA58752F9E0221F1FCFF" box="[875,1007,832,855]" italics="true" pageId="4" pageNumber="405">Dryosaurus</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5936DA5875BA9E7C26FCFCFF" box="[1022,1250,830,855]" italics="true" pageId="4" pageNumber="405">Macrogryphosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5872B59E02278BFCFF" box="[1265,1429,832,855]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5872B59E02278BFCFF" box="[1265,1429,832,855]" italics="true" pageId="4" pageNumber="405">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5876D39E20235BFCD3" box="[151,325,866,891]" class="Reptilia" family="Hypsilophodontidae" genus="Thescelosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5876D39E20235BFCD3" box="[151,325,866,891]" italics="true" pageId="4" pageNumber="405">Thescelosaurus</emphasis>
|
||
</taxonomicName>
|
||
and a set of dorsal vertebrae recovered in Cretaceous outcrops of the Bajo Barreal Formation (Cenomanian-Turonian; UNPSJB-PV 960 see
|
||
<bibRefCitation id="875A79AC5936DA5874E59EC4216EFC37" author="Ibiricu" box="[673,880,902,927]" pageId="4" pageNumber="405" refString="Ibiricu, L. M., Martinez, R. D., Lamanna, M. C., Casal, G. A., Luna, M., Harris, J. D. & Lacovara, K. J. (2010) A medium-sized ornithopod (Dinosauria: Ornithischia) from the Upper Cretaceous Bajo Barreal Formation of Lago Colhue Huapi, southern Chubut Province, Argentina. Annals of the Carnegie Museum, 79, 39 - 50. http: // dx. doi. org / 10.2992 / 007.079.0103" type="journal article" year="2010">
|
||
Ibiricu
|
||
<emphasis id="D1BFD84F5936DA5874B79EC52132FC37" box="[755,812,902,927]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). Ventrally, the centra of
|
||
<taxonomicName id="24CB7FDE5936DA5872CD9EC4277BFC37" box="[1161,1381,902,927]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5872CD9EC4277BFC37" box="[1161,1381,902,927]" italics="true" pageId="4" pageNumber="405">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
bear a midline keel as in the cervical vertebrae; however, this structure is not as sharp or as well defined as in the cervicals. A similar ventral keel is also seen in
|
||
<taxonomicName id="24CB7FDE5936DA5874839E8F218DFC4E" box="[711,915,973,998]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5874839E8F218DFC4E" box="[711,915,973,998]" italics="true" pageId="4" pageNumber="405">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5875E09E8F278CFC4E" authority="Trinisaura and Yueosaurus" authorityName="Trinisaura and Yueosaurus" box="[932,1426,973,998]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5875E09E8F2651FC4E" box="[932,1103,973,998]" italics="true" pageId="4" pageNumber="405">Hypsilophodon</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5936DA5872259E8D26C8FC4E" box="[1121,1238,975,998]" italics="true" pageId="4" pageNumber="405">Trinisaura</emphasis>
|
||
and
|
||
<emphasis id="D1BFD84F5936DA5873519E8D278CFC4E" box="[1301,1426,975,998]" italics="true" pageId="4" pageNumber="405">Yueosaurus</emphasis>
|
||
</taxonomicName>
|
||
, whereas non-hadrosaurid iguanodontians (e.g.,
|
||
<emphasis id="D1BFD84F5936DA5874F19EB02139FBA1" box="[693,807,1010,1033]" italics="true" pageId="4" pageNumber="405">Cumnoria</emphasis>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5875739EB321AEFBA2" box="[823,944,1009,1034]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5875739EB321AEFBA2" box="[823,944,1009,1034]" italics="true" pageId="4" pageNumber="405">Iguanodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5875869EB32675FBA2" box="[962,1131,1009,1034]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5875869EB32675FBA2" box="[962,1131,1009,1034]" italics="true" pageId="4" pageNumber="405">Mantellisaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5872389EB0273EFBA1" box="[1148,1312,1010,1033]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5872389EB0273EFBA1" box="[1148,1312,1010,1033]" italics="true" pageId="4" pageNumber="405">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5873759EB32782FBA2" box="[1329,1436,1009,1034]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5873759EB32782FBA2" box="[1329,1436,1009,1034]" italics="true" pageId="4" pageNumber="405">Zalmoxes</emphasis>
|
||
</taxonomicName>
|
||
and UNPSJB-PV 960) have centrum ventral surfaces that lack keels and which are more concave and rounded. The posterior surfaces of the centra are suboval in outline, a feature present in basal neornithischians (e.g.,
|
||
<emphasis id="D1BFD84F5936DA5876D3991E2324FBDD" box="[151,314,1116,1141]" italics="true" pageId="4" pageNumber="405">Hexinlusaurus</emphasis>
|
||
and
|
||
<taxonomicName id="24CB7FDE5936DA587730991E201AFBDD" box="[372,516,1116,1141]" class="Reptilia" family="Hypsilophodontidae" genus="Yandusaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA587730991E201AFBDD" box="[372,516,1116,1141]" italics="true" pageId="4" pageNumber="405">Yandusaurus</emphasis>
|
||
</taxonomicName>
|
||
) and basal iguanodontians (e.g.,
|
||
<taxonomicName id="24CB7FDE5936DA587538991F2601FBDC" box="[892,1055,1117,1140]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA587538991F2601FBDC" box="[892,1055,1117,1140]" italics="true" pageId="4" pageNumber="405">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA58726B991E26B6FBDD" box="[1071,1192,1116,1141]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA58726B991E26B6FBDD" box="[1071,1192,1116,1141]" italics="true" pageId="4" pageNumber="405">Iguanodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE5936DA5872A6991E2792FBDD" box="[1250,1420,1116,1141]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5872A6991E2792FBDD" box="[1250,1420,1116,1141]" italics="true" pageId="4" pageNumber="405">Mantellisaurus</emphasis>
|
||
</taxonomicName>
|
||
). The transverse processes preserved in
|
||
<taxonomicName id="24CB7FDE5936DA58740D99C2213AFB31" box="[585,804,1152,1177]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA58740D99C2213AFB31" box="[585,804,1152,1177]" italics="true" pageId="4" pageNumber="405">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
project laterally, at approximately the same level as the zygapophyses, as is characteristic of basal ornithopods (Norman 2004;
|
||
<bibRefCitation id="875A79AC5936DA58759299E626FDFB15" author="Makovicky" box="[982,1251,1188,1213]" pageId="4" pageNumber="405" refString="Makovicky, P. J., Kilbourne, B. M., Sadleir, R. W. & Norell, M. A. (2011) A new basal ornithopod (Dinosauria, Ornithischia) from the Late Cretaceous of Mongolia. Journal of Vertebrate Paleontology, 31, 626 - 640. http: // dx. doi. org / 10.1080 / 02724634.2011.557114" type="journal article" year="2011">
|
||
Makovicky
|
||
<emphasis id="D1BFD84F5936DA58722499E72682FB15" box="[1120,1180,1188,1213]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
). The preserved dorsal neural spine appears to be relatively low and anteroposteriorly narrow. This feature is comparable to the anatomy of basal ornithopods and differs from that seen in the more derived members of the group (i.e., nonhadrosaurid iguanodontians), where the dorsal neural spines are elevated and rectangular in shape (Norman
|
||
<emphasis id="D1BFD84F5936DA5873279852278BFA80" box="[1379,1429,1295,1320]" italics="true" pageId="4" pageNumber="405">et al</emphasis>
|
||
. 2004).
|
||
</paragraph>
|
||
<paragraph id="E374045D5936DA58768398142319F919" blockId="4.[151,1436,151,2035]" pageId="4" pageNumber="405">
|
||
<emphasis id="D1BFD84F5936DA58768398142386FAC7" bold="true" box="[199,408,1366,1391]" pageId="4" pageNumber="405">Sacral vertebrae.</emphasis>
|
||
Due to the disarticulated nature of the sacral vertebrae, it is difficult to determine with confidence the positions of two of the sacral vertebrae recovered. Nevertheless, the sacral vertebra possessing a posterior articular surface that is laterally wider than its anterior counterpart is considered the dorsosacral (
|
||
<figureCitation id="7BF018D85936DA58731598DC2789FA1F" box="[1361,1431,1438,1463]" captionStart="FIGURE 4" captionStartId="5.[151,250,1786,1808]" captionTargetBox="[151,1436,193,1764]" captionTargetId="figure@5.[151,1436,193,1765]" captionTargetPageId="5" captionText="FIGURE 4. Notohypsilophodon comodorensis. Sacral vertebrae (UNPSJB-PV 942 / 12,13,14,15,16 respectively), in left lateral (A and B), in dorsal (C and D) and ventral (E and F) views. Abbreviations: ns, neural spine; rg, rugosities; tp, transverse process." httpUri="https://zenodo.org/record/226297/files/figure.png" pageId="4" pageNumber="405">Fig. 4</figureCitation>
|
||
, also see Butler
|
||
<emphasis id="D1BFD84F5936DA58770898812396FA73" box="[332,392,1474,1499]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
|
||
2011). Unfortunately, it is not possible to determine the full sacral number (including the sacrodorsal vertebra) due to the absence of complete articulation. However, based on the preserved elements, we considered a putative number of five or six, as also known in many other ornithischians (e.g.,
|
||
<taxonomicName id="24CB7FDE5936DA5873559B48278BF989" box="[1297,1429,1546,1569]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5873559B48278BF989" box="[1297,1429,1546,1569]" italics="true" pageId="4" pageNumber="405">Dryosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5876D39B6C204EF9EE" authority="Haya, Hexinlusaurus" authorityName="Haya, Hexinlusaurus" box="[151,592,1581,1606]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5876D39B6C2356F9ED" box="[151,328,1582,1605]" italics="true" pageId="4" pageNumber="405">Gasparinisaura</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5936DA5877189B6C2387F9ED" box="[348,409,1582,1605]" italics="true" pageId="4" pageNumber="405">Haya</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5936DA5877E99B6F204EF9EE" box="[429,592,1581,1606]" italics="true" pageId="4" pageNumber="405">Hexinlusaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5936DA5874209B6F2111F9EE" box="[612,783,1581,1606]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5874209B6F2111F9EE" box="[612,783,1581,1606]" italics="true" pageId="4" pageNumber="405">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
: also see
|
||
<bibRefCitation id="875A79AC5936DA5875CB9B6F262CF9EE" author="Galton" box="[911,1074,1581,1606]" pageId="4" pageNumber="405" refString="Galton, P. M. (1974 a) The ornithischian dinosaur Hypsilophodon from the Wealden of the Isle of Wight. Bulletin of the British Museum (Natural History), Geology, 25, 1 - 152." type="journal article" year="1974" yearSuffix="a">Galton 1974a</bibRefCitation>
|
||
). On the other hand, with the exception of
|
||
<taxonomicName id="24CB7FDE5936DA5877769B132119F9C1" authority="Varricchio et al., 2007" authorityName="Varricchio et al." authorityYear="2007" box="[306,775,1617,1642]" class="Reptilia" family="Hypsilophodontidae" genus="Oryctodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5877769B1323C1F9C2" box="[306,479,1617,1642]" italics="true" pageId="4" pageNumber="405">Oryctodromeus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5936DA5877B69B1320E1F9C1" author="Varricchio" box="[498,767,1617,1642]" pageId="4" pageNumber="405" refString="Varricchio, D. J., Martin, A. J. & Katsura, Y. (2007) First trace and body fossil evidence of a burrowing, denning dinosaur. Proceedings of the Royal Society, 274, 1361 - 1368. http: // dx. doi. org / 10.1098 / rspb. 2006.0443" type="journal article" year="2007">
|
||
Varricchio
|
||
<emphasis id="D1BFD84F5936DA5874309B1020AFF9C2" box="[628,689,1617,1642]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
|
||
, 2007
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, some derived ornithopods, such as rhabdodontids (e.g.,
|
||
<taxonomicName id="24CB7FDE5936DA5876D39B36231DF925" box="[151,259,1652,1677]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5876D39B36231DF925" box="[151,259,1652,1677]" italics="true" pageId="4" pageNumber="405">Zalmoxes</emphasis>
|
||
</taxonomicName>
|
||
and probably
|
||
<emphasis id="D1BFD84F5936DA5877E19B362030F925" box="[421,558,1652,1677]" italics="true" pageId="4" pageNumber="405">Mochlodon;</emphasis>
|
||
see
|
||
<bibRefCitation id="875A79AC5936DA5874259B362117F925" author="Osi" box="[609,777,1652,1677]" pageId="4" pageNumber="405" refString="Osi, A., Prondvai, E., Butler, R. J. & Weishampel, D. B. (2012) Phylogeny, histology and inferred body size evolution in a new rhabdodontid dinosaur from the Late Cretaceous of Hungary. PLoS ONE, 7 (9), e 44318. http: // dx. doi. org / 10.1371 / journal. pone. 0044318" type="journal article" year="2012">
|
||
Ösi
|
||
<emphasis id="D1BFD84F5936DA5874CB9B3420D6F925" box="[655,712,1652,1677]" italics="true" pageId="4" pageNumber="405">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
) and derived iguanodontians, possess more than six sacral vertebrae.
|
||
</paragraph>
|
||
<paragraph id="E374045D5936DA5876839BFE26C5F85A" blockId="4.[151,1436,151,2035]" pageId="4" pageNumber="405">
|
||
Of the two sets of sacral vertebrae recovered with
|
||
<taxonomicName id="24CB7FDE5936DA5875409BFE21C1F97D" box="[772,991,1724,1749]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5875409BFE21C1F97D" box="[772,991,1724,1749]" italics="true" pageId="4" pageNumber="405">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
, three vertebrae, probably representing the anterior ones, are co-ossified (
|
||
<figureCitation id="7BF018D85936DA5874529B9D2047F950" box="[534,601,1759,1784]" captionStart="FIGURE 4" captionStartId="5.[151,250,1786,1808]" captionTargetBox="[151,1436,193,1764]" captionTargetId="figure@5.[151,1436,193,1765]" captionTargetPageId="5" captionText="FIGURE 4. Notohypsilophodon comodorensis. Sacral vertebrae (UNPSJB-PV 942 / 12,13,14,15,16 respectively), in left lateral (A and B), in dorsal (C and D) and ventral (E and F) views. Abbreviations: ns, neural spine; rg, rugosities; tp, transverse process." httpUri="https://zenodo.org/record/226297/files/figure.png" pageId="4" pageNumber="405">Fig. 4</figureCitation>
|
||
A–C–E), whereas the other set, of two sacral vertebrae, are articulated but not co-ossified (
|
||
<figureCitation id="7BF018D85936DA58770F9A412392F8B4" box="[331,396,1795,1820]" captionStart="FIGURE 4" captionStartId="5.[151,250,1786,1808]" captionTargetBox="[151,1436,193,1764]" captionTargetId="figure@5.[151,1436,193,1765]" captionTargetPageId="5" captionText="FIGURE 4. Notohypsilophodon comodorensis. Sacral vertebrae (UNPSJB-PV 942 / 12,13,14,15,16 respectively), in left lateral (A and B), in dorsal (C and D) and ventral (E and F) views. Abbreviations: ns, neural spine; rg, rugosities; tp, transverse process." httpUri="https://zenodo.org/record/226297/files/figure.png" pageId="4" pageNumber="405">Fig. 4</figureCitation>
|
||
B–D–F). The sacral rib preserved (which is considered to be the first true sacral rib) is robust. The centra are slightly laterally compressed and are cotton-reel-shaped in ventral view. Circumscribing the articular surfaces, well-developed longitudinal striations are present. These striations continue onto the ventral surfaces of the centra, where they become more pronounced. The preserved sacral neural spine is wider anteroposteriorly than transversely. The distal end of the neural spine is posteriorly extended. In ventral view, the centra are comparable to those in
|
||
<taxonomicName id="24CB7FDE5936DA5874499AF520A0F866" box="[525,702,1975,1998]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5874499AF520A0F866" box="[525,702,1975,1998]" italics="true" pageId="4" pageNumber="405">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
, in possessing slightly concave ventral surfaces, whereas in many other basal ornithopods a longitudinal keel or ridge is present (e.g.,
|
||
<taxonomicName id="24CB7FDE5936DA5875C89A982646F85B" box="[908,1112,2010,2035]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="405" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5936DA5875C89A982646F85B" box="[908,1112,2010,2035]" italics="true" pageId="4" pageNumber="405">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D1BFD84F5936DA5872CA9A9926D4F85A" box="[1166,1226,2011,2034]" italics="true" pageId="4" pageNumber="405">Haya</emphasis>
|
||
).
