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<document id="62A31F24A42E013D377062D56DF3897B" ID-CLB-Dataset="27431" ID-DOI="10.11646/zootaxa.4560.1.3" ID-GBIF-Dataset="66d86ea3-e671-4144-b8be-7704010c10e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2627372" ID-ZooBank="1057AA8B-9221-4045-99A5-8F0CC0F7395B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554322446059" checkinUser="plazi" docAuthor="Roux, Michel, Eléaume, Marc &amp; Améziane, Nadia" docDate="2019" docId="913887B7FFE7FF9FD098FB67FDA4FB9A" docLanguage="en" docName="zootaxa.4560.1.3.pdf" docOrigin="Zootaxa 4560 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Conocrinus cahuzaci Roux &amp; Eléaume &amp; Améziane 2019, n. sp." docType="treatment" docVersion="8" lastPageNumber="77" masterDocId="6D01FFCFFFFEFF85D00FFFD9FFD3FFF8" masterDocTitle="A revision of the genus Conocrinus dOrbigny, 1850 (Echinodermata, Crinoidea, Rhizocrinidae) and its place among extant and fossil crinoids with a xenomorphic stalk" masterLastPageNumber="84" masterPageNumber="51" pageNumber="76" updateTime="1698717837333" updateUser="ExternalLinkService">
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<mods:title id="99B078F1BE48B7AAD7B7324DA2F224B3">A revision of the genus Conocrinus dOrbigny, 1850 (Echinodermata, Crinoidea, Rhizocrinidae) and its place among extant and fossil crinoids with a xenomorphic stalk</mods:title>
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<mods:namePart id="682E622E647472B4CF5D25618C5231E3">Roux, Michel</mods:namePart>
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<mods:namePart id="F7468C7B8ADB1362AEAE62EA01E45F18">Eléaume, Marc</mods:namePart>
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<paragraph id="192E36A1FFE7FF9CD098FB67FE32FB20" blockId="25.[151,481,1214,1274]" box="[151,481,1214,1240]" pageId="25" pageNumber="76">
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<emphasis id="2BE5EAB3FFE7FF9CD098FB67FE32FB20" bold="true" box="[151,481,1214,1240]" pageId="25" pageNumber="76">
<taxonomicName id="DE914D22FFE7FF9CD098FB67FE44FB20" authority="Roux &amp; Eléaume &amp; Améziane, 2019" authorityName="Roux &amp; Eléaume &amp; Améziane" authorityYear="2019" box="[151,407,1214,1240]" class="Crinoidea" family="Bathycrinidae" genus="Conocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="25" pageNumber="76" phylum="Echinodermata" rank="species" species="cahuzaci" status="sp. nov.">
<emphasis id="2BE5EAB3FFE7FF9CD098FB67FE44FB20" bold="true" box="[151,407,1214,1240]" italics="true" pageId="25" pageNumber="76">Conocrinus cahuzaci</emphasis>
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<taxonomicNameLabel id="30D657C8FFE7FF9CD191FB67FE32FB20" box="[414,481,1214,1240]" pageId="25" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="192E36A1FFE7FF9CD098FB38FE40FB02" blockId="25.[151,481,1214,1274]" box="[151,403,1249,1274]" pageId="25" pageNumber="76">
<figureCitation id="81AA2A24FFE7FF9CD098FB38FF04FB02" box="[151,215,1249,1274]" captionStart="FIGURE 5" captionStartId="11.[151,250,1675,1697]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@11.[151,1436,193,1653]" captionTargetPageId="11" captionText="FIGURE 5. Aboral cups of Conocrinus thorenti (A-D) and other Eocene species from southern France closely related to it. A- D: Conocrinus thorenti, lectotype MNHN.F.R54426, AB: oblique lateral view, CD: adoral view, D: closeup of arm socket indicated by arrow in C; EG: Conocrinus archiaci, Villa Marbella, Biarritz, EF: specimen MNHN.F.A69934a, E: lateral view, F: oblique adoral view with closeup of arm socket, G: specimen MNHN.F.A69934b, oblique adoral view; H: Conocrinus aff. archiaci, Les Scaffarels near Annot, MHN Nice, Campredon Collection, specimen no. 6, flat arm socket (synostosis); IK: Conocrinus cahuzaci n. sp., holotype, MNHN.F.A69935, I oblique side adoral view, JK: adoral views; LO: Conocrinus duperrieri, marnes à Pentacrines, Côte des Basques, Biarritz, LN: holotype MNHN.F.R06934 (arrow head indicating segment of radial suture), O: side view of paratype MNHN.F.R06935; bip: broken interradial process, ip: interradial process, ir: interradial rib. See Figure 1 for other abbreviations." httpUri="https://zenodo.org/record/2627382/files/figure.png" pageId="25" pageNumber="76">Fig. 5</figureCitation>
IK,
<tableCitation id="5413031AFFE7FF9CD11DFB38FE5CFB02" box="[274,399,1249,1274]" captionStart-0="TABLE 5" captionStart-1="TABLE 6" captionStartId-0="25.[151,239,1609,1632]" captionStartId-1="26.[151,239,404,427]" captionText-0="TABLE 5. Main quantitative characters of external cup morphology in the type series of Conocrinus cahuzaci n. sp. Hc: cup height, Hb: basal ring height, Hpr: maximum height of interradial processes, Ws: maximum width of arm socket, Wr: distal radial width at level of arm socket, Dr: radial ring diameter at level of arm sockets, Drb: diameter at basal/ radial transition, Db: basal ring diameter at cup base. Values in mm." captionText-1="TABLE 6. Main morphological ratios in aboral cups of the type series of Conocrinus cahuzaci n. sp. See Table 5 for abbreviations." pageId="25" pageNumber="76">Tables 56</tableCitation>
.
</paragraph>
<paragraph id="192E36A1FFE7FF9CD098FAF0FCE1FABA" blockId="25.[151,818,1321,1346]" box="[151,818,1321,1346]" pageId="25" pageNumber="76">
<emphasis id="2BE5EAB3FFE7FF9CD098FAF0FEC0FABA" bold="true" box="[151,275,1321,1346]" pageId="25" pageNumber="76">Synonymy</emphasis>
.
