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<document id="5FFEA603CB5E812E2DFAFDC3E31EF32B" ID-DOI="10.1649/0010-065X-77.2.255" ID-ISSN="1938-4394" ID-Zenodo-Dep="13242980" ID-ZooBank="urn:lsid:zoobank.org:pub:2FAD679F-567B-4EE8-AE74-EFD324C7A926" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722967636808" checkinUser="felipe" docAuthor="Cognato, Anthony I. &amp; Smith, Sarah M." docDate="2023" docId="454A87D7FFAB357017BCFE1DFC0FFA72" docLanguage="en" docName="TheColeopBull.77.2.255-263.pdf" docOrigin="The Coleopterists Bulletin 77 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-77.2.255" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Carphobius caterinoi Cognato and Smith 2023, new species" docType="treatment" docVersion="2" lastPageNumber="259" masterDocId="B973FFAFFFAE35751772FFF1FFEDFFA5" masterDocTitle="Taxonomic Review of Carphobius Blackman, 1943 (Coleoptera: Curculionidae: Scolytinae) and a New Species from San Clemente Island, California, USA" masterLastPageNumber="263" masterPageNumber="255" pageNumber="259" updateTime="1722970493295" updateUser="ExternalLinkService">
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<mods:title id="3BB51ED9B3ADC9AE5ED36B4074678B07">Taxonomic Review of Carphobius Blackman, 1943 (Coleoptera: Curculionidae: Scolytinae) and a New Species from San Clemente Island, California, USA</mods:title>
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<mods:namePart id="313F98445099BEBEDD3D474F9EFB916F">Cognato, Anthony I.</mods:namePart>
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<mods:namePart id="F5E208F787E69FC2880F4829B59B311B">Smith, Sarah M.</mods:namePart>
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<mods:title id="4A6E135831A1362C8442C747E1ABF6C0">The Coleopterists Bulletin</mods:title>
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<mods:part id="6BBFA1A2390ACFDD53E404AAFBD6427F">
<mods:date id="A90D4783CE16A33A640F86D8BB3D1D48">2023</mods:date>
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<mods:number id="0A9F7EFAB9F4BA2F5A2D8E4D22D99907">2023-06-16</mods:number>
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<mods:identifier id="A2C302F0CEEA6533ADD5E5B028840818" type="DOI">10.1649/0010-065X-77.2.255</mods:identifier>
<mods:identifier id="2852688645EACB45780E98C0BB0149BF" type="ISSN">1938-4394</mods:identifier>
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<treatment id="454A87D7FFAB357017BCFE1DFC0FFA72" LSID="urn:lsid:plazi:treatment:454A87D7FFAB357017BCFE1DFC0FFA72" httpUri="http://treatment.plazi.org/id/454A87D7FFAB357017BCFE1DFC0FFA72" lastPageNumber="259" pageId="5" pageNumber="259">
<subSubSection id="85F9654AFFAB357017BCFE1DFDE7FDC9" pageId="5" pageNumber="259" type="nomenclature">
<paragraph id="CD5C36C1FFAB357017BCFE1DFDE7FDC9" blockId="5.[177,655,492,621]" pageId="5" pageNumber="259">
<heading id="961481ADFFAB357017BCFE1DFE36FDB8" centered="true" fontSize="9" level="2" pageId="5" pageNumber="259" reason="2">
<emphasis id="FF97EAD3FFAB357017BCFE1DFE36FDB8" bold="true" pageId="5" pageNumber="259">
<taxonomicName id="0AE34D42FFAB357017BCFE1DFD83FDA7" authority="Cognato and Smith" authorityName="Cognato and Smith" authorityYear="2023" box="[206,622,492,514]" class="Insecta" family="Curculionidae" genus="Carphobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="259" phylum="Arthropoda" rank="species" species="caterinoi" status="sp. nov.">
<emphasis id="FF97EAD3FFAB357017BCFE1DFE4DFDA4" bold="true" box="[206,416,492,514]" italics="true" pageId="5" pageNumber="259">Carphobius caterinoi</emphasis>
Cognato and Smith
</taxonomicName>
,
<taxonomicNameLabel id="E4A457A8FFAB35701617FDF6FE36FDB8" box="[357,475,519,541]" pageId="5" pageNumber="259" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