|
||
</paragraph>
|
||
<caption id="B7B454D55937DA5976D39BB822F4F8E5" httpUri="https://zenodo.org/record/226297/files/figure.png" pageId="5" pageNumber="406" targetBox="[151,1436,193,1764]" targetPageId="5">
|
||
<paragraph id="E374045D5937DA5976D39BB822F4F8E5" blockId="5.[151,1436,1786,1869]" pageId="5" pageNumber="406">
|
||
<emphasis id="D1BFD84F5937DA5976D39BB8230DF8A7" bold="true" box="[151,275,1786,1808]" pageId="5" pageNumber="406">FIGURE 4.</emphasis>
|
||
<taxonomicName id="24CB7FDE5937DA59775E9BB8206CF8B8" box="[282,626,1786,1808]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="comodorensis">
|
||
<emphasis id="D1BFD84F5937DA59775E9BB8206CF8B8" box="[282,626,1786,1808]" italics="true" pageId="5" pageNumber="406">Notohypsilophodon comodorensis</emphasis>
|
||
</taxonomicName>
|
||
. Sacral vertebrae (UNPSJB-PV 942/12,13,14,15,16 respectively), in left lateral (A and B), in dorsal (C and D) and ventral (E and F) views. Abbreviations: ns, neural spine; rg, rugosities; tp, transverse process.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E374045D5937DA5A76839A372701FE5D" blockId="5.[151,1436,1909,2006]" lastBlockId="6.[151,1436,151,501]" lastPageId="6" lastPageNumber="407" pageId="5" pageNumber="406">
|
||
<emphasis id="D1BFD84F5937DA5976839A3723BEF826" bold="true" box="[199,416,1909,1934]" pageId="5" pageNumber="406">Caudal vertebrae.</emphasis>
|
||
The caudal vertebral centra (
|
||
<figureCitation id="7BF018D85937DA5974A99A37212EF826" box="[749,816,1909,1934]" captionStart="FIGURE 5" captionStartId="6.[151,250,1450,1472]" captionTargetBox="[340,1247,543,1429]" captionTargetId="figure@6.[340,1248,543,1429]" captionTargetPageId="6" captionText="FIGURE 5. Notohypsilophodon comodorensis. Anterior caudal vertebrae (UNPSJB-PV 942 / 17,18), in right lateral (A and C) and ventral (B and D) views. Abbreviations: cf, chevron facet; lgr, longitudinal groove; ncs, neurocentral suture; str, striations. Scale bar equals 2 cm." httpUri="https://zenodo.org/record/226298/files/figure.png" pageId="5" pageNumber="406">Fig. 5</figureCitation>
|
||
) are slightly amphicoelous, as in
|
||
<emphasis id="D1BFD84F5937DA5972EB9A372775F826" box="[1199,1387,1909,1934]" italics="true" pageId="5" pageNumber="406">Gideonmantellia</emphasis>
|
||
and UNPSJB-PV 960, and they are significantly longer anteroposteriorly than wide transversely. Additionally, comparisons with well-preserved caudal sequences (e.g.,
|
||
<taxonomicName id="24CB7FDE5937DA59756E9AFF21CBF87E" box="[810,981,1981,2006]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5937DA59756E9AFF21CBF87E" box="[810,981,1981,2006]" italics="true" pageId="5" pageNumber="406">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5937DA5975A19AFC2697F87D" box="[997,1161,1982,2005]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5937DA5975A19AFC2697F87D" box="[997,1161,1982,2005]" italics="true" pageId="5" pageNumber="406">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
) demonstrate that these caudal vertebrae were located in the anterior portion of the tail.
|
||
<taxonomicName id="24CB7FDE5934DA5A75DC9DD5266AFF18" box="[920,1140,151,176]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A75DC9DD5266AFF18" box="[920,1140,151,176]" italics="true" pageId="6" pageNumber="407">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
possesses subrectangular intercentral articular surfaces, a feature shared with
|
||
<taxonomicName id="24CB7FDE5934DA5A74989DFF219EFF7C" box="[732,896,189,212]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A74989DFF219EFF7C" box="[732,896,189,212]" italics="true" pageId="6" pageNumber="407">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
and UNPSJB-Pv 960. There are relatively welldeveloped longitudinal striations toward each lateral and ventral margin. This feature may be related to the insertion of soft tissue (e.g., the caudofemoralis musculature). Ventrally, the caudal centra of
|
||
<emphasis id="D1BFD84F5934DA5A72989C462785FEB5" box="[1244,1435,260,285]" italics="true" pageId="6" pageNumber="407">Notohypsilophon</emphasis>
|
||
exhibit a marked longitudinal groove and well-developed facets for the insertion of the haemal arches. These last two features are also seen in several basal ornithopods and basal non-hadrosaurid iguanodontians (e.g.,
|
||
<taxonomicName id="24CB7FDE5934DA5A76D39C322356FE2F" box="[151,328,368,391]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A76D39C322356FE2F" box="[151,328,368,391]" italics="true" pageId="6" pageNumber="407">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5934DA5A771C9C2D20E6FE20" authority="Macrogryphosaurus" authorityName="Macrogryphosaurus" box="[344,760,367,392]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A771C9C2D201DFE20" box="[344,515,367,392]" italics="true" pageId="6" pageNumber="407">Hypsilophodon</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5934DA5A74509C2D20E6FE20" box="[532,760,367,392]" italics="true" pageId="6" pageNumber="407">Macrogryphosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D1BFD84F5934DA5A75439C2D2167FE20" box="[775,889,367,392]" italics="true" pageId="6" pageNumber="407">
|
||
<taxonomicName id="24CB7FDE5934DA5A75439C2D216BFE20" box="[775,885,367,392]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">Zalmoxes</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
UNPSJB-Pv 960 and, although less developed, in the basal ornithopod
|
||
<taxonomicName id="24CB7FDE5934DA5A77E19CD62024FE05" box="[421,570,404,429]" class="Reptilia" family="Thescelosauridae" genus="Jeholosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A77E19CD62024FE05" box="[421,570,404,429]" italics="true" pageId="6" pageNumber="407">Jeholosaurus</emphasis>
|
||
</taxonomicName>
|
||
). The presence of the marked longitudinal groove could be related to either ontogenetic status (
|
||
<bibRefCitation id="875A79AC5934DA5A77349CFA203CFE78" author="Han" box="[368,546,439,464]" pageId="6" pageNumber="407" refString="Han, F. L., Barrett, P. M., Butler, R. J. & Xu, X. (2012) Postcranial anatomy of Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Late Cretaceous Yixian Formation of China. Journal of Vertebrate Paleontology, 32, 1370 - 1395. http: // dx. doi. org / 10.1080 / 02724634.2012.694385" type="journal article" year="2012">
|
||
Han
|
||
<emphasis id="D1BFD84F5934DA5A77EC9CFB23FEFE78" box="[424,480,439,464]" italics="true" pageId="6" pageNumber="407">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
) or sexual dimorphism (
|
||
<bibRefCitation id="875A79AC5934DA5A75719CFA21D1FE78" author="Nopcsa" box="[821,975,440,464]" pageId="6" pageNumber="407" refString="Nopcsa, F. (1929) Sexual differences in ornithopodous dinosaurs. Palaeobiologica, 2, 187 - 200." type="journal article" year="1929">Nopcsa 1929</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5934DA5A759E9CF526B7FE78" author="Ibiricu" box="[986,1193,439,464]" pageId="6" pageNumber="407" refString="Ibiricu, L. M., Martinez, R. D., Lamanna, M. C., Casal, G. A., Luna, M., Harris, J. D. & Lacovara, K. J. (2010) A medium-sized ornithopod (Dinosauria: Ornithischia) from the Upper Cretaceous Bajo Barreal Formation of Lago Colhue Huapi, southern Chubut Province, Argentina. Annals of the Carnegie Museum, 79, 39 - 50. http: // dx. doi. org / 10.2992 / 007.079.0103" type="journal article" year="2010">
|
||
Ibiricu
|
||
<emphasis id="D1BFD84F5934DA5A72689CFB267BFE78" box="[1068,1125,439,464]" italics="true" pageId="6" pageNumber="407">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). No ossified tendons are preserved in
|
||
<taxonomicName id="24CB7FDE5934DA5A77159C9E2032FE5D" box="[337,556,476,501]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A77159C9E2032FE5D" box="[337,556,476,501]" italics="true" pageId="6" pageNumber="407">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
, which may be related to preservation rather than genuine absence.
|
||
</paragraph>
|
||
<caption id="B7B454D55934DA5A76D398E82365FA56" httpUri="https://zenodo.org/record/226298/files/figure.png" pageId="6" pageNumber="407" targetBox="[340,1247,543,1429]" targetPageId="6">
|
||
<paragraph id="E374045D5934DA5A76D398E82365FA56" blockId="6.[151,1437,1450,1534]" pageId="6" pageNumber="407">
|
||
<emphasis id="D1BFD84F5934DA5A76D398E8230AFA68" bold="true" box="[151,276,1450,1472]" pageId="6" pageNumber="407">FIGURE 5.</emphasis>
|
||
<taxonomicName id="24CB7FDE5934DA5A775F98E8206AFA68" box="[283,628,1450,1472]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="comodorensis">
|
||
<emphasis id="D1BFD84F5934DA5A775F98E8206AFA68" box="[283,628,1450,1472]" italics="true" pageId="6" pageNumber="407">Notohypsilophodon comodorensis</emphasis>
|
||
</taxonomicName>
|
||
. Anterior caudal vertebrae (UNPSJB-PV 942/17,18), in right lateral (A and C) and ventral (B and D) views. Abbreviations: cf, chevron facet; lgr, longitudinal groove; ncs, neurocentral suture; str, striations. Scale bar equals 2 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E374045D5934DA5A76839B672702F846" blockId="6.[151,1436,1573,2030]" pageId="6" pageNumber="407">
|
||
<emphasis id="D1BFD84F5934DA5A76839B672333F996" bold="true" box="[199,301,1573,1598]" pageId="6" pageNumber="407">Scapula.</emphasis>
|
||
The left scapula of
|
||
<taxonomicName id="24CB7FDE5934DA5A74619B67211FF996" box="[549,769,1573,1598]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A74619B67211FF996" box="[549,769,1573,1598]" italics="true" pageId="6" pageNumber="407">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="7BF018D85934DA5A75519B672142F996" box="[789,860,1573,1598]" captionStart="FIGURE 6" captionStartId="7.[151,250,1595,1617]" captionTargetBox="[340,1247,808,1572]" captionTargetId="figure@7.[340,1247,808,1573]" captionTargetPageId="7" captionText="FIGURE 6. Notohypsilophodon comodorensis. Left scapula (UNPSJB-PV 942 / 23), in lateral (A) and proximal (B) views. Right coracoids (UNPSJB-PV 942 / 24), in lateral view (C). Abbreviation: cof, coracoids foramen; gl, glenoid; scb, scapular blade; spb, supraglenoidal buttress; spf, supraglenoid fossa; stp, sternal process." httpUri="https://zenodo.org/record/226299/files/figure.png" pageId="6" pageNumber="407">Fig. 6</figureCitation>
|
||
A–B;
|
||
<tableCitation id="AE4931E65934DA5A75E19B6721E4F996" box="[933,1018,1573,1598]" captionStart="TABLE 1" captionStartId="7.[151,239,153,176]" captionText="TABLE 1. Selected measurements (in mm) of Notohypsilophodon comodorensis (UNPSJB-PV 942)." pageId="6" pageNumber="407">Table 1</tableCitation>
|
||
) preserves the proximal end and a portion of shaft, which is transversally narrow as in
|
||
<taxonomicName id="24CB7FDE5934DA5A74AC9B082187F9C9" box="[744,921,1610,1633]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A74AC9B082187F9C9" box="[744,921,1610,1633]" italics="true" pageId="6" pageNumber="407">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
. The scapula and coracoids are unfused as in the majority of small ornithischians (
|
||
<bibRefCitation id="875A79AC5934DA5A74119B2C2105F92E" author="Han" box="[597,795,1645,1670]" pageId="6" pageNumber="407" refString="Han, F. L., Barrett, P. M., Butler, R. J. & Xu, X. (2012) Postcranial anatomy of Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Late Cretaceous Yixian Formation of China. Journal of Vertebrate Paleontology, 32, 1370 - 1395. http: // dx. doi. org / 10.1080 / 02724634.2012.694385" type="journal article" year="2012">
|
||
Han
|
||
<emphasis id="D1BFD84F5934DA5A74D79B2C20CCF92E" box="[659,722,1645,1670]" italics="true" pageId="6" pageNumber="407">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
), and unlike
|
||
<taxonomicName id="24CB7FDE5934DA5A75FB9B2C2675F92D" box="[959,1131,1646,1669]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A75FB9B2C2675F92D" box="[959,1131,1646,1669]" italics="true" pageId="6" pageNumber="407">Koreanosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE5934DA5A72EA9B2F2745F92E" box="[1198,1371,1645,1670]" class="Reptilia" family="Hypsilophodontidae" genus="Oryctodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A72EA9B2F2745F92E" box="[1198,1371,1645,1670]" italics="true" pageId="6" pageNumber="407">Oryctodromeus</emphasis>
|
||
</taxonomicName>
|
||
. The proximal end is expanded as in basal ornithopods (e.g.,
|
||
<taxonomicName id="24CB7FDE5934DA5A75429BD021A9F901" box="[774,951,1682,1705]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A75429BD021A9F901" box="[774,951,1682,1705]" italics="true" pageId="6" pageNumber="407">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5934DA5A75819BD3266EF902" box="[965,1136,1681,1706]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A75819BD3266EF902" box="[965,1136,1681,1706]" italics="true" pageId="6" pageNumber="407">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
) and basal iguanodontians (e.g.,
|
||
<taxonomicName id="24CB7FDE5934DA5A769D9BF42343F965" box="[217,349,1718,1741]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A769D9BF42343F965" box="[217,349,1718,1741]" italics="true" pageId="6" pageNumber="407">Dryosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5934DA5A77299BF4200FF965" box="[365,529,1718,1741]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A77299BF4200FF965" box="[365,529,1718,1741]" italics="true" pageId="6" pageNumber="407">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
). The acromial process is not complete; therefore, it cannot be determined if it process was well developed, as in
|
||
<taxonomicName id="24CB7FDE5934DA5A74539B9B2091F95A" box="[535,655,1753,1778]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A74539B9B2091F95A" box="[535,655,1753,1778]" italics="true" pageId="6" pageNumber="407">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
, or only moderately developed, as in
|
||
<taxonomicName id="24CB7FDE5934DA5A72719B9822C2F8BE" authority="Coria & Calvo 2002" authorityName="Coria & Calvo" authorityYear="2002" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A72719B9826F8F959" box="[1077,1254,1754,1777]" italics="true" pageId="6" pageNumber="407">Gasparinisaura</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5934DA5A72B19B9B22CAF8BE" author="Coria" pageId="6" pageNumber="407" refString="Coria, R. A. & Calvo, J. O. (2002) A new iguanodontian ornithopod from Neuquen Basin, Patagonia, Argentina. Journal of Vertebrate Paleontology, 22, 503 - 509. http: // dx. doi. org / 10.1671 / 0272 - 4634 (2002) 022 [0503: aniofn] 2.0. co; 2" type="journal article" year="2002">Coria & Calvo 2002</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. In dorsal view, the scapula of
|
||
<taxonomicName id="24CB7FDE5934DA5A74709BBF210EF8BE" box="[564,784,1789,1814]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A74709BBF210EF8BE" box="[564,784,1789,1814]" italics="true" pageId="6" pageNumber="407">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
exhibits two asymmetrical surfaces, the glenoid fossa and the coracoid articulation. The former is concave, whereas the latter is gently convex. The glenoid fossa is subrectangular in outline. Likewise, dorsally to this fossa, a marked concavity (= supraglenoid fossa;
|
||
<bibRefCitation id="875A79AC5934DA5A73509A0722CFF82A" author="Zheng" pageId="6" pageNumber="407" refString="Zheng, W. J., Jin, X. S., Shibata, M., Azuma, Y. & Yu, F. M. (2012) A new ornithischian dinosaur from the Cretaceous Liangtoutang Formation of Tiantai, Zhejiang Province, China. Cretaceous Research, 34, 208 - 219. http: // dx. doi. org / 10.1016 / j. cretres. 2011.11.001" type="journal article" year="2012">
|
||
Zheng
|
||
<emphasis id="D1BFD84F5934DA5A73279A042782F8F6" box="[1379,1436,1861,1886]" italics="true" pageId="6" pageNumber="407">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
) is also present. This last feature is shared with various ornithischians including:
|
||
<emphasis id="D1BFD84F5934DA5A72B69A2B278BF82A" box="[1266,1429,1897,1922]" italics="true" pageId="6" pageNumber="407">Hexinlusaurus</emphasis>
|
||
,
|
||
<taxonomicName id="24CB7FDE5934DA5A76D39ACF235CF80E" box="[151,322,1933,1958]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A76D39ACF235CF80E" box="[151,322,1933,1958]" italics="true" pageId="6" pageNumber="407">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5934DA5A771D9ACF20CFF80E" authority="Thulborn 1972" authorityName="Thulborn" authorityYear="1972" box="[345,721,1933,1958]" class="Reptilia" family="Fabrosauridae" genus="Lesothosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A771D9ACF23E3F80E" box="[345,509,1933,1958]" italics="true" pageId="6" pageNumber="407">Lesothosaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5934DA5A74579ACF20D7F80E" author="Thulborn" box="[531,713,1933,1958]" pageId="6" pageNumber="407" refString="Thulborn, R. A. (1972) The postcranial skeleton of the Triassic ornithischian dinosaur Fabrosaurus australis. Palaeontology, 15, 29 - 60." type="journal article" year="1972">Thulborn 1972</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5934DA5A74A19ACF218DF80E" box="[741,915,1933,1958]" class="Reptilia" family="Hypsilophodontidae" genus="Thescelosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A74A19ACF218DF80E" box="[741,915,1933,1958]" italics="true" pageId="6" pageNumber="407">Thescelosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5934DA5A75EF9ACC26DBF80E" authority="Uteodon" authorityName="Uteodon" box="[939,1221,1933,1958]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A75EF9ACC2651F80D" box="[939,1103,1934,1957]" italics="true" pageId="6" pageNumber="407">Tenontosaurus</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5934DA5A72219ACF26DBF80E" box="[1125,1221,1933,1958]" italics="true" pageId="6" pageNumber="407">Uteodon</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="D1BFD84F5934DA5A73519ACC278CF80D" box="[1301,1426,1934,1957]" italics="true" pageId="6" pageNumber="407">Yueosaurus</emphasis>