<emphasis id="2BE5EAB3FFE7FF9CD12EFAF3FE7CFAB9" box="[289,431,1322,1345]" italics="true" pageId="25" pageNumber="76">Tormocrinus</emphasis>
(g)
<bibRefCitation id="7D004B50FFE7FF9CD1D1FAF3FD4FFABA" author="Roux, M. &amp; Cahuzac, B. &amp; Sztrakos, K." box="[478,668,1321,1346]" pageId="25" pageNumber="76" pagination="262 - 271" refId="ref23064" refString="Roux, M., Cahuzac, B. &amp; Sztrakos, K. (2006) Les paleoenvironnements eocenes a Crinoides pedoncules des marnes de Miretrain (Angoume, SW de la France): interpretations paleobathymetriques. Comptes Rendus Geoscience, 338 (4), 262 - 271. https: // doi. org / 10.1016 / j. crte. 2006.01.001" type="journal article" year="2006">
Roux
<emphasis id="2BE5EAB3FFE7FF9CD22EFAF3FD89FABA" box="[545,602,1321,1346]" italics="true" pageId="25" pageNumber="76">et al.</emphasis>
2006
</bibRefCitation>
: 267, table 1.
</paragraph>
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<paragraph id="192E36A1FFE7FF9CD098FAA8FE25FA29" blockId="25.[151,1436,1393,1562]" pageId="25" pageNumber="76">
<emphasis id="2BE5EAB3FFE7FF9CD098FAA8FECDFA72" bold="true" box="[151,286,1393,1418]" pageId="25" pageNumber="76">Etymology.</emphasis>
Species dedicated to Bruno Cahuzac (University of Bordeaux), who collected a crinoid fauna (including the
<typeStatus id="C62A8803FFE7FF9CD130FA4FFEBCFA56" box="[319,367,1430,1454]" pageId="25" pageNumber="76">type</typeStatus>
series) that is more diversified than previously expected during his investigations of Eocene sediments in southwest
<collectingCountry id="61867631FFE7FF9CD1ADFA63FE21FA29" box="[418,498,1466,1489]" name="France" pageId="25" pageNumber="76">France</collectingCountry>
.
</paragraph>
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<materialsCitation id="A9F93CFCFFE7FF9CD0C8FA04FD61F9E1" ID-GBIF-Occurrence="2234214344" country="France" county="Dax" location="Miretrain" municipality="Angoume" pageId="25" pageNumber="76" specimenCount="2" typeStatus="holotype">
<emphasis id="2BE5EAB3FFE7FF9CD0C8FA04FDE9FA0E" bold="true" box="[199,570,1501,1526]" pageId="25" pageNumber="76">
<typeStatus id="C62A8803FFE7FF9CD0C8FA04FED1FA0E" box="[199,258,1501,1526]" pageId="25" pageNumber="76">Type</typeStatus>
locality and
<typeStatus id="C62A8803FFE7FF9CD195FA04FE1DFA0E" box="[410,462,1501,1526]" pageId="25" pageNumber="76">type</typeStatus>
stratum.
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<location id="1C4E607AFFE7FF9CD24EFA04FD78FA0E" LSID="urn:lsid:plazi:treatment:913887B7FFE7FF9FD098FB67FDA4FB9A:1C4E607AFFE7FF9CD24EFA04FD78FA0E" box="[577,683,1501,1526]" country="France" county="Dax" municipality="Angoume" name="Miretrain" pageId="25" pageNumber="76">Miretrain</location>
quarry; dark grey marls C6a of
<bibRefCitation id="7D004B50FFE7FF9CD41EFA07FB33FA0E" author="Roux, M. &amp; Cahuzac, B. &amp; Sztrakos, K." box="[1041,1248,1501,1526]" pageId="25" pageNumber="76" pagination="262 - 271" refId="ref23064" refString="Roux, M., Cahuzac, B. &amp; Sztrakos, K. (2006) Les paleoenvironnements eocenes a Crinoides pedoncules des marnes de Miretrain (Angoume, SW de la France): interpretations paleobathymetriques. Comptes Rendus Geoscience, 338 (4), 262 - 271. https: // doi. org / 10.1016 / j. crte. 2006.01.001" type="journal article" year="2006">
Roux
<emphasis id="2BE5EAB3FFE7FF9CD45BFA07FB5EFA0E" box="[1108,1165,1501,1526]" italics="true" pageId="25" pageNumber="76">et al.</emphasis>
(2006)
</bibRefCitation>
,
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(base NP17) at
<collectingMunicipality id="F94AACDBFFE7FF9CD10DF9DBFEA3F9E2" box="[258,368,1538,1562]" pageId="25" pageNumber="76">Angoumé</collectingMunicipality>
near
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, southwest
<collectingCountry id="61867631FFE7FF9CD26DF9DBFD61F9E1" box="[610,690,1538,1561]" name="France" pageId="25" pageNumber="76">France</collectingCountry>
</materialsCitation>
.
</paragraph>
<caption id="4DEE6629FFE7FF9CD098F990FCB8F938" pageId="25" pageNumber="76">
<paragraph id="192E36A1FFE7FF9CD098F990FCB8F938" blockId="25.[151,1436,1609,1728]" pageId="25" pageNumber="76">
<emphasis id="2BE5EAB3FFE7FF9CD098F990FEDBF998" bold="true" box="[151,264,1609,1632]" pageId="25" pageNumber="76">TABLE 5.</emphasis>
Main quantitative characters of external cup morphology in the type series of
<taxonomicName id="DE914D22FFE7FF9CD444F990FAFEF998" authority="Roux &amp; Eléaume &amp; Améziane, 2019" authorityName="Roux &amp; Eléaume &amp; Améziane" authorityYear="2019" box="[1099,1325,1609,1632]" class="Crinoidea" family="Bathycrinidae" genus="Conocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="25" pageNumber="76" phylum="Echinodermata" rank="species" species="cahuzaci" status="sp. nov.">
<emphasis id="2BE5EAB3FFE7FF9CD444F990FAFEF998" box="[1099,1325,1609,1632]" italics="true" pageId="25" pageNumber="76">Conocrinus cahuzaci</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="30D657C8FFE7FF9CD53CF990FABCF998" box="[1331,1391,1609,1632]" pageId="25" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Hc: cup height, Hb: basal ring height, Hpr: maximum height of interradial processes, Ws: maximum width of arm socket, Wr: distal radial width at level of arm socket, Dr: radial ring diameter at level of arm sockets, Drb: diameter at basal/ radial transition, Db: basal ring diameter at cup base. Values in mm.