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urn:lsid:zoobank.org:act:
<uuid id="B9450C14FFAB357017C3FDCDFD62FDF7" box="[177,655,572,594]" pageId="5" pageNumber="259">8AEDDD42-D7C0-45CB-87A1-58AADA2CF82A</uuid>
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(
<figureCitation id="55D82A44FFAB3570164EFDA6FE0CFDC9" box="[316,481,599,621]" captionStart="Figs" captionStartId="4.[176,215,1627,1646]" captionTargetBox="[192,1130,193,1610]" captionTargetId="figure-6@4.[192,1130,193,1610]" captionTargetPageId="4" captionText="Figs. 213. Carphobius species. C. arizonicus, male (USNMENT_01595074); 2) Habitus, dorsal view; 3) Habitus, lateral view; 8) Declivital view; 11) Frontal view. C. arizonicus, female (USNMENT_01595075): 4) Habitus, dorsal view; 5) Habitus, lateral view; 9) Declivital view; 12) Frontal view. C. caterinoi, new species, male (holotype; CBP0079193): 6) Habitus, dorsal view; 7) Habitus, lateral view; 10) Declivital view; 13) Frontal view." figureDoi="http://doi.org/10.5281/zenodo.13242984" httpUri="https://zenodo.org/record/13242984/files/figure.png" pageId="5" pageNumber="259">Figs. 6, 7, 10, 13</figureCitation>
,
<figureCitation id="55D82A44FFAB35701699FDA6FDE9FDC9" box="[491,516,599,620]" captionStart="Fig" captionStartId="5.[197,228,1663,1682]" captionTargetBox="[210,625,1156,1646]" captionTargetId="figure-603@5.[173,663,1156,1646]" captionTargetPageId="5" captionText="Fig. 14. Carphobius caterinoi, new species, paratype male, genitalia and spiculum gastrale (rotated 180 degrees)." figureDoi="http://doi.org/10.5281/zenodo.13242986" httpUri="https://zenodo.org/record/13242986/files/figure.png" pageId="5" pageNumber="259">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="85F9654AFFAB357017B7FD70FE49FCF7" pageId="5" pageNumber="259" type="diagnosis">
<paragraph id="CD5C36C1FFAB357017B7FD70FE49FCF7" blockId="5.[173,659,641,1090]" pageId="5" pageNumber="259">
<emphasis id="FF97EAD3FFAB357017B7FD70FEC3FD32" bold="true" box="[197,302,641,663]" pageId="5" pageNumber="259">Diagnosis.</emphasis>
This species is distinguished from other
<taxonomicName id="0AE34D42FFAB35701796FD6DFEB8FD14" authorityName="Blackman" authorityYear="1943" box="[228,341,668,689]" class="Insecta" family="Curculionidae" genus="Carphobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="FF97EAD3FFAB35701796FD6DFEB8FD14" box="[228,341,668,689]" italics="true" pageId="5" pageNumber="259">Carphobius</emphasis>
</taxonomicName>
by the acuminate, scale-like vestiture and the two or three small, acute spines on the first and third elytral declivity interstriae.
<taxonomicName id="0AE34D42FFAB357017DFFD1DFE96FCA7" authorityName="Cognato and Smith" authorityYear="2023" box="[173,379,748,770]" class="Insecta" family="Curculionidae" genus="Carphobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="259" phylum="Arthropoda" rank="species" species="caterinoi">
<emphasis id="FF97EAD3FFAB357017DFFD1DFE96FCA7" box="[173,379,748,770]" italics="true" pageId="5" pageNumber="259">Carphobius caterinoi</emphasis>
</taxonomicName>
is superficially similar to the monotypic
<taxonomicName id="0AE34D42FFAB3570166AFCF6FDBDFCB9" authority="(Bright, 1972)" baseAuthorityName="Bright" baseAuthorityYear="1972" box="[280,592,775,797]" class="Insecta" family="Curculionidae" genus="Carphotoreus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="259" phylum="Arthropoda" rank="species" species="alni">
<emphasis id="FF97EAD3FFAB3570166AFCF6FE28FCB9" box="[280,453,775,796]" italics="true" pageId="5" pageNumber="259">Carphotoreus alni</emphasis>
(Bright, 1972)
</taxonomicName>
except for the emarginate eye, concealed scutellar shield, and contiguous procoxae.