|
||
. However, the supraglenoid fossa in
|
||
<taxonomicName id="24CB7FDE5934DA5A747A9AF32104F862" box="[574,794,1969,1994]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A747A9AF32104F862" box="[574,794,1969,1994]" italics="true" pageId="6" pageNumber="407">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
is more pronounced than in all of these taxa with the exception of
|
||
<taxonomicName id="24CB7FDE5934DA5A776D9A9423D3F845" box="[297,461,2006,2029]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A776D9A9423D3F845" box="[297,461,2006,2029]" italics="true" pageId="6" pageNumber="407">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
, whose fossa is similar to that seen in
|
||
<taxonomicName id="24CB7FDE5934DA5A75389A972709F846" authority="Martinez 1998" authorityName="Martinez" authorityYear="1998" box="[892,1303,2005,2030]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5934DA5A75389A972646F846" box="[892,1112,2005,2030]" italics="true" pageId="6" pageNumber="407">Notohypsilophodon</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC5934DA5A72229A972711F846" author="Martinez" box="[1126,1295,2005,2030]" pageId="6" pageNumber="407" refString="Martinez, R. D. (1998) Notohypsilophodon comodorensis gen. et sp. nov. Un Hypsilophodontidae (Ornithischia: Ornithopoda) del Cretacico Superior de Chubut, Patagonia central, Argentina. Acta Geologica Leopoldensia, 21, 119 - 135." type="journal article" year="1998">Martinez 1998</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="B7B454D55935DA5B76D39DDB26D1FF18" box="[151,1231,152,176]" pageId="7" pageNumber="408">
|
||
<paragraph id="E374045D5935DA5B76D39DDB26D1FF18" blockId="7.[151,1231,152,176]" box="[151,1231,152,176]" pageId="7" pageNumber="408">
|
||
<emphasis id="D1BFD84F5935DA5B76D39DDB2317FF18" bold="true" box="[151,265,153,176]" pageId="7" pageNumber="408">TABLE 1.</emphasis>
|
||
Selected measurements (in mm) of
|
||
<taxonomicName id="24CB7FDE5935DA5B74CF9DDA21EBFF18" box="[651,1013,152,176]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="comodorensis">
|
||
<emphasis id="D1BFD84F5935DA5B74CF9DDA21EBFF18" box="[651,1013,152,176]" italics="true" pageId="7" pageNumber="408">Notohypsilophodon comodorensis</emphasis>
|
||
</taxonomicName>
|
||
(UNPSJB-PV 942).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E374045D5935DA5B76DB9D8F263BFEC9" blockId="7.[159,1155,205,731]" pageId="7" pageNumber="408">Element Measurement Humerus Total length 127.3 Ulna Total length 91.3 Femur (proximal end) Maximun anteroposterior width 44.9</paragraph>
|
||
<paragraph id="E374045D5935DA5B77A39C37262FFE77" blockId="7.[159,1155,205,731]" pageId="7" pageNumber="408">Maximun lateral width 47.7 Femur (distal end) Maximun anteroposterior width 45.5 Tibia Total length 211.5</paragraph>
|
||
<paragraph id="E374045D5935DA5B77A39CB1263BFDA1" blockId="7.[159,1155,205,731]" box="[487,1061,499,521]" pageId="7" pageNumber="408">Greatest proximal width 56.5</paragraph>
|
||
<paragraph id="E374045D5935DA5B77A39F5F2601FDF5" blockId="7.[159,1155,205,731]" pageId="7" pageNumber="408">Greatest distal width 45.5 Fibula Total length 192</paragraph>
|
||
<paragraph id="E374045D5935DA5B77A39F332604F862" blockId="7.[159,1155,205,731]" lastBlockId="7.[151,1436,1717,1994]" pageId="7" pageNumber="408">
|
||
Greatest proximal width 29.6 Greatest distal width 14.1 Phalanx (?-1) Total length 36.1
|
||
<emphasis id="D1BFD84F5935DA5B76839BF72359F966" bold="true" box="[199,327,1717,1742]" pageId="7" pageNumber="408">Coracoids.</emphasis>
|
||
The semicircular shape of the right coracoid (
|
||
<figureCitation id="7BF018D85935DA5B751C9BF72182F966" box="[856,924,1717,1742]" captionStart="FIGURE 6" captionStartId="7.[151,250,1595,1617]" captionTargetBox="[340,1247,808,1572]" captionTargetId="figure@7.[340,1247,808,1573]" captionTargetPageId="7" captionText="FIGURE 6. Notohypsilophodon comodorensis. Left scapula (UNPSJB-PV 942 / 23), in lateral (A) and proximal (B) views. Right coracoids (UNPSJB-PV 942 / 24), in lateral view (C). Abbreviation: cof, coracoids foramen; gl, glenoid; scb, scapular blade; spb, supraglenoidal buttress; spf, supraglenoid fossa; stp, sternal process." httpUri="https://zenodo.org/record/226299/files/figure.png" pageId="7" pageNumber="408">Fig. 6</figureCitation>
|
||
C), with a slightly convex lateral surface and medially concave surface, is comparable to that of
|
||
<taxonomicName id="24CB7FDE5935DA5B74929B982199F959" box="[726,903,1754,1777]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B74929B982199F959" box="[726,903,1754,1777]" italics="true" pageId="7" pageNumber="408">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5935DA5B75D29B9B265FF95A" box="[918,1089,1753,1778]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B75D29B9B265FF95A" box="[918,1089,1753,1778]" italics="true" pageId="7" pageNumber="408">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE5935DA5B723D9B9B2716F95A" box="[1145,1288,1753,1778]" class="Reptilia" family="Hypsilophodontidae" genus="Yandusaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B723D9B9B2716F95A" box="[1145,1288,1753,1778]" italics="true" pageId="7" pageNumber="408">Yandusaurus</emphasis>
|
||
</taxonomicName>
|
||
. Conversely, in
|
||
<taxonomicName id="24CB7FDE5935DA5B76FE9BBF2398F8BE" box="[186,390,1789,1814]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B76FE9BBF2398F8BE" box="[186,390,1789,1814]" italics="true" pageId="7" pageNumber="408">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5935DA5B77DE9BBF2031F8BE" box="[410,559,1789,1814]" class="Reptilia" family="Thescelosauridae" genus="Jeholosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B77DE9BBF2031F8BE" box="[410,559,1789,1814]" italics="true" pageId="7" pageNumber="408">Jeholosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE5935DA5B74379BBC2101F8BD" box="[627,799,1790,1813]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B74379BBC2101F8BD" box="[627,799,1790,1813]" italics="true" pageId="7" pageNumber="408">Koreanosaurus</emphasis>
|
||
</taxonomicName>
|
||
, the coracoids are sub-rectangular in shape with flat surfaces. In
|
||
<emphasis id="D1BFD84F5935DA5B775A9A60238AF891" box="[286,404,1826,1849]" italics="true" pageId="7" pageNumber="408">Trinisaura</emphasis>
|
||
, the coracoid is semicircular in outline, as in
|
||
<taxonomicName id="24CB7FDE5935DA5B75C89A632679F892" box="[908,1127,1825,1850]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B75C89A632679F892" box="[908,1127,1825,1850]" italics="true" pageId="7" pageNumber="408">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
; however, in the former the dorsal margin is interrupted by a marked concavity (see
|
||
<bibRefCitation id="875A79AC5935DA5B750C9A072602F8F6" author="Coria" box="[840,1052,1861,1886]" pageId="7" pageNumber="408" refString="Coria, R. A., Moly, J. J., Reguero, M., Santillana, S. & Marenssi, S. (2013) A new ornithopod (Dinosauria; Ornithischia) from Antarctica. Cretaceous Research, 41, 186 - 193. http: // dx. doi. org / 10.1016 / j. cretres. 2012.12.004" type="journal article" year="2013">
|
||
Coria
|
||
<emphasis id="D1BFD84F5935DA5B75D09A0421CCF8F6" box="[916,978,1861,1886]" italics="true" pageId="7" pageNumber="408">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
: fig. 3). The sternal process, in
|
||
<taxonomicName id="24CB7FDE5935DA5B76D39A2B236DF82A" box="[151,371,1897,1922]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B76D39A2B236DF82A" box="[151,371,1897,1922]" italics="true" pageId="7" pageNumber="408">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
is not as well-developed as those of
|
||
<taxonomicName id="24CB7FDE5935DA5B75019A2B21A3F82A" box="[837,957,1897,1922]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B75019A2B21A3F82A" box="[837,957,1897,1922]" italics="true" pageId="7" pageNumber="408">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5935DA5B75969A2826C6F829" authority="Haya" authorityName="Haya" box="[978,1240,1898,1921]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B75969A28269DF829" box="[978,1155,1898,1921]" italics="true" pageId="7" pageNumber="408">Gasparinisaura</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5935DA5B72DC9A2826C6F829" box="[1176,1240,1898,1921]" italics="true" pageId="7" pageNumber="408">Haya</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5935DA5B72AD9A28278BF829" box="[1257,1429,1898,1921]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">Koreanosaurus</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5935DA5B76D39ACD2325F80E" box="[151,315,1935,1958]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B76D39ACD2325F80E" box="[151,315,1935,1958]" italics="true" pageId="7" pageNumber="408">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<emphasis id="D1BFD84F5935DA5B77269ACC23C6F80D" box="[354,472,1934,1957]" italics="true" pageId="7" pageNumber="408">Trinisaura</emphasis>
|
||
. Although, it is taphonomically affected on its lateral size, in
|
||
<taxonomicName id="24CB7FDE5935DA5B72D29ACF276CF80E" box="[1174,1394,1933,1958]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5935DA5B72D29ACF276CF80E" box="[1174,1394,1933,1958]" italics="true" pageId="7" pageNumber="408">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
the sternal process is markedly more reduced than in those ornithopods (see below).
|
||
</paragraph>
|
||
<caption id="B7B454D55935DA5B76D39B7921A1F926" httpUri="https://zenodo.org/record/226299/files/figure.png" pageId="7" pageNumber="408" targetBox="[340,1247,808,1572]" targetPageId="7">
|
||
<paragraph id="E374045D5935DA5B76D39B7921A1F926" blockId="7.[151,1437,1595,1678]" pageId="7" pageNumber="408">
|
||
<emphasis id="D1BFD84F5935DA5B76D39B792308F9F8" bold="true" box="[151,278,1595,1617]" pageId="7" pageNumber="408">FIGURE 6.</emphasis>
|
||
<taxonomicName id="24CB7FDE5935DA5B775B9B792065F9F9" box="[287,635,1595,1617]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="comodorensis">
|
||
<emphasis id="D1BFD84F5935DA5B775B9B792065F9F9" box="[287,635,1595,1617]" italics="true" pageId="7" pageNumber="408">Notohypsilophodon comodorensis</emphasis>
|
||
</taxonomicName>
|
||
. Left scapula (UNPSJB-PV 942/23), in lateral (A) and proximal (B) views. Right coracoids (UNPSJB-PV 942/24), in lateral view (C). Abbreviation: cof, coracoids foramen; gl, glenoid; scb, scapular blade; spb, supraglenoidal buttress; spf, supraglenoid fossa; stp, sternal process.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B7B454D5593ADA5476D399CF20D0FB57" httpUri="https://zenodo.org/record/226300/files/figure.png" pageId="8" pageNumber="409" targetBox="[151,1436,193,1143]" targetPageId="8">
|
||
<paragraph id="E374045D593ADA5476D399CF20D0FB57" blockId="8.[151,1436,1165,1279]" pageId="8" pageNumber="409">
|
||
<emphasis id="D1BFD84F593ADA5476D399CF230DFB0B" bold="true" box="[151,275,1165,1187]" pageId="8" pageNumber="409">FIGURE 7.</emphasis>
|
||
<taxonomicName id="24CB7FDE593ADA54775E99CF206DFB0B" box="[282,627,1165,1187]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="species" species="comodorensis">
|
||
<emphasis id="D1BFD84F593ADA54775E99CF206DFB0B" box="[282,627,1165,1187]" italics="true" pageId="8" pageNumber="409">Notohypsilophodon comodorensis</emphasis>
|
||
</taxonomicName>
|
||
. Right humerus (UNPSJB-PV 942/25), in anterior (A), proximal (B) and distal (C) views. Left ulna (UNPSJB-PV 942/27), in medial (D), proximal (E) and distal (F) views. Abbreviation: asr, articular surface radius; dc, deltopectoral crest; hh, humeral head; lsh, lateral shoulder; ms, muscle scars; msh, medial shoulder; op, olecranon process; rc, radial condyle; uc, ulnar condyle.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E374045D593ADA547683986E2733F9B5" blockId="8.[151,1436,1324,2033]" pageId="8" pageNumber="409">
|
||
The coracoid foramen is circular in outline, proportionately large and completely enclosed. The coracoid foramen in
|
||
<taxonomicName id="24CB7FDE593ADA54775E981223EBFAC1" box="[282,501,1360,1385]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA54775E981223EBFAC1" box="[282,501,1360,1385]" italics="true" pageId="8" pageNumber="409">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
is placed well above the scapula-coracoid suture (i.e., there is no contact between the foramen and scapula). The shape and position of the coracoid foramen is similar to those of
|
||
<taxonomicName id="24CB7FDE593ADA5473599836278BFA25" box="[1309,1429,1396,1421]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5473599836278BFA25" box="[1309,1429,1396,1421]" italics="true" pageId="8" pageNumber="409">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA5476D398DB23CFFA18" authority="Trinisaura" authorityName="Trinisaura" box="[151,465,1433,1456]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5476D398DB2356FA18" box="[151,328,1433,1456]" italics="true" pageId="8" pageNumber="409">Gasparinisaura</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F593ADA54771E98DB23CFFA18" box="[346,465,1433,1456]" italics="true" pageId="8" pageNumber="409">Trinisaura</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE593ADA54744A98DB20ACFA18" box="[526,690,1433,1456]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA54744A98DB20ACFA18" box="[526,690,1433,1456]" italics="true" pageId="8" pageNumber="409">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="875A79AC593ADA5474B098DA21C5FA19" author="Winkler" box="[756,987,1432,1457]" pageId="8" pageNumber="409" refString="Winkler, D. A., Murry, P. A. & Jacobs, L. L. (1997) A new species of Tenontosaurus (Dinosauria: Ornithopoda) from the Early Cretaceous of Texas. Journal of Vertebrate Paleontology, 17, 330 - 348. http: // dx. doi. org / 10.1080 / 02724634.1997.10010978" type="journal article" year="1997">
|
||
Winkler
|
||
<emphasis id="D1BFD84F593ADA54751D98DB218BFA19" box="[857,917,1432,1457]" italics="true" pageId="8" pageNumber="409">et al.</emphasis>
|
||
1997
|
||
</bibRefCitation>
|
||
: fig. 16a), and unlike that observed in
|
||
<taxonomicName id="24CB7FDE593ADA5476D398FE235CFA7D" box="[151,322,1468,1493]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5476D398FE235CFA7D" box="[151,322,1468,1493]" italics="true" pageId="8" pageNumber="409">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
. In the latter taxon, the foramen is positioned close to the suture and connected to the scapula by a groove (
|
||
<bibRefCitation id="875A79AC593ADA5476BD98A22382FA51" author="Galton" box="[249,412,1504,1529]" pageId="8" pageNumber="409" refString="Galton, P. M. (1974 a) The ornithischian dinosaur Hypsilophodon from the Wealden of the Isle of Wight. Bulletin of the British Museum (Natural History), Geology, 25, 1 - 152." type="journal article" year="1974" yearSuffix="a">Galton 1974a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC593ADA5477E998A2206DFA51" author="Huh" box="[429,627,1504,1529]" pageId="8" pageNumber="409" refString="Huh, M., Lee, D. G., Kim, J. K., Lim, J. D. & Godefroit, P. (2010) A new basal ornithopod dinosaur from the Upper Cretaceous of South Korea. Neues Jahrbuch fur Geologie Palaontologie, Abhandlungen, 259, 1 - 24. http: // dx. doi. org / 10.1127 / 0077 - 7749 / 2010 / 0102" type="journal article" year="2010">
|
||
Huh
|
||
<emphasis id="D1BFD84F593ADA5477AF98A32037FA51" box="[491,553,1504,1529]" italics="true" pageId="8" pageNumber="409">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). In more basal forms (e.g.,
|
||
<taxonomicName id="24CB7FDE593ADA54759598A22683FA51" box="[977,1181,1504,1529]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA54759598A22683FA51" box="[977,1181,1504,1529]" italics="true" pageId="8" pageNumber="409">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA5472F598A32743FA50" box="[1201,1373,1505,1528]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5472F598A32743FA50" box="[1201,1373,1505,1528]" italics="true" pageId="8" pageNumber="409">Koreanosaurus</emphasis>
|
||
</taxonomicName>
|
||
), the foramen is smaller than in
|
||
<taxonomicName id="24CB7FDE593ADA5477869B462083F9B5" box="[450,669,1540,1565]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5477869B462083F9B5" box="[450,669,1540,1565]" italics="true" pageId="8" pageNumber="409">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
and located more towards the center of the coracoid plate.