</paragraph>
</caption>
<paragraph id="192E36A1FFE7FF9CD090F905FA90F86F" pageId="25" pageNumber="76">
<table id="6B91C401FFE7007AD090F905FA90F86F" box="[159,1347,1756,1943]" gridcols="9" gridrows="5" pageId="25" pageNumber="76" rowsContinueIn="26.[159,1347,486,674]">
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<th id="E4705D9FFFE7007AD090F905FEC3F90A" box="[159,272,1756,1778]" gridcol="0" gridrow="0" pageId="25" pageNumber="76">Characters</th>
<th id="E4705D9FFFE7007AD14AF905FEADF90A" box="[325,382,1756,1778]" gridcol="1" gridrow="0" pageId="25" pageNumber="76">Hc</th>
<th id="E4705D9FFFE7007AD1DEF905FE2EF90A" box="[465,509,1756,1778]" gridcol="2" gridrow="0" pageId="25" pageNumber="76">Hb</th>
<th id="E4705D9FFFE7007AD252F905FD5AF90A" box="[605,649,1756,1778]" gridcol="3" gridrow="0" pageId="25" pageNumber="76">Hpr</th>
<th id="E4705D9FFFE7007AD2E7F905FCC7F90A" box="[744,788,1756,1778]" gridcol="4" gridrow="0" pageId="25" pageNumber="76">Wr</th>
<th id="E4705D9FFFE7007AD37BF905FC73F90A" box="[884,928,1756,1778]" gridcol="5" gridrow="0" pageId="25" pageNumber="76">Ws</th>
<th id="E4705D9FFFE7007AD40FF905FBFFF90A" box="[1024,1068,1756,1778]" gridcol="6" gridrow="0" pageId="25" pageNumber="76">Dr</th>
<th id="E4705D9FFFE7007AD484F905FB64F90A" box="[1163,1207,1756,1778]" gridcol="7" gridrow="0" pageId="25" pageNumber="76">Drb</th>
<th id="E4705D9FFFE7007AD518F905FA90F90A" box="[1303,1347,1756,1778]" gridcol="8" gridrow="0" pageId="25" pageNumber="76">Db</th>
</tr>
<tr id="A7A134E3FFE7007AD090F8DCFA90F8E3" box="[159,1347,1797,1819]" gridrow="1" pageId="25" pageNumber="76">
<th id="E4705D9FFFE7007AD090F8DCFEC3F8E3" box="[159,272,1797,1819]" gridcol="0" gridrow="1" pageId="25" pageNumber="76">Holotype</th>
<td id="E4705D9FFFE7007AD14AF8DCFEADF8E3" box="[325,382,1797,1819]" gridcol="1" gridrow="1" pageId="25" pageNumber="76">11.55</td>
<td id="E4705D9FFFE7007AD1DEF8DCFE2EF8E3" box="[465,509,1797,1819]" gridcol="2" gridrow="1" pageId="25" pageNumber="76">8.60</td>
<td id="E4705D9FFFE7007AD252F8DCFD5AF8E3" box="[605,649,1797,1819]" gridcol="3" gridrow="1" pageId="25" pageNumber="76">1.45</td>
<td id="E4705D9FFFE7007AD2E7F8DCFCC7F8E3" box="[744,788,1797,1819]" gridcol="4" gridrow="1" pageId="25" pageNumber="76">2.35</td>
<td id="E4705D9FFFE7007AD37BF8DCFC73F8E3" box="[884,928,1797,1819]" gridcol="5" gridrow="1" pageId="25" pageNumber="76">0.68</td>
<td id="E4705D9FFFE7007AD40FF8DCFBFFF8E3" box="[1024,1068,1797,1819]" gridcol="6" gridrow="1" pageId="25" pageNumber="76">3.90</td>
<td id="E4705D9FFFE7007AD484F8DCFB64F8E3" box="[1163,1207,1797,1819]" gridcol="7" gridrow="1" pageId="25" pageNumber="76">2.50</td>
<td id="E4705D9FFFE7007AD518F8DCFA90F8E3" box="[1303,1347,1797,1819]" gridcol="8" gridrow="1" pageId="25" pageNumber="76">1.40</td>
</tr>
<tr id="A7A134E3FFE7007AD090F8F7FA90F8BC" box="[159,1347,1838,1860]" gridrow="2" pageId="25" pageNumber="76">
<th id="E4705D9FFFE7007AD090F8F7FEC3F8BC" box="[159,272,1838,1860]" gridcol="0" gridrow="2" pageId="25" pageNumber="76">Paratype A</th>
<td id="E4705D9FFFE7007AD14AF8F7FEADF8BC" box="[325,382,1838,1860]" gridcol="1" gridrow="2" pageId="25" pageNumber="76">10.70</td>
<td id="E4705D9FFFE7007AD1DEF8F7FE2EF8BC" box="[465,509,1838,1860]" gridcol="2" gridrow="2" pageId="25" pageNumber="76">7.90</td>
<td id="E4705D9FFFE7007AD252F8F7FD5AF8BC" box="[605,649,1838,1860]" gridcol="3" gridrow="2" pageId="25" pageNumber="76">1.10</td>
<td id="E4705D9FFFE7007AD2E7F8F7FCC7F8BC" box="[744,788,1838,1860]" gridcol="4" gridrow="2" pageId="25" pageNumber="76">1.95</td>
<td id="E4705D9FFFE7007AD37BF8F7FC73F8BC" box="[884,928,1838,1860]" gridcol="5" gridrow="2" pageId="25" pageNumber="76">0.56</td>
<td id="E4705D9FFFE7007AD40FF8F7FBFFF8BC" box="[1024,1068,1838,1860]" gridcol="6" gridrow="2" pageId="25" pageNumber="76">3.60</td>
<td id="E4705D9FFFE7007AD484F8F7FB64F8BC" box="[1163,1207,1838,1860]" gridcol="7" gridrow="2" pageId="25" pageNumber="76">2.70</td>
<td id="E4705D9FFFE7007AD518F8F7FA90F8BC" box="[1303,1347,1838,1860]" gridcol="8" gridrow="2" pageId="25" pageNumber="76">1.50</td>
</tr>
<tr id="A7A134E3FFE7007AD090F881FA90F896" box="[159,1347,1880,1902]" gridrow="3" pageId="25" pageNumber="76">
<th id="E4705D9FFFE7007AD090F881FEC3F896" box="[159,272,1880,1902]" gridcol="0" gridrow="3" pageId="25" pageNumber="76">Paratype B</th>
<td id="E4705D9FFFE7007AD14AF881FEADF896" box="[325,382,1880,1902]" gridcol="1" gridrow="3" pageId="25" pageNumber="76">11.75</td>