</paragraph>
</subSubSection>
<subSubSection id="85F9654AFFAB357017B7FCA6FCE4FC5D" pageId="5" pageNumber="259" type="description">
<paragraph id="CD5C36C1FFAB357017B7FCA6FB44FF7D" blockId="5.[173,659,641,1090]" lastBlockId="5.[706,1194,194,1496]" pageId="5" pageNumber="259">
<emphasis id="FF97EAD3FFAB357017B7FCA6FDC5FCC9" bold="true" box="[197,552,855,877]" pageId="5" pageNumber="259">
Description (Male
<typeStatus id="12588863FFAB357016F2FCA6FE0DFCC8" box="[384,480,855,877]" pageId="5" pageNumber="259" type="holotype">Holotype</typeStatus>
). Body.
</emphasis>
Length 2.0 mm; 2.8 times as long as wide; vestiture of scale-like acuminate setae.
<emphasis id="FF97EAD3FFAB3570160EFC7DFE54FC04" bold="true" box="[380,441,908,929]" pageId="5" pageNumber="259">Head.</emphasis>
Frons with medial depression, vestiture of acuminate, scale-like setae, uniformly distributed from epistoma to vertex; epistomal margin with abundant, long, hair-like setae; eyes shallowly emarginate; antennal funicle of six antennomeres, club elongate-conical and flattened with three transverse sutures, club longer than scape and funicle combined, scape shorter then funicle.
</paragraph>
<paragraph id="CD5C36C1FFAB357015B1FF2DFB44FD27" blockId="5.[706,1194,194,1496]" pageId="5" pageNumber="259">
<emphasis id="FF97EAD3FFAB357015B1FF2DFCDDFF54" bold="true" box="[707,816,220,241]" pageId="5" pageNumber="259">Pronotum.</emphasis>
Widest at base; anterior quarter of pronotum constricted; anterior margin with coarse asperities; patch of submarginal coarse asperities reaching summit but not extending to lateral margins; vestiture a mix of acuminate scale-like and smaller fine setae.
<emphasis id="FF97EAD3FFAB35701405FE93FC53FED2" bold="true" box="[887,958,354,375]" pageId="5" pageNumber="259">Elytra.</emphasis>
Scutellar shield not visible, each basal margin with eight coarse asperities; submarginal asperities restricted to interstriae 1 and 2; discal striae uniseriately punctured with fine, hair-like setae; interstriae 3
<emphasis id="FF97EAD3FFAB357014B1FE3DFC3CFE44" bold="true" box="[963,977,460,481]" pageId="5" pageNumber="259">×</emphasis>
wider than striae, with confused, scale-like, acuminate setae; declivity gradually sloping, occupying 50% of the elytra, posterolateral angles rounded, interstria 2 impressed, all interstriae with uniseriate and biseriate, scale-like, acuminate setae, and two widely spaced acute denticles on interstriae 1 and 3; striae obscure.
</paragraph>
<paragraph id="CD5C36C1FFAB357015B1FD76FC61FCAD" blockId="5.[706,1194,194,1496]" pageId="5" pageNumber="259">
<emphasis id="FF97EAD3FFAB357015B1FD76FD15FD39" bold="true" box="[707,760,647,668]" pageId="5" pageNumber="259">Legs.</emphasis>
Pro-, meso-, and metacoxae contiguous; pro-, meso-, and metatibiae obliquely triangular, broadest at apical one-fourth; apical one-half of outer margin with five large, socketed denticles, their length longer than basal width.