|
||
</paragraph>
|
||
<paragraph id="E374045D593ADA5576839B6A2795FE5C" blockId="8.[151,1436,1324,2033]" lastBlockId="9.[151,1436,151,2013]" lastPageId="9" lastPageNumber="410" pageId="8" pageNumber="409">
|
||
<emphasis id="D1BFD84F593ADA5476839B6A2320F9E9" bold="true" box="[199,318,1576,1601]" pageId="8" pageNumber="409">Humerus.</emphasis>
|
||
The right humerus (
|
||
<figureCitation id="7BF018D8593ADA5474629B6A2077F9E9" box="[550,617,1576,1601]" captionStart="FIGURE 7" captionStartId="8.[151,250,1165,1187]" captionTargetBox="[151,1436,193,1143]" captionTargetId="figure@8.[151,1436,193,1144]" captionTargetPageId="8" captionText="FIGURE 7. Notohypsilophodon comodorensis. Right humerus (UNPSJB-PV 942 / 25), in anterior (A), proximal (B) and distal (C) views. Left ulna (UNPSJB-PV 942 / 27), in medial (D), proximal (E) and distal (F) views. Abbreviation: asr, articular surface radius; dc, deltopectoral crest; hh, humeral head; lsh, lateral shoulder; ms, muscle scars; msh, medial shoulder; op, olecranon process; rc, radial condyle; uc, ulnar condyle." httpUri="https://zenodo.org/record/226300/files/figure.png" pageId="8" pageNumber="409">Fig. 7</figureCitation>
|
||
A–C;
|
||
<tableCitation id="AE4931E6593ADA5474E99B6A20E1F9E9" box="[685,767,1576,1601]" captionStart="TABLE 1" captionStartId="7.[151,239,153,176]" captionText="TABLE 1. Selected measurements (in mm) of Notohypsilophodon comodorensis (UNPSJB-PV 942)." pageId="8" pageNumber="409">Table 1</tableCitation>
|
||
) of
|
||
<taxonomicName id="24CB7FDE593ADA5475759B6A2613F9E9" box="[817,1037,1576,1601]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5475759B6A2613F9E9" box="[817,1037,1576,1601]" italics="true" pageId="8" pageNumber="409">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
is gracile (
|
||
<quantity id="2433A9B8593ADA5472CF9B6A26C2F9E8" box="[1163,1244,1576,1601]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="409" unit="cm" value="1.5">1.5 cm</quantity>
|
||
, measurement of the shaft taken in anterior view below the deltopectoral crest) as in
|
||
<taxonomicName id="24CB7FDE593ADA5475E59B0E2607F9CD" box="[929,1049,1612,1637]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5475E59B0E2607F9CD" box="[929,1049,1612,1637]" italics="true" pageId="8" pageNumber="409">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA54726D9B0F26C4F9CC" box="[1065,1242,1613,1636]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA54726D9B0F26C4F9CC" box="[1065,1242,1613,1636]" italics="true" pageId="8" pageNumber="409">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA5472AE9B0E2774F9CD" box="[1258,1386,1612,1637]" class="Reptilia" family="Hadrosauridae" genus="Talenkauen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5472AE9B0E2774F9CD" box="[1258,1386,1612,1637]" italics="true" pageId="8" pageNumber="409">Talenkauen</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D1BFD84F593ADA5476D39B332313F920" box="[151,269,1649,1672]" italics="true" pageId="8" pageNumber="409">Trinisaura</emphasis>
|
||
, whereas in the rhabdodontid
|
||
<taxonomicName id="24CB7FDE593ADA5474229B3220CCF921" box="[614,722,1648,1673]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5474229B3220CCF921" box="[614,722,1648,1673]" italics="true" pageId="8" pageNumber="409">Zalmoxes</emphasis>
|
||
</taxonomicName>
|
||
(approximately
|
||
<quantity id="2433A9B8593ADA5475C99B3321D6F920" box="[909,968,1649,1673]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="409" unit="cm" value="3.0">3 cm</quantity>
|
||
, measurement taken from
|
||
<bibRefCitation id="875A79AC593ADA5472B29B3222E7F905" author="Weishampel" pageId="8" pageNumber="409" refString="Weishampel, D. B., Jianu, C. M., Csiki, Z. & Norman, D. B. (2003) Osteology and phylogeny of Zalmoxes (n. g.), an unusual euornithopod dinosaur from the latest Cretaceous of Romania. Journal of Systematic Palaeontology, 1, 65 - 123. http: // dx. doi. org / 10.1017 / s 1477201903001032" type="journal article" year="2003">
|
||
Weishampel
|
||
<emphasis id="D1BFD84F593ADA5473CC9B3322AAF905" italics="true" pageId="8" pageNumber="409">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
: fig. 20d) and the basal iguanodontian
|
||
<taxonomicName id="24CB7FDE593ADA5474FE9BD72140F904" box="[698,862,1685,1708]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5474FE9BD72140F904" box="[698,862,1685,1708]" italics="true" pageId="8" pageNumber="409">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
, the humerus is robust (Norman
|
||
<emphasis id="D1BFD84F593ADA5472989BD72708F905" box="[1244,1302,1684,1709]" italics="true" pageId="8" pageNumber="409">et al.</emphasis>
|
||
2004). The shaft is slightly twisted, as is characteristic of a variety of basal ornithopods (Norman
|
||
<emphasis id="D1BFD84F593ADA5472359BFB26B5F979" box="[1137,1195,1720,1745]" italics="true" pageId="8" pageNumber="409">et al.</emphasis>
|
||
2004). Nevertheless, the rotation of the humeral shaft appears to be less than in
|
||
<taxonomicName id="24CB7FDE593ADA54752C9B9E260DF95D" box="[872,1043,1756,1781]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA54752C9B9E260DF95D" box="[872,1043,1756,1781]" italics="true" pageId="8" pageNumber="409">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC593ADA5472639B9E2741F95D" author="Galton" box="[1063,1375,1756,1781]" pageId="8" pageNumber="409" refString="Galton, P. M. (1974 a) The ornithischian dinosaur Hypsilophodon from the Wealden of the Isle of Wight. Bulletin of the British Museum (Natural History), Geology, 25, 1 - 152." type="journal article" year="1974" yearSuffix="a">Galton 1974a: figs. 38–39</bibRefCitation>
|
||
) and resembles the condition of
|
||
<taxonomicName id="24CB7FDE593ADA54778F9A42205DF8B1" box="[459,579,1792,1817]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA54778F9A42205DF8B1" box="[459,579,1792,1817]" italics="true" pageId="8" pageNumber="409">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
. In anterior view, the shaft is bowed as in
|
||
<taxonomicName id="24CB7FDE593ADA54726E9A4226A4F8B1" box="[1066,1210,1792,1817]" class="Reptilia" family="Hypsilophodontidae" genus="Yandusaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA54726E9A4226A4F8B1" box="[1066,1210,1792,1817]" italics="true" pageId="8" pageNumber="409">Yandusaurus</emphasis>
|
||
</taxonomicName>
|
||
and unlike those of
|
||
<taxonomicName id="24CB7FDE593ADA5476D39A662311F895" box="[151,271,1828,1853]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5476D39A662311F895" box="[151,271,1828,1853]" italics="true" pageId="8" pageNumber="409">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D1BFD84F593ADA54770F9A6723DFF894" box="[331,449,1829,1852]" italics="true" pageId="8" pageNumber="409">Trinisaura</emphasis>
|
||
where the bowing is less marked. The deltopectoral crest is reduced (i.e., to shallow rugosities for the pectoral muscle insertions) and is positioned in the proximal portion of the humerus as in
|
||
<taxonomicName id="24CB7FDE593ADA5476D39A2E2311F82D" box="[151,271,1900,1925]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5476D39A2E2311F82D" box="[151,271,1900,1925]" italics="true" pageId="8" pageNumber="409">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA5477669A2F20DCF82D" authority="Norman et al. 2004" authorityName="Norman et al." authorityYear="2004" box="[290,706,1900,1925]" class="Reptilia" family="Hypsilophodontidae" genus="Bugenasaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5477669A2F23A9F82C" box="[290,439,1901,1924]" italics="true" pageId="8" pageNumber="409">Bugenasaura</emphasis>
|
||
(Norman
|
||
<emphasis id="D1BFD84F593ADA5474709A2F206CF82D" box="[564,626,1900,1925]" italics="true" pageId="8" pageNumber="409">et al.</emphasis>
|
||
2004)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA5474969A2F219DF82C" box="[722,899,1901,1924]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5474969A2F219DF82C" box="[722,899,1901,1924]" italics="true" pageId="8" pageNumber="409">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA5475D39A2E2609F82D" box="[919,1047,1900,1925]" class="Reptilia" family="Hadrosauridae" genus="Talenkauen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5475D39A2E2609F82D" box="[919,1047,1900,1925]" italics="true" pageId="8" pageNumber="409">Talenkauen</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D1BFD84F593ADA54721C9A2F26D0F82C" box="[1112,1230,1901,1924]" italics="true" pageId="8" pageNumber="409">Trinisaura</emphasis>
|
||
. By contrast, the deltopectoral crests of basal neornithischians and ornithopods (e.g.,
|
||
<emphasis id="D1BFD84F593ADA5475A29AD3263CF800" box="[998,1058,1937,1960]" italics="true" pageId="8" pageNumber="409">Haya</emphasis>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA5472789AD226F9F801" box="[1084,1255,1936,1961]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5472789AD226F9F801" box="[1084,1255,1936,1961]" italics="true" pageId="8" pageNumber="409">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA5473449AD2278BF801" box="[1280,1429,1936,1961]" class="Reptilia" family="Thescelosauridae" genus="Jeholosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5473449AD2278BF801" box="[1280,1429,1936,1961]" italics="true" pageId="8" pageNumber="409">Jeholosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA5476D39AF7235DF864" box="[151,323,1973,1996]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5476D39AF7235DF864" box="[151,323,1973,1996]" italics="true" pageId="8" pageNumber="409">Koreanosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593ADA5477149AF623FEF865" box="[336,480,1972,1997]" class="Reptilia" family="Hypsilophodontidae" genus="Yandusaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="409" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593ADA5477149AF623FEF865" box="[336,480,1972,1997]" italics="true" pageId="8" pageNumber="409">Yandusaurus</emphasis>
|
||
</taxonomicName>
|
||
, among others) are well developed and angular in outline. Likewise, the deltopectoral crest in basal (non-hadrosaurid) iguanodontians is robust and it occupies approximately 50% of total humerus length (Norman
|
||
<emphasis id="D1BFD84F593BDA5577149DDB2397FF18" box="[336,393,151,176]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2004). The proximal end of the humeral head is transversally expanded as in
|
||
<taxonomicName id="24CB7FDE593BDA5572B09DD52772FF18" box="[1268,1388,151,176]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572B09DD52772FF18" box="[1268,1388,151,176]" italics="true" pageId="9" pageNumber="410">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D1BFD84F593BDA5576D39DFF2351FF7C" box="[151,335,189,212]" italics="true" pageId="9" pageNumber="410">
|
||
<taxonomicName id="24CB7FDE593BDA5576D39DFF2355FF7C" box="[151,331,189,212]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">Gasparinisaura</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and exhibits a marked tuberosity, similar to that described in
|
||
<taxonomicName id="24CB7FDE593BDA55724F9DFF274FFF7C" authority="Martinez 1998" authorityName="Martinez" authorityYear="1998" box="[1035,1361,188,212]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55724F9DFF2691FF7C" box="[1035,1167,189,212]" italics="true" pageId="9" pageNumber="410">Dryosaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="875A79AC593BDA5572E49DFE2757FF7C" author="Martinez" box="[1184,1353,188,212]" pageId="9" pageNumber="410" refString="Martinez, R. D. (1998) Notohypsilophodon comodorensis gen. et sp. nov. Un Hypsilophodontidae (Ornithischia: Ornithopoda) del Cretacico Superior de Chubut, Patagonia central, Argentina. Acta Geologica Leopoldensia, 21, 119 - 135." type="journal article" year="1998">Martínez 1998</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Thus, the proximal end is wider than the distal end as in
|
||
<taxonomicName id="24CB7FDE593BDA5574BE9D9D216CFF50" box="[762,882,223,248]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5574BE9D9D216CFF50" box="[762,882,223,248]" italics="true" pageId="9" pageNumber="410">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D1BFD84F593BDA5575F49DA32638FF50" box="[944,1062,225,248]" italics="true" pageId="9" pageNumber="410">Trinisaura</emphasis>
|
||
. Conversely, in
|
||
<taxonomicName id="24CB7FDE593BDA5572AE9D9D278BFF50" box="[1258,1429,223,248]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572AE9D9D278BFF50" box="[1258,1429,223,248]" italics="true" pageId="9" pageNumber="410">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA5576D39C462339FEB5" box="[151,295,260,285]" class="Reptilia" family="Hypsilophodontidae" genus="Yandusaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5576D39C462339FEB5" box="[151,295,260,285]" italics="true" pageId="9" pageNumber="410">Yandusaurus</emphasis>
|
||
</taxonomicName>
|
||
and basal iguanodontians (e.g.,
|
||
<taxonomicName id="24CB7FDE593BDA5574E19C472157FEB4" box="[677,841,261,284]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5574E19C472157FEB4" box="[677,841,261,284]" italics="true" pageId="9" pageNumber="410">Tenontosaurus</emphasis>
|
||
</taxonomicName>
|
||
), the proximal and distal ends of the humerus are approximately equal in mediolateral width. The distal end of the humerus exhibits moderately developed radial and ulnar condyles, as well as an intercondylar groove (=fossa) on its anterior surface. This condition differs from that seen in
|
||
<taxonomicName id="24CB7FDE593BDA5576B39C2D23BCFE20" box="[247,418,367,392]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5576B39C2D23BCFE20" box="[247,418,367,392]" italics="true" pageId="9" pageNumber="410">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA5577F19C2D207DFE20" box="[437,611,367,392]" class="Reptilia" family="Hypsilophodontidae" genus="Thescelosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5577F19C2D207DFE20" box="[437,611,367,392]" italics="true" pageId="9" pageNumber="410">Thescelosaurus</emphasis>
|
||
</taxonomicName>
|
||
and non-hadrosaurid iguanodontians (e.g.,
|
||
<taxonomicName id="24CB7FDE593BDA5572219C322717FE2F" box="[1125,1289,368,391]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572219C322717FE2F" box="[1125,1289,368,391]" italics="true" pageId="9" pageNumber="410">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA5573589C2D278BFE20" box="[1308,1429,367,392]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5573589C2D278BFE20" box="[1308,1429,367,392]" italics="true" pageId="9" pageNumber="410">Iguanodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA5576D39CD6235FFE05" box="[151,321,404,429]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5576D39CD6235FFE05" box="[151,321,404,429]" italics="true" pageId="9" pageNumber="410">Mantellisaurus</emphasis>
|
||
</taxonomicName>
|
||
) where the condyles and the groove are more strongly developed. The lateral radial condyle is expanded anteroposteriorly and the medial ulnar condyle is rounded, both of which are features that
|
||
<taxonomicName id="24CB7FDE593BDA5576D39C9E236DFE5D" box="[151,371,476,501]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5576D39C9E236DFE5D" box="[151,371,476,501]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
shares with basal neornithischians and ornithopods such as
|
||
<taxonomicName id="24CB7FDE593BDA5572579C9E26BDFE5D" box="[1043,1187,476,501]" class="Reptilia" family="Hypsilophodontidae" genus="Yandusaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572579C9E26BDFE5D" box="[1043,1187,476,501]" italics="true" pageId="9" pageNumber="410">Yandusaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE593BDA55729D9C9F279BFE5C" box="[1241,1413,477,500]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55729D9C9F279BFE5C" box="[1241,1413,477,500]" italics="true" pageId="9" pageNumber="410">Koreanosaurus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E374045D593BDA5576839CBD20E7FCF5" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="410">