<td id="E4705D9FFFE7007AD1DEF881FE2EF896" box="[465,509,1880,1902]" gridcol="2" gridrow="3" pageId="25" pageNumber="76">9.20</td>
<td id="E4705D9FFFE7007AD252F881FD5AF896" box="[605,649,1880,1902]" gridcol="3" gridrow="3" pageId="25" pageNumber="76">2.30</td>
<td id="E4705D9FFFE7007AD2E7F881FCC7F896" box="[744,788,1880,1902]" gridcol="4" gridrow="3" pageId="25" pageNumber="76">2.10</td>
<td id="E4705D9FFFE7007AD37BF881FC73F896" box="[884,928,1880,1902]" gridcol="5" gridrow="3" pageId="25" pageNumber="76">0.84</td>
<td id="E4705D9FFFE7007AD40FF881FBFFF896" box="[1024,1068,1880,1902]" gridcol="6" gridrow="3" pageId="25" pageNumber="76">4.90</td>
<td id="E4705D9FFFE7007AD484F881FB64F896" box="[1163,1207,1880,1902]" gridcol="7" gridrow="3" pageId="25" pageNumber="76">3.60</td>
<td id="E4705D9FFFE7007AD518F881FA90F896" box="[1303,1347,1880,1902]" gridcol="8" gridrow="3" pageId="25" pageNumber="76">1.80</td>
</tr>
<tr id="A7A134E3FFE7007AD090F858FA90F86F" box="[159,1347,1921,1943]" gridrow="4" pageId="25" pageNumber="76">
<th id="E4705D9FFFE7007AD090F858FEC3F86F" box="[159,272,1921,1943]" gridcol="0" gridrow="4" pageId="25" pageNumber="76">Paratype C</th>
<td id="E4705D9FFFE7007AD14AF858FEADF86F" box="[325,382,1921,1943]" gridcol="1" gridrow="4" pageId="25" pageNumber="76">11.80</td>
<td id="E4705D9FFFE7007AD1DEF858FE2EF86F" box="[465,509,1921,1943]" gridcol="2" gridrow="4" pageId="25" pageNumber="76">9.30</td>
<td id="E4705D9FFFE7007AD252F858FD5AF86F" box="[605,649,1921,1943]" gridcol="3" gridrow="4" pageId="25" pageNumber="76">1.20</td>
<td id="E4705D9FFFE7007AD2E7F858FCC7F86F" box="[744,788,1921,1943]" gridcol="4" gridrow="4" pageId="25" pageNumber="76">2.50</td>
<td id="E4705D9FFFE7007AD37BF858FC73F86F" box="[884,928,1921,1943]" gridcol="5" gridrow="4" pageId="25" pageNumber="76">0.77</td>
<td id="E4705D9FFFE7007AD40FF858FBFFF86F" box="[1024,1068,1921,1943]" gridcol="6" gridrow="4" pageId="25" pageNumber="76">4.30</td>
<td id="E4705D9FFFE7007AD484F858FB64F86F" box="[1163,1207,1921,1943]" gridcol="7" gridrow="4" pageId="25" pageNumber="76">3.40</td>
<td id="E4705D9FFFE7007AD518F858FA90F86F" box="[1303,1347,1921,1943]" gridcol="8" gridrow="4" pageId="25" pageNumber="76">2.20</td>
</tr>
</table>
</paragraph>
<paragraph id="192E36A1FFE4FF9FD0C8FF4EFC1BFF2D" blockId="26.[151,1437,151,357]" pageId="26" pageNumber="77">
<emphasis id="2BE5EAB3FFE4FF9FD0C8FF4EFEE5FF48" bold="true" box="[199,310,151,176]" pageId="26" pageNumber="77">Material.</emphasis>
Four aboral cups, including the
<typeStatus id="C62A8803FFE4FF9FD32FFF4EFC51FF48" box="[800,898,151,176]" pageId="26" pageNumber="77" type="holotype">holotype</typeStatus>
(MNHN.F.A69935) and three
<typeStatus id="C62A8803FFE4FF9FD53FFF41FA4FFF48" box="[1328,1436,152,176]" pageId="26" pageNumber="77" type="paratype">paratypes</typeStatus>
(MNHN.F.A69936). Quantitative parameters are listed in
<tableCitation id="5413031AFFE4FF9FD32FFF65FCAEFF2D" box="[800,893,188,213]" captionStart="TABLE 5" captionStartId="25.[151,239,1609,1632]" captionText="TABLE 5. Main quantitative characters of external cup morphology in the type series of Conocrinus cahuzaci n. sp. Hc: cup height, Hb: basal ring height, Hpr: maximum height of interradial processes, Ws: maximum width of arm socket, Wr: distal radial width at level of arm socket, Dr: radial ring diameter at level of arm sockets, Drb: diameter at basal/ radial transition, Db: basal ring diameter at cup base. Values in mm." pageId="26" pageNumber="77">Tables 5</tableCitation>
and
<tableCitation id="5413031AFFE4FF9FD3BBFF65FC16FF2D" box="[948,965,188,213]" captionStart="TABLE 6" captionStartId="26.[151,239,404,427]" captionText="TABLE 6. Main morphological ratios in aboral cups of the type series of Conocrinus cahuzaci n. sp. See Table 5 for abbreviations." pageId="26" pageNumber="77">6</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="518B652AFFE4FF9FD0C8FF06FC93FE9C" pageId="26" pageNumber="77" type="diagnosis">
<paragraph id="192E36A1FFE4FF9FD0C8FF06FC93FE9C" blockId="26.[151,1437,151,357]" pageId="26" pageNumber="77">
<emphasis id="2BE5EAB3FFE4FF9FD0C8FF06FE92FF00" bold="true" box="[199,321,223,248]" pageId="26" pageNumber="77">Diagnosis.</emphasis>
Species with an abrupt enlargement of cup diameter at the level of basal/radial transition, five conspicuous and stout hypermorphic interradial processes, coarse and well-developed interradial ribs, ratio of central depression to radial ring diameter 0.44; arm articulatiom of muscular synarthry oval to semi-circular in inner position, ratio of arm socket to distal radial width 0.29.