</paragraph>
<paragraph id="CD5C36C1FFAB357015A9FCFCFCE4FC5D" blockId="5.[706,1194,194,1496]" pageId="5" pageNumber="259">
<typeStatus id="12588863FFAB357015A9FCFCFCD5FC87" box="[731,824,781,802]" pageId="5" pageNumber="259" type="paratype">Paratypes</typeStatus>
(males) similar to
<typeStatus id="12588863FFAB35701498FCFDFBD3FC87" box="[1002,1086,780,802]" pageId="5" pageNumber="259" type="holotype">holotype</typeStatus>
with slight variation in size (mean = 2.0 mm, range
<quantity id="0A1B9B24FFAB35701343FCD6FB4FFC99" box="[1073,1186,807,828]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="5" pageNumber="259" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.92.1 mm</quantity>
) and number (2 or 3) of elytral spines. Aedeagus of
<specimenCount id="DBE5FD48FFAB357015B1FCACFCAFFCD7" box="[707,834,861,882]" count="1" pageId="5" pageNumber="259" type="generic" typeStatus="paratype">one paratype</specimenCount>
examined (
<figureCitation id="55D82A44FFAB357014CBFCADFBE8FCD4" box="[953,1029,860,882]" captionStart="Fig" captionStartId="5.[197,228,1663,1682]" captionTargetBox="[210,625,1156,1646]" captionTargetId="figure-603@5.[173,663,1156,1646]" captionTargetPageId="5" captionText="Fig. 14. Carphobius caterinoi, new species, paratype male, genitalia and spiculum gastrale (rotated 180 degrees)." figureDoi="http://doi.org/10.5281/zenodo.13242986" httpUri="https://zenodo.org/record/13242986/files/figure.png" pageId="5" pageNumber="259">Fig. 14</figureCitation>
): aedeagal body rectangular and distally narrowed, apophyses length 2
<emphasis id="FF97EAD3FFAB357015BDFC63FD30FC02" bold="true" box="[719,733,914,935]" pageId="5" pageNumber="259">×</emphasis>
as long as aedeagal body, tegmen C-shaped (open dorsally), manubrium absent, flagellum presence uncertain, spiculum gastrale rod apically forked.
</paragraph>
</subSubSection>
<subSubSection id="85F9654AFFAB357015A9FC0DFB76FAC9" pageId="5" pageNumber="259" type="materials_examined">
<paragraph id="CD5C36C1FFAB357015A9FC0DFB76FAC9" blockId="5.[706,1194,194,1496]" pageId="5" pageNumber="259">
<emphasis id="FF97EAD3FFAB357015A9FC0DFC7EFBB7" bold="true" box="[731,915,1020,1042]" pageId="5" pageNumber="259">Type Specimens.</emphasis>
<materialsCitation id="7D8B3C9CFFAB357014D3FC0DFCC4FB32" collectionCode="CA, BEETLE, PROJ, SBMNH" collectorName="San Clemente Isl &amp; Upper Thirst C &amp; Caterino &amp; Chatzimanolis. Blue" country="United States of America" county="Los Angeles Co." latitude="32.8813" location="Upper Thirst" longLatPrecision="6" longitude="-118.4445" pageId="5" pageNumber="259" specimenCode="CBP0079193" specimenCount="1" stateProvince="California" typeStatus="holotype">
<typeStatus id="12588863FFAB357014D3FC0DFBECFBB7" box="[929,1025,1020,1042]" pageId="5" pageNumber="259" type="holotype">Holotype</typeStatus>
, male, labeled:
<collectingCountry id="B5F47651FFAB357015B1FBE6FD14FB89" box="[707,761,1047,1068]" name="United States of America" pageId="5" pageNumber="259">USA</collectingCountry>
:
<collectingRegion id="0F27F823FFAB35701478FBE6FC81FB88" box="[778,876,1047,1069]" country="United States of America" name="California" pageId="5" pageNumber="259">California</collectingRegion>
,
<collectingCounty id="243D4E4DFFAB3570140EFBE6FBC6FB89" box="[892,1067,1047,1069]" pageId="5" pageNumber="259">Los Angeles Co.</collectingCounty>
,
<geoCoordinate id="A8D75006FFAB35701349FBE6FB48FB89" box="[1083,1189,1047,1068]" degrees="32.8813" direction="north" orientation="latitude" pageId="5" pageNumber="259" precision="5" value="32.8813">32.8813°N</geoCoordinate>
,
<geoCoordinate id="A8D75006FFAB357015B1FBC0FCD7FBE2" box="[707,826,1073,1095]" degrees="118.4445" direction="west" orientation="longitude" pageId="5" pageNumber="259" precision="5" value="-118.4445">118.4445°W</geoCoordinate>
,
<collectorName id="60165317FFAB35701434FBC3FC03FBED" box="[838,1006,1074,1096]" pageId="5" pageNumber="259">San Clemente Isl</collectorName>
[and],
<collectorName id="60165317FFAB3570135FFBC3FD39FBC4" pageId="5" pageNumber="259">