|
||
<emphasis id="D1BFD84F593BDA5576839CBD2319FDB0" bold="true" box="[199,263,511,536]" pageId="9" pageNumber="410">Ulna.</emphasis>
|
||
The ulna (
|
||
<figureCitation id="7BF018D8593BDA5577C19CBD23D6FDBF" box="[389,456,511,536]" captionStart="FIGURE 7" captionStartId="8.[151,250,1165,1187]" captionTargetBox="[151,1436,193,1143]" captionTargetId="figure@8.[151,1436,193,1144]" captionTargetPageId="8" captionText="FIGURE 7. Notohypsilophodon comodorensis. Right humerus (UNPSJB-PV 942 / 25), in anterior (A), proximal (B) and distal (C) views. Left ulna (UNPSJB-PV 942 / 27), in medial (D), proximal (E) and distal (F) views. Abbreviation: asr, articular surface radius; dc, deltopectoral crest; hh, humeral head; lsh, lateral shoulder; ms, muscle scars; msh, medial shoulder; op, olecranon process; rc, radial condyle; uc, ulnar condyle." httpUri="https://zenodo.org/record/226300/files/figure.png" pageId="9" pageNumber="410">Fig. 7</figureCitation>
|
||
D–E–F;
|
||
<tableCitation id="AE4931E6593BDA55746E9CBD2062FDB0" box="[554,636,511,536]" captionStart="TABLE 1" captionStartId="7.[151,239,153,176]" captionText="TABLE 1. Selected measurements (in mm) of Notohypsilophodon comodorensis (UNPSJB-PV 942)." pageId="9" pageNumber="410">Table 1</tableCitation>
|
||
) of
|
||
<taxonomicName id="24CB7FDE593BDA5574F49CBD2192FDB0" box="[688,908,511,536]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5574F49CBD2192FDB0" box="[688,908,511,536]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
is slender, as in
|
||
<taxonomicName id="24CB7FDE593BDA5572089CBD26E9FDB0" box="[1100,1271,511,536]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572089CBD26E9FDB0" box="[1100,1271,511,536]" italics="true" pageId="9" pageNumber="410">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
, in contrast to the more robust ulnae of non-hadrosaurid iguanodontians. In proximal view, the ulna is triangular in outline (formed by the head of the ulna and the lateral and medial processes according to Butler
|
||
<emphasis id="D1BFD84F593BDA5572349F0B26B6FDC8" box="[1136,1192,583,608]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2011) with the lateral border convex, the medial slightly concave and the articular surface for the radius gently concave. The olecranon process is reduced, and comparable to that observed in
|
||
<taxonomicName id="24CB7FDE593BDA5575429FCD21AFFD00" box="[774,945,655,680]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5575429FCD21AFFD00" box="[774,945,655,680]" italics="true" pageId="9" pageNumber="410">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE593BDA5575AD9FCD269DFD00" box="[1001,1155,655,680]" class="Reptilia" family="Hypsilophodontidae" genus="Parksosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5575AD9FCD269DFD00" box="[1001,1155,655,680]" italics="true" pageId="9" pageNumber="410">Parksosaurus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="24CB7FDE593BDA5572D59FCD2717FD00" box="[1169,1289,655,680]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572D59FCD2717FD00" box="[1169,1289,655,680]" italics="true" pageId="9" pageNumber="410">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA55735C9FD32782FD00" box="[1304,1436,657,680]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55735C9FD32782FD00" box="[1304,1436,657,680]" italics="true" pageId="9" pageNumber="410">Dryosaurus</emphasis>
|
||
</taxonomicName>
|
||
and a variety of basal ornithopods (
|
||
<bibRefCitation id="875A79AC593BDA5574649FF620CCFD64" author="Huh" box="[544,722,692,717]" pageId="9" pageNumber="410" refString="Huh, M., Lee, D. G., Kim, J. K., Lim, J. D. & Godefroit, P. (2010) A new basal ornithopod dinosaur from the Upper Cretaceous of South Korea. Neues Jahrbuch fur Geologie Palaontologie, Abhandlungen, 259, 1 - 24. http: // dx. doi. org / 10.1127 / 0077 - 7749 / 2010 / 0102" type="journal article" year="2010">
|
||
Huh
|
||
<emphasis id="D1BFD84F593BDA55741C9FF7208EFD65" box="[600,656,692,717]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC593BDA5574999FF621C5FD64" author="Makovicky" box="[733,987,692,717]" pageId="9" pageNumber="410" refString="Makovicky, P. J., Kilbourne, B. M., Sadleir, R. W. & Norell, M. A. (2011) A new basal ornithopod (Dinosauria, Ornithischia) from the Late Cretaceous of Mongolia. Journal of Vertebrate Paleontology, 31, 626 - 640. http: // dx. doi. org / 10.1080 / 02724634.2011.557114" type="journal article" year="2011">
|
||
Makovicky
|
||
<emphasis id="D1BFD84F593BDA5575279FF72185FD65" box="[867,923,692,717]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
) possess relatively prominent olecranon processes that are more pronounced than that present in
|
||
<taxonomicName id="24CB7FDE593BDA5575499F9521F6FD58" box="[781,1000,727,752]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5575499F9521F6FD58" box="[781,1000,727,752]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
, but reduced in comparison with those of heterodontosaurids (Norman
|
||
<emphasis id="D1BFD84F593BDA5577BA9FBF2029FCBD" box="[510,567,764,789]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2004). The shaft of the ulna is narrow toward the medial section and slightly expanded distally. The ulna is approximately 75% of the length of the humerus. These last two features in
|
||
<taxonomicName id="24CB7FDE593BDA5576D39E06236DFCF5" box="[151,371,836,861]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5576D39E06236DFCF5" box="[151,371,836,861]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
are shared with basal ornithopods.
|
||
</paragraph>
|
||
<paragraph id="E374045D593BDA5776839E2523ABFEE8" blockId="9.[151,1436,151,2013]" lastBlockId="11.[151,1436,151,320]" lastPageId="11" lastPageNumber="412" pageId="9" pageNumber="410">
|
||
<emphasis id="D1BFD84F593BDA5576839E252303FC28" bold="true" box="[199,285,871,896]" pageId="9" pageNumber="410">Femur.</emphasis>
|
||
The proximal and distal ends of the left femur (
|
||
<figureCitation id="7BF018D8593BDA55757B9E25219DFC28" box="[831,899,871,896]" captionStart="FIGURE 8" captionStartId="10.[151,250,1729,1751]" captionTargetBox="[151,1436,193,1707]" captionTargetId="figure@10.[151,1436,193,1707]" captionTargetPageId="10" captionText="FIGURE 8. Notohypsilophodon comodorensis. Left femur (UNPSJB-PV 942 / 28), in medial (A- D), proximal (B), distal (C), posterolateral (E), lateral (F), anterior (G) and posterior (H) views. Abbreviation: fh, femoral head; gth, greater trochanter; ict, intertrochanteric cleft; lc, lateral condyle; lis, ligament sulcus; lth, lesser trochanter; mc, medial condyle; pgr, posterior groove; tub, tubercle. Note: dotted lines delimit the reconstructed femoral shaft." httpUri="https://zenodo.org/record/226301/files/figure.png" pageId="9" pageNumber="410">Fig. 8</figureCitation>
|
||
,
|
||
<tableCitation id="AE4931E6593BDA5575D59E2521FAFC28" box="[913,996,871,896]" captionStart="TABLE 1" captionStartId="7.[151,239,153,176]" captionText="TABLE 1. Selected measurements (in mm) of Notohypsilophodon comodorensis (UNPSJB-PV 942)." pageId="9" pageNumber="410">Table 1</tableCitation>
|
||
), and a portion of the shaft, have been recovered in
|
||
<taxonomicName id="24CB7FDE593BDA5577709ECE2011FC0D" box="[308,527,908,933]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5577709ECE2011FC0D" box="[308,527,908,933]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
. The preserved portion of the shaft strongly suggests that it was slender (a constricted neck is present) as in
|
||
<taxonomicName id="24CB7FDE593BDA5574099EF220E0FC6F" box="[589,766,944,967]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5574099EF220E0FC6F" box="[589,766,944,967]" italics="true" pageId="9" pageNumber="410">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
and other basal ornithopods such as
|
||
<taxonomicName id="24CB7FDE593BDA5572AE9EED278BFC60" box="[1258,1429,943,968]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572AE9EED278BFC60" box="[1258,1429,943,968]" italics="true" pageId="9" pageNumber="410">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA5576D39E962332FC45" box="[151,300,980,1005]" class="Reptilia" family="Thescelosauridae" genus="Jeholosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5576D39E962332FC45" box="[151,300,980,1005]" italics="true" pageId="9" pageNumber="410">Jeholosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA55777F9E9723F9FC44" box="[315,487,981,1004]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55777F9E9723F9FC44" box="[315,487,981,1004]" italics="true" pageId="9" pageNumber="410">Koreanosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D1BFD84F593BDA55745B9E972081FC44" box="[543,671,981,1004]" italics="true" pageId="9" pageNumber="410">Yueosaurus</emphasis>
|
||
. The femoral head is bulbous (i.e., with a well-developed spherical articular condyle), a feature shared with
|
||
<taxonomicName id="24CB7FDE593BDA5574289EBA2103FBA7" box="[620,797,1016,1039]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5574289EBA2103FBA7" box="[620,797,1016,1039]" italics="true" pageId="9" pageNumber="410">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="24CB7FDE593BDA5575149EB52635FBB8" box="[848,1067,1015,1040]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5575149EB52635FBB8" box="[848,1067,1015,1040]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
, as in
|
||
<taxonomicName id="24CB7FDE593BDA5572389EB52716FBB8" box="[1148,1288,1015,1040]" class="Reptilia" family="Hypsilophodontidae" genus="Orodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572389EB52716FBB8" box="[1148,1288,1015,1040]" italics="true" pageId="9" pageNumber="410">Orodromeus</emphasis>
|
||
</taxonomicName>
|
||
, the femoral head is oriented slightly dorsally with respect to the shaft, forming an angle greater than 100°, unlike the angle of approximately 90° found in most basal ornithopods (Norman
|
||
<emphasis id="D1BFD84F593BDA55750399032161FBF0" box="[839,895,1087,1112]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2004). On the other hand, overall, the femora of basal iguanodontians (see
|
||
<bibRefCitation id="875A79AC593BDA55778D992620D4FBD4" author="Naish" box="[457,714,1124,1149]" pageId="9" pageNumber="410" refString="Naish, D. & Martill, D. M. (2001) Ornithopod dinosaurs. In: Martill, D. M. & Naish, D. (Eds.), Dinosaurs of the Isle of Wight. The Palaeontological Association, London, pp. 60 - 132. http: // dx. doi. org / 10.1017 / s 0016756801276237" type="book chapter" year="2001">Naish & Martill 2001</bibRefCitation>
|
||
: figs. 1-3), including rhabdodontids and ankylopollexians, are more robust than those of
|
||
<taxonomicName id="24CB7FDE593BDA55778499C52085FB08" box="[448,667,1159,1184]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55778499C52085FB08" box="[448,667,1159,1184]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
, with the exception of those in dryosaurids, which are slender (e.g.,
|
||
<taxonomicName id="24CB7FDE593BDA5576D399EE2325FB6D" box="[151,315,1196,1221]" class="Reptilia" family="Camptosauridae" genus="Callovosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5576D399EE2325FB6D" box="[151,315,1196,1221]" italics="true" pageId="9" pageNumber="410">Callovosaurus</emphasis>
|
||
</taxonomicName>
|
||
[see
|
||
<bibRefCitation id="875A79AC593BDA55773099EE2080FB6C" author="Ruiz-Omenaca" box="[372,670,1196,1221]" pageId="9" pageNumber="410" refString="Ruiz-Omenaca, J. I., Pereda Suberbiola, X. & Galton, P. M. (2007) Callovosaurus leedsi, the earliest dryosaurid dinosaur (Ornithischia: Euornithopoda) from the middle Jurassic of England. In: Carpenter, K. (Ed.), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Indiana University Press, Bloomington, pp. 3 - 16." type="book chapter" year="2007">
|
||
Ruiz-Omeñaca
|
||
<emphasis id="D1BFD84F593BDA55746799EF2045FB6D" box="[547,603,1196,1221]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
],
|
||
<taxonomicName id="24CB7FDE593BDA5574F499EF212AFB6C" box="[688,820,1197,1220]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5574F499EF212AFB6C" box="[688,820,1197,1220]" italics="true" pageId="9" pageNumber="410">Dryosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE593BDA55752E99EE21EAFB6D" box="[874,1012,1196,1221]" class="Reptilia" family="Dryosauridae" genus="Valdosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55752E99EE21EAFB6D" box="[874,1012,1196,1221]" italics="true" pageId="9" pageNumber="410">Valdosaurus</emphasis>
|
||
</taxonomicName>
|
||
[see
|
||
<bibRefCitation id="875A79AC593BDA55726B99EF271EFB6C" author="Barrett" box="[1071,1280,1196,1221]" pageId="9" pageNumber="410" refString="Barrett, P. M., Butler, R. J., Twitchett, R. J. & Hutt, S. (2011) New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of Southern England. Special Papers in Palaeontology, 86, 131 - 163." type="journal article" year="2011">
|
||
Barrett
|
||
<emphasis id="D1BFD84F593BDA5572C799EF26A5FB6D" box="[1155,1211,1196,1221]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
]). In proximal view, the head is compressed anteroposteriorly and the trochanters are expanded anteroposteriorly, as in
|
||
<taxonomicName id="24CB7FDE593BDA5576D399B6237DFAA5" box="[151,355,1268,1293]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5576D399B6237DFAA5" box="[151,355,1268,1293]" italics="true" pageId="9" pageNumber="410">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
. The femoral head is separated from the trochanters by a shallow depression, the
|
||
<emphasis id="D1BFD84F593BDA55732699B62301FA98" italics="true" pageId="9" pageNumber="410">fossa trochanteris</emphasis>
|
||
. This fossa is absent in basal ornithischians such as
|
||
<emphasis id="D1BFD84F593BDA55752898552611FA98" box="[876,1039,1303,1328]" italics="true" pageId="9" pageNumber="410">Hexinlusaurus</emphasis>
|
||
and heterodontosaurids, whereas it is usually present in cerapodans (Butler
|
||
<emphasis id="D1BFD84F593BDA557423987F20BFFAFD" box="[615,673,1340,1365]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2011). Likewise, in proximal view, a well-marked convexity (i.e., “tuberosity” of
|
||
<bibRefCitation id="875A79AC593BDA55770E982223CBFAD0" author="Novas" box="[330,469,1375,1400]" pageId="9" pageNumber="410" refString="Novas, F. E. (1996) Dinosaur monophyly. Journal of Vertebrate Paleontology, 16, 723 - 741." type="journal article" year="1996">Novas 1996</bibRefCitation>
|
||
) and a shallow notch (=sulcus, sensu Butler
|
||
<emphasis id="D1BFD84F593BDA55759B98232604FAD0" box="[991,1050,1375,1400]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2011) for the
|
||
<emphasis id="D1BFD84F593BDA557287981D22F5FA34" italics="true" pageId="9" pageNumber="410">ligamentum capitis femoris</emphasis>
|
||
are present in
|
||
<taxonomicName id="24CB7FDE593BDA5577E698C62063FA35" box="[418,637,1412,1437]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5577E698C62063FA35" box="[418,637,1412,1437]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
. The greater trochanter is anteroposteriorly expanded and its dorsal surface is slightly convex, whereas the “finger-like” lesser (=anterior) trochanter is also gently convex in the same direction. The greater and lesser trochanters are unfused and separated by a marked cleft (the intertrochanteric cleft for some authors, for the m.