</paragraph>
</subSubSection>
<caption id="4DEE6629FFE4FF9FD098FE4DFEFDFE33" pageId="26" pageNumber="77">
<paragraph id="192E36A1FFE4FF9FD098FE4DFEFDFE33" blockId="26.[151,1436,404,459]" pageId="26" pageNumber="77">
<emphasis id="2BE5EAB3FFE4FF9FD098FE4DFEDFFE53" bold="true" box="[151,268,404,427]" pageId="26" pageNumber="77">TABLE 6.</emphasis>
Main morphological ratios in aboral cups of the type series of
<taxonomicName id="DE914D22FFE4FF9FD3CDFE4DFB76FE53" authority="Roux &amp; Eléaume &amp; Améziane, 2019" authorityName="Roux &amp; Eléaume &amp; Améziane" authorityYear="2019" box="[962,1189,404,427]" class="Crinoidea" family="Bathycrinidae" genus="Conocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="26" pageNumber="77" phylum="Echinodermata" rank="species" species="cahuzaci" status="sp. nov.">
<emphasis id="2BE5EAB3FFE4FF9FD3CDFE4DFB76FE53" box="[962,1189,404,427]" italics="true" pageId="26" pageNumber="77">Conocrinus cahuzaci</emphasis>
</taxonomicName>
<emphasis id="2BE5EAB3FFE4FF9FD4A1FE4DFB3FFE53" bold="true" box="[1198,1260,404,427]" pageId="26" pageNumber="77">
<taxonomicNameLabel id="30D657C8FFE4FF9FD4A1FE4DFB3FFE53" box="[1198,1260,404,427]" pageId="26" pageNumber="77" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
See Table 5 for abbreviations.
</paragraph>
</caption>
<paragraph id="192E36A1FFE4FF9FD090FE3FFAFCFD5A" pageId="26" pageNumber="77">
<table id="6B91C401FFE4007AD090FE3FFA90FD5A" box="[159,1347,486,674]" gridcols="8" gridrows="5" pageId="26" pageNumber="77" rowsContinueFrom="25.[159,1347,1756,1943]">
<tr id="A7A134E3FFE4007AD090FE3FFA90FE04" box="[159,1347,486,508]" gridrow="0" pageId="26" pageNumber="77">
<th id="E4705D9FFFE4007AD090FE3FFEC3FE04" box="[159,272,486,508]" gridcol="0" gridrow="0" pageId="26" pageNumber="77">Ratios</th>
<th id="E4705D9FFFE4007AD130FE3FFE50FE04" box="[319,387,486,508]" gridcol="1" gridrow="0" pageId="26" pageNumber="77">Hb/Hc</th>
<th id="E4705D9FFFE4007AD1EFFE3FFDCCFE04" box="[480,543,486,508]" gridcol="2" gridrow="0" pageId="26" pageNumber="77">Dr/Hc</th>
<th id="E4705D9FFFE4007AD28FFE3FFD1EFE04" box="[640,717,486,508]" gridcol="3" gridrow="0" pageId="26" pageNumber="77">Drb/Hb</th>
<th id="E4705D9FFFE4007AD32EFE3FFCB9FE04" box="[801,874,486,508]" gridcol="4" gridrow="0" pageId="26" pageNumber="77">Drb/Dr</th>
<th id="E4705D9FFFE4007AD3CEFE3FFBDDFE04" box="[961,1038,486,508]" gridcol="5" gridrow="0" pageId="26" pageNumber="77">Db/Drb</th>
<th id="E4705D9FFFE4007AD46DFE3FFB79FE04" box="[1122,1194,486,508]" gridcol="6" gridrow="0" pageId="26" pageNumber="77">Ws/Wr</th>
<th id="E4705D9FFFE4007AD50CFE3FFA90FE04" box="[1283,1347,486,508]" gridcol="7" gridrow="0" pageId="26" pageNumber="77">Db/Dr</th>
</tr>
<tr id="A7A134E3FFE4007AD090FDC9FA90FDDE" box="[159,1347,528,550]" gridrow="1" pageId="26" pageNumber="77">
<th id="E4705D9FFFE4007AD090FDC9FEC3FDDE" box="[159,272,528,550]" gridcol="0" gridrow="1" pageId="26" pageNumber="77">Holotype</th>
<td id="E4705D9FFFE4007AD130FDC9FE50FDDE" box="[319,387,528,550]" gridcol="1" gridrow="1" pageId="26" pageNumber="77">0.74</td>
<td id="E4705D9FFFE4007AD1EFFDC9FDCCFDDE" box="[480,543,528,550]" gridcol="2" gridrow="1" pageId="26" pageNumber="77">0.34</td>
<td id="E4705D9FFFE4007AD28FFDC9FD1EFDDE" box="[640,717,528,550]" gridcol="3" gridrow="1" pageId="26" pageNumber="77">0.29</td>
<td id="E4705D9FFFE4007AD32EFDC9FCB9FDDE" box="[801,874,528,550]" gridcol="4" gridrow="1" pageId="26" pageNumber="77">0.64</td>
<td id="E4705D9FFFE4007AD3CEFDC9FBDDFDDE" box="[961,1038,528,550]" gridcol="5" gridrow="1" pageId="26" pageNumber="77">0.56</td>
<td id="E4705D9FFFE4007AD46DFDC9FB79FDDE" box="[1122,1194,528,550]" gridcol="6" gridrow="1" pageId="26" pageNumber="77">0.29</td>
<td id="E4705D9FFFE4007AD50CFDC9FA90FDDE" box="[1283,1347,528,550]" gridcol="7" gridrow="1" pageId="26" pageNumber="77">0.36</td>
</tr>
<tr id="A7A134E3FFE4007AD090FDE0FA90FDB7" box="[159,1347,569,591]" gridrow="2" pageId="26" pageNumber="77">