<location id="C83C601AFFAB3570135FFBC3FB45FBED" LSID="urn:lsid:plazi:treatment:454A87D7FFAB357017BCFE1DFC0FFA72:C83C601AFFAB3570135FFBC3FB45FBED" box="[1069,1192,1074,1096]" country="United States of America" county="Los Angeles Co." latitude="32.8813" longLatPrecision="6" longitude="-118.4445" name="Upper Thirst" pageId="5" pageNumber="259" stateProvince="California">Upper Thirst</location>
C
</collectorName>
[an]y[o]n, vi.15.2008,
<collectorName id="60165317FFAB357014DBFBBDFC17FBC7" box="[937,1018,1100,1122]" pageId="5" pageNumber="259">Caterino</collectorName>
&amp;
<collectorName id="60165317FFAB35701369FBBDFD03FBD8" pageId="5" pageNumber="259">Chatzimanolis. Blue</collectorName>
second label:
<collectionCode id="ABF2AE04FFAB3570141CFB96FC60FBD9" box="[878,909,1127,1148]" country="USA" name="Chicago Academy of Sciences" pageId="5" pageNumber="259" type="Museum">CA</collectionCode>
<collectionCode id="ABF2AE04FFAB357014E3FB96FBCEFBD9" box="[913,1059,1127,1148]" pageId="5" pageNumber="259">BEETLE PROJ</collectionCode>
,
<specimenCode id="9D459EBAFFAB35701359FB96FB44FBD9" box="[1067,1193,1127,1148]" collectionCode="CBP" pageId="5" pageNumber="259">CBP0079193</specimenCode>
(
<collectionCode id="ABF2AE04FFAB357015BBFB73FCCFFB32" box="[713,802,1154,1175]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="5" pageNumber="259" type="Museum">SBMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="7D8B3C9CFFAB3570144EFB73FCCDFA92" collectingDate="2019-04-13" collectionCode="SBMNH, MSUC" country="United States of America" county="Los Angeles Co." location="Lat" municipality="San Clemente Island" pageId="5" pageNumber="259" specimenCount="2" specimenCount-male="2" stateProvince="California" typeStatus="paratype">
<typeStatus id="12588863FFAB3570144EFB73FC77FB32" box="[828,922,1154,1175]" pageId="5" pageNumber="259" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="DBE5FD48FFAB357014DCFB73FBEEFB3D" box="[942,1027,1154,1176]" count="2" pageId="5" pageNumber="259" type="male">2 males</specimenCount>
, labeled:
<collectingCountry id="B5F47651FFAB35701300FB73FB48FB32" box="[1138,1189,1154,1175]" name="United States of America" pageId="5" pageNumber="259">USA</collectingCountry>
:
<collectingRegion id="0F27F823FFAB357015B1FB6DFCC8FB17" box="[707,805,1180,1202]" country="United States of America" name="California" pageId="5" pageNumber="259">California</collectingRegion>
,
<collectingCounty id="243D4E4DFFAB3570145DFB6CFC3CFB14" box="[815,977,1180,1202]" pageId="5" pageNumber="259">Los Angeles Co.</collectingCounty>
,
<collectingMunicipality id="2D38ACBBFFAB357014AEFB6DFB48FB17" box="[988,1189,1180,1202]" pageId="5" pageNumber="259">San Clemente Island</collectingMunicipality>
,
<location id="C83C601AFFAB357015B0FB46FD08FB69" LSID="urn:lsid:plazi:treatment:454A87D7FFAB357017BCFE1DFC0FFA72:C83C601AFFAB357015B0FB46FD08FB69" box="[706,741,1207,1228]" country="United States of America" county="Los Angeles Co." municipality="San Clemente Island" name="Lat" pageId="5" pageNumber="259" stateProvince="California">Lat</location>
: 32.8776
<location id="C83C601AFFAB3570144EFB46FC88FB69" LSID="urn:lsid:plazi:treatment:454A87D7FFAB357017BCFE1DFC0FFA72:C83C601AFFAB3570144EFB46FC88FB69" box="[828,869,1207,1228]" country="United States of America" county="Los Angeles Co." municipality="San Clemente Island" name="Lon" pageId="5" pageNumber="259" stateProvince="California">Lon</location>
: 118.4351,
<date id="B95D1001FFAB357014A8FB46FBB6FB69" box="[986,1115,1207,1229]" pageId="5" pageNumber="259" value="2019-04-13">
<collectingDate id="A919E9E9FFAB357014A8FB46FBB6FB69" box="[986,1115,1207,1229]" pageId="5" pageNumber="259" value="2019-04-13">13April 2019</collectingDate>
</date>
, collect- ed from a steep, forested canyon via beat sheet from
<taxonomicName id="0AE34D42FFAB357015B0FB1CFBC0FAA7" authority=", C. H. Richart Coll." authorityName="C. H. Richart Coll." box="[706,1069,1260,1282]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="259" phylum="Tracheophyta" rank="species" species="tomentella">
<emphasis id="FF97EAD3FFAB357015B0FB1CFC9AFAA4" box="[706,887,1260,1282]" italics="true" pageId="5" pageNumber="259">Quercus tomentella</emphasis>
, C. H. Richart Coll.