|
||
<emphasis id="D1BFD84F593BDA55744698AD208FF9A0" box="[514,657,1519,1544]" italics="true" pageId="9" pageNumber="410">iliofemoralis</emphasis>
|
||
insertion:
|
||
<bibRefCitation id="875A79AC593BDA55755C98AD267AF9A0" author="Hutchinson" box="[792,1124,1519,1544]" pageId="9" pageNumber="410" refString="Hutchinson, J. R. & Gatesy, S. M. (2000) Adductors, abductors, and the evolution of archosaur locomotion. Paleobiology, 26,734 - 751. http: // dx. doi. org / 10.1666 / 0094 - 8373 (2000) 026 <0734: aaateo> 2.0. co; 2" type="book" year="2000">Hutchinson & Gatesy 2000</bibRefCitation>
|
||
), a feature that occurs in
|
||
<emphasis id="D1BFD84F593BDA5576D39B56234DF985" box="[151,339,1556,1581]" italics="true" pageId="9" pageNumber="410">Gideonmantellia</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F593BDA5577249B56201DF985" box="[352,515,1556,1581]" italics="true" pageId="9" pageNumber="410">Hexinlusaurus</emphasis>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA5574559B5620A2F985" box="[529,700,1556,1581]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5574559B5620A2F985" box="[529,700,1556,1581]" italics="true" pageId="9" pageNumber="410">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA55748E9B56214BF985" box="[714,853,1556,1581]" class="Reptilia" family="Dryosauridae" genus="Valdosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55748E9B56214BF985" box="[714,853,1556,1581]" italics="true" pageId="9" pageNumber="410">Valdosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE593BDA5575CE9B562604F985" box="[906,1050,1556,1581]" class="Reptilia" family="Hypsilophodontidae" genus="Yandusaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5575CE9B562604F985" box="[906,1050,1556,1581]" italics="true" pageId="9" pageNumber="410">Yandusaurus</emphasis>
|
||
</taxonomicName>
|
||
(Norman
|
||
<emphasis id="D1BFD84F593BDA5572C89B5726DDF985" box="[1164,1219,1556,1581]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2004;
|
||
<bibRefCitation id="875A79AC593BDA5573549B5722CFF9F8" author="Barrett" pageId="9" pageNumber="410" refString="Barrett, P. M., Butler, R. J., Twitchett, R. J. & Hutt, S. (2011) New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of Southern England. Special Papers in Palaeontology, 86, 131 - 163." type="journal article" year="2011">
|
||
Barrett
|
||
<emphasis id="D1BFD84F593BDA5573209B57278AF985" box="[1380,1428,1556,1581]" italics="true" pageId="9" pageNumber="410">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
; Ruiz- Omeñaca
|
||
<emphasis id="D1BFD84F593BDA5577D99B7B23C7F9F8" box="[413,473,1591,1616]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2012). Additionally, this feature is variable in
|
||
<taxonomicName id="24CB7FDE593BDA5572449B7526B5F9F8" box="[1024,1195,1591,1616]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572449B7526B5F9F8" box="[1024,1195,1591,1616]" italics="true" pageId="9" pageNumber="410">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
(P. M. Barrett, pers. comm.,
|
||
<date id="9775229D593BDA5576B09B1E23B3F9DC" box="[244,429,1628,1653]" pageId="9" pageNumber="410" value="2013-12">December 2013</date>
|
||
). In contrast, in
|
||
<taxonomicName id="24CB7FDE593BDA55742E9B1F2105F9DC" box="[618,795,1629,1652]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55742E9B1F2105F9DC" box="[618,795,1629,1652]" italics="true" pageId="9" pageNumber="410">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA55756F9B1F21C9F9DC" box="[811,983,1629,1652]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55756F9B1F21C9F9DC" box="[811,983,1629,1652]" italics="true" pageId="9" pageNumber="410">Koreanosaurus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="D1BFD84F593BDA55725C9B1F2686F9DC" box="[1048,1176,1629,1652]" italics="true" pageId="9" pageNumber="410">Yueosaurus</emphasis>
|
||
, the greater and lesser trochanters are fused. Although in
|
||
<taxonomicName id="24CB7FDE593BDA5574069B3D215BF930" authority="Trinisaura" authorityName="Trinisaura" box="[578,837,1663,1688]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5574069B3D20A4F930" box="[578,698,1663,1688]" italics="true" pageId="9" pageNumber="410">Anabisetia</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F593BDA5574949BC3215BF930" box="[720,837,1665,1688]" italics="true" pageId="9" pageNumber="410">Trinisaura</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="875A79AC593BDA5575D59B3D2676F930" author="Coria" box="[913,1128,1663,1688]" pageId="9" pageNumber="410" refString="Coria, R. A., Moly, J. J., Reguero, M., Santillana, S. & Marenssi, S. (2013) A new ornithopod (Dinosauria; Ornithischia) from Antarctica. Cretaceous Research, 41, 186 - 193. http: // dx. doi. org / 10.1016 / j. cretres. 2012.12.004" type="journal article" year="2013">
|
||
Coria
|
||
<emphasis id="D1BFD84F593BDA55759A9BC32600F930" box="[990,1054,1663,1688]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
), a cleft separating those trochanters is present, it is narrow in comparison with that seen in
|
||
<taxonomicName id="24CB7FDE593BDA55753E9BE6264BF915" box="[890,1109,1700,1725]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55753E9BE6264BF915" box="[890,1109,1700,1725]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
. In lateral view, the tip of the lesser trochanter is set below the level of the greater trochanter, as in most basal ornithischians (
|
||
<bibRefCitation id="875A79AC593BDA5572BD9B8522CAF8AC" author="Agnolin" pageId="9" pageNumber="410" refString="Agnolin, F. L., Ezcurra, M. D., Pais, D. F. & Salisbury, W. (2010) A reappraisal of the Cretaceous non-avian dinosaur faunas from Australia and New Zealand: evidence for their Gondwanan affinities. Journal of Systematic Palaeontology, 8, 257 - 300. http: // dx. doi. org / 10.1080 / 14772011003594870" type="journal article" year="2010">
|
||
Agnolin
|
||
<emphasis id="D1BFD84F593BDA5573259B8B2782F948" box="[1377,1436,1735,1760]" italics="true" pageId="9" pageNumber="410">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). This differs, for example, from those see in
|
||
<taxonomicName id="24CB7FDE593BDA5574A29BAE2140F8AD" box="[742,862,1772,1797]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5574A29BAE2140F8AD" box="[742,862,1772,1797]" italics="true" pageId="9" pageNumber="410">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA55752A9BAE2624F8AD" box="[878,1082,1772,1797]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55752A9BAE2624F8AD" box="[878,1082,1772,1797]" italics="true" pageId="9" pageNumber="410">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA55720F9BAE26FEF8AD" box="[1099,1248,1772,1797]" class="Reptilia" family="Thescelosauridae" genus="Jeholosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA55720F9BAE26FEF8AD" box="[1099,1248,1772,1797]" italics="true" pageId="9" pageNumber="410">Jeholosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593BDA5572B49BAF2782F8AC" box="[1264,1436,1773,1796]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572B49BAF2782F8AC" box="[1264,1436,1773,1796]" italics="true" pageId="9" pageNumber="410">Koreanosaurus</emphasis>
|
||
</taxonomicName>
|
||
and dryosaurids (both trochanters are nearly to the same level, also see
|
||
<bibRefCitation id="875A79AC593BDA5575F49A4D26BBF880" author="Naish" box="[944,1189,1807,1832]" pageId="9" pageNumber="410" refString="Naish, D. & Martill, D. M. (2001) Ornithopod dinosaurs. In: Martill, D. M. & Naish, D. (Eds.), Dinosaurs of the Isle of Wight. The Palaeontological Association, London, pp. 60 - 132. http: // dx. doi. org / 10.1017 / s 0016756801276237" type="book chapter" year="2001">Naish & Martill 2001</bibRefCitation>
|
||
: fig. 1-6, plate 8). The distal end of the femur in
|
||
<taxonomicName id="24CB7FDE593BDA5577869A762080F8E5" box="[450,670,1844,1869]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5577869A762080F8E5" box="[450,670,1844,1869]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
exhibits two asymmetrical condyles, the tibial (medial) and fibular (lateral). The former is larger than the latter and the rounded articular surfaces are covered with well-developed rugosities for soft tissue insertions. The lateral condyle projects slightly laterally, like those of
|
||
<taxonomicName id="24CB7FDE593BDA5572FF9A3F2773F83C" box="[1211,1389,1917,1940]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572FF9A3F2773F83C" box="[1211,1389,1917,1940]" italics="true" pageId="9" pageNumber="410">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D1BFD84F593BDA5576D39AE32313F810" box="[151,269,1953,1976]" italics="true" pageId="9" pageNumber="410">Trinisaura</emphasis>
|
||
. Lateral to the medial condyle, a marked and slightly concave surface is present, as in
|
||
<taxonomicName id="24CB7FDE593BDA5572B59AE22782F811" box="[1265,1436,1952,1977]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572B59AE22782F811" box="[1265,1436,1952,1977]" italics="true" pageId="9" pageNumber="410">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
(i.e., the extension of the medial surface occupies all the lateral aspect of the tibial condyle).
|
||
<taxonomicName id="24CB7FDE593BDA5572E19A86279FF875" box="[1189,1409,1988,2013]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="410" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593BDA5572E19A86279FF875" box="[1189,1409,1988,2013]" italics="true" pageId="9" pageNumber="410">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
possesses a shallow anterior (extensor) groove as in basal euornithopods (Norman
|
||
<emphasis id="D1BFD84F5938DA5672189A192689F8DB" box="[1116,1175,1882,1907]" italics="true" pageId="10" pageNumber="411">et al.</emphasis>
|
||
2004). This groove is absent in some other taxa (e.g.,
|
||
<taxonomicName id="24CB7FDE5938DA56740E9A3D20E5F83E" box="[586,763,1919,1942]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="411" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5938DA56740E9A3D20E5F83E" box="[586,763,1919,1942]" italics="true" pageId="10" pageNumber="411">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
) but well-developed in
|
||
<taxonomicName id="24CB7FDE5938DA5672789A3C26AAF83F" box="[1084,1204,1918,1943]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="411" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5938DA5672789A3C26AAF83F" box="[1084,1204,1918,1943]" italics="true" pageId="10" pageNumber="411">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5938DA56728A9A3C2782F83F" box="[1230,1436,1918,1943]" class="Reptilia" family="Camptosauridae" genus="Muttaburrasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="411" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5938DA56728A9A3C2782F83F" box="[1230,1436,1918,1943]" italics="true" pageId="10" pageNumber="411">Muttaburrasaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="875A79AC5938DA5676DA9AE023F5F812" author="Bartholomai" box="[158,491,1954,1979]" pageId="10" pageNumber="411" refString="Bartholomai, A. & Molnar, R. E. (1981) Muttaburrasaurus, a new iguanodontid (Ornithischia: Ornithopoda) dinosaur from the Lower Cretaceous of Queensland. Memory Queensland Museum, 20, 319 - 349." type="journal article" year="1981">Bartholomai & Molnar 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="875A79AC5938DA5677BD9AE02093F813" author="Molnar" box="[505,653,1954,1979]" pageId="10" pageNumber="411" refString="Molnar, R. E. (1996) Observations on the Australian ornithopod dinosaur Muttaburrasaurus. Memoirs of the Queensland Museum, 39, 639 - 652." type="journal article" year="1996">Molnar 1996</bibRefCitation>
|
||
also see
|
||
<bibRefCitation id="875A79AC5938DA5674BE9AE021C2F812" author="Agnolin" box="[762,988,1954,1979]" pageId="10" pageNumber="411" refString="Agnolin, F. L., Ezcurra, M. D., Pais, D. F. & Salisbury, W. (2010) A reappraisal of the Cretaceous non-avian dinosaur faunas from Australia and New Zealand: evidence for their Gondwanan affinities. Journal of Systematic Palaeontology, 8, 257 - 300. http: // dx. doi. org / 10.1080 / 14772011003594870" type="journal article" year="2010">
|
||
Agnolin
|
||
<emphasis id="D1BFD84F5938DA56751A9AE12187F813" box="[862,921,1954,1979]" italics="true" pageId="10" pageNumber="411">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
) and
|
||
<emphasis id="D1BFD84F5938DA56725A9AE1268DF812" box="[1054,1171,1955,1978]" italics="true" pageId="10" pageNumber="411">Trinisaura</emphasis>
|
||
. The posterior (flexor) intercondylar groove is comparable to those observed in basal ornithopods (Norman
|
||
<emphasis id="D1BFD84F5938DA5672169A852695F877" box="[1106,1163,1990,2015]" italics="true" pageId="10" pageNumber="411">et al.</emphasis>
|
||
2004), whereas in basal non-hadrosaurid iguanodontians it is wider. Lastly, the distal end of the “Victoria femur
|
||
<typeStatus id="3C70BAFF5939DA57722F9DDA2685FF18" box="[1131,1179,152,176]" pageId="11" pageNumber="412">type</typeStatus>
|
||
I” (
|
||
<bibRefCitation id="875A79AC5939DA5772829DD5278AFF18" author="Rich" box="[1222,1428,151,176]" pageId="11" pageNumber="412" refString="Rich, T. H. & Vickers-Rich, P. (1999) The Hypsilophodontidae from southeastern Australia. In: Tomida, Y., Rich, T. H. & Vickers-Rich P. (Eds.), Proceedings of the Second Gondwanan Dinosaur Symposium National Science Museum Monographs, Tokyo, pp. 167 - 180." type="book chapter" year="1999">Rich & Rich 1999</bibRefCitation>
|
||
) is similar in morphology to that of
|
||
<taxonomicName id="24CB7FDE5939DA5774729DFE210CFF7D" box="[566,786,188,213]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="412" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5939DA5774729DFE210CFF7D" box="[566,786,188,213]" italics="true" pageId="11" pageNumber="412">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
, in having a laterally placed fibular condyle, transverse compression and a medial condyle that is markedly larger than the lateral (
|
||
<bibRefCitation id="875A79AC5939DA5775B69D9D26C7FF50" author="Agnolin" box="[1010,1241,223,248]" pageId="11" pageNumber="412" refString="Agnolin, F. L., Ezcurra, M. D., Pais, D. F. & Salisbury, W. (2010) A reappraisal of the Cretaceous non-avian dinosaur faunas from Australia and New Zealand: evidence for their Gondwanan affinities. Journal of Systematic Palaeontology, 8, 257 - 300. http: // dx. doi. org / 10.1080 / 14772011003594870" type="journal article" year="2010">
|
||
Agnolin
|
||
<emphasis id="D1BFD84F5939DA57721C9DA3268DFF50" box="[1112,1171,223,248]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). However, these features probably are due to taphonomic aspects rather than genuine anatomical characteristics (P. M. Barrett, pers. comm.,
|
||
<date id="9775229D5939DA5776B69C6523B7FEE8" box="[242,425,295,320]" pageId="11" pageNumber="412" value="2013-12">December 2013</date>
|
||
).