<th id="E4705D9FFFE4007AD090FDE0FEC3FDB7" box="[159,272,569,591]" gridcol="0" gridrow="2" pageId="26" pageNumber="77">Paratype A</th>
<td id="E4705D9FFFE4007AD130FDE0FE50FDB7" box="[319,387,569,591]" gridcol="1" gridrow="2" pageId="26" pageNumber="77">0.74</td>
<td id="E4705D9FFFE4007AD1EFFDE0FDCCFDB7" box="[480,543,569,591]" gridcol="2" gridrow="2" pageId="26" pageNumber="77">0.34</td>
<td id="E4705D9FFFE4007AD28FFDE0FD1EFDB7" box="[640,717,569,591]" gridcol="3" gridrow="2" pageId="26" pageNumber="77">0.34</td>
<td id="E4705D9FFFE4007AD32EFDE0FCB9FDB7" box="[801,874,569,591]" gridcol="4" gridrow="2" pageId="26" pageNumber="77">0.75</td>
<td id="E4705D9FFFE4007AD3CEFDE0FBDDFDB7" box="[961,1038,569,591]" gridcol="5" gridrow="2" pageId="26" pageNumber="77">0.56</td>
<td id="E4705D9FFFE4007AD46DFDE0FB79FDB7" box="[1122,1194,569,591]" gridcol="6" gridrow="2" pageId="26" pageNumber="77">0.27</td>
<td id="E4705D9FFFE4007AD50CFDE0FA90FDB7" box="[1283,1347,569,591]" gridcol="7" gridrow="2" pageId="26" pageNumber="77">0.42</td>
</tr>
<tr id="A7A134E3FFE4007AD090FDBBFA90FD80" box="[159,1347,610,632]" gridrow="3" pageId="26" pageNumber="77">
<th id="E4705D9FFFE4007AD090FDBBFEC3FD80" box="[159,272,610,632]" gridcol="0" gridrow="3" pageId="26" pageNumber="77">Paratype B</th>
<td id="E4705D9FFFE4007AD130FDBBFE50FD80" box="[319,387,610,632]" gridcol="1" gridrow="3" pageId="26" pageNumber="77">0.78</td>
<td id="E4705D9FFFE4007AD1EFFDBBFDCCFD80" box="[480,543,610,632]" gridcol="2" gridrow="3" pageId="26" pageNumber="77">0.42</td>
<td id="E4705D9FFFE4007AD28FFDBBFD1EFD80" box="[640,717,610,632]" gridcol="3" gridrow="3" pageId="26" pageNumber="77">0.39</td>
<td id="E4705D9FFFE4007AD32EFDBBFCB9FD80" box="[801,874,610,632]" gridcol="4" gridrow="3" pageId="26" pageNumber="77">0.73</td>
<td id="E4705D9FFFE4007AD3CEFDBBFBDDFD80" box="[961,1038,610,632]" gridcol="5" gridrow="3" pageId="26" pageNumber="77">0.50</td>
<td id="E4705D9FFFE4007AD46DFDBBFB79FD80" box="[1122,1194,610,632]" gridcol="6" gridrow="3" pageId="26" pageNumber="77">0.40</td>
<td id="E4705D9FFFE4007AD50CFDBBFA90FD80" box="[1283,1347,610,632]" gridcol="7" gridrow="3" pageId="26" pageNumber="77">0.37</td>
</tr>
<tr id="A7A134E3FFE4007AD090FD55FA90FD5A" box="[159,1347,652,674]" gridrow="4" pageId="26" pageNumber="77">
<th id="E4705D9FFFE4007AD090FD55FEC3FD5A" box="[159,272,652,674]" gridcol="0" gridrow="4" pageId="26" pageNumber="77">Paratype C</th>
<td id="E4705D9FFFE4007AD130FD55FE50FD5A" box="[319,387,652,674]" gridcol="1" gridrow="4" pageId="26" pageNumber="77">0.79</td>
<td id="E4705D9FFFE4007AD1EFFD55FDCCFD5A" box="[480,543,652,674]" gridcol="2" gridrow="4" pageId="26" pageNumber="77">0.36</td>
<td id="E4705D9FFFE4007AD28FFD55FD1EFD5A" box="[640,717,652,674]" gridcol="3" gridrow="4" pageId="26" pageNumber="77">0.37</td>
<td id="E4705D9FFFE4007AD32EFD55FCB9FD5A" box="[801,874,652,674]" gridcol="4" gridrow="4" pageId="26" pageNumber="77">0.79</td>
<td id="E4705D9FFFE4007AD3CEFD55FBDDFD5A" box="[961,1038,652,674]" gridcol="5" gridrow="4" pageId="26" pageNumber="77">0.65</td>
<td id="E4705D9FFFE4007AD46DFD55FB79FD5A" box="[1122,1194,652,674]" gridcol="6" gridrow="4" pageId="26" pageNumber="77">0.31</td>
<td id="E4705D9FFFE4007AD50CFD55FA90FD5A" box="[1283,1347,652,674]" gridcol="7" gridrow="4" pageId="26" pageNumber="77">0.36</td>
</tr>
</table>
</paragraph>
<subSubSection id="518B652AFFE4FF9FD0C8FD3BFD01FC2B" pageId="26" pageNumber="77" type="description">
<paragraph id="192E36A1FFE4FF9FD0C8FD3BFD01FC2B" blockId="26.[151,1436,738,1123]" pageId="26" pageNumber="77">
<emphasis id="2BE5EAB3FFE4FF9FD0C8FD3BFD45FD03" bold="true" box="[199,662,738,763]" pageId="26" pageNumber="77">
Description of
<typeStatus id="C62A8803FFE4FF9FD178FD3BFE0DFD03" box="[375,478,738,763]" pageId="26" pageNumber="77" type="holotype">holotype</typeStatus>
and
<typeStatus id="C62A8803FFE4FF9FD216FD3BFD42FD03" box="[537,657,738,763]" pageId="26" pageNumber="77" type="paratype">paratypes</typeStatus>
.