</taxonomicName>
SBBG-SCL- TIC-001193 (1
<collectionCode id="ABF2AE04FFAB3570143CFAF6FC4EFAB9" box="[846,931,1287,1308]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="5" pageNumber="259" type="Museum">SBMNH</collectionCode>
, 1
<collectingRegion id="0F27F823FFAB357014C8FAF6FBA9FAB9" box="[954,1092,1287,1309]" country="United States of America" name="Michigan" pageId="5" pageNumber="259">Michigan State</collectingRegion>
<location id="C83C601AFFAB35701335FAF6FB45FAB8" LSID="urn:lsid:plazi:treatment:454A87D7FFAB357017BCFE1DFC0FFA72:C83C601AFFAB35701335FAF6FB45FAB8" box="[1095,1192,1287,1309]" country="United States of America" county="Los Angeles Co." municipality="San Clemente Island" name="University" pageId="5" pageNumber="259" stateProvince="California">University</location>
[
<collectionCode id="ABF2AE04FFAB357015BAFAD3FCFEFA92" box="[712,787,1314,1335]" pageId="5" pageNumber="259">MSUC</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="7D8B3C9CFFAB3570145AFAD3FB7AFAC9" collectingDate="2019-04-13" collectionCode="MSUC" collectorName="A. I. Cognato" country="United States of America" county="Los Angeles Co." location="Lat" municipality="San Clemente Island" pageId="5" pageNumber="259" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="paratype">
<specimenCount id="DBE5FD48FFAB3570145AFAD3FC8BFA9D" box="[808,870,1314,1336]" count="1" pageId="5" pageNumber="259" type="male">1 male</specimenCount>
(dissected) glued to card, genitalia in vial, same locality data and second label: DNA voucher, SMS400,
<collectorName id="60165317FFAB35701404FAA6FC1BFAC9" box="[886,1014,1367,1388]" pageId="5" pageNumber="259">A. I. Cognato</collectorName>
2022 (1
<collectionCode id="ABF2AE04FFAB3570133BFAA6FB7DFAC9" box="[1097,1168,1367,1388]" pageId="5" pageNumber="259">MSUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="85F9654AFFAB35701597FA83FC0FFA72" pageId="5" pageNumber="259" type="etymology">
<paragraph id="CD5C36C1FFAB35701597FA83FC0FFA72" blockId="5.[706,1194,194,1496]" pageId="5" pageNumber="259">
<emphasis id="FF97EAD3FFAB35701597FA83FCB5FA22" bold="true" box="[741,856,1394,1415]" pageId="5" pageNumber="259">Etymology.</emphasis>
Named in honor of Dr. Michael Caterino who began and directed the
<collectingRegion id="0F27F823FFAB35701335FA7DFB45FA07" box="[1095,1192,1420,1442]" country="United States of America" name="California" pageId="5" pageNumber="259">California</collectingRegion>
Beetle Project at the Santa Barbara Museum of Natural History (20012014).
</paragraph>
</subSubSection>
</treatment>
</document>