|
||
</paragraph>
|
||
<caption id="B7B454D55938DA5676D39B832172F89A" httpUri="https://zenodo.org/record/226301/files/figure.png" pageId="10" pageNumber="411" targetBox="[151,1436,193,1707]" targetPageId="10">
|
||
<paragraph id="E374045D5938DA5676D39B832172F89A" blockId="10.[151,1436,1729,1842]" pageId="10" pageNumber="411">
|
||
<emphasis id="D1BFD84F5938DA5676D39B83230AF97E" bold="true" box="[151,276,1729,1751]" pageId="10" pageNumber="411">FIGURE 8.</emphasis>
|
||
<taxonomicName id="24CB7FDE5938DA56775F9B83206BF97F" box="[283,629,1729,1751]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="comodorensis">
|
||
<emphasis id="D1BFD84F5938DA56775F9B83206BF97F" box="[283,629,1729,1751]" italics="true" pageId="10" pageNumber="411">Notohypsilophodon comodorensis</emphasis>
|
||
</taxonomicName>
|
||
. Left femur (UNPSJB-PV 942/28), in medial (A- D), proximal (B), distal (C), posterolateral (E), lateral (F), anterior (G) and posterior (H) views. Abbreviation: fh, femoral head; gth, greater trochanter; ict, intertrochanteric cleft; lc, lateral condyle; lis, ligament sulcus; lth, lesser trochanter; mc, medial condyle; pgr, posterior groove; tub, tubercle. Note: dotted lines delimit the reconstructed femoral shaft.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B7B454D55939DA5776D39BE2278AF899" httpUri="https://zenodo.org/record/226302/files/figure.png" pageId="11" pageNumber="412" targetBox="[151,1436,373,1674]" targetPageId="11">
|
||
<paragraph id="E374045D5939DA5776D39BE2278AF899" blockId="11.[151,1436,1696,1841]" pageId="11" pageNumber="412">
|
||
<emphasis id="D1BFD84F5939DA5776D39BE2230BF91D" bold="true" box="[151,277,1696,1718]" pageId="11" pageNumber="412">FIGURE 9.</emphasis>
|
||
<taxonomicName id="24CB7FDE5939DA5777599BE22066F91E" box="[285,632,1696,1718]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="comodorensis">
|
||
<emphasis id="D1BFD84F5939DA5777599BE22066F91E" box="[285,632,1696,1718]" italics="true" pageId="11" pageNumber="412">Notohypsilophodon comodorensis</emphasis>
|
||
</taxonomicName>
|
||
. Left tibia (UNPSJB-PV 942/29), in medial (A), proximal (B) and distal (C) views. Right tibia (UNPSJB-PV 942/30), in distal (D) view. Left fibula (UNPSJB-PV 942/32), in medial (E) view. Right astragalus (UNPSJB-PV 942/33), in posterior (F) and anterior (G) views. Left calcaneum (UNPSJB-PV 942/34), in lateral (H) and medial (I) views. Abbreviation: ac, astragalar contact; cc, cnemial crest; dbl, distal bulge; dpe, distal perforation; fc, fibular condyle; fs, fibular surface; ic, inner condyle; it, incisura tibialis; ppr, proximal process; prb, proximal bulge; ts, tibial surface.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E374045D5939DA5076839A1A2683FD00" blockId="11.[151,1436,1880,2013]" lastBlockId="12.[151,1437,151,2013]" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412">
|
||
<emphasis id="D1BFD84F5939DA5776839A1A2313F8D9" bold="true" box="[199,269,1880,1905]" pageId="11" pageNumber="412">Tibia.</emphasis>
|
||
The straight right tibia (
|
||
<figureCitation id="7BF018D85939DA5774689A1A206EF8D9" box="[556,624,1880,1905]" captionStart="FIGURE 9" captionStartId="11.[151,250,1696,1718]" captionTargetBox="[151,1436,373,1674]" captionTargetId="figure@11.[151,1436,373,1675]" captionTargetPageId="11" captionText="FIGURE 9. Notohypsilophodon comodorensis. Left tibia (UNPSJB-PV 942 / 29), in medial (A), proximal (B) and distal (C) views. Right tibia (UNPSJB-PV 942 / 30), in distal (D) view. Left fibula (UNPSJB-PV 942 / 32), in medial (E) view. Right astragalus (UNPSJB-PV 942 / 33), in posterior (F) and anterior (G) views. Left calcaneum (UNPSJB-PV 942 / 34), in lateral (H) and medial (I) views. Abbreviation: ac, astragalar contact; cc, cnemial crest; dbl, distal bulge; dpe, distal perforation; fc, fibular condyle; fs, fibular surface; ic, inner condyle; it, incisura tibialis; ppr, proximal process; prb, proximal bulge; ts, tibial surface." httpUri="https://zenodo.org/record/226302/files/figure.png" pageId="11" pageNumber="412">Fig. 9</figureCitation>
|
||
A–B–C–D;
|
||
<tableCitation id="AE4931E65939DA5774BE9A1A2150F8D9" box="[762,846,1880,1905]" captionStart="TABLE 1" captionStartId="7.[151,239,153,176]" captionText="TABLE 1. Selected measurements (in mm) of Notohypsilophodon comodorensis (UNPSJB-PV 942)." pageId="11" pageNumber="412">Table 1</tableCitation>
|
||
) of
|
||
<taxonomicName id="24CB7FDE5939DA5775C09A1A267EF8D9" box="[900,1120,1880,1905]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="412" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5939DA5775C09A1A267EF8D9" box="[900,1120,1880,1905]" italics="true" pageId="11" pageNumber="412">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
is slender, a feature shared with a variety of ornithopods (e.g.,
|
||
<taxonomicName id="24CB7FDE5939DA5774789A3E20AAF83D" box="[572,692,1916,1941]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="412" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5939DA5774789A3E20AAF83D" box="[572,692,1916,1941]" italics="true" pageId="11" pageNumber="412">Anabisetia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5939DA5774819A3F265BF83D" authority="Gideonmantellia" authorityName="Gideonmantellia" box="[709,1093,1916,1941]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="412" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5939DA5774819A3F2168F83C" box="[709,886,1917,1940]" italics="true" pageId="11" pageNumber="412">Gasparinisaura</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5939DA5775CD9A3E265BF83D" box="[905,1093,1916,1941]" italics="true" pageId="11" pageNumber="412">Gideonmantellia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE5939DA5772139A3E278CF83C" authority="Yueosaurus" authorityName="Yueosaurus" box="[1111,1426,1916,1941]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="412" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5939DA5772139A3E271CF83D" box="[1111,1282,1916,1941]" italics="true" pageId="11" pageNumber="412">Hypsilophodon</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F5939DA5773519A3F278CF83C" box="[1301,1426,1917,1940]" italics="true" pageId="11" pageNumber="412">Yueosaurus</emphasis>
|
||
</taxonomicName>
|
||
, among others). The proximal end is anteroposteriorly expanded as in
|
||
<taxonomicName id="24CB7FDE5939DA5775E89AE22666F811" box="[940,1144,1952,1977]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="412" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5939DA5775E89AE22666F811" box="[940,1144,1952,1977]" italics="true" pageId="11" pageNumber="412">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE5939DA5772F49AE3277FF810" box="[1200,1377,1953,1976]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="412" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F5939DA5772F49AE3277FF810" box="[1200,1377,1953,1976]" italics="true" pageId="11" pageNumber="412">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
. The cnemial crest is modestly developed and slightly laterally and dorsally projected as is usual in basal ornithopods (
|
||
<bibRefCitation id="875A79AC593EDA5076DA9DD52344FF18" author="Huh" box="[158,346,151,176]" pageId="12" pageNumber="413" refString="Huh, M., Lee, D. G., Kim, J. K., Lim, J. D. & Godefroit, P. (2010) A new basal ornithopod dinosaur from the Upper Cretaceous of South Korea. Neues Jahrbuch fur Geologie Palaontologie, Abhandlungen, 259, 1 - 24. http: // dx. doi. org / 10.1127 / 0077 - 7749 / 2010 / 0102" type="journal article" year="2010">
|
||
Huh
|
||
<emphasis id="D1BFD84F593EDA50769E9DDB230AFF18" box="[218,276,151,176]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). The cnemial crest is separated from the fibular condyle by a groove (the
|
||
<emphasis id="D1BFD84F593EDA5072F89DDB2771FF18" box="[1212,1391,151,176]" italics="true" pageId="12" pageNumber="413">incisura tibialis</emphasis>
|
||
see Butler
|
||
<emphasis id="D1BFD84F593EDA5076A19DFF2300FF7D" box="[229,286,188,213]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
|
||
2011) as in
|
||
<taxonomicName id="24CB7FDE593EDA5077E89DFF2046FF7C" box="[428,600,189,212]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5077E89DFF2046FF7C" box="[428,600,189,212]" italics="true" pageId="12" pageNumber="413">Koreanosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593EDA5074239DFE20E2FF7D" box="[615,764,188,213]" class="Reptilia" family="Thescelosauridae" genus="Jeholosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5074239DFE20E2FF7D" box="[615,764,188,213]" italics="true" pageId="12" pageNumber="413">Jeholosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE593EDA5075719DFE21FEFF7D" box="[821,992,188,213]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5075719DFE21FEFF7D" box="[821,992,188,213]" italics="true" pageId="12" pageNumber="413">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
. Both condyles of
|
||
<taxonomicName id="24CB7FDE593EDA5072FE9DFE278BFF7D" box="[1210,1429,188,213]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5072FE9DFE278BFF7D" box="[1210,1429,188,213]" italics="true" pageId="12" pageNumber="413">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
, the rounded inner and fibular are anteroposteriorly expanded, particularly the latter. The proximal expansion of the condyles is more strongly developed than in basal forms such as
|
||
<emphasis id="D1BFD84F593EDA5075399C4721A4FEB4" box="[893,954,261,284]" italics="true" pageId="12" pageNumber="413">Haya</emphasis>
|
||
and
|
||
<taxonomicName id="24CB7FDE593EDA5075B79C472681FEB4" box="[1011,1183,261,284]" class="Reptilia" family="Dromaeosauridae" genus="Koreanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5075B79C472681FEB4" box="[1011,1183,261,284]" italics="true" pageId="12" pageNumber="413">Koreanosaurus</emphasis>
|
||
</taxonomicName>
|
||
. Proximally, the tibial surface exhibits a well-developed bulge. The distal end of the tibia in
|
||
<taxonomicName id="24CB7FDE593EDA50759F9C6526A8FEE8" box="[987,1206,295,320]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA50759F9C6526A8FEE8" box="[987,1206,295,320]" italics="true" pageId="12" pageNumber="413">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
, is subtriangular in outline as in
|
||
<taxonomicName id="24CB7FDE593EDA5077729C0F23A4FECC" box="[310,442,333,356]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5077729C0F23A4FECC" box="[310,442,333,356]" italics="true" pageId="12" pageNumber="413">Dryosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D1BFD84F593EDA5077B89C0F2062FECC" box="[508,636,333,356]" italics="true" pageId="12" pageNumber="413">Yueosaurus</emphasis>
|
||
. Additionally, it is transversely expanded as in basal ornithischians (
|
||
<bibRefCitation id="875A79AC593EDA5076DA9C2D23B3FE20" author="Makovicky" box="[158,429,367,392]" pageId="12" pageNumber="413" refString="Makovicky, P. J., Kilbourne, B. M., Sadleir, R. W. & Norell, M. A. (2011) A new basal ornithopod (Dinosauria, Ornithischia) from the Late Cretaceous of Mongolia. Journal of Vertebrate Paleontology, 31, 626 - 640. http: // dx. doi. org / 10.1080 / 02724634.2011.557114" type="journal article" year="2011">
|
||
Makovicky
|
||
<emphasis id="D1BFD84F593EDA50776F9C332379FE20" box="[299,359,367,392]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
). The outer malleolus is larger than the inner malleolus, a condition reported in basal ornithopods (
|
||
<bibRefCitation id="875A79AC593EDA50776A9CD623E0FE04" author="Zheng" box="[302,510,404,429]" pageId="12" pageNumber="413" refString="Zheng, W. J., Jin, X. S., Shibata, M., Azuma, Y. & Yu, F. M. (2012) A new ornithischian dinosaur from the Cretaceous Liangtoutang Formation of Tiantai, Zhejiang Province, China. Cretaceous Research, 34, 208 - 219. http: // dx. doi. org / 10.1016 / j. cretres. 2011.11.001" type="journal article" year="2012">
|
||
Zheng
|
||
<emphasis id="D1BFD84F593EDA5077C49CD723A5FE05" box="[384,443,404,429]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
) and unlike the subequally-sized malleoli in
|
||
<taxonomicName id="24CB7FDE593EDA5072439CD726A6FE04" box="[1031,1208,405,428]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5072439CD726A6FE04" box="[1031,1208,405,428]" italics="true" pageId="12" pageNumber="413">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
. With regard to the distal end, the subtriangular shape, prominently extended anteromedial flange, presence of a shallow lateral groove, and the similarly-shaped astragalar articular surface, are features that
|
||
<taxonomicName id="24CB7FDE593EDA5072759C9E2712FE5D" box="[1073,1292,476,501]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5072759C9E2712FE5D" box="[1073,1292,476,501]" italics="true" pageId="12" pageNumber="413">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="24CB7FDE593EDA5076D39F432345FDB0" box="[151,347,511,536]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="altus">
|
||
<emphasis id="D1BFD84F593EDA5076D39F432345FDB0" box="[151,347,511,536]" italics="true" pageId="12" pageNumber="413">Dryosaurus altus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="875A79AC593EDA5077DD9F422038FDB0" author="Novas" box="[409,550,511,536]" pageId="12" pageNumber="413" refString="Novas, F. E. (1996) Dinosaur monophyly. Journal of Vertebrate Paleontology, 16, 723 - 741." type="journal article" year="1996">Novas 1996</bibRefCitation>
|
||
: fig.
|
||
<date id="9775229D593EDA5074269F422062FDB0" box="[610,636,512,536]" pageId="12" pageNumber="413">4I</date>
|
||
). Also in distal view, the articular surface of the tibia exhibits a welldeveloped perforation similar to that drawn in
|
||
<taxonomicName id="24CB7FDE593EDA5074909F672146FD94" box="[724,856,549,572]" class="Reptilia" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5074909F672146FD94" box="[724,856,549,572]" italics="true" pageId="12" pageNumber="413">Dryosaurus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="875A79AC593EDA5075E69F67262AFD95" author="Novas" box="[930,1076,548,573]" pageId="12" pageNumber="413" refString="Novas, F. E. (1996) Dinosaur monophyly. Journal of Vertebrate Paleontology, 16, 723 - 741." type="journal article" year="1996">Novas 1996</bibRefCitation>
|
||
: fig.
|
||
<date id="9775229D593EDA50723E9F66268AFD94" box="[1146,1172,548,572]" pageId="12" pageNumber="413">4I</date>
|
||
). This sub-oval distal perforation, which could be related with insertion of soft tissue (probably variable ossification of the cartilage capping the tibia), is placed closer to the external margin of the inner malleolus than the outer malleolus. Likewise, its counterpart, in the same position, exhibits a well-developed distal bulge (
|
||
<figureCitation id="7BF018D8593EDA5075A99FCD2631FD00" box="[1005,1071,655,680]" captionStart="FIGURE 9" captionStartId="11.[151,250,1696,1718]" captionTargetBox="[151,1436,373,1674]" captionTargetId="figure@11.[151,1436,373,1675]" captionTargetPageId="11" captionText="FIGURE 9. Notohypsilophodon comodorensis. Left tibia (UNPSJB-PV 942 / 29), in medial (A), proximal (B) and distal (C) views. Right tibia (UNPSJB-PV 942 / 30), in distal (D) view. Left fibula (UNPSJB-PV 942 / 32), in medial (E) view. Right astragalus (UNPSJB-PV 942 / 33), in posterior (F) and anterior (G) views. Left calcaneum (UNPSJB-PV 942 / 34), in lateral (H) and medial (I) views. Abbreviation: ac, astragalar contact; cc, cnemial crest; dbl, distal bulge; dpe, distal perforation; fc, fibular condyle; fs, fibular surface; ic, inner condyle; it, incisura tibialis; ppr, proximal process; prb, proximal bulge; ts, tibial surface." httpUri="https://zenodo.org/record/226302/files/figure.png" pageId="12" pageNumber="413">Fig. 9</figureCitation>
|
||
C and D).
|
||
</paragraph>
|
||
<paragraph id="E374045D593EDA5076839FF62783FC90" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="413">
|
||
<emphasis id="D1BFD84F593EDA5076839FF62302FD65" bold="true" box="[199,284,692,717]" pageId="12" pageNumber="413">Fibula.</emphasis>
|
||
The left fibula (
|
||
<figureCitation id="7BF018D8593EDA5077AF9FF6202DFD64" box="[491,563,692,717]" captionStart="FIGURE 9" captionStartId="11.[151,250,1696,1718]" captionTargetBox="[151,1436,373,1674]" captionTargetId="figure@11.[151,1436,373,1675]" captionTargetPageId="11" captionText="FIGURE 9. Notohypsilophodon comodorensis. Left tibia (UNPSJB-PV 942 / 29), in medial (A), proximal (B) and distal (C) views. Right tibia (UNPSJB-PV 942 / 30), in distal (D) view. Left fibula (UNPSJB-PV 942 / 32), in medial (E) view. Right astragalus (UNPSJB-PV 942 / 33), in posterior (F) and anterior (G) views. Left calcaneum (UNPSJB-PV 942 / 34), in lateral (H) and medial (I) views. Abbreviation: ac, astragalar contact; cc, cnemial crest; dbl, distal bulge; dpe, distal perforation; fc, fibular condyle; fs, fibular surface; ic, inner condyle; it, incisura tibialis; ppr, proximal process; prb, proximal bulge; ts, tibial surface." httpUri="https://zenodo.org/record/226302/files/figure.png" pageId="12" pageNumber="413">Fig. 9</figureCitation>
|
||
E) of
|
||
<taxonomicName id="24CB7FDE593EDA5074C49FF62142FD65" box="[640,860,692,717]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5074C49FF62142FD65" box="[640,860,692,717]" italics="true" pageId="12" pageNumber="413">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
is slender. Its proximal end is anteroposteriorly expanded and its surface is slightly concave. The shaft narrows somewhat to the mid-length of the bone. At this point, the shaft exhibits a thin well-developed flange (flange on medial surface of fibula according to Butler
|
||
<emphasis id="D1BFD84F593EDA5073279FBF2782FCBD" box="[1379,1436,764,789]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
|
||
2011) that extends medially. This flange reaches the distal end of the fibula. The distal end is moderately expanded.
|
||
</paragraph>
|
||
<paragraph id="E374045D593EDA5076839E062136FC44" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="413">
|
||
<emphasis id="D1BFD84F593EDA5076839E062351FCF5" bold="true" box="[199,335,836,861]" pageId="12" pageNumber="413">Astragalus.</emphasis>
|
||
The partially preserved right astragalus (
|
||
<figureCitation id="7BF018D8593EDA50755A9E062141FCF4" box="[798,863,836,861]" captionStart="FIGURE 9" captionStartId="11.[151,250,1696,1718]" captionTargetBox="[151,1436,373,1674]" captionTargetId="figure@11.[151,1436,373,1675]" captionTargetPageId="11" captionText="FIGURE 9. Notohypsilophodon comodorensis. Left tibia (UNPSJB-PV 942 / 29), in medial (A), proximal (B) and distal (C) views. Right tibia (UNPSJB-PV 942 / 30), in distal (D) view. Left fibula (UNPSJB-PV 942 / 32), in medial (E) view. Right astragalus (UNPSJB-PV 942 / 33), in posterior (F) and anterior (G) views. Left calcaneum (UNPSJB-PV 942 / 34), in lateral (H) and medial (I) views. Abbreviation: ac, astragalar contact; cc, cnemial crest; dbl, distal bulge; dpe, distal perforation; fc, fibular condyle; fs, fibular surface; ic, inner condyle; it, incisura tibialis; ppr, proximal process; prb, proximal bulge; ts, tibial surface." httpUri="https://zenodo.org/record/226302/files/figure.png" pageId="12" pageNumber="413">Fig. 9</figureCitation>
|
||
F–G) has a sub-rectangular shape. In distal view, it is convex, whereas in dorsal view, it is concave and divided into two articular surfaces divided by a ridge. The surface that received the distal end of tibia is wide, whereas the laterally placed surface for contact with the calcaneum is small. The ascending process is sub-triangular in shape and it is moderately developed, as also described in other basal ornithopods (Norman
|
||
<emphasis id="D1BFD84F593EDA5074E49E9720C6FC45" box="[672,728,980,1005]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
|
||
2004).