</emphasis>
Aboral cup of the
<typeStatus id="C62A8803FFE4FF9FD366FD3BFC18FD03" box="[873,971,738,763]" pageId="26" pageNumber="77" type="holotype">holotype</typeStatus>
has an abrupt enlargement of diameter at the level of the presumed basal/radial transition, five conspicuous and stout hypermorphic interradial processes of equal size (
<figureCitation id="81AA2A24FFE4FF9FD114FCF3FEB8FCBB" box="[283,363,810,835]" captionStart="FIGURE 5" captionStartId="11.[151,250,1675,1697]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@11.[151,1436,193,1653]" captionTargetPageId="11" captionText="FIGURE 5. Aboral cups of Conocrinus thorenti (A-D) and other Eocene species from southern France closely related to it. A- D: Conocrinus thorenti, lectotype MNHN.F.R54426, AB: oblique lateral view, CD: adoral view, D: closeup of arm socket indicated by arrow in C; EG: Conocrinus archiaci, Villa Marbella, Biarritz, EF: specimen MNHN.F.A69934a, E: lateral view, F: oblique adoral view with closeup of arm socket, G: specimen MNHN.F.A69934b, oblique adoral view; H: Conocrinus aff. archiaci, Les Scaffarels near Annot, MHN Nice, Campredon Collection, specimen no. 6, flat arm socket (synostosis); IK: Conocrinus cahuzaci n. sp., holotype, MNHN.F.A69935, I oblique side adoral view, JK: adoral views; LO: Conocrinus duperrieri, marnes à Pentacrines, Côte des Basques, Biarritz, LN: holotype MNHN.F.R06934 (arrow head indicating segment of radial suture), O: side view of paratype MNHN.F.R06935; bip: broken interradial process, ip: interradial process, ir: interradial rib. See Figure 1 for other abbreviations." httpUri="https://zenodo.org/record/2627382/files/figure.png" pageId="26" pageNumber="77">
Fig.
<date id="6D2F1061FFE4FF9FD15DFCF3FEB8FCBB" box="[338,363,810,835]" pageId="26" pageNumber="77">5I</date>
</figureCitation>
), coarse and well-developed interradial ribs and a hemispherical central depression (
<figureCitation id="81AA2A24FFE4FF9FD534FCF3FA5DFCBB" box="[1339,1422,810,835]" captionStart="FIGURE 5" captionStartId="11.[151,250,1675,1697]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@11.[151,1436,193,1653]" captionTargetPageId="11" captionText="FIGURE 5. Aboral cups of Conocrinus thorenti (A-D) and other Eocene species from southern France closely related to it. A- D: Conocrinus thorenti, lectotype MNHN.F.R54426, AB: oblique lateral view, CD: adoral view, D: closeup of arm socket indicated by arrow in C; EG: Conocrinus archiaci, Villa Marbella, Biarritz, EF: specimen MNHN.F.A69934a, E: lateral view, F: oblique adoral view with closeup of arm socket, G: specimen MNHN.F.A69934b, oblique adoral view; H: Conocrinus aff. archiaci, Les Scaffarels near Annot, MHN Nice, Campredon Collection, specimen no. 6, flat arm socket (synostosis); IK: Conocrinus cahuzaci n. sp., holotype, MNHN.F.A69935, I oblique side adoral view, JK: adoral views; LO: Conocrinus duperrieri, marnes à Pentacrines, Côte des Basques, Biarritz, LN: holotype MNHN.F.R06934 (arrow head indicating segment of radial suture), O: side view of paratype MNHN.F.R06935; bip: broken interradial process, ip: interradial process, ir: interradial rib. See Figure 1 for other abbreviations." httpUri="https://zenodo.org/record/2627382/files/figure.png" pageId="26" pageNumber="77">Fig. 5J</figureCitation>
); ratio of central depression to radial ring diameter 0.44; arm articulatiom of muscular synarthry, oval to semicircular in inner position regarding outer edge of the radial circlet (
<figureCitation id="81AA2A24FFE4FF9FD3AFFCABFC2FFC73" box="[928,1020,882,907]" captionStart="FIGURE 5" captionStartId="11.[151,250,1675,1697]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@11.[151,1436,193,1653]" captionTargetPageId="11" captionText="FIGURE 5. Aboral cups of Conocrinus thorenti (A-D) and other Eocene species from southern France closely related to it. A- D: Conocrinus thorenti, lectotype MNHN.F.R54426, AB: oblique lateral view, CD: adoral view, D: closeup of arm socket indicated by arrow in C; EG: Conocrinus archiaci, Villa Marbella, Biarritz, EF: specimen MNHN.F.A69934a, E: lateral view, F: oblique adoral view with closeup of arm socket, G: specimen MNHN.F.A69934b, oblique adoral view; H: Conocrinus aff. archiaci, Les Scaffarels near Annot, MHN Nice, Campredon Collection, specimen no. 6, flat arm socket (synostosis); IK: Conocrinus cahuzaci n. sp., holotype, MNHN.F.A69935, I oblique side adoral view, JK: adoral views; LO: Conocrinus duperrieri, marnes à Pentacrines, Côte des Basques, Biarritz, LN: holotype MNHN.F.R06934 (arrow head indicating segment of radial suture), O: side view of paratype MNHN.F.R06935; bip: broken interradial process, ip: interradial process, ir: interradial rib. See Figure 1 for other abbreviations." httpUri="https://zenodo.org/record/2627382/files/figure.png" pageId="26" pageNumber="77">Fig. 5K</figureCitation>
), ratio of arm socket to distal radial width 0.29. Two
<typeStatus id="C62A8803FFE4FF9FD15CFC4EFE6DFC57" box="[339,446,919,943]" pageId="26" pageNumber="77" type="paratype">paratypes</typeStatus>
(A and C) with five shorter interradial processes of equal size, and the third (B) with one interradial process substantially longer than others.
</paragraph>
</subSubSection>
<subSubSection id="518B652AFFE4FF9FD0C8FC07FDA4FB9A" pageId="26" pageNumber="77" type="discussion">
<paragraph id="192E36A1FFE4FF9FD0C8FC07FDA4FB9A" blockId="26.[151,1436,738,1123]" pageId="26" pageNumber="77">
<emphasis id="2BE5EAB3FFE4FF9FD0C8FC07FEE8FC0F" bold="true" box="[199,315,990,1015]" pageId="26" pageNumber="77">Remarks.</emphasis>
Cups of
<taxonomicName id="DE914D22FFE4FF9FD1A2FC06FD4CFC0F" authority="Roux &amp; Eléaume &amp; Améziane, 2019" authorityName="Roux &amp; Eléaume &amp; Améziane" authorityYear="2019" box="[429,671,990,1015]" class="Crinoidea" family="Bathycrinidae" genus="Conocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="26" pageNumber="77" phylum="Echinodermata" rank="species" species="cahuzaci" status="sp. nov.">
<emphasis id="2BE5EAB3FFE4FF9FD1A2FC06FD4CFC0F" box="[429,671,990,1015]" italics="true" pageId="26" pageNumber="77">Conocrinus cahuzaci</emphasis>
</taxonomicName>
<emphasis id="2BE5EAB3FFE4FF9FD2A5FC07FD3EFC0F" bold="true" box="[682,749,990,1015]" pageId="26" pageNumber="77">
<taxonomicNameLabel id="30D657C8FFE4FF9FD2A5FC07FD3EFC0F" box="[682,749,990,1015]" pageId="26" pageNumber="77" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, one of the
<typeStatus id="C62A8803FFE4FF9FD38CFC06FC3CFC0F" box="[899,1007,991,1015]" pageId="26" pageNumber="77" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="DE914D22FFE4FF9FD413FC06FB67FC0F" authorityName="Roux" authorityYear="1978" box="[1052,1204,990,1015]" class="Crinoidea" family="Bathycrinidae" genus="Conocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="26" pageNumber="77" phylum="Echinodermata" rank="species" species="duperrieri">
<emphasis id="2BE5EAB3FFE4FF9FD413FC06FB67FC0F" box="[1052,1204,990,1015]" italics="true" pageId="26" pageNumber="77">C. duperrieri</emphasis>
</taxonomicName>
,
<taxonomicName id="DE914D22FFE4FF9FD4C8FC06FA97FC0F" authorityName="d'Archiac" authorityYear="1846" box="[1223,1348,990,1015]" class="Crinoidea" family="Bourgueticrinidae" genus="Bourgueticrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="26" pageNumber="77" phylum="Echinodermata" rank="species" species="thorenti">
<emphasis id="2BE5EAB3FFE4FF9FD4C8FC06FA97FC0F" box="[1223,1348,990,1015]" italics="true" pageId="26" pageNumber="77">C. thorenti</emphasis>
</taxonomicName>
and
<emphasis id="2BE5EAB3FFE4FF9FD58DFC06FEDDFBE3" italics="true" pageId="26" pageNumber="77">C. veronensis</emphasis>
share a rounded to semi-circular arm socket in inner position.
<taxonomicName id="DE914D22FFE4FF9FD3D3FBDAFB10FBE3" authorityName="Roux" authorityYear="1978" box="[988,1219,1026,1051]" class="Crinoidea" family="Bathycrinidae" genus="Conocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="26" pageNumber="77" phylum="Echinodermata" rank="species" species="archiaci">
<emphasis id="2BE5EAB3FFE4FF9FD3D3FBDAFB10FBE3" box="[988,1219,1026,1051]" italics="true" pageId="26" pageNumber="77">Conocrinus archiaci</emphasis>
</taxonomicName>
differs in having a trapezoidal socket adjacent to the outer edge of the radial circlet. Interradial processes are substantially less developed in
<taxonomicName id="DE914D22FFE4FF9FD122FB92FE12FB9B" authorityName="Roux" authorityYear="1978" box="[301,449,1098,1123]" class="Crinoidea" family="Bathycrinidae" genus="Conocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="26" pageNumber="77" phylum="Echinodermata" rank="species" species="duperrieri">
<emphasis id="2BE5EAB3FFE4FF9FD122FB92FE12FB9B" box="[301,449,1098,1123]" italics="true" pageId="26" pageNumber="77">C. duperrieri</emphasis>
</taxonomicName>
and
<taxonomicName id="DE914D22FFE4FF9FD1F7FB92FDA3FB9B" authorityName="d'Archiac" authorityYear="1846" box="[504,624,1098,1123]" class="Crinoidea" family="Bourgueticrinidae" genus="Bourgueticrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="26" pageNumber="77" phylum="Echinodermata" rank="species" species="thorenti">
<emphasis id="2BE5EAB3FFE4FF9FD1F7FB92FDA3FB9B" box="[504,624,1098,1123]" italics="true" pageId="26" pageNumber="77">C. thorenti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>