|
||
</paragraph>
|
||
<paragraph id="E374045D593EDA5076839EB52019F93F" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="413">
|
||
<emphasis id="D1BFD84F593EDA5076839EB5234BFBB8" bold="true" box="[199,341,1015,1040]" pageId="12" pageNumber="413">Calcaneum.</emphasis>
|
||
The left calcaneum (
|
||
<figureCitation id="7BF018D8593EDA5074239EB520AFFBB8" box="[615,689,1015,1040]" captionStart="FIGURE 9" captionStartId="11.[151,250,1696,1718]" captionTargetBox="[151,1436,373,1674]" captionTargetId="figure@11.[151,1436,373,1675]" captionTargetPageId="11" captionText="FIGURE 9. Notohypsilophodon comodorensis. Left tibia (UNPSJB-PV 942 / 29), in medial (A), proximal (B) and distal (C) views. Right tibia (UNPSJB-PV 942 / 30), in distal (D) view. Left fibula (UNPSJB-PV 942 / 32), in medial (E) view. Right astragalus (UNPSJB-PV 942 / 33), in posterior (F) and anterior (G) views. Left calcaneum (UNPSJB-PV 942 / 34), in lateral (H) and medial (I) views. Abbreviation: ac, astragalar contact; cc, cnemial crest; dbl, distal bulge; dpe, distal perforation; fc, fibular condyle; fs, fibular surface; ic, inner condyle; it, incisura tibialis; ppr, proximal process; prb, proximal bulge; ts, tibial surface." httpUri="https://zenodo.org/record/226302/files/figure.png" pageId="12" pageNumber="413">Fig. 9</figureCitation>
|
||
H–I) is semicircular in shape, slightly concave laterally, and mediolaterally compressed. The articular surfaces for contact with the distal ends of tibia and fibula are divided by a moderately developed and partially oblique proximal projection. The fibular and tibial articular surfaces are concave. The tibial articular surface is open medially. In contrast, the fibular surface is delimited by a thin lip. The medial surface of the calcaneum displays a well-developed astragalar process or contact, as well as a medial groove. Distally, this element exhibits well-developed rugosities indicating soft tissue insertions. The semicircular shape is shared with other ornithopods including
|
||
<taxonomicName id="24CB7FDE593EDA5074A39992260EFB4F" authority="Cumnoria" authorityName="Cumnoria" box="[743,1040,1232,1255]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5074A399922194FB4F" box="[743,906,1232,1255]" italics="true" pageId="12" pageNumber="413">Camptosaurus</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F593EDA5075DA9992260EFB4F" box="[926,1040,1232,1255]" italics="true" pageId="12" pageNumber="413">Cumnoria</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="24CB7FDE593EDA507260999226CBFB4F" box="[1060,1237,1232,1255]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA507260999226CBFB4F" box="[1060,1237,1232,1255]" italics="true" pageId="12" pageNumber="413">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593EDA5072AE998D278BFB40" box="[1258,1429,1231,1256]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5072AE998D278BFB40" box="[1258,1429,1231,1256]" italics="true" pageId="12" pageNumber="413">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593EDA5076D399B6230EFAA5" box="[151,272,1268,1293]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5076D399B6230EFAA5" box="[151,272,1268,1293]" italics="true" pageId="12" pageNumber="413">Iguanodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593EDA50776499B723DCFAA4" box="[288,450,1269,1292]" class="Reptilia" family="Iguanodontidae" genus="Ouranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA50776499B723DCFAA4" box="[288,450,1269,1292]" italics="true" pageId="12" pageNumber="413">Ouranosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593EDA50779599B62042FAA5" box="[465,604,1268,1293]" class="Reptilia" family="Dryosauridae" genus="Valdosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA50779599B62042FAA5" box="[465,604,1268,1293]" italics="true" pageId="12" pageNumber="413">Valdosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593EDA50742899B620C9FAA5" box="[620,727,1268,1293]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA50742899B620C9FAA5" box="[620,727,1268,1293]" italics="true" pageId="12" pageNumber="413">Zalmoxes</emphasis>
|
||
</taxonomicName>
|
||
, and UNPSJB-PV 960, among others. Likewise, the proximal ridge that separates the fibular and tibial articular surfaces, the well-developed astragalar process, and the medial groove are also anatomical characteristics shared with these other ornithopods. However, by contrast, the medial surface of the calcaneum in
|
||
<taxonomicName id="24CB7FDE593EDA507795981D2083FAD0" box="[465,669,1375,1400]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA507795981D2083FAD0" box="[465,669,1375,1400]" italics="true" pageId="12" pageNumber="413">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
is flat, lacking the groove and process. The tibial articular surface in
|
||
<taxonomicName id="24CB7FDE593EDA5076D398C6236DFA35" box="[151,371,1412,1437]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5076D398C6236DFA35" box="[151,371,1412,1437]" italics="true" pageId="12" pageNumber="413">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
is concave in lateral view and it may have distally enveloped the anterior portion of the tibial lateral malleolus, this condition, when in natural articulation, is comparable to those seen in
|
||
<taxonomicName id="24CB7FDE593EDA5072A098EA278BFA17" box="[1252,1429,1448,1471]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5072A098EA278BFA17" box="[1252,1429,1448,1471]" italics="true" pageId="12" pageNumber="413">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593EDA5076D3988E235CFA4D" box="[151,322,1484,1509]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5076D3988E235CFA4D" box="[151,322,1484,1509]" italics="true" pageId="12" pageNumber="413">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593EDA507716988F23EAFA4C" box="[338,500,1485,1508]" class="Reptilia" family="Iguanodontidae" genus="Ouranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA507716988F23EAFA4C" box="[338,500,1485,1508]" italics="true" pageId="12" pageNumber="413">Ouranosaurus</emphasis>
|
||
</taxonomicName>
|
||
and UNPSJB-PV 960. In
|
||
<taxonomicName id="24CB7FDE593EDA507560988E261EFA4D" box="[804,1024,1484,1509]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA507560988E261EFA4D" box="[804,1024,1484,1509]" italics="true" pageId="12" pageNumber="413">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
there is a marked difference in size between the fibular and tibial articular surfaces, the former being smaller than the latter in lateral view. This feature is shared with
|
||
<taxonomicName id="24CB7FDE593EDA5077079B5723EBF984" box="[323,501,1557,1580]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5077079B5723EBF984" box="[323,501,1557,1580]" italics="true" pageId="12" pageNumber="413">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
and UNPSJB-PV 960, but in
|
||
<taxonomicName id="24CB7FDE593EDA5075119B5726B3F985" authority="Hexinlusaurus" authorityName="Hexinlusaurus" box="[853,1197,1556,1581]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5075119B5721E6F984" box="[853,1016,1557,1580]" italics="true" pageId="12" pageNumber="413">Camptosaurus</emphasis>
|
||
,
|
||
<emphasis id="D1BFD84F593EDA50724E9B5626B3F985" box="[1034,1197,1556,1581]" italics="true" pageId="12" pageNumber="413">Hexinlusaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="24CB7FDE593EDA5072FA9B562777F985" box="[1214,1385,1556,1581]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5072FA9B562777F985" box="[1214,1385,1556,1581]" italics="true" pageId="12" pageNumber="413">Hypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE593EDA5076D39B75231DF9F8" box="[151,259,1591,1616]" class="Reptilia" family="Rhabdodontidae" genus="Zalmoxes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5076D39B75231DF9F8" box="[151,259,1591,1616]" italics="true" pageId="12" pageNumber="413">Zalmoxes</emphasis>
|
||
</taxonomicName>
|
||
these fossae are subequal in size in the same view. Likewise, in iguanodontians such as
|
||
<taxonomicName id="24CB7FDE593EDA5072B69B752775F9F8" box="[1266,1387,1591,1616]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5072B69B752775F9F8" box="[1266,1387,1591,1616]" italics="true" pageId="12" pageNumber="413">Iguanodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24CB7FDE593EDA5076D39B1E235FF9DD" box="[151,321,1628,1653]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5076D39B1E235FF9DD" box="[151,321,1628,1653]" italics="true" pageId="12" pageNumber="413">Mantellisaurus</emphasis>
|
||
</taxonomicName>
|
||
, the fibular articular surface is clearly longer than the tibial (see
|
||
<bibRefCitation id="875A79AC593EDA5072789B1E270DF9DC" author="Ibiricu" box="[1084,1299,1628,1653]" pageId="12" pageNumber="413" refString="Ibiricu, L. M., Martinez, R. D., Lamanna, M. C., Casal, G. A., Luna, M., Harris, J. D. & Lacovara, K. J. (2010) A medium-sized ornithopod (Dinosauria: Ornithischia) from the Upper Cretaceous Bajo Barreal Formation of Lago Colhue Huapi, southern Chubut Province, Argentina. Annals of the Carnegie Museum, 79, 39 - 50. http: // dx. doi. org / 10.2992 / 007.079.0103" type="journal article" year="2010">
|
||
Ibiricu
|
||
<emphasis id="D1BFD84F593EDA5072D59B1F26D3F9DD" box="[1169,1229,1628,1653]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
), unlike the condition in
|
||
<taxonomicName id="24CB7FDE593EDA5077609B3D23E1F930" box="[292,511,1663,1688]" class="Sauropsida" family="Hypsilophodontidae" genus="Notohypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5077609B3D23E1F930" box="[292,511,1663,1688]" italics="true" pageId="12" pageNumber="413">Notohypsilophodon</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E374045D593EDA5176839BE6201DFF50" blockId="12.[151,1437,151,2013]" lastBlockId="13.[151,1436,151,429]" lastPageId="13" lastPageNumber="414" pageId="12" pageNumber="413">
|
||
<emphasis id="D1BFD84F593EDA5076839BE623D8F915" bold="true" box="[199,454,1700,1725]" pageId="12" pageNumber="413">Non-ungual phalanx.</emphasis>
|
||
One of the best-preserved proximal non-ungual phalanges (
|
||
<figureCitation id="7BF018D8593EDA5072339BE626D6F914" box="[1143,1224,1700,1725]" captionStart="FIGURE 10" captionStartId="13.[151,250,1327,1349]" captionTargetBox="[340,1247,483,1305]" captionTargetId="figure@13.[340,1247,483,1305]" captionTargetPageId="13" captionText="FIGURE 10. Notohypsilophodon comodorensis. Non-ungual phalanx (UNPSJB-PV 942 / 35), in medial (A), dorsal (B) and proximal (C) views. Non-ungual phalanx (UNPSJB-PV 942 / 37), in medial (B) view. Non-ungual phalanx (UNPSJB-PV 942 / 39), in dorsal (C) view. Ungual phalanges (UNPSJB-PV 942 / 45,46,47), in dorsal view. Abbreviation: clp, collateral ligament pit; ugr, ungula groove." httpUri="https://zenodo.org/record/226303/files/figure.png" pageId="12" pageNumber="413">Fig. 10</figureCitation>
|
||
A–B–C), probably the first phalanx, exhibits a concave surface for articulation with the metatarsal. The shaft is slightly constricted as in
|
||
<taxonomicName id="24CB7FDE593EDA5076F29BAF2379F8AC" box="[182,359,1773,1796]" class="Reptilia" family="Hypsilophodontidae" genus="Gasparinisaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5076F29BAF2379F8AC" box="[182,359,1773,1796]" italics="true" pageId="12" pageNumber="413">Gasparinisaura</emphasis>
|
||
</taxonomicName>
|
||
. The distal articulation of the phalanx possesses two almost symmetrical condyles. These are separated by a marked intercondylar groove. The phalanx has ovoid collateral ligament pits on the external surfaces of the lateral and medial condyles (although the latter is not as strongly developed as the former). Another well preserved non-ungual phalanx (
|
||
<figureCitation id="7BF018D8593EDA5074449A15204CF8D8" box="[512,594,1879,1904]" captionStart="FIGURE 10" captionStartId="13.[151,250,1327,1349]" captionTargetBox="[340,1247,483,1305]" captionTargetId="figure@13.[340,1247,483,1305]" captionTargetPageId="13" captionText="FIGURE 10. Notohypsilophodon comodorensis. Non-ungual phalanx (UNPSJB-PV 942 / 35), in medial (A), dorsal (B) and proximal (C) views. Non-ungual phalanx (UNPSJB-PV 942 / 37), in medial (B) view. Non-ungual phalanx (UNPSJB-PV 942 / 39), in dorsal (C) view. Ungual phalanges (UNPSJB-PV 942 / 45,46,47), in dorsal view. Abbreviation: clp, collateral ligament pit; ugr, ungula groove." httpUri="https://zenodo.org/record/226303/files/figure.png" pageId="12" pageNumber="413">Fig. 10</figureCitation>
|
||
D–E; a second phalanx) is shorter than the previously described phalanx. The proximal articular surface is divided by a marked sagittal ridge (= intercondylar ridge, according to
|
||
<bibRefCitation id="875A79AC593EDA50737C9A3E22E6F810" author="Zheng" pageId="12" pageNumber="413" refString="Zheng, W. J., Jin, X. S., Shibata, M., Azuma, Y. & Yu, F. M. (2012) A new ornithischian dinosaur from the Cretaceous Liangtoutang Formation of Tiantai, Zhejiang Province, China. Cretaceous Research, 34, 208 - 219. http: // dx. doi. org / 10.1016 / j. cretres. 2011.11.001" type="journal article" year="2012">
|
||
Zheng
|
||
<emphasis id="D1BFD84F593EDA5073CC9A3F22AAF811" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
). This strong dorsal process delimits two concave surfaces. This suggests that it is not a proximal phalanx [(based on anatomical comparisons with well represented pedes; e.g.,
|
||
<taxonomicName id="24CB7FDE593EDA5075E59A862673F875" box="[929,1133,1988,2013]" class="Reptilia" family="Thescelosauridae" genus="Changchunsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="413" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1BFD84F593EDA5075E59A862673F875" box="[929,1133,1988,2013]" italics="true" pageId="12" pageNumber="413">Changchunsaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="D1BFD84F593EDA5072E69A872701F874" box="[1186,1311,1989,2012]" italics="true" pageId="12" pageNumber="413">Yueosaurus</emphasis>
|
||
), probably phalanx?-2]. The well-developed collateral ligament pits and the presence of dorsal and sagittal ridges on the proximal end of the non-proximal phalanx are features that are common to all basal neornithischians and basal ornithopods (
|
||
<bibRefCitation id="875A79AC593FDA51776F9D9D23E9FF50" author="Zheng" box="[299,503,223,248]" pageId="13" pageNumber="414" refString="Zheng, W. J., Jin, X. S., Shibata, M., Azuma, Y. & Yu, F. M. (2012) A new ornithischian dinosaur from the Cretaceous Liangtoutang Formation of Tiantai, Zhejiang Province, China. Cretaceous Research, 34, 208 - 219. http: // dx. doi. org / 10.1016 / j. cretres. 2011.11.001" type="journal article" year="2012">
|
||
Zheng
|
||
<emphasis id="D1BFD84F593FDA51773F9DA323ADFF50" box="[379,435,223,248]" italics="true" pageId="13" pageNumber="414">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E374045D593FDA5176839C46232BFE05" blockId="13.[151,1436,151,429]" pageId="13" pageNumber="414">
|
||
<emphasis id="D1BFD84F593FDA5176839C4623B5FEB5" bold="true" box="[199,427,260,285]" pageId="13" pageNumber="414">Ungual phalanges.</emphasis>
|
||
The ungual phalanges are subtriangular in shape (
|
||
<figureCitation id="7BF018D8593FDA5172509C462674FEB4" box="[1044,1130,260,285]" captionStart="FIGURE 10" captionStartId="13.[151,250,1327,1349]" captionTargetBox="[340,1247,483,1305]" captionTargetId="figure@13.[340,1247,483,1305]" captionTargetPageId="13" captionText="FIGURE 10. Notohypsilophodon comodorensis. Non-ungual phalanx (UNPSJB-PV 942 / 35), in medial (A), dorsal (B) and proximal (C) views. Non-ungual phalanx (UNPSJB-PV 942 / 37), in medial (B) view. Non-ungual phalanx (UNPSJB-PV 942 / 39), in dorsal (C) view. Ungual phalanges (UNPSJB-PV 942 / 45,46,47), in dorsal view. Abbreviation: clp, collateral ligament pit; ugr, ungula groove." httpUri="https://zenodo.org/record/226303/files/figure.png" pageId="13" pageNumber="414">Fig. 10</figureCitation>
|
||
F–G–H). They bear welldeveloped grooves on both the lateral and medial sides and the ventral surface is planar. The elongate grooves extend for almost the full length of the unguals. The morphology of these elements resembles to those described in MOZ-PV 0 20 (
|
||
<bibRefCitation id="875A79AC593FDA51770D9C2D202EFE20" author="Canudo" box="[329,560,367,392]" pageId="13" pageNumber="414" refString="Canudo, J. I., Salgado, L., Garrido, A. & Carballido, J. (2013) Primera evidencia de dinosaurios ornitopodos en la base de la Formacion Huincul (Cenomaniense Superior-Turoniense, Cuenca Nuequina, Argentina). Geogaceta, 53, 9 - 12." type="journal article" year="2013">
|
||
Canudo
|
||
<emphasis id="D1BFD84F593FDA5177E99C3323F7FE20" box="[429,489,367,392]" italics="true" pageId="13" pageNumber="414">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
). In contrast, those seen in styracosternans are dorsoventrally flattened and spade-shaped.